199 lines
21 KiB
XML
199 lines
21 KiB
XML
<document id="F9A7D14D78B1C0524D9E8D675D4BF71B" ID-DOI="10.11646/zootaxa.2435.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5315673" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629671386604" checkinUser="felipe" docAuthor="Oláh, János & Johanson, Kjell Arne" docDate="2010" docId="0393CE26FFC7FFC67CFB89D4FBA4FC21" docLanguage="en" docName="zt02435p063.pdf" docOrigin="Zootaxa 2435 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2435.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Pahamunaya wamana Oláh & Johanson 2010, new species" docType="treatment" docVersion="5" lastPageNumber="59" masterDocId="FFAAB65EFFFEFFFC7C6C8801FFE1FFE5" masterDocTitle="Generic review of Polycentropodidae with description of 32 new species and 19 new species records from the Oriental, Australian and Afrotropical Biogeographical Regions 2435" masterLastPageNumber="63" masterPageNumber="1" pageNumber="58" updateTime="1699093793393" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E3BFBA892458E5222C98FC0DDD4E00CA">Generic review of Polycentropodidae with description of 32 new species and 19 new species records from the Oriental, Australian and Afrotropical Biogeographical Regions 2435</mods:title>
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<mods:namePart id="2AEB0D7A7BCEA41721A72B1890FF28B6">Oláh, János</mods:namePart>
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<mods:namePart id="F38A40831F27225E180C0EE5F064BF6A">Johanson, Kjell Arne</mods:namePart>
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<mods:title id="83B7A7D0BF41885DBAF2F6D7A1B56E10">Zootaxa</mods:title>
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<mods:part id="B195B0228520C7B5BFF493A43BBDEEC2">
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<mods:date id="FB9A50E775DADD02D9466C57578BF50F">2010</mods:date>
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<mods:number id="7016A9700D0FC113208C539C1D94C636">2010-04-23</mods:number>
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<mods:number id="6C1C1E0434BBCDAEDE4DA0EDE2B31FD0">2435</mods:number>
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<mods:url id="EC7B0381D5A5B3DA1174BFB0CEE79C0F">https://biotaxa.org/Zootaxa/article/view/zootaxa.2435.1.1</mods:url>
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<mods:classification id="53FBCBF3DA2C648F2765B147E1BB0F34">journal article</mods:classification>
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<mods:identifier id="1B2D765755EF0C2E6496AAAFBC3221D4" type="DOI">10.11646/zootaxa.2435.1.1</mods:identifier>
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<paragraph id="8B857F30FFC7FFC57CFB89D4FDA5FE0A" blockId="57.[151,580,469,530]" box="[151,580,469,495]" pageId="57" pageNumber="58">
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<heading id="D0CDC85CFFC7FFC57CFB89D4FDA5FE0A" bold="true" box="[151,580,469,495]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
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<taxonomicName id="4C3A04B3FFC7FFC57CFB89D4FE44FE0A" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[151,421,469,495]" class="Insecta" family="Polycentropodidae" genus="Pahamunaya" kingdom="Animalia" order="Trichoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="wamana" status="sp. nov.">
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<emphasis id="B94EA322FFC7FFC57CFB89D4FE44FE0A" bold="true" box="[151,421,469,495]" italics="true" pageId="57" pageNumber="58">Pahamunaya wamana</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27D1E59FFC7FFC57DD989D4FDA5FE0A" box="[437,580,469,495]" pageId="57" pageNumber="58" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C3202CBBFFC7FFC57CFB89F9FEBCFDF7" box="[151,349,504,530]" pageId="57" pageNumber="58" type="description">
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<paragraph id="8B857F30FFC7FFC57CFB89F9FEBCFDF7" blockId="57.[151,580,469,530]" box="[151,349,504,530]" pageId="57" pageNumber="58">
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<figureCitation id="130163B5FFC7FFC57CFB89F9FEBCFDF7" box="[151,349,504,530]" captionStart="FIGURES 106–108" captionStartId="57.[151,272,1886,1910]" captionTargetBox="[175,1407,790,1856]" captionTargetId="figure-131@57.[171,1415,773,1865]" captionTargetPageId="57" captionText="FIGURES 106–108. Pahamunaya wamana, new species, holotype. 106—genitalia, left lateral; 107—left gonopod, ventral; 108—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361570" httpUri="https://zenodo.org/record/5361570/files/figure.png" pageId="57" pageNumber="58">Figures 106–108</figureCitation>
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</paragraph>
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<subSubSection id="C3202CBBFFC7FFC57CFB8A43FD5EFD29" pageId="57" pageNumber="58" type="diagnosis">
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<paragraph id="8B857F30FFC7FFC57CFB8A43FD5EFD29" blockId="57.[151,1436,578,716]" pageId="57" pageNumber="58">
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<emphasis id="B94EA322FFC7FFC57CFB8A43FEF8FDB9" bold="true" box="[151,281,578,604]" pageId="57" pageNumber="58">Diagnosis:</emphasis>
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This species is similar to
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<taxonomicName id="4C3A04B3FFC7FFC57E2B8A43FCB8FDB9" authority="Malicky" authorityName="Malicky" box="[583,857,578,604]" class="Insecta" family="Polycentropodidae" genus="Pahamunaya" kingdom="Animalia" order="Trichoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nacesiveci">
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<emphasis id="B94EA322FFC7FFC57E2B8A43FD10FDBE" box="[583,753,578,603]" italics="true" pageId="57" pageNumber="58">Ph. nacesiveci</emphasis>
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Malicky
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</taxonomicName>
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from
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<collectingCountry id="F32D3FA0FFC7FFC57FCC8A43FBE4FDB9" box="[928,1029,578,604]" name="Malaysia" pageId="57" pageNumber="58">Sarawak</collectingCountry>
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but it is easily separated by details in the genitalia, particularly by the shorter sternite IX; the dorsal paraproctal processes that are each fused with its paraproctal base and its ventral paraproctal process; the smaller cerci; and the gonopods that each have a subapicodorsal mesal bidentate process.
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<caption id="DF452FB8FFC7FFC57CFB8F5FFD88F873" ID-DOI="http://doi.org/10.5281/zenodo.5361570" ID-Zenodo-Dep="5361570" httpUri="https://zenodo.org/record/5361570/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,272,1886,1910]" targetBox="[175,1407,790,1856]" targetPageId="57">
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<paragraph id="8B857F30FFC7FFC57CFB8F5FFD88F873" blockId="57.[151,1436,1886,1942]" pageId="57" pageNumber="58">
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<emphasis id="B94EA322FFC7FFC57CFB8F5FFE9EF893" bold="true" box="[151,383,1886,1910]" pageId="57" pageNumber="58">FIGURES 106–108.</emphasis>
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<taxonomicName id="4C3A04B3FFC7FFC57DEB8F5EFD9DF893" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[391,636,1887,1910]" class="Insecta" family="Polycentropodidae" genus="Pahamunaya" kingdom="Animalia" order="Trichoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="wamana" status="sp. nov.">
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<emphasis id="B94EA322FFC7FFC57DEB8F5EFD9DF893" box="[391,636,1887,1910]" italics="true" pageId="57" pageNumber="58">Pahamunaya wamana</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27D1E59FFC7FFC57EE78F5EFCF5F893" box="[651,788,1887,1910]" pageId="57" pageNumber="58" rank="species">new species</taxonomicNameLabel>
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, holotype. 106—genitalia, left lateral; 107—left gonopod, ventral; 108—phallic apparatus, left lateral.
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</paragraph>
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</caption>
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<subSubSection id="C3202CBBFFC4FFC67CA88896FB67FD07" pageId="58" pageNumber="59" type="description">
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<paragraph id="8B857F30FFC4FFC67CA88896FCC6FEC7" blockId="58.[151,1436,151,964]" pageId="58" pageNumber="59">
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<emphasis id="B94EA322FFC4FFC67CA88896FEBFFF54" bold="true" box="[196,350,151,177]" pageId="58" pageNumber="59">Description:</emphasis>
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Male. Body pale with almost whitish antennae and head; and pale brown legs and mesonotal and metanotal sclerites. Maxillary palp formula (I,II)-IV-III-V. Forewings each 4.0 mm long; with 3 isolated hyaline areas: (1) narrow line on R behind crossvein
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<emphasis id="B94EA322FFC4FFC67F6F88E3FCC3FF1E" box="[771,802,226,252]" italics="true" pageId="58" pageNumber="59">r -s</emphasis>
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; (2) small spot on crossvein
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<emphasis id="B94EA322FFC4FFC6781888E3FB7FFF1E" box="[1140,1182,226,252]" italics="true" pageId="58" pageNumber="59">s -m</emphasis>
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; and (3) long vertical line on crossvein
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<emphasis id="B94EA322FFC4FFC67D088909FE41FEC4" box="[356,416,264,290]" italics="true" pageId="58" pageNumber="59">m -cu</emphasis>
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and on 1st branching point of M.
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</paragraph>
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<paragraph id="8B857F30FFC4FFC67CA8892CFB67FD07" blockId="58.[151,1436,151,964]" pageId="58" pageNumber="59">
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Male genitalia. Segment IX forming quadrangular, short sternite IX with apical rim indicated by subapical constriction (
|
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<figureCitation id="130163B5FFC4FFC67D5A8953FE40FE89" box="[310,417,338,364]" captionStart="FIGURES 106–108" captionStartId="57.[151,272,1886,1910]" captionTargetBox="[175,1407,790,1856]" captionTargetId="figure-131@57.[171,1415,773,1865]" captionTargetPageId="57" captionText="FIGURES 106–108. Pahamunaya wamana, new species, holotype. 106—genitalia, left lateral; 107—left gonopod, ventral; 108—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361570" httpUri="https://zenodo.org/record/5361570/files/figure.png" pageId="58" pageNumber="59">Fig. 106</figureCitation>
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); constriction pronounced ventrally by presence of blunt posteroapical median keel; posterior margins of sternite IX above and below base of each gonopod turned mesad and fused into very short sternal tube surrounding basal part of each gonopod. Basal half of sternite IX on each side connected to cercus and paraproctal complex, possibly via rudimentary tergite IX. Segment X well-developed, weakly sclerotized, monolobed, broadly ligulate, as long as dorsal paraproctal plate; apex broad, with median excision; dorsoapical half setose. Cerci elongated, foliaceous, uniformly setose. Paraproctal complexes each including dorsal and ventral processes fused into mesal vertical plate with lateral horizontal carina; mesal vertical plate encircling phallic apparatus. Gonopods originating from sternal tubes (
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<figureCitation id="130163B5FFC4FFC6781B8A59FB3FFD97" box="[1143,1246,600,626]" captionStart="FIGURES 106–108" captionStartId="57.[151,272,1886,1910]" captionTargetBox="[175,1407,790,1856]" captionTargetId="figure-131@57.[171,1415,773,1865]" captionTargetPageId="57" captionText="FIGURES 106–108. Pahamunaya wamana, new species, holotype. 106—genitalia, left lateral; 107—left gonopod, ventral; 108—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361570" httpUri="https://zenodo.org/record/5361570/files/figure.png" pageId="58" pageNumber="59">Fig. 106</figureCitation>
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); in lateral view almost parallel-sided, slightly geniculate, turning dorsad on mid-length; each with bidentate subapicodorsal lobe pointing mesad (
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<figureCitation id="130163B5FFC4FFC67DF88AA3FE1AFD59" box="[404,507,674,700]" captionStart="FIGURES 106–108" captionStartId="57.[151,272,1886,1910]" captionTargetBox="[175,1407,790,1856]" captionTargetId="figure-131@57.[171,1415,773,1865]" captionTargetPageId="57" captionText="FIGURES 106–108. Pahamunaya wamana, new species, holotype. 106—genitalia, left lateral; 107—left gonopod, ventral; 108—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361570" httpUri="https://zenodo.org/record/5361570/files/figure.png" pageId="58" pageNumber="59">Fig. 107</figureCitation>
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). Phallic apparatus (
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<figureCitation id="130163B5FFC4FFC67E848AA3FCADFD59" box="[744,844,674,700]" captionStart="FIGURES 106–108" captionStartId="57.[151,272,1886,1910]" captionTargetBox="[175,1407,790,1856]" captionTargetId="figure-131@57.[171,1415,773,1865]" captionTargetPageId="57" captionText="FIGURES 106–108. Pahamunaya wamana, new species, holotype. 106—genitalia, left lateral; 107—left gonopod, ventral; 108—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361570" httpUri="https://zenodo.org/record/5361570/files/figure.png" pageId="58" pageNumber="59">Fig. 108</figureCitation>
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) constituting simple tube with slightly broadening phallobase and straight phallicata; apical membranes broadening and curving ventrad.
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</paragraph>
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<subSubSection id="C3202CBBFFC4FFC67CA88AECFEE6FCB1" pageId="58" pageNumber="59" type="materials_examined">
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<paragraph id="8B857F30FFC4FFC67CA88AECFEE6FCB1" blockId="58.[151,1436,151,964]" pageId="58" pageNumber="59">
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<materialsCitation id="3B52756DFFC4FFC67CA88AECFEE6FCB1" collectingDate="2007-12-09" collectingDateMax="2007-12-14" collectingDateMin="2007-12-09" collectionCode="NHMS" collectorName="B. Viklund & N. Jonsson" country="Malaysia" county="Tawau" elevation="994" latitude="4.733028" location="Nepenthes Camp" longLatPrecision="1" longitude="116.877975" municipality="Maliau Basin" pageId="58" pageNumber="59" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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<typeStatus id="5481C192FFC4FFC67CA88AECFEDFFCE2" box="[196,318,749,775]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
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male:
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:
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<collectingRegion id="49FEB1D2FFC4FFC67E3B8AECFD4EFCE2" box="[599,687,749,775]" country="Malaysia" name="Sabah" pageId="58" pageNumber="59">Sabah</collectingRegion>
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:
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</emphasis>
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<collectingCounty id="62E407BCFFC4FFC67ED38AECFCF4FCE2" box="[703,789,749,775]" pageId="58" pageNumber="59">Tawau</collectingCounty>
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,
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<collectingMunicipality id="6BE1E54AFFC4FFC67F488AECFC30FCE2" box="[804,977,749,775]" pageId="58" pageNumber="59">Maliau Basin</collectingMunicipality>
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,
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<location id="8EE529EBFFC4FFC67F8C8AECFB5DFCE2" LSID="urn:lsid:plazi:treatment:0393CE26FFC7FFC67CFB89D4FBA4FC21:8EE529EBFFC4FFC67F8C8AECFB5DFCE2" box="[992,1212,749,775]" country="Malaysia" county="Tawau" latitude="4.733028" longLatPrecision="1" longitude="116.877975" municipality="Maliau Basin" name="Nepenthes Camp" pageId="58" pageNumber="59" stateProvince="Sabah">Nepenthes Camp</location>
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, crossing stream,
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4
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43
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58.9
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<superScript id="7C4FD278FFC4FFC67D688B13FEEFFCC5" attach="right" box="[260,270,786,800]" fontSize="6" pageId="58" pageNumber="59">’’</superScript>
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N
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</geoCoordinate>
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,
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<geoCoordinate id="EE0E19F7FFC4FFC67D5F8B14FE3AFCCA" box="[307,475,786,815]" degrees="116" direction="east" minutes="52" orientation="longitude" pageId="58" pageNumber="59" precision="1" seconds="40.7" value="116.877975">
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116
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52
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40.7
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E
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,
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|
||
<elevation id="00179803FFC4FFC67D848B14FDD6FCCA" box="[488,567,789,815]" metricMagnitude="2" metricUnit="m" metricValue="9.94" pageId="58" pageNumber="59" unit="m" value="994.0">994 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF8459F0FFC4FFC67E2F8B14FCFBFCCA" box="[579,794,789,815]" pageId="58" pageNumber="59" value="2007-12-09" valueMax="2007-12-14" valueMin="2007-12-09">
|
||
<collectingDate id="EFC0A018FFC4FFC67E2F8B14FCFBFCCA" box="[579,794,789,815]" pageId="58" pageNumber="59" value="2007-12-09" valueMax="2007-12-14" valueMin="2007-12-09">9.xii.–14.xii.2007</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="527B0727FFC4FFC67F4B8B14FBF1FCCA" box="[807,1040,789,815]" pageId="58" pageNumber="59">Mega Malaise Trap</collectingMethod>
|
||
(
|
||
<quantity id="4CC2D2D5FFC4FFC6784D8B14FBA8FCCA" box="[1057,1097,789,815]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="58" pageNumber="59" unit="m" value="6.0">
|
||
<elevation id="00179803FFC4FFC6784D8B14FBA8FCCA" box="[1057,1097,789,815]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="58" pageNumber="59" unit="m" value="6.0">6m</elevation>
|
||
</quantity>
|
||
) [
|
||
<collectorName id="26CF1AE6FFC4FFC6780F8B14FB09FCCA" box="[1123,1256,789,815]" pageId="58" pageNumber="59">B. Viklund</collectorName>
|
||
&
|
||
<collectorName id="26CF1AE6FFC4FFC679638B14FA74FCCA" box="[1295,1429,789,815]" pageId="58" pageNumber="59">N. Jönsson</collectorName>
|
||
] (
|
||
<collectionCode id="ED2BE7F5FFC4FFC67CF38B3BFF1BFCB1" box="[159,250,826,852]" pageId="58" pageNumber="59">NHMS</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3202CBBFFC4FFC67CA88B61FDA2FC9F" box="[196,579,864,890]" pageId="58" pageNumber="59" type="distribution">
|
||
<paragraph id="8B857F30FFC4FFC67CA88B61FDA2FC9F" blockId="58.[151,1436,151,964]" box="[196,579,864,890]" pageId="58" pageNumber="59">
|
||
<emphasis id="B94EA322FFC4FFC67CA88B61FE87FC9F" bold="true" box="[196,358,864,890]" pageId="58" pageNumber="59">Distribution:</emphasis>
|
||
<collectingCountry id="F32D3FA0FFC4FFC67D028B61FE3BFC9F" box="[366,474,864,890]" name="Malaysia" pageId="58" pageNumber="59">Malaysia</collectingCountry>
|
||
(
|
||
<collectingRegion id="49FEB1D2FFC4FFC67D858B61FDD6FC9F" box="[489,567,864,890]" country="Malaysia" name="Sabah" pageId="58" pageNumber="59">Sabah</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3202CBBFFC4FFC67CA88B84FBA4FC21" pageId="58" pageNumber="59" type="etymology">
|
||
<paragraph id="8B857F30FFC4FFC67CA88B84FBA4FC21" blockId="58.[151,1436,151,964]" pageId="58" pageNumber="59">
|
||
<emphasis id="B94EA322FFC4FFC67CA88B84FEB7FC7A" bold="true" box="[196,342,901,927]" pageId="58" pageNumber="59">Etymology:</emphasis>
|
||
<emphasis id="B94EA322FFC4FFC67D338B87FE27FC7A" box="[351,454,902,927]" italics="true" pageId="58" pageNumber="59">Wamana</emphasis>
|
||
, “short”, in Sanscrit; the name refers to the short sternite IX, contrasting with the relatively long sternites of the 2 similar species,
|
||
<taxonomicName id="4C3A04B3FFC4FFC67EA28BAAFC8EFC21" authorityName="Oláh & Johanson" authorityYear="2010" box="[718,879,939,964]" class="Insecta" family="Polycentropodidae" genus="Pahamunaya" kingdom="Animalia" order="Trichoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="espelandae">
|
||
<emphasis id="B94EA322FFC4FFC67EA28BAAFC8EFC21" box="[718,879,939,964]" italics="true" pageId="58" pageNumber="59">P. espelandae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A04B3FFC4FFC67FCB8BAAFBDFFC21" box="[935,1086,939,964]" class="Insecta" family="Polycentropodidae" genus="Pahamunaya" kingdom="Animalia" order="Trichoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="nacesiveci">
|
||
<emphasis id="B94EA322FFC4FFC67FCB8BAAFBDFFC21" box="[935,1086,939,964]" italics="true" pageId="58" pageNumber="59">P. nacesiveci</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |