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<document id="0EF18AF02E9EE10A814C3E40DC58AED1" ID-DOI="10.1080/00222930500145123" ID-ISSN="1464-5262" ID-Zenodo-Dep="4581703" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614874722613" checkinUser="carolina" docAuthor="Huber, Bernhard A. &amp; Benjamin, Suresh P." docDate="2005" docId="039387D9FFCBFFF5C2D81C4DFC7D6D06" docLanguage="en" docName="JNATHIST.39.37-3305-3319.pdf" docOrigin="Journal of Natural History 39 (37)" docSource="http://dx.doi.org/10.1080/00222930500145123" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Wanniyala agrabopath Huber &amp; Benjamin 2005, new species" docType="treatment" docVersion="3" lastPageNumber="3317" masterDocId="FFAAFFA1FFC2FFFBC3731D01FFC96C16" masterDocTitle="The pholcid spiders from Sri Lanka: redescription of Pholcus ceylonicus and description of a new genus (Araneae: Pholcidae)" masterLastPageNumber="3319" masterPageNumber="3305" pageNumber="3312" updateTime="1698880134788" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="88C36875730B886085CE934638690E9C">The pholcid spiders from Sri Lanka: redescription of Pholcus ceylonicus and description of a new genus (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="1766A54D12DDA9814FDF789441E73CA4">Huber, Bernhard A.</mods:namePart>
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<mods:namePart id="76D653F61618A2F0F3B2BD33B1FB4D97">Benjamin, Suresh P.</mods:namePart>
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<mods:title id="FB2A6685C7D37EE983A681B79B7485BF">Journal of Natural History</mods:title>
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<mods:date id="6ADFA41BEF00C5377FBF9DF3E902F79D">2005</mods:date>
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<mods:number id="508B1FDBABFFE762972C4D62E0D5996F">2005-11-25</mods:number>
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<treatment id="039387D9FFCBFFF5C2D81C4DFC7D6D06" LSID="urn:lsid:plazi:treatment:039387D9FFCBFFF5C2D81C4DFC7D6D06" httpUri="http://treatment.plazi.org/id/039387D9FFCBFFF5C2D81C4DFC7D6D06" lastPageId="14" lastPageNumber="3317" pageId="9" pageNumber="3312">
<subSubSection id="C3206544FFCBFFF2C2D81C4DFCAF6D72" box="[427,870,332,356]" pageId="9" pageNumber="3312" type="nomenclature">
<paragraph id="8B8536CFFFCBFFF2C2D81C4DFCAF6D72" blockId="9.[427,870,332,388]" box="[427,870,332,356]" pageId="9" pageNumber="3312">
<heading id="D0CD81A3FFCBFFF2C2D81C4DFCAF6D72" box="[427,870,332,356]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="3312" reason="2">
<taxonomicName id="4C3A4D4CFFCBFFF2C2D81C4DFD1C6D72" ID-CoL="7FTSW" authority="Huber &amp; Benjamin, 2005" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[427,725,332,356]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="3312" phylum="Arthropoda" rank="species" species="agrabopath" status="sp. nov.">
<emphasis id="B94EEADDFFCBFFF2C2D81C4DFD1C6D72" bold="true" box="[427,725,332,356]" italics="true" pageId="9" pageNumber="3312">Wanniyala agrabopath</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57A6FFCBFFF2C1AD1C4DFCAF6D72" box="[734,870,332,356]" pageId="9" pageNumber="3312" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3206544FFCBFFF5C1701C6DFC7D6D06" lastPageId="14" lastPageNumber="3317" pageId="9" pageNumber="3312" type="description">
<paragraph id="8B8536CFFFCBFFF2C1701C6DFCC66D92" blockId="9.[427,870,332,388]" box="[515,783,364,388]" pageId="9" pageNumber="3312">
(
<figureCitation id="13012A4AFFCBFFF2C1791C6DFDBD6D92" box="[522,628,364,388]" captionStart="Figure 5" captionStartId="9.[125,185,1636,1655]" captionTargetBox="[125,1173,566,1600]" captionTargetId="figure-82@9.[125,1173,565,1601]" captionTargetPageId="9" captionText="Figure 5. (ae) Wanniyala agrabopath; (fi) W. hakgala. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="9" pageNumber="3312">Figures 5</figureCitation>
ae, 6, 7ad)
</paragraph>
<paragraph id="8B8536CFFFCBFFF2C30E1CA9FF7A6DA9" blockId="9.[125,179,424,447]" box="[125,179,424,447]" pageId="9" pageNumber="3312">
<emphasis id="B94EEADDFFCBFFF2C30E1CA9FF7A6DA9" box="[125,179,424,447]" italics="true" pageId="9" pageNumber="3312">Type</emphasis>
</paragraph>
<paragraph id="8B8536CFFFCBFFF2C30E1CD9FDBA6E07" blockId="9.[125,1172,472,529]" pageId="9" pageNumber="3312">
<materialsCitation id="3B523C92FFCBFFF2C30E1CD9FDA66E07" collectingDate="2000-03-07" collectionCode="MHNG" collectorName="S. P. Benjamin" country="Sri Lanka" county="Agrapatana" location="Male" municipality="Agrabopath Forest" pageId="9" pageNumber="3312" specimenCount="1" typeStatus="holotype">
<location id="8EE56014FFCBFFF2C30E1CD9FF716DE6" LSID="urn:lsid:plazi:treatment:039387D9FFCBFFF5C2D81C4DFC7D6D06:8EE56014FFCBFFF2C30E1CD9FF716DE6" box="[125,184,472,496]" country="Sri Lanka" county="Agrapatana" municipality="Agrabopath Forest" name="Male" pageId="9" pageNumber="3312">Male</location>
<typeStatus id="5481886DFFCBFFF2C3B51CD9FEE36DE6" box="[198,298,472,496]" pageId="9" pageNumber="3312" type="holotype">holotype</typeStatus>
from
<collectingMunicipality id="6BE1ACB5FFCBFFF2C20E1CD9FDA96DE6" box="[381,608,472,496]" pageId="9" pageNumber="3312">Agrabopath Forest</collectingMunicipality>
,
<collectingCounty id="62E44E43FFCBFFF2C1011CD9FD3F6DE6" box="[626,758,472,496]" pageId="9" pageNumber="3312">Agrapatana</collectingCounty>
(6
<emphasis id="B94EEADDFFCBFFF2C06E1CD9FCEE6DE6" bold="true" box="[797,807,472,496]" italics="true" pageId="9" pageNumber="3312">°</emphasis>
529N, 80
<emphasis id="B94EEADDFFCBFFF2C0E41CD9FC686DE6" bold="true" box="[919,929,472,496]" italics="true" pageId="9" pageNumber="3312">°</emphasis>
439E),
<collectingCountry id="F32D765FFFCBFFF2C08A1CD9FBBD6DE6" box="[1017,1140,472,496]" name="Sri Lanka" pageId="9" pageNumber="3312">Sri Lanka</collectingCountry>
,
<date id="FF84100FFFCBFFF2C7F61CD9FEC76E07" pageId="9" pageNumber="3312" value="2000-03-07">
<collectingDate id="EFC0E9E7FFCBFFF2C7F61CD9FEC76E07" pageId="9" pageNumber="3312" value="2000-03-07">7 March 2000</collectingDate>
</date>
(
<collectorName id="26CF5319FFCBFFF2C2511CF8FE116E07" box="[290,472,505,529]" pageId="9" pageNumber="3312">S. P. Benjamin</collectorName>
), in
<collectionCode id="ED2BAE0AFFCBFFF2C17E1CF8FDA66E07" box="[525,623,505,529]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="3312" type="Museum">MHNG</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="DF456647FFCBFFF2C30E1B65FC1F6A84" ID-DOI="http://doi.org/10.5281/zenodo.4581719" ID-Zenodo-Dep="4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="9" pageNumber="3312" startId="9.[125,185,1636,1655]" targetBox="[125,1173,566,1600]" targetPageId="9">
<paragraph id="8B8536CFFFCBFFF2C30E1B65FC1F6A84" blockId="9.[125,1172,1636,1682]" pageId="9" pageNumber="3312">
Figure 5. (ae)
<taxonomicName id="4C3A4D4CFFCBFFF2C2601B65FE156A61" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[275,476,1636,1655]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="3312" phylum="Arthropoda" rank="species" species="agrabopath">
<emphasis id="B94EEADDFFCBFFF2C2601B65FE156A61" box="[275,476,1636,1655]" italics="true" pageId="9" pageNumber="3312">Wanniyala agrabopath</emphasis>
</taxonomicName>
; (fi)
<taxonomicName id="4C3A4D4CFFCBFFF2C16B1B65FD496A61" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[536,640,1636,1655]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="3312" phylum="Arthropoda" rank="species" species="hakgala">
<emphasis id="B94EEADDFFCBFFF2C16B1B65FD496A61" box="[536,640,1636,1655]" italics="true" pageId="9" pageNumber="3312">W. hakgala</emphasis>
</taxonomicName>
. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views.
</paragraph>
</caption>
<caption id="DF456647FFC8FFF1C3F91860FCB869BF" ID-DOI="http://doi.org/10.5281/zenodo.4581723" ID-Zenodo-Dep="4581723" httpUri="https://zenodo.org/record/4581723/files/figure.png" pageId="10" pageNumber="3313" startId="10.[138,198,1377,1396]" targetBox="[148,1175,133,1327]" targetPageId="10">
<paragraph id="8B8536CFFFC8FFF1C3F91860FCB869BF" blockId="10.[138,1185,1377,1449]" pageId="10" pageNumber="3313">
Figure 6.
<taxonomicName id="4C3A4D4CFFC8FFF1C3981860FE7C6962" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[235,437,1377,1396]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="3313" phylum="Arthropoda" rank="species" species="agrabopath">
<emphasis id="B94EEADDFFC8FFF1C3981860FE7C6962" box="[235,437,1377,1396]" italics="true" pageId="10" pageNumber="3313">Wanniyala agrabopath</emphasis>
</taxonomicName>
. (a, b) Left male palp, prolateral and retrolateral views; (c) male prosoma and chelicerae, dorsal view; (d) tip of male cheliceral apophysis; (e) male chelicerae, frontal view. b, genital bulb; ba, bulbal apophysis; e, embolus. Scale bars: 0.5 mm (ac); 50 Mm (d); 0.2 mm (e).
</paragraph>
</caption>
<paragraph id="8B8536CFFFC8FFF1C3F91B4AFF326A75" blockId="10.[138,251,1611,1635]" box="[138,251,1611,1635]" pageId="10" pageNumber="3313">
<emphasis id="B94EEADDFFC8FFF1C3F91B4AFF326A75" box="[138,251,1611,1635]" italics="true" pageId="10" pageNumber="3313">Etymology</emphasis>
</paragraph>
<paragraph id="8B8536CFFFC8FFF1C3F91B7DFC1B6A82" blockId="10.[138,978,1660,1684]" box="[138,978,1660,1684]" pageId="10" pageNumber="3313">
The specific name is a noun in apposition, taken from the
<typeStatus id="5481886DFFC8FFF1C04E1B7DFCA76A82" box="[829,878,1660,1684]" pageId="10" pageNumber="3313">type</typeStatus>
locality.
</paragraph>
<caption id="DF456647FFC9FFF0C30E19E4FC0C693B" ID-DOI="http://doi.org/10.5281/zenodo.4581725" ID-Zenodo-Dep="4581725" httpUri="https://zenodo.org/record/4581725/files/figure.png" pageId="11" pageNumber="3314" startId="11.[125,185,1253,1272]" targetBox="[133,1168,138,1203]" targetPageId="11">
<paragraph id="8B8536CFFFC9FFF0C30E19E4FC0C693B" blockId="11.[125,1172,1253,1325]" pageId="11" pageNumber="3314">
Figure 7. (ad)
<taxonomicName id="4C3A4D4CFFC9FFF0C26B19E4FE2A68EE" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[280,483,1253,1272]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="3314" phylum="Arthropoda" rank="species" species="agrabopath">
<emphasis id="B94EEADDFFC9FFF0C26B19E4FE2A68EE" box="[280,483,1253,1272]" italics="true" pageId="11" pageNumber="3314">Wanniyala agrabopath</emphasis>
</taxonomicName>
; (eh)
<taxonomicName id="4C3A4D4CFFC9FFF0C15919E4FD5D68EE" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[554,660,1253,1272]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="3314" phylum="Arthropoda" rank="species" species="hakgala">
<emphasis id="B94EEADDFFC9FFF0C15919E4FD5D68EE" box="[554,660,1253,1272]" italics="true" pageId="11" pageNumber="3314">W. hakgala</emphasis>
</taxonomicName>
. (a, b, e, f) Left procursus, prolateral and retrolateral views; (c, g) cleared epigyna, ventral views; (d, h) cleared epigyna, dorsal views. ds, dorsal sclerite; pp, proximal projection; t13, transparent projections. Scale bars: 0.3 mm (a, b, e, f: all to same scale).
</paragraph>
</caption>
<paragraph id="8B8536CFFFC9FFF0C30E1868FF2F6997" blockId="11.[125,230,1385,1409]" box="[125,230,1385,1409]" pageId="11" pageNumber="3314">
<emphasis id="B94EEADDFFC9FFF0C30E1868FF2F6997" box="[125,230,1385,1409]" italics="true" pageId="11" pageNumber="3314">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B8536CFFFC9FFF0C30E189BFEED6A05" blockId="11.[125,1172,1434,1555]" pageId="11" pageNumber="3314">
Closely related to
<taxonomicName id="4C3A4D4CFFC9FFF0C222189AFE1A69A4" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[337,467,1434,1458]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="3314" phylum="Arthropoda" rank="species" species="hakgala">
<emphasis id="B94EEADDFFC9FFF0C222189AFE1A69A4" box="[337,467,1434,1458]" italics="true" pageId="11" pageNumber="3314">W. hakgala</emphasis>
</taxonomicName>
; easily distinguished by shape of dorsal sclerite on procursus (ds in
<figureCitation id="13012A4AFFC9FFF0C3B918BAFEF269C5" box="[202,315,1467,1491]" captionStart="Figure 7" captionStartId="11.[125,185,1253,1272]" captionTargetBox="[133,1168,138,1203]" captionTargetId="figure-96@11.[125,1173,133,1217]" captionTargetPageId="11" captionText="Figure 7. (ad) Wanniyala agrabopath; (eh) W. hakgala. (a, b, e, f) Left procursus, prolateral and retrolateral views; (c, g) cleared epigyna, ventral views; (d, h) cleared epigyna, dorsal views. ds, dorsal sclerite; pp, proximal projection; t13, transparent projections. Scale bars: 0.3 mm (a, b, e, f: all to same scale)." figureDoi="http://doi.org/10.5281/zenodo.4581725" httpUri="https://zenodo.org/record/4581725/files/figure.png" pageId="11" pageNumber="3314">Figure 7b</figureCitation>
), by epigynum without pointed projection (cf.
<figureCitation id="13012A4AFFC9FFF0C01518BAFBE369C5" box="[870,1066,1467,1491]" captionStart="Figure 5" captionStartId="9.[125,185,1636,1655]" captionTargetBox="[125,1173,566,1600]" captionTargetId="figure-82@9.[125,1173,565,1601]" captionTargetPageId="9" captionText="Figure 5. (ae) Wanniyala agrabopath; (fi) W. hakgala. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="11" pageNumber="3314">Figure 5d and 5i</figureCitation>
); also by shorter clypeus projections, shorter cheliceral apophyses, and apparently by colour pattern (
<figureCitation id="13012A4AFFC9FFF0C3F618FAFEDF6A05" box="[133,278,1531,1555]" captionStart="Figure 5" captionStartId="9.[125,185,1636,1655]" captionTargetBox="[125,1173,566,1600]" captionTargetId="figure-82@9.[125,1173,565,1601]" captionTargetPageId="9" captionText="Figure 5. (ae) Wanniyala agrabopath; (fi) W. hakgala. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="11" pageNumber="3314">Figure 5a, b</figureCitation>
).
</paragraph>
<paragraph id="8B8536CFFFC9FFF6C30E1B3DFD4B689B" blockId="11.[125,1176,1594,1685]" lastBlockId="13.[125,1172,1108,1165]" lastPageId="13" lastPageNumber="3316" pageId="11" pageNumber="3314">
<emphasis id="B94EEADDFFC9FFF0C30E1B3DFE816A42" box="[125,328,1596,1620]" italics="true" pageId="11" pageNumber="3314">
Male (
<typeStatus id="5481886DFFC9FFF0C3AB1B3DFEF56A42" box="[216,316,1596,1620]" pageId="11" pageNumber="3314" type="holotype">holotype</typeStatus>
).
</emphasis>
Total length 1.9, carapace width 0.86. Leg 1: 15.7 (3.75+ 0.4+3.95+5.8+1.8), tibia 2: 2.25, tibia 3: 1.7, tibia 4: 2.3; tibia 1 L/d: 47. Habitus as in
<figureCitation id="13012A4AFFC9FFF0C30E1B7CFEC26A83" box="[125,267,1661,1685]" captionStart="Figure 5" captionStartId="9.[125,185,1636,1655]" captionTargetBox="[125,1173,566,1600]" captionTargetId="figure-82@9.[125,1173,565,1601]" captionTargetPageId="9" captionText="Figure 5. (ae) Wanniyala agrabopath; (fi) W. hakgala. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="11" pageNumber="3314">Figure 5a, b</figureCitation>
. Carapace ochre-yellow with wide blackish median band, without lateral dark margins, clypeus only slightly darkened; sternum pale ochre yellow. Legs pale ochre-yellow, dark rings on tibiae (subdistally). Opisthosoma grey with black pattern, ventrally without pattern. Ocular area slightly elevated, thoracic furrow very distinct frontally; distance PMEPME 70 Mm; diameter PME 100 Mm; distance PMEALE 30 Mm; AME absent. Clypeus with pair of short projections 35 Mm long and indistinct median hump. Sternum wider than long (0.65/0.50). Chelicerae as in
<figureCitation id="13012A4AFFCEFFF7C1C618BAFC8A69C5" box="[693,835,1467,1491]" captionStart="Figure 6" captionStartId="10.[138,198,1377,1396]" captionTargetBox="[148,1175,133,1327]" captionTargetId="figure-15@10.[138,1186,133,1341]" captionTargetPageId="10" captionText="Figure 6. Wanniyala agrabopath. (a, b) Left male palp, prolateral and retrolateral views; (c) male prosoma and chelicerae, dorsal view; (d) tip of male cheliceral apophysis; (e) male chelicerae, frontal view. b, genital bulb; ba, bulbal apophysis; e, embolus. Scale bars: 0.5 mm (ac); 50 Mm (d); 0.2 mm (e)." figureDoi="http://doi.org/10.5281/zenodo.4581723" httpUri="https://zenodo.org/record/4581723/files/figure.png" pageId="12" pageNumber="3315">Figure 6c, e</figureCitation>
, with pair of long apophyses provided with two modified hairs on each tip (
<figureCitation id="13012A4AFFCEFFF7C1BB18DAFCF569E5" box="[712,828,1499,1523]" captionStart="Figure 6" captionStartId="10.[138,198,1377,1396]" captionTargetBox="[148,1175,133,1327]" captionTargetId="figure-15@10.[138,1186,133,1341]" captionTargetPageId="10" captionText="Figure 6. Wanniyala agrabopath. (a, b) Left male palp, prolateral and retrolateral views; (c) male prosoma and chelicerae, dorsal view; (d) tip of male cheliceral apophysis; (e) male chelicerae, frontal view. b, genital bulb; ba, bulbal apophysis; e, embolus. Scale bars: 0.5 mm (ac); 50 Mm (d); 0.2 mm (e)." figureDoi="http://doi.org/10.5281/zenodo.4581723" httpUri="https://zenodo.org/record/4581723/files/figure.png" pageId="12" pageNumber="3315">Figure 6d</figureCitation>
), tips 0.52 apart. Palps as in
<figureCitation id="13012A4AFFCEFFF7C3F918FDFED36A02" box="[138,282,1532,1556]" captionStart="Figure 6" captionStartId="10.[138,198,1377,1396]" captionTargetBox="[148,1175,133,1327]" captionTargetId="figure-15@10.[138,1186,133,1341]" captionTargetPageId="10" captionText="Figure 6. Wanniyala agrabopath. (a, b) Left male palp, prolateral and retrolateral views; (c) male prosoma and chelicerae, dorsal view; (d) tip of male cheliceral apophysis; (e) male chelicerae, frontal view. b, genital bulb; ba, bulbal apophysis; e, embolus. Scale bars: 0.5 mm (ac); 50 Mm (d); 0.2 mm (e)." figureDoi="http://doi.org/10.5281/zenodo.4581723" httpUri="https://zenodo.org/record/4581723/files/figure.png" pageId="12" pageNumber="3315">Figure 6a, b</figureCitation>
; trochanter with several short projections, femur with small proximo-dorsal apophysis, procursus very complex distally, apparently with two hinged processes, with three transparent processes (t
<quantity id="4CC29B2AFFCEFFF7C2961B3DFDF96A42" box="[485,560,1596,1620]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="12" pageNumber="3315" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">13 in</quantity>
<figureCitation id="13012A4AFFCEFFF7C1491B3DFD606A42" box="[570,681,1596,1620]" captionStart="Figure 7" captionStartId="11.[125,185,1253,1272]" captionTargetBox="[133,1168,138,1203]" captionTargetId="figure-96@11.[125,1173,133,1217]" captionTargetPageId="11" captionText="Figure 7. (ad) Wanniyala agrabopath; (eh) W. hakgala. (a, b, e, f) Left procursus, prolateral and retrolateral views; (c, g) cleared epigyna, ventral views; (d, h) cleared epigyna, dorsal views. ds, dorsal sclerite; pp, proximal projection; t13, transparent projections. Scale bars: 0.3 mm (a, b, e, f: all to same scale)." figureDoi="http://doi.org/10.5281/zenodo.4581725" httpUri="https://zenodo.org/record/4581725/files/figure.png" pageId="12" pageNumber="3315">Figure 7a</figureCitation>
); bulb with distinctive hooked sclerite and membranous embolus (
<figureCitation id="13012A4AFFCEFFF7C2D71B5CFDDF6A63" box="[420,534,1629,1653]" captionStart="Figure 6" captionStartId="10.[138,198,1377,1396]" captionTargetBox="[148,1175,133,1327]" captionTargetId="figure-15@10.[138,1186,133,1341]" captionTargetPageId="10" captionText="Figure 6. Wanniyala agrabopath. (a, b) Left male palp, prolateral and retrolateral views; (c) male prosoma and chelicerae, dorsal view; (d) tip of male cheliceral apophysis; (e) male chelicerae, frontal view. b, genital bulb; ba, bulbal apophysis; e, embolus. Scale bars: 0.5 mm (ac); 50 Mm (d); 0.2 mm (e)." figureDoi="http://doi.org/10.5281/zenodo.4581723" httpUri="https://zenodo.org/record/4581723/files/figure.png" pageId="12" pageNumber="3315">Figure 6a</figureCitation>
). Retrolateral trichobothrium of tibia 1 at 12%; most hairs on legs missing, but apparently without spines, without curved hairs, with several rows of vertical hairs on all metatarsi (clearly present in other male seen); tarsus 1 with.20 pseudosegments, only distally quite distinct.
</paragraph>
<caption id="DF456647FFCEFFF7C3F91943FF33689C" ID-DOI="http://doi.org/10.5281/zenodo.4581727" ID-Zenodo-Dep="4581727" httpUri="https://zenodo.org/record/4581727/files/figure.png" pageId="12" pageNumber="3315" startId="12.[138,198,1090,1109]" targetBox="[138,1186,133,1054]" targetPageId="12">
<paragraph id="8B8536CFFFCEFFF7C3F91943FF33689C" blockId="12.[138,1186,1090,1162]" pageId="12" pageNumber="3315">
Figure 8.
<taxonomicName id="4C3A4D4CFFCEFFF7C3981943FE536843" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[235,410,1090,1109]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="3315" phylum="Arthropoda" rank="species" species="hakgala">
<emphasis id="B94EEADDFFCEFFF7C3981943FE536843" box="[235,410,1090,1109]" italics="true" pageId="12" pageNumber="3315">Wanniyala hakgala</emphasis>
</taxonomicName>
. (a) Male gonopore with four epiandrous spigots; (b, c) female and male anterior lateral spinnerets with widened and pointed spigots; (d) female abdomen, ventral view. Scale bars: 20 Mm (ac); 0.5 mm (d).
</paragraph>
</caption>
<caption id="DF456647FFCFFFF6C30E1EACFE746FCD" ID-DOI="http://doi.org/10.5281/zenodo.4581729" ID-Zenodo-Dep="4581729" httpUri="https://zenodo.org/record/4581729/files/figure.png" pageId="13" pageNumber="3316" startId="13.[125,185,941,960]" targetBox="[365,932,133,906]" targetPageId="13">
<paragraph id="8B8536CFFFCFFFF6C30E1EACFE746FCD" blockId="13.[125,1172,941,987]" pageId="13" pageNumber="3316">
Figure 9. Known geographic distributions of
<taxonomicName id="4C3A4D4CFFCFFFF6C14C1EACFD296FD6" authorityName="O. Pickard-Cambridge" authorityYear="1869" box="[575,736,941,960]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="3316" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B94EEADDFFCFFFF6C14C1EACFD296FD6" box="[575,736,941,960]" italics="true" pageId="13" pageNumber="3316">Pholcus ceylonicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D4CFFCFFFF6C1811EACFC096FD6" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[754,960,941,960]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="3316" phylum="Arthropoda" rank="species" species="agrabopath">
<emphasis id="B94EEADDFFCFFFF6C1811EACFC096FD6" box="[754,960,941,960]" italics="true" pageId="13" pageNumber="3316">Wanniyala agrabopath</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D4CFFCFFFF6C0A21EACFBF76FD6" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[977,1086,941,960]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="3316" phylum="Arthropoda" rank="species" species="hakgala">
<emphasis id="B94EEADDFFCFFFF6C0A21EACFBF76FD6" box="[977,1086,941,960]" italics="true" pageId="13" pageNumber="3316">W. hakgala</emphasis>
</taxonomicName>
, and an undescribed species of
<taxonomicName id="4C3A4D4CFFCFFFF6C2261EC9FE7F6FCD" authorityName="Huber &amp; Benjamin" authorityYear="2005" box="[341,438,968,987]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="3316" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEADDFFCFFFF6C2261EC9FE7F6FCD" box="[341,438,968,987]" italics="true" pageId="13" pageNumber="3316">Wanniyala</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B8536CFFFCFFFF6C30E19B7FDC56939" blockId="13.[125,1172,1206,1327]" pageId="13" pageNumber="3316">
<emphasis id="B94EEADDFFCFFFF6C30E19B7FF2768D8" box="[125,238,1206,1230]" italics="true" pageId="13" pageNumber="3316">Variation.</emphasis>
Tibia
<quantity id="4CC29B2AFFCFFFF6C23219B7FEA568D8" box="[321,364,1206,1230]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="3316" unit="in" value="1.0">1 in</quantity>
male from Horton Plains: 3.2. This male differs also in colour pattern and is therefore assigned tentatively: lateral margins on carapace darkened, three pairs of black spots on sternum (cf.
<figureCitation id="13012A4AFFCFFFF6C2B219F6FDE66919" box="[449,559,1271,1295]" captionStart="Figure 5" captionStartId="9.[125,185,1636,1655]" captionTargetBox="[125,1173,566,1600]" captionTargetId="figure-82@9.[125,1173,565,1601]" captionTargetPageId="9" captionText="Figure 5. (ae) Wanniyala agrabopath; (fi) W. hakgala. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="13" pageNumber="3316">Figure 5e</figureCitation>
), with dark pattern on abdomen also ventrally, black rings also on femora (subdistally).
</paragraph>
<paragraph id="8B8536CFFFCFFFF6C30E1859FF3A6A04" blockId="13.[125,1172,1368,1554]" pageId="13" pageNumber="3316">
<emphasis id="B94EEADDFFCFFFF6C30E1859FF1C6966" box="[125,213,1368,1392]" italics="true" pageId="13" pageNumber="3316">Female.</emphasis>
In general similar to male, but clypeus unmodified and metatarsi without rows of vertical hairs; the female from Horton Plains has the same colour pattern as the male from this locality (see above). Tibia
<quantity id="4CC29B2AFFCFFFF6C1F21898FD7B69A7" box="[641,690,1433,1457]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="3316" unit="in" value="1.0">1 in</quantity>
female from
<typeStatus id="5481886DFFCFFFF6C0281898FC4569A7" box="[859,908,1433,1457]" pageId="13" pageNumber="3316">type</typeStatus>
locality: 2.85; female from Horton Plains: 2.55. Epigynum large but rather simple externally, without conical elevation (
<figureCitation id="13012A4AFFCFFFF6C38918DBFE9369E4" box="[250,346,1498,1522]" captionStart="Figure 5" captionStartId="9.[125,185,1636,1655]" captionTargetBox="[125,1173,566,1600]" captionTargetId="figure-82@9.[125,1173,565,1601]" captionTargetPageId="9" captionText="Figure 5. (ae) Wanniyala agrabopath; (fi) W. hakgala. (a, b, f, g) Male habitus, dorsal and lateral views; (c, h) female abdomen, ventral views; (e) female, ventral view; (d, i) female habitus, lateral views." figureDoi="http://doi.org/10.5281/zenodo.4581719" httpUri="https://zenodo.org/record/4581719/files/figure.png" pageId="13" pageNumber="3316">Figure 5</figureCitation>
ce), apparently with pair of pockets 0.65 apart. Dorsal view as in
<figureCitation id="13012A4AFFCFFFF6C30E18FBFF266A04" box="[125,239,1530,1554]" captionStart="Figure 7" captionStartId="11.[125,185,1253,1272]" captionTargetBox="[133,1168,138,1203]" captionTargetId="figure-96@11.[125,1173,133,1217]" captionTargetPageId="11" captionText="Figure 7. (ad) Wanniyala agrabopath; (eh) W. hakgala. (a, b, e, f) Left procursus, prolateral and retrolateral views; (c, g) cleared epigyna, ventral views; (d, h) cleared epigyna, dorsal views. ds, dorsal sclerite; pp, proximal projection; t13, transparent projections. Scale bars: 0.3 mm (a, b, e, f: all to same scale)." figureDoi="http://doi.org/10.5281/zenodo.4581725" httpUri="https://zenodo.org/record/4581725/files/figure.png" pageId="13" pageNumber="3316">Figure 7d</figureCitation>
.
</paragraph>
<paragraph id="8B8536CFFFCFFFF6C30E1B4AFF346A75" blockId="13.[125,253,1611,1635]" box="[125,253,1611,1635]" pageId="13" pageNumber="3316">
<emphasis id="B94EEADDFFCFFFF6C30E1B4AFF346A75" box="[125,253,1611,1635]" italics="true" pageId="13" pageNumber="3316">Distribution</emphasis>
</paragraph>
<paragraph id="8B8536CFFFCFFFF6C30E1B7DFD1A6A82" blockId="13.[125,723,1660,1684]" box="[125,723,1660,1684]" pageId="13" pageNumber="3316">
Known from two localities in
<collectingCountry id="F32D765FFFCFFFF6C2AE1B7DFD876A82" box="[477,590,1660,1684]" name="Sri Lanka" pageId="13" pageNumber="3316">Sri Lanka</collectingCountry>
(
<figureCitation id="13012A4AFFCFFFF6C1131B7DFD0D6A82" box="[608,708,1660,1684]" captionStart="Figure 9" captionStartId="13.[125,185,941,960]" captionTargetBox="[365,932,133,906]" captionTargetId="figure-152@13.[365,932,133,906]" captionTargetPageId="13" captionText="Figure 9. Known geographic distributions of Pholcus ceylonicus, Wanniyala agrabopath, W. hakgala, and an undescribed species of Wanniyala." figureDoi="http://doi.org/10.5281/zenodo.4581729" httpUri="https://zenodo.org/record/4581729/files/figure.png" pageId="13" pageNumber="3316">Figure 9</figureCitation>
).
</paragraph>
<paragraph id="8B8536CFFFCCFFF5C3F91D89FE916CB6" blockId="14.[138,344,136,160]" box="[138,344,136,160]" pageId="14" pageNumber="3317">
<emphasis id="B94EEADDFFCCFFF5C3F91D89FE916CB6" box="[138,344,136,160]" italics="true" pageId="14" pageNumber="3317">Material examined</emphasis>
</paragraph>
<paragraph id="8B8536CFFFCCFFF5C3F91DB9FC7D6D06" blockId="14.[138,1185,184,272]" pageId="14" pageNumber="3317">
<materialsCitation id="3B523C92FFCCFFF5C3F91DB9FC796D06" collectingDate="1981-08-13" collectingDateMax="1981-08-15" collectingDateMin="1981-08-13" collectionCode="R, RMNH" collectorName="C. L. Deeleman" country="Sri Lanka" elevation="2200" location="Agrapatana" municipality="Agrabopath Forest" pageId="14" pageNumber="3317" specimenCount="2" specimenCount-female="2">
<collectingCountry id="F32D765FFFCCFFF5C3F91DB9FECC6CC6" box="[138,261,184,208]" name="Sri Lanka" pageId="14" pageNumber="3317">Sri Lanka</collectingCountry>
:
<location id="8EE56014FFCCFFF5C2661DB9FE576CC6" LSID="urn:lsid:plazi:treatment:039387D9FFCBFFF5C2D81C4DFC7D6D06:8EE56014FFCCFFF5C2661DB9FE576CC6" box="[277,414,184,208]" country="Sri Lanka" municipality="Agrabopath Forest" name="Agrapatana" pageId="14" pageNumber="3317">Agrapatana</location>
:
<collectingMunicipality id="6BE1ACB5FFCCFFF5C2DC1DB9FD5B6CC6" box="[431,658,184,208]" pageId="14" pageNumber="3317">Agrabopath Forest</collectingMunicipality>
:
<typeStatus id="5481886DFFCCFFF5C1D11DB9FD1A6CC6" box="[674,723,184,208]" pageId="14" pageNumber="3317">type</typeStatus>
above, together with 1
<emphasis id="B94EEADDFFCCFFF5C0801DB8FBCC6CC6" box="[1011,1029,185,208]" italics="true" pageId="14" pageNumber="3317"></emphasis>
,
<specimenCount id="9D3CFD46FFCCFFF5C7691DB9FBFE6CC7" box="[1050,1079,184,209]" pageId="14" pageNumber="3317" type="female">1♀</specimenCount>
; Horton Plains (6
<emphasis id="B94EEADDFFCCFFF5C3821DD9FF326CE6" bold="true" box="[241,251,216,240]" italics="true" pageId="14" pageNumber="3317">°</emphasis>
489N, 80
<emphasis id="B94EEADDFFCCFFF5C2161DD9FEA66CE6" bold="true" box="[357,367,216,240]" italics="true" pageId="14" pageNumber="3317">°</emphasis>
489E),
<elevation id="0017D1FCFFCCFFF5C2B11DD9FD026CE6" box="[450,715,216,240]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="14" pageNumber="3317" unit="m" value="2200.0">
<quantity id="4CC29B2AFFCCFFF5C2B11DD9FDD16CE6" box="[450,536,216,240]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="14" pageNumber="3317" unit="m" value="2200.0">2200 m</quantity>
above sea level
</elevation>
(a.s.l.), forest litter,
<date id="FF84100FFFCCFFF5C0CF1DD9FB686CE6" box="[956,1185,216,240]" pageId="14" pageNumber="3317" value="1981-08-13" valueMax="1981-08-15" valueMin="1981-08-13">
<collectingDate id="EFC0E9E7FFCCFFF5C0CF1DD9FB686CE6" box="[956,1185,216,240]" pageId="14" pageNumber="3317" value="1981-08-13" valueMax="1981-08-15" valueMin="1981-08-13">1315 August 1981</collectingDate>
</date>
(P.
<collectionCode id="ED2BAE0AFFCCFFF5C3C51DF9FF046D06" box="[182,205,248,272]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="3317">R</collectionCode>
. and
<collectorName id="26CF5319FFCCFFF5C27D1DF9FE1B6D06" box="[270,466,248,272]" pageId="14" pageNumber="3317">C. L. Deeleman</collectorName>
), 1
<emphasis id="B94EEADDFFCCFFF5C2851DF8FDC16D06" box="[502,520,249,272]" italics="true" pageId="14" pageNumber="3317"></emphasis>
,
<specimenCount id="9D3CFD46FFCCFFF5C16B1DF9FDFC6D06" box="[536,565,247,272]" pageId="14" pageNumber="3317" type="female">1♀</specimenCount>
(and one epigynum!) (
<collectionCode id="ED2BAE0AFFCCFFF5C0371DF9FC6E6D06" box="[836,935,248,272]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="3317" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>