171 lines
19 KiB
XML
171 lines
19 KiB
XML
<document id="17269116E538C76DCCF5BD8332791246" ID-DOI="10.11646/zootaxa.2435.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5315673" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629671386604" checkinUser="felipe" docAuthor="Oláh, János & Johanson, Kjell Arne" docDate="2010" docId="0393CE26FFD0FFD37CFB8E07FEDCFDA9" docLanguage="en" docName="zt02435p063.pdf" docOrigin="Zootaxa 2435 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2435.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Nyctiophylax (Paranyctiophylax) kilah Oláh & Johanson 2010, new species" docType="treatment" docVersion="5" lastPageNumber="48" masterDocId="FFAAB65EFFFEFFFC7C6C8801FFE1FFE5" masterDocTitle="Generic review of Polycentropodidae with description of 32 new species and 19 new species records from the Oriental, Australian and Afrotropical Biogeographical Regions 2435" masterLastPageNumber="63" masterPageNumber="1" pageNumber="47" updateTime="1699093793393" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="6B7318C686EEAD01E260612C79439A20">
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<mods:title id="FFF0BFD4F68C13C8BEF529D455803102">Generic review of Polycentropodidae with description of 32 new species and 19 new species records from the Oriental, Australian and Afrotropical Biogeographical Regions 2435</mods:title>
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<mods:namePart id="386DB7BF3698134EDA9042572804D9A5">Oláh, János</mods:namePart>
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<mods:namePart id="57441BC9FDB64EAF898E8A66599483BD">Johanson, Kjell Arne</mods:namePart>
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<mods:title id="AEBC57915D8349842FAECC63C6C533F3">Zootaxa</mods:title>
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<mods:part id="E35566D6E7860EB26EF1A734454A2C79">
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<mods:date id="4999F37502AC314BF34EB8DFDE551090">2010</mods:date>
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<mods:number id="F5B53A360F0CC0F3D34555A4302AD81F">2010-04-23</mods:number>
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<mods:detail id="D8A8805EFADD326E537779620BB145DA" type="volume">
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<mods:number id="2ADC02933F675523E1839ACCF6FDAEBD">2435</mods:number>
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<mods:url id="5482E869388138E121441E0C74CF5901">https://biotaxa.org/Zootaxa/article/view/zootaxa.2435.1.1</mods:url>
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<mods:classification id="3CD9ED4D96B9EEDB666C403426736E36">journal article</mods:classification>
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<mods:identifier id="931F06A5953192B48225627E65958C91" type="DOI">10.11646/zootaxa.2435.1.1</mods:identifier>
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<mods:identifier id="D986CDDB15EA5F5CD007AA8B2D0F6032" type="ISSN">11755334</mods:identifier>
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<subSubSection id="C3202CBBFFD0FFD27CFB8E07FCEBF9C5" box="[151,778,1542,1568]" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="8B857F30FFD0FFD27CFB8E07FCEBF9C5" blockId="46.[151,778,1542,1603]" box="[151,778,1542,1568]" pageId="46" pageNumber="47">
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<heading id="D0CDC85CFFD0FFD27CFB8E07FCEBF9C5" bold="true" box="[151,778,1542,1568]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
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<emphasis id="B94EA322FFD0FFD27CFB8E07FCEBF9C5" bold="true" box="[151,778,1542,1568]" pageId="46" pageNumber="47">
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<taxonomicName id="4C3A04B3FFD0FFD27CFB8E07FD8DF9C5" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[151,620,1542,1568]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="kilah" status="sp. nov." subGenus="Paranyctiophylax">
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<emphasis id="B94EA322FFD0FFD27CFB8E07FED6F9C5" bold="true" box="[151,311,1542,1568]" italics="true" pageId="46" pageNumber="47">Nyctiophylax</emphasis>
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(
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<emphasis id="B94EA322FFD0FFD27D2B8E07FDFDF9C5" bold="true" box="[327,540,1542,1568]" italics="true" pageId="46" pageNumber="47">Paranyctiophylax</emphasis>
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)
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<emphasis id="B94EA322FFD0FFD27E428E07FD8DF9C5" bold="true" box="[558,620,1542,1568]" italics="true" pageId="46" pageNumber="47">kilah</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27D1E59FFD0FFD27E168E07FCEBF9C5" box="[634,778,1542,1568]" pageId="46" pageNumber="47" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3202CBBFFD0FFD27CFB8E28FEDEF9A6" box="[151,319,1577,1603]" pageId="46" pageNumber="47" type="description">
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<paragraph id="8B857F30FFD0FFD27CFB8E28FEDEF9A6" blockId="46.[151,778,1542,1603]" box="[151,319,1577,1603]" pageId="46" pageNumber="47">
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<figureCitation id="130163B5FFD0FFD27CFB8E28FEDEF9A6" box="[151,319,1577,1603]" captionStart="FIGURES 83–86" captionStartId="47.[151,273,1636,1660]" captionTargetBox="[173,1411,661,1605]" captionTargetId="figure-256@47.[167,1419,645,1615]" captionTargetPageId="47" captionText="FIGURES 83–86. Nyctiophylax (Paranyctiophylax) kilah, new species, holotype. 83—genitalia, left lateral; 84— genitalia, dorsal; 85—left gonopod, ventral; 86—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361558" httpUri="https://zenodo.org/record/5361558/files/figure.png" pageId="46" pageNumber="47">Figures 83–86</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3202CBBFFD0FFD27CFB8E72FC02F93D" pageId="46" pageNumber="47" type="diagnosis">
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<paragraph id="8B857F30FFD0FFD27CFB8E72FC02F93D" blockId="46.[151,1436,1651,2013]" pageId="46" pageNumber="47">
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<emphasis id="B94EA322FFD0FFD27CFB8E72FEF8F968" bold="true" box="[151,281,1651,1677]" pageId="46" pageNumber="47">Diagnosis:</emphasis>
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This species has triangular gonopods as in
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<taxonomicName id="4C3A04B3FFD0FFD27F7A8E75FBADF968" authority="Kimmins" authorityName="Kimmins" authorityYear="1957" box="[790,1100,1651,1677]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="africanus" subGenus="Paranyctiophylax">
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<emphasis id="B94EA322FFD0FFD27F7A8E75FC36F968" box="[790,983,1652,1677]" italics="true" pageId="46" pageNumber="47">N. (P.) africanus</emphasis>
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Kimmins
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</taxonomicName>
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from East Africa and
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<taxonomicName id="4C3A04B3FFD0FFD2793D8E75FEA4F956" authority="Gibon" authorityName="Gibon" authorityYear="1986" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="aurorae" subGenus="Paranyctiophylax">
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<emphasis id="B94EA322FFD0FFD2793D8E75FF15F957" italics="true" pageId="46" pageNumber="47">N. (P.) aurorae</emphasis>
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Gibon
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</taxonomicName>
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and
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<taxonomicName id="4C3A04B3FFD0FFD27DEC8E98FD7AF956" authority="Gibon" authorityName="Gibon" authorityYear="1986" box="[384,667,1689,1715]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="brenguesi" subGenus="Paranyctiophylax">
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<emphasis id="B94EA322FFD0FFD27DEC8E98FDA6F957" box="[384,583,1689,1714]" italics="true" pageId="46" pageNumber="47">N. (P.) brenguesi</emphasis>
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Gibon
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</taxonomicName>
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, from West Africa. This new species is easily distinguished from those 3 species by its bifurcate apex of each ventral paraproctal process.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3202CBBFFD0FFD37CA88EE2FB84FE77" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
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<paragraph id="8B857F30FFD0FFD27CA88EE2FCECF8C6" blockId="46.[151,1436,1651,2013]" pageId="46" pageNumber="47">
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<emphasis id="B94EA322FFD0FFD27CA88EE2FEBFF918" bold="true" box="[196,350,1763,1789]" pageId="46" pageNumber="47">Description:</emphasis>
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Male. Body uniformly dark brown. Maxillary palp formula (I,II)-IV-III-V. Forewing length 6.0 mm. Forewing anal veins
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<emphasis id="B94EA322FFD0FFD27D988F08FDF5F8C7" box="[500,532,1801,1826]" italics="true" pageId="46" pageNumber="47">A1</emphasis>
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,
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<emphasis id="B94EA322FFD0FFD27E4F8F08FDA2F8C7" box="[547,579,1801,1826]" italics="true" pageId="46" pageNumber="47">A2</emphasis>
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, and
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<emphasis id="B94EA322FFD0FFD27EE88F08FD44F8C7" box="[644,677,1801,1826]" italics="true" pageId="46" pageNumber="47">A3</emphasis>
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looping.
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</paragraph>
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<paragraph id="8B857F30FFD0FFD37CA88F2FFB84FE77" blockId="46.[151,1436,1651,2013]" lastBlockId="47.[151,1436,152,588]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
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Male genitalia. Abdominal segment IX reduced to robust sternite, nearly triangular in lateral view, with pronounced ventroapical mesal process (
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<figureCitation id="130163B5FFD0FFD27EF88F52FD13F888" box="[660,754,1875,1901]" captionStart="FIGURES 83–86" captionStartId="47.[151,273,1636,1660]" captionTargetBox="[173,1411,661,1605]" captionTargetId="figure-256@47.[167,1419,645,1615]" captionTargetPageId="47" captionText="FIGURES 83–86. Nyctiophylax (Paranyctiophylax) kilah, new species, holotype. 83—genitalia, left lateral; 84— genitalia, dorsal; 85—left gonopod, ventral; 86—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361558" httpUri="https://zenodo.org/record/5361558/files/figure.png" pageId="46" pageNumber="47">Fig. 83</figureCitation>
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); sclerotized tergite IX narrow, tall, each side fused to paraproctal base. Dorsal and ventral paraproctal processes fusing at paraproctal base. Segment X membranous, well discernible; small and triangular in lateral view (
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<figureCitation id="130163B5FFD0FFD27FD88F9FFBEDF85D" box="[948,1036,1950,1976]" captionStart="FIGURES 83–86" captionStartId="47.[151,273,1636,1660]" captionTargetBox="[173,1411,661,1605]" captionTargetId="figure-256@47.[167,1419,645,1615]" captionTargetPageId="47" captionText="FIGURES 83–86. Nyctiophylax (Paranyctiophylax) kilah, new species, holotype. 83—genitalia, left lateral; 84— genitalia, dorsal; 85—left gonopod, ventral; 86—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361558" httpUri="https://zenodo.org/record/5361558/files/figure.png" pageId="46" pageNumber="47">Fig. 83</figureCitation>
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); apically cut, with shallow mesal concavity in dorsal view. Cerci setose, narrow, elongate in lateral view, slightly curving mesad (
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<figureCitation id="130163B5FFD0FFD2795D8FC2FA6CF838" box="[1329,1421,1987,2013]" captionStart="FIGURES 83–86" captionStartId="47.[151,273,1636,1660]" captionTargetBox="[173,1411,661,1605]" captionTargetId="figure-256@47.[167,1419,645,1615]" captionTargetPageId="47" captionText="FIGURES 83–86. Nyctiophylax (Paranyctiophylax) kilah, new species, holotype. 83—genitalia, left lateral; 84— genitalia, dorsal; 85—left gonopod, ventral; 86—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361558" httpUri="https://zenodo.org/record/5361558/files/figure.png" pageId="46" pageNumber="47">Fig. 84</figureCitation>
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); slightly longer than dorsal paraproctal processes (
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<figureCitation id="130163B5FFD1FFD37E868899FCA2FF57" box="[746,835,152,178]" captionStart="FIGURES 83–86" captionStartId="47.[151,273,1636,1660]" captionTargetBox="[173,1411,661,1605]" captionTargetId="figure-256@47.[167,1419,645,1615]" captionTargetPageId="47" captionText="FIGURES 83–86. Nyctiophylax (Paranyctiophylax) kilah, new species, holotype. 83—genitalia, left lateral; 84— genitalia, dorsal; 85—left gonopod, ventral; 86—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361558" httpUri="https://zenodo.org/record/5361558/files/figure.png" pageId="47" pageNumber="48">Fig. 83</figureCitation>
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). Paraproctal complex highly sclerotized, smooth; each composed of (1) paraproctal base narrow, fused to narrow tergite IX; (2) dorsal paraproctal process slender, long, and laterad-curving; (3) ventral paraproctal process short, arching ventrad, with bifurcate apex for which ventral arm tapered, spine-like. Gonopods (
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<figureCitation id="130163B5FFD1FFD37F658909FCBFFEC7" box="[777,862,264,290]" captionStart="FIGURES 83–86" captionStartId="47.[151,273,1636,1660]" captionTargetBox="[173,1411,661,1605]" captionTargetId="figure-256@47.[167,1419,645,1615]" captionTargetPageId="47" captionText="FIGURES 83–86. Nyctiophylax (Paranyctiophylax) kilah, new species, holotype. 83—genitalia, left lateral; 84— genitalia, dorsal; 85—left gonopod, ventral; 86—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361558" httpUri="https://zenodo.org/record/5361558/files/figure.png" pageId="47" pageNumber="48">Fig. 83</figureCitation>
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) robust, broad, nearly triangular, each with small mesoventral branch; dorsal branch long, curving caudad; apices of dorsal and ventral branches tapering. Phallic apparatus (
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<figureCitation id="130163B5FFD1FFD37D188953FE2AFE89" box="[372,459,338,364]" captionStart="FIGURES 83–86" captionStartId="47.[151,273,1636,1660]" captionTargetBox="[173,1411,661,1605]" captionTargetId="figure-256@47.[167,1419,645,1615]" captionTargetPageId="47" captionText="FIGURES 83–86. Nyctiophylax (Paranyctiophylax) kilah, new species, holotype. 83—genitalia, left lateral; 84— genitalia, dorsal; 85—left gonopod, ventral; 86—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361558" httpUri="https://zenodo.org/record/5361558/files/figure.png" pageId="47" pageNumber="48">Fig. 86</figureCitation>
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) located dorsally in genitalia, guided by paraproctal processes; simple, with thick phallobase, slender distal membranes without visible sclerites or spines; apex dark.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3202CBBFFD1FFD37CA8899CFC1BFE39" pageId="47" pageNumber="48" type="materials_examined">
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<paragraph id="8B857F30FFD1FFD37CA8899CFB1BFE52" blockId="47.[151,1436,152,588]" box="[196,1274,413,439]" pageId="47" pageNumber="48">
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||
<materialsCitation id="3B52756DFFD1FFD37CA8899CFB1BFE52" box="[196,1274,413,439]" collectionCode="MNHN" collectorName="R. Paulian" country="Madagascar" location="MADAGASCAR" pageId="47" pageNumber="48" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="B94EA322FFD1FFD37CA8899CFD8AFE52" bold="true" box="[196,619,413,439]" pageId="47" pageNumber="48">
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||
<typeStatus id="5481C192FFD1FFD37CA8899CFED7FE52" box="[196,310,413,439]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
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||
male:
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||
<collectingCountry id="F32D3FA0FFD1FFD37DE6899CFD87FE52" box="[394,614,413,439]" name="Madagascar" pageId="47" pageNumber="48">MADAGASCAR</collectingCountry>
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||
:
|
||
</emphasis>
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Anosibe, “km 57 route”, 1955 [
|
||
<collectorName id="26CF1AE6FFD1FFD37F8E899CFB83FE52" box="[994,1122,413,439]" pageId="47" pageNumber="48">R. Paulian</collectorName>
|
||
]—(
|
||
<collectionCode id="ED2BE7F5FFD1FFD378FC899CFB0FFE52" box="[1168,1262,413,439]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="47" pageNumber="48">MNHN</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
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||
<paragraph id="8B857F30FFD1FFD37CA889C3FC1BFE39" blockId="47.[151,1436,152,588]" box="[196,1018,450,476]" pageId="47" pageNumber="48">
|
||
<materialsCitation id="3B52756DFFD1FFD37CA889C3FC14FE39" box="[196,1013,450,476]" collectionCode="MNHN, OPC" collectorName="R. Paulian" country="Madagascar" location="MADAGASCAR" pageId="47" pageNumber="48" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
|
||
<emphasis id="B94EA322FFD1FFD37CA889C3FEAAFE39" bold="true" box="[196,331,450,476]" pageId="47" pageNumber="48">
|
||
<typeStatus id="5481C192FFD1FFD37CA889C3FEA7FE39" box="[196,326,450,476]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
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||
:
|
||
</emphasis>
|
||
Same data as holotype —
|
||
<specimenCount id="9D3CB4B9FFD1FFD37EEE89C3FD2DFE39" box="[642,716,450,476]" pageId="47" pageNumber="48" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="ED2BE7F5FFD1FFD37EB089C3FCDDFE39" box="[732,828,450,476]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="47" pageNumber="48">MNHN</collectionCode>
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),
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||
<specimenCount id="9D3CB4B9FFD1FFD37F2389C3FC7FFE39" box="[847,926,450,476]" pageId="47" pageNumber="48" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="ED2BE7F5FFD1FFD37FC289C3FC0CFE39" box="[942,1005,450,476]" pageId="47" pageNumber="48">OPC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3202CBBFFD1FFD37CA889E6FDE3FDE7" box="[196,514,487,514]" pageId="47" pageNumber="48" type="distribution">
|
||
<paragraph id="8B857F30FFD1FFD37CA889E6FDE3FDE7" blockId="47.[151,1436,152,588]" box="[196,514,487,514]" pageId="47" pageNumber="48">
|
||
<emphasis id="B94EA322FFD1FFD37CA889E6FE87FDE4" bold="true" box="[196,358,487,513]" pageId="47" pageNumber="48">Distribution:</emphasis>
|
||
<collectingCountry id="F32D3FA0FFD1FFD37D0289E9FE1FFDE7" box="[366,510,488,514]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3202CBBFFD1FFD37CA88A0CFEDCFDA9" pageId="47" pageNumber="48" type="etymology">
|
||
<paragraph id="8B857F30FFD1FFD37CA88A0CFEDCFDA9" blockId="47.[151,1436,152,588]" pageId="47" pageNumber="48">
|
||
<emphasis id="B94EA322FFD1FFD37CA88A0CFEB5FDC2" bold="true" box="[196,340,525,551]" pageId="47" pageNumber="48">Etymology:</emphasis>
|
||
<emphasis id="B94EA322FFD1FFD37D308A0CFE7CFDC3" box="[348,413,525,550]" italics="true" pageId="47" pageNumber="48">Kilah</emphasis>
|
||
, road in Sanscrit, referring to Road
|
||
<quantity id="4CC2D2D5FFD1FFD37F288A0CFC6FFDC2" box="[836,910,525,551]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="47" pageNumber="48" unit="km" value="57.0">57 km</quantity>
|
||
from the city of Anosibe, where this species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |