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<document id="2082304187B00BA2B79FF8432FA4F4DA" ID-CLB-Dataset="9255" ID-DOI="10.5252/zoosystema2018v40a26" ID-GBIF-Dataset="2697b081-f3fa-4256-a62d-f084ee352b1c" ID-ISSN="1638-9387" ID-Zenodo-Dep="4336858" ID-ZooBank="urn:lsid:zoobank.org:pub:E018714D-7CCF-4AB8-A88A-EF033530CA75" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1608156727094" checkinUser="felipe" docAuthor="Guinot, Danièle, Ng, Ngan Kee &amp; Rodríguez Moreno, Paula A." docDate="2018" docId="039387B2FF9C2624FF2E1AC5FAB1FDC9" docLanguage="en" docName="Zoosystema.40.26.547-604.pdf" docOrigin="Zoosystema 40 (26)" docStyle="DocumentStyle:AFAF64E709DD7C8340B70A24A5804402.4:Zoosystema.2018-2019.journal_article" docStyleId="AFAF64E709DD7C8340B70A24A5804402" docStyleName="Zoosystema.2018-2019.journal_article" docStyleVersion="4" docTitle="Cardisoma Latreille, in Latreille, Le Peletier, Serville &amp; Guerin 1828, sensu stricto" docType="treatment" docVersion="8" lastPageNumber="564" masterDocId="FFAAFFCAFF8F2637FFFB1A12FFECFFD9" masterDocTitle="Review of grapsoid families for the establishment of a new family for Leptograpsodes Montgomery, 1931, and a new genus of Gecarcinidae H. Milne Edwards, 1837 (Crustacea, Decapoda, Brachyura, Grapsoidea MacLeay, 1838)" masterLastPageNumber="604" masterPageNumber="547" pageNumber="564" updateTime="1698847703829" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="26C12371CC19545AAF06DBE2AF63BD9B">Review of grapsoid families for the establishment of a new family for Leptograpsodes Montgomery, 1931, and a new genus of Gecarcinidae H. Milne Edwards, 1837 (Crustacea, Decapoda, Brachyura, Grapsoidea MacLeay, 1838)</mods:title>
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<mods:namePart id="19BA6BDE2AFF013C2EC2389F19D035C7">Guinot, Danièle</mods:namePart>
<mods:affiliation id="FC258B450763370249B3ED83627E4032">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) guinot @ mnhn. fr (corresponding author)</mods:affiliation>
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<mods:namePart id="3C2BEB8F0D2E31BA6DF7FDA4697961B6">Ng, Ngan Kee</mods:namePart>
<mods:affiliation id="A1C64D0F8AD399B8FF3F3655F2C6EDDD">Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543 (Republic of Singapore) dbsngnk @ nus. edu. sg</mods:affiliation>
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<mods:namePart id="FEC18EADFAE3DF88AAB7B5B208881924">Rodríguez Moreno, Paula A.</mods:namePart>
<mods:affiliation id="A4CACC5AFD6DF5F54FA7258BAFE0C6CE">Direction générale déléguée aux Collections, Muséum national dHistoire naturelle, case postale 30, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) martin-lefevre @ mnhn. fr Dedicated to the memory of Michael Türkay (1948 - 2015)</mods:affiliation>
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<mods:title id="D889815933CA014315DF71EFB9844318">Zoosystema</mods:title>
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<mods:date id="5EDCF9765DA44C2E20CB21CD377EDED1">2018</mods:date>
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<mods:number id="6469A7085C15BE382A10B3DED929CEDC">40</mods:number>
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<subSubSection id="C320652FFF9C2624FF2E1AC5FD5AFF28" box="[213,694,215,241]" pageId="19" pageNumber="564" type="nomenclature">
<paragraph id="8B8536A4FF9C2624FF2E1AC5FD5AFF28" blockId="19.[213,694,215,241]" box="[213,694,215,241]" pageId="19" pageNumber="564">
<heading id="D0CD81C8FF9C2624FF2E1AC5FD5AFF28" box="[213,694,215,241]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="564" reason="2">
Genus
<taxonomicName id="4C3A4D27FF9C2624FED91AC5FDDAFF28" ID-CoL="7NRWM" authority="Latreille, 1828" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[290,566,215,241]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B94EEAB6FF9C2624FED91AC5FE7EFF28" box="[290,402,215,241]" italics="true" pageId="19" pageNumber="564">Cardisoma</emphasis>
Latreille, 1828
</taxonomicName>
<emphasis id="B94EEAB6FF9C2624FDC61ACAFD5AFF28" box="[573,694,216,241]" italics="true" pageId="19" pageNumber="564">
<taxonomicNameLabel id="A27D57CDFF9C2624FDC61ACAFD5AFF28" box="[573,694,216,241]" pageId="19" pageNumber="564" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320652FFF9C2624FF7F1B03FD47FE54" pageId="19" pageNumber="564" type="materials_examined">
<paragraph id="8B8536A4FF9C2624FF7F1B03FE21FE9A" blockId="19.[132,774,272,323]" pageId="19" pageNumber="564">
<typeStatus id="54818806FF9C2624FF7F1B03FF5AFEFF" box="[132,182,273,296]" pageId="19" pageNumber="564">TYPE</typeStatus>
SPECIES. —
<taxonomicName id="4C3A4D27FF9C2624FECC1B02FE26FE9A" authority="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin, 1828" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FECC1B02FDE5FEF1" box="[311,521,272,296]" italics="true" pageId="19" pageNumber="564">Cardisoma guanhumi</emphasis>
Latreille,
<emphasis id="B94EEAB6FF9C2624FD891B03FD6AFEF1" box="[626,646,273,296]" italics="true" pageId="19" pageNumber="564">in</emphasis>
Latreille, Le Peletier, Serville &amp; Guérin, 1828
</taxonomicName>
.
</paragraph>
<paragraph id="8B8536A4FF9C2624FF7F1B4EFD47FE54" blockId="19.[131,775,346,397]" pageId="19" pageNumber="564">
OTHER SPECIES. —
<taxonomicName id="4C3A4D27FF9C2624FEC41B48FD75FEAB" authority="Herklots, 1861" authorityName="Herklots" authorityYear="1861" box="[319,665,346,370]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="armatum">
<emphasis id="B94EEAB6FF9C2624FEC41B48FDECFEAB" box="[319,512,346,370]" italics="true" pageId="19" pageNumber="564">Cardisoma armatum</emphasis>
Herklots, 1861
</taxonomicName>
;
<taxonomicName id="4C3A4D27FF9C2624FD5A1B49FEFBFE54" authority="(Herbst, 1796)" baseAuthorityName="Herbst" baseAuthorityYear="1796" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FD5A1B49FCEBFEAB" box="[673,775,346,370]" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
(Herbst, 1796)
</taxonomicName>
;
<taxonomicName id="4C3A4D27FF9C2624FED91B67FDE4FE54" authority="Smith, 1870" authorityName="Smith" authorityYear="1870" box="[290,520,373,397]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="crassum">
<emphasis id="B94EEAB6FF9C2624FED91B67FE6BFE55" box="[290,391,373,396]" italics="true" pageId="19" pageNumber="564">C. crassum</emphasis>
Smith, 1870
</taxonomicName>
(not examined).
</paragraph>
</subSubSection>
<subSubSection id="C320652FFF9C2624FF7F1BA5FE80FD09" pageId="19" pageNumber="564" type="discussion">
<paragraph id="8B8536A4FF9C2624FF7F1BA5FF04FE16" blockId="19.[130,775,439,720]" box="[132,232,439,465]" pageId="19" pageNumber="564">REMARKS</paragraph>
<paragraph id="8B8536A4FF9C2624FF7F1BC5FE80FD09" blockId="19.[130,775,439,720]" pageId="19" pageNumber="564">
Two species traditionally assigned to
<taxonomicName id="4C3A4D27FF9C2624FDE11BC4FD66FE29" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[538,650,470,496]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAB6FF9C2624FDE11BC4FD66FE29" box="[538,650,470,496]" italics="true" pageId="19" pageNumber="564">Cardisoma</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D27FF9C2624FD621BC5FCEBFE29" authorityName="Guinot &amp; Ng &amp; Rodríguez Moreno" authorityYear="2018" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[665,775,470,497]" class="Malacostraca" family="Gecarcinidae" genus="Discoplax" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="hirtipes">
<emphasis id="B94EEAB6FF9C2624FD621BC5FCEBFE29" box="[665,775,470,497]" italics="true" pageId="19" pageNumber="564">C. hirtipes</emphasis>
</taxonomicName>
(see Appendix) and
<taxonomicName id="4C3A4D27FF9C2624FEA81BE5FE34FDC9" authorityName="Guinot &amp; Ng &amp; Rodríguez Moreno" authorityYear="2018" baseAuthorityName="Guinot &amp; Ng &amp; Rodríguez Moreno" baseAuthorityYear="2018" box="[339,472,502,529]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="rotundum">
<emphasis id="B94EEAB6FF9C2624FEA81BE5FE34FDC9" box="[339,472,502,529]" italics="true" pageId="19" pageNumber="564">C. rotundum</emphasis>
</taxonomicName>
, are hereby assigned to a new genus
<taxonomicName id="4C3A4D27FF9C2624FF3D1804FED3FDE9" authority="Guinot &amp; Ng &amp; Rodríguez Moreno, 2018" authorityName="Guinot &amp; Ng &amp; Rodríguez Moreno" authorityYear="2018" box="[198,319,534,560]" class="Malacostraca" family="Gecarcinidae" genus="Tuerkayana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94EEAB6FF9C2624FF3D1804FED3FDE9" box="[198,319,534,560]" italics="true" pageId="19" pageNumber="564">Tuerkayana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57CDFF9C2624FEBF1805FE62FDE8" box="[324,398,535,561]" pageId="19" pageNumber="564" rank="genus">n. gen.</taxonomicNameLabel>
, to which are added
<taxonomicName id="4C3A4D27FF9C2624FD981804FCEBFDE9" authorityName="Guinot &amp; Ng &amp; Rodríguez Moreno" authorityYear="2018" baseAuthorityName="Ng &amp; Davie" baseAuthorityYear="2012" box="[611,775,534,560]" class="Malacostraca" family="Gecarcinidae" genus="Discoplax" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="celeste">
<emphasis id="B94EEAB6FF9C2624FD981804FCEBFDE9" box="[611,775,534,560]" italics="true" pageId="19" pageNumber="564">Discoplax celeste</emphasis>
</taxonomicName>
of
<bibRefCitation id="EFAB4B55FF9C2624FF5A1825FE9BFD88" author="NG P. K. L. &amp; DAVIE P. J. F." box="[161,375,566,593]" pageId="19" pageNumber="564" pagination="89 - 100" refId="ref59312" refString="NG P. K. L. &amp; DAVIE P. J. F. 2012. - The blue crab of Christmas Island, Discoplax celeste, new species (Crustacea: Decapoda: Brachyura: Gecarcinidae). Raffles Bulletin of Zoology 60 (1): 89 - 100." type="journal article" year="2012">Ng &amp; Davie (2012)</bibRefCitation>
and
<taxonomicName id="4C3A4D27FF9C2624FE571825FDF9FD89" authorityName="P.K.L.Ng &amp; Shih" authorityYear="2014" box="[428,533,567,592]" class="Malacostraca" family="Gecarcinidae" genus="Discoplax" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="B94EEAB6FF9C2624FE571825FDF9FD89" box="[428,533,567,592]" italics="true" pageId="19" pageNumber="564">D. magna</emphasis>
</taxonomicName>
of
<bibRefCitation id="EFAB4B55FF9C2624FDCC1825FCEEFD88" author="NG P. K. L. &amp; SHIH H. - T." box="[567,770,566,593]" pageId="19" pageNumber="564" pagination="109 - 135" refId="ref59569" refString="NG P. K. L. &amp; SHIH H. - T. 2014. - The systematics of the land crabs of the Discoplax hirtipes (Dana, 1851) species-group (Crustacea: Decapoda: Brachyura: Gecarcinidae), with description of a new species from the eastern Indian Ocean. Raffles Bulletin of Zoology Supplement 30: 109 - 135." type="journal article" year="2014">Ng &amp; Shih (2014)</bibRefCitation>
. The more significant features that characterise
<emphasis id="B94EEAB6FF9C2624FD6C1844FF12FD49" italics="true" pageId="19" pageNumber="564">
<taxonomicName id="4C3A4D27FF9C2624FD6C1844FF54FD49" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="sensu">
Cardisoma
<taxonomicNameLabel id="A27D57CDFF9C2624FF7F1865FF54FD49" box="[132,184,631,656]" pageId="19" pageNumber="564" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
are the traditionally used ones that identify the
<typeStatus id="54818806FF9C2624FF7F1884FF5EFD69" box="[132,178,662,688]" pageId="19" pageNumber="564">type</typeStatus>
species
<taxonomicName id="4C3A4D27FF9C2624FEFC1884FE7CFD69" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[263,400,662,688]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FEFC1884FE7CFD69" box="[263,400,662,688]" italics="true" pageId="19" pageNumber="564">C. guanhumi</emphasis>
</taxonomicName>
and, in addition, features based on the thoracic sternum.
</paragraph>
</subSubSection>
<subSubSection id="C320652FFF9C2624FF7F18E4FAB1FDC9" pageId="19" pageNumber="564" type="diagnosis">
<paragraph id="8B8536A4FF9C2624FF7F18E4FF16FCD7" blockId="19.[130,777,758,2028]" box="[132,250,758,784]" pageId="19" pageNumber="564">DIAGNOSIS</paragraph>
<paragraph id="8B8536A4FF9C2624FF7F1907FAB1FDC9" blockId="19.[130,777,758,2028]" lastBlockId="19.[811,1456,215,529]" pageId="19" pageNumber="564">
Carapace inflated and thick (
<figureCitation id="13012A21FF9C2624FE4F1907FE1DFCF6" box="[436,497,789,815]" captionStart="FIG" captionStartId="20.[132,143,1847,1865]" captionTargetBox="[132,1455,212,1794]" captionTargetId="figure-250@20.[703,1280,727,1299]" captionTargetPageId="20" captionText="FIG. 4. — Carapaces of Gecarcinidae:Cardisoma Latreille,1828 (A-C), Discoplax A. Milne-Edwards,1867 (D, E) and Tuerkayana n. gen. (F-I): A, Cardisoma guanhumi Latreille, in Latreille,Le Peletier, Serville &amp; Guérin,1828, ♂ 56.5× 68.0 mm, Antilles,MNHN-IU-2013-14983 (=MNHN-B12270);B, C. carnifex (Herbst, 1796), ♂ 71.2 ×88.0 mm, French Polynesia, Society Islands, MNHN-IU-2013-14798; C, C. armatum Herklots, 1861, ♂ 51.6× 65.2 mm, Benin, MNHN-IU-2017-11710 (=MNHN-B21986);D, Discoplax longipes A.Milne-Edwards,1867,♂ 32.4× 37.2 mm,Loyalty Islands,Lifou,Inegoj Cave,MNHN-IU-2008-11402 (= MNHN-B24815); E, D. gracilipes Ng &amp; Guinot,2001, paratype,♂ 29.6 ×26.3 mm, Panglao Island,Bohol,Tawala Cave,MNHN-IU-2014-11217 (= MNHN-B27770);F, Tuerkayana aff. hirtipes (Dana,1851), ♂ 61.0× 79.0 mm,Loyalty Islands,Lifou Island,We cave,MNHN-IU-2017-8397 (=MNHN-B24811);G, Tuerkayana rotundum (Quoy &amp; Gaimard, 1824) n. comb.,♀ 32.7 ×40.1 mm,Loyalty Islands,MNHN-IU-2013-3740 (=MNHN-B13141);H, T. celeste (Ng &amp; Davie,2012) n. comb., ♂ 44.4 ×52.2 mm,Australia, Christmas Island (ZRC 2012.0171); I, T. magnum (Ng &amp; Shih, 2014) n. comb., ♂ 49.0 ×58.7 mm, Indonesia, Java, Cilacap (ZRC 2017.0252). Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4336876" httpUri="https://zenodo.org/record/4336876/files/figure.png" pageId="19" pageNumber="564">Fig. 4</figureCitation>
A-C:
<taxonomicName id="4C3A4D27FF9C2624FDD01904FD58FCF6" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[555,692,789,816]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FDD01904FDD1FCE9" box="[555,573,790,816]" italics="true" pageId="19" pageNumber="564">C</emphasis>
.
<emphasis id="B94EEAB6FF9C2624FDB11907FD58FCF6" box="[586,692,789,815]" italics="true" pageId="19" pageNumber="564">guanhumi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D27FF9C2624FD3B1904FF56FC96" baseAuthorityName="Herbst" baseAuthorityYear="1796" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FD3B1904FF56FC96" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D27FF9C2624FF371924FEBAFC96" authorityName="Herklots" authorityYear="1861" box="[204,342,822,848]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="armatum">
<emphasis id="B94EEAB6FF9C2624FF371924FEBAFC96" box="[204,342,822,848]" italics="true" pageId="19" pageNumber="564">C. armatum</emphasis>
</taxonomicName>
, respectively) (
<bibRefCitation id="EFAB4B55FF9C2624FDF21927FD13FC96" author="BOUCHARD J. - M. &amp; POUPIN J. &amp; CLEVA R. &amp; DUMAS J. &amp; DINHUT V." box="[521,767,821,847]" pageId="19" pageNumber="564" pagination="1 - 69" refId="ref48888" refString="BOUCHARD J. - M., POUPIN J., CLEVA R., DUMAS J. &amp; DINHUT V. 2013. - Land, mangrove and freshwater decapod crustaceans of Mayotte region (Crustacea Decapoda). Atoll Research Bulletin 592: 1 - 69. https: // doi. org / 10.5479 / si. 00775630.592" type="journal article" year="2013">
Bouchard
<emphasis id="B94EEAB6FF9C2624FD791924FD57FC96" box="[642,699,821,847]" italics="true" pageId="19" pageNumber="564">et al.</emphasis>
2013
</bibRefCitation>
: fig. 12C,
<taxonomicName id="4C3A4D27FF9C2624FF091947FE86FCB6" baseAuthorityName="Herbst" baseAuthorityYear="1796" box="[242,362,853,879]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FF091947FE86FCB6" box="[242,362,853,879]" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
</taxonomicName>
); dorsal surface with regions weakly or hardly demarcated, smooth and without posterolateral striae; anterolateral margin not delimited, unarmed, except for one notch just behind the exorbital angle in
<taxonomicName id="4C3A4D27FF9C2624FD8519A7FCEAFC16" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[638,774,949,975]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FD8519A7FCEAFC16" box="[638,774,949,975]" italics="true" pageId="19" pageNumber="564">C. guanhumi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFAB4B55FF9C2624FF7619C6FEC7FC36" author="RATHBUN M. J." box="[141,299,980,1007]" pageId="19" pageNumber="564" pagination="1 - 161" refId="ref61063" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. United States National Museum Bulletin 97: xxii + 461 p., pls 1 - 161. https: // doi. org / 10.5479 / si. 03629236.97. i" type="book chapter" year="1918">Rathbun 1918</bibRefCitation>
: pl. 106,
<taxonomicName id="4C3A4D27FF9C2624FE6A19C7FDF6FC37" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[401,538,980,1007]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FE6A19C7FDF6FC37" box="[401,538,980,1007]" italics="true" pageId="19" pageNumber="564">C. guanhumi</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FDDD19C6FD5DFC36" author="TURKAY M." box="[550,689,980,1007]" pageId="19" pageNumber="564" pagination="333 - 354" refId="ref63882" refString="TURKAY M. 1970. - Die Gecarcinidae Amerikas. Mit einem Anhang uber Ucides Rathbun (Crustacea; Decapoda). Senckenbergiana biologica 51: 333 - 354." type="journal article" year="1970">Türkay 1970</bibRefCitation>
: fig. 8a,
<taxonomicName id="4C3A4D27FF9C2624FF7819E7FEE5FBD7" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[131,265,1012,1039]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FF7819E7FEE5FBD7" box="[131,265,1012,1039]" italics="true" pageId="19" pageNumber="564">C. guanhumi</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FEEE19E6FE70FBD6" author="TURKAY M." box="[277,412,1012,1039]" pageId="19" pageNumber="564" pagination="81 - 103" refId="ref63912" refString="TURKAY M. 1973. - Die Gecarcinidae Afrikas (Crustacea: Decapoda). Senckenbergiana Biologica 54 (1 - 3): 81 - 103." type="journal article" year="1973">Türkay 1973</bibRefCitation>
: fig. 7,
<taxonomicName id="4C3A4D27FF9C2624FE1C19E7FD88FBD7" authorityName="Herklots" authorityYear="1861" box="[487,612,1013,1039]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="armatum">
<emphasis id="B94EEAB6FF9C2624FE1C19E7FD88FBD7" box="[487,612,1013,1039]" italics="true" pageId="19" pageNumber="564">C. armatum</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FD9519E6FCECFBD6" author="TURKAY M." box="[622,768,1012,1039]" pageId="19" pageNumber="564" pagination="223 - 259" refId="ref63940" refString="TURKAY M. 1974 a. - Die Gecarcinidae Asiens und Ozeaniens (Crustacea: Decapoda). Senckenbergiana Biologcia 55 (4 - 6): 223 - 259." type="journal article" year="1974">Türkay 1974a</bibRefCitation>
: fig. 11,
<taxonomicName id="4C3A4D27FF9C2624FF2B1E07FED7FBF7" baseAuthorityName="Herbst" baseAuthorityYear="1796" box="[208,315,1044,1071]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FF2B1E07FED7FBF7" box="[208,315,1044,1071]" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FEBE1E07FE27FBF7" author="TAVARES M. D. S." box="[325,459,1044,1071]" pageId="19" pageNumber="564" refId="ref63319" refString="TAVARES M. D. S. 1989. - Sistematica filogenetica dos Gecarcinidae H. Milne Edwards (Crustacea, Brachyura). Unpublished master thesis, Universidade Federal do Rio de Janeiro, 163 p." type="book" year="1989">Tavares 1989</bibRefCitation>
: fig.3,
<taxonomicName id="4C3A4D27FF9C2624FDEF1E07FD7BFBF7" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[532,663,1044,1071]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FDEF1E07FD7BFBF7" box="[532,663,1044,1071]" italics="true" pageId="19" pageNumber="564">C. guanhumi</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FD591E06FF10FB97" author="BOUCHARD J. - M. &amp; POUPIN J. &amp; CLEVA R. &amp; DUMAS J. &amp; DINHUT V." pageId="19" pageNumber="564" pagination="1 - 69" refId="ref48888" refString="BOUCHARD J. - M., POUPIN J., CLEVA R., DUMAS J. &amp; DINHUT V. 2013. - Land, mangrove and freshwater decapod crustaceans of Mayotte region (Crustacea Decapoda). Atoll Research Bulletin 592: 1 - 69. https: // doi. org / 10.5479 / si. 00775630.592" type="journal article" year="2013">
Bouchard
<emphasis id="B94EEAB6FF9C2624FF7F1E27FF56FB97" box="[132,186,1076,1102]" italics="true" pageId="19" pageNumber="564">et al.</emphasis>
2013
</bibRefCitation>
: fig. 12C,
<taxonomicName id="4C3A4D27FF9C2624FE8C1E27FE00FB97" baseAuthorityName="Herbst" baseAuthorityYear="1796" box="[375,492,1076,1103]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FE8C1E27FE00FB97" box="[375,492,1076,1103]" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FE061E26FD4CFB97" author="RATHBUN M. J." box="[509,672,1076,1102]" pageId="19" pageNumber="564" pagination="1 - 161" refId="ref61063" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. United States National Museum Bulletin 97: xxii + 461 p., pls 1 - 161. https: // doi. org / 10.5479 / si. 03629236.97. i" type="book chapter" year="1918">Rathbun 1918</bibRefCitation>
: pl. 108,
<taxonomicName id="4C3A4D27FF9C2624FF7D1E47FF17FBB7" authorityName="Smith" authorityYear="1870" box="[134,251,1109,1135]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="crassum">
<emphasis id="B94EEAB6FF9C2624FF7D1E47FF17FBB7" box="[134,251,1109,1135]" italics="true" pageId="19" pageNumber="564">C. crassum</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FEF21E46FE14FBB6" author="DIEZ Y. L. &amp; CAPOTE A. J." box="[265,504,1108,1135]" pageId="19" pageNumber="564" pagination="1 - 22" refId="ref51848" refString="DIEZ Y. L. &amp; CAPOTE A. J. 2015. - List of marine crabs (Decapoda: Anomura and Brachyura) of shallow littoral of Santiago de Cuba, Cuba. Check List 11 (2) 1601: 1 - 22. https: // doi. org / 10.15560 / 11.2.1601" type="journal article" year="2015">Diez &amp; Capote 2015</bibRefCitation>
: fig. 7K,
<taxonomicName id="4C3A4D27FF9C2624FD9C1E47FD1BFBB7" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[615,759,1108,1135]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FD9C1E47FD1BFBB7" box="[615,759,1108,1135]" italics="true" pageId="19" pageNumber="564">C. guanhumi</emphasis>
</taxonomicName>
). Proepistome prominently dome-shaped (
<figureCitation id="13012A21FF9C2624FDCC1E66FD64FB56" box="[567,648,1140,1167]" captionStart="FIG" captionStartId="24.[813,823,1804,1822]" captionTargetBox="[822,1441,215,1751]" captionTargetId="figure-725@24.[782,1305,754,1300]" captionTargetPageId="24" captionText="FIG. 6. — Ventral view of Gecarcinidae to show front, proepistome, cephalic appendages, orbit and mxp3: A, Discoplax longipes A. Milne-Edwards, 1867, ♂ 32.4× 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);B, Cardisoma guanhumi Latreille in Latreille,Le Peletier,Serville &amp; Guérin,1828,♂ 56.5×68.0 mm,Antilles,MNHN-IU-2013-14983 (=MNHN- B12270); C, Tuerkayana aff. hirtipes (Dana, 1851) n. comb., ♂ 61.0× 79.0 mm, Loyalty Islands,Lifou Island,We cave,MNHN-IU-2017-8397 (=MNHN-B24811); D, T. celeste (Ng &amp; Davie, 2012) n. comb., ♂ 44.4× 52.2 mm, Australia, Christmas Island, ZRC 2012.0171; E, T. magnum (Ng &amp; Shih, 2014) n. comb., ♂ 49.0× 58.7 mm, Indonesia, Java, ZRC 2017.0252. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4336884" httpUri="https://zenodo.org/record/4336884/files/figure.png" pageId="19" pageNumber="564">Fig. 6B</figureCitation>
); suborbital margin entirely joining exorbital tooth; orbit closed laterally by right-angled margin; no suborbital crest (
<figureCitation id="13012A21FF9C2624FDA81EA6FD4DFB16" box="[595,673,1204,1231]" captionStart="FIG" captionStartId="24.[813,823,1804,1822]" captionTargetBox="[822,1441,215,1751]" captionTargetId="figure-725@24.[782,1305,754,1300]" captionTargetPageId="24" captionText="FIG. 6. — Ventral view of Gecarcinidae to show front, proepistome, cephalic appendages, orbit and mxp3: A, Discoplax longipes A. Milne-Edwards, 1867, ♂ 32.4× 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);B, Cardisoma guanhumi Latreille in Latreille,Le Peletier,Serville &amp; Guérin,1828,♂ 56.5×68.0 mm,Antilles,MNHN-IU-2013-14983 (=MNHN- B12270); C, Tuerkayana aff. hirtipes (Dana, 1851) n. comb., ♂ 61.0× 79.0 mm, Loyalty Islands,Lifou Island,We cave,MNHN-IU-2017-8397 (=MNHN-B24811); D, T. celeste (Ng &amp; Davie, 2012) n. comb., ♂ 44.4× 52.2 mm, Australia, Christmas Island, ZRC 2012.0171; E, T. magnum (Ng &amp; Shih, 2014) n. comb., ♂ 49.0× 58.7 mm, Indonesia, Java, ZRC 2017.0252. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4336884" httpUri="https://zenodo.org/record/4336884/files/figure.png" pageId="19" pageNumber="564">Fig. 6B</figureCitation>
) (
<bibRefCitation id="EFAB4B55FF9C2624FD431EA6FF50FB37" author="TAVARES M. D. S." pageId="19" pageNumber="564" refId="ref63319" refString="TAVARES M. D. S. 1989. - Sistematica filogenetica dos Gecarcinidae H. Milne Edwards (Crustacea, Brachyura). Unpublished master thesis, Universidade Federal do Rio de Janeiro, 163 p." type="book" year="1989">Tavares 1989</bibRefCitation>
: fig. 3). Subhepatic and subbranchial areas not striated (
<figureCitation id="13012A21FF9C2624FF701EE1FF0DFAD7" box="[139,225,1267,1294]" captionStart="FIG" captionStartId="24.[813,823,1804,1822]" captionTargetBox="[822,1441,215,1751]" captionTargetId="figure-725@24.[782,1305,754,1300]" captionTargetPageId="24" captionText="FIG. 6. — Ventral view of Gecarcinidae to show front, proepistome, cephalic appendages, orbit and mxp3: A, Discoplax longipes A. Milne-Edwards, 1867, ♂ 32.4× 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815);B, Cardisoma guanhumi Latreille in Latreille,Le Peletier,Serville &amp; Guérin,1828,♂ 56.5×68.0 mm,Antilles,MNHN-IU-2013-14983 (=MNHN- B12270); C, Tuerkayana aff. hirtipes (Dana, 1851) n. comb., ♂ 61.0× 79.0 mm, Loyalty Islands,Lifou Island,We cave,MNHN-IU-2017-8397 (=MNHN-B24811); D, T. celeste (Ng &amp; Davie, 2012) n. comb., ♂ 44.4× 52.2 mm, Australia, Christmas Island, ZRC 2012.0171; E, T. magnum (Ng &amp; Shih, 2014) n. comb., ♂ 49.0× 58.7 mm, Indonesia, Java, ZRC 2017.0252. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4336884" httpUri="https://zenodo.org/record/4336884/files/figure.png" pageId="19" pageNumber="564">Fig. 6B</figureCitation>
). Setose pterygostomial area usually narrow. Male chelipeds with strong heterochely and heterodonty: major chela very stout; fingers markedly gaping, with one main tooth on both prehensile margins; minor chela narrow, long; fingers elongated, weakly gaping. Thoracic sternum (
<figureCitation id="13012A21FF9C2624FD511F61FD0AFA57" box="[682,742,1395,1422]" captionStart="FIG" captionStartId="22.[132,143,1894,1912]" captionTargetBox="[237,1384,214,1839]" captionTargetId="figure-233@22.[693,1154,664,1119]" captionTargetPageId="22" captionText="FIG. 5. — Pleon,thoracic sternum and genital region of Gecarcinidae.A, D, G, J, M, thoracic sternum (brushed) with pleon;B, E, H, K, N, thoracic sternum (brushed) without pleon; C, F, I, L, O, genital region with gonopore and penis: A-C, Discoplax longipes A. Milne-Edwards, 1867, ♂ 32.4× 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815); D-F, Cardisoma guanhumi Latreille in Latreille, Le Peletier, Serville &amp; Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN-IU-2013-14983 (=MNHN-B12270); G-I, Tuerkayana aff. hirtipes (Dana, 1851) n. comb., ♂ 61.0 ×79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (=MNHN-B24811); J-L, Tuerkayana celeste (Ng &amp; Davie, 2012) n. comb., ♂ 44.4 × 52.2 mm, Australia, Christmas Island, ZRC 2012.0171; M-O, T. magnum (Ng &amp; Shih, 2014) n. comb., ♂ 49.0× 58.7 mm, Indonesia, Java, ZRC 2017.0252. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4336880" httpUri="https://zenodo.org/record/4336880/files/figure.png" pageId="19" pageNumber="564">Fig. 5</figureCitation>
D- F) inclined posteriorly, proportionally narrow, especially at level of sternite 4 that is restricted between the P1; anterior sternites forming a proportionally developed plate; sternite 2 semi-ovate, bluntly triangular; no suture 3/4, without lateral trace; sternite 4 anteriorly narrow and long, with more or less concave margins (
<bibRefCitation id="EFAB4B55FF9C2624FEB91C20FE20F994" author="TURKAY M." box="[322,460,1586,1613]" pageId="19" pageNumber="564" pagination="333 - 354" refId="ref63882" refString="TURKAY M. 1970. - Die Gecarcinidae Amerikas. Mit einem Anhang uber Ucides Rathbun (Crustacea; Decapoda). Senckenbergiana biologica 51: 333 - 354." type="journal article" year="1970">Türkay 1970</bibRefCitation>
: fig. 8b,
<taxonomicName id="4C3A4D27FF9C2624FDDC1C21FD43F995" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[551,687,1586,1613]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FDDC1C21FD43F995" box="[551,687,1586,1613]" italics="true" pageId="19" pageNumber="564">C. guanhumi</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FD411C20FF50F9B5" author="TURKAY M." pageId="19" pageNumber="564" pagination="81 - 103" refId="ref63912" refString="TURKAY M. 1973. - Die Gecarcinidae Afrikas (Crustacea: Decapoda). Senckenbergiana Biologica 54 (1 - 3): 81 - 103." type="journal article" year="1973">Türkay 1973</bibRefCitation>
: fig. 8,
<taxonomicName id="4C3A4D27FF9C2624FEF11C41FE66F9B5" authorityName="Herklots" authorityYear="1861" box="[266,394,1619,1645]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="armatum">
<emphasis id="B94EEAB6FF9C2624FEF11C41FE66F9B5" box="[266,394,1619,1645]" italics="true" pageId="19" pageNumber="564">C. armatum</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FE6E1C40FDC6F9B5" author="TURKAY M." box="[405,554,1618,1644]" pageId="19" pageNumber="564" pagination="223 - 259" refId="ref63940" refString="TURKAY M. 1974 a. - Die Gecarcinidae Asiens und Ozeaniens (Crustacea: Decapoda). Senckenbergiana Biologcia 55 (4 - 6): 223 - 259." type="journal article" year="1974">Türkay 1974a</bibRefCitation>
: fig. 11,
<taxonomicName id="4C3A4D27FF9C2624FD7D1C41FD1BF9B5" baseAuthorityName="Herbst" baseAuthorityYear="1796" box="[646,759,1618,1645]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FD7D1C41FD1BF9B5" box="[646,759,1618,1645]" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
</taxonomicName>
); on sternite 4 a thick ridge forming the rim of the sterno-pleonal cavity at level of telson; a thick bridge at level of suture 6/7; sternite 8 rather large, developed medially; no visible portion when pleon is folded; suture 7/8 rather short. Median line on sternites 8, 7 and extending on the whole sternite 6, in front of transverse bridge (
<figureCitation id="13012A21FF9C2624FEA51D00FE2BF8F5" box="[350,455,1809,1836]" captionStart="FIG" captionStartId="22.[132,143,1894,1912]" captionTargetBox="[237,1384,214,1839]" captionTargetId="figure-233@22.[693,1154,664,1119]" captionTargetPageId="22" captionText="FIG. 5. — Pleon,thoracic sternum and genital region of Gecarcinidae.A, D, G, J, M, thoracic sternum (brushed) with pleon;B, E, H, K, N, thoracic sternum (brushed) without pleon; C, F, I, L, O, genital region with gonopore and penis: A-C, Discoplax longipes A. Milne-Edwards, 1867, ♂ 32.4× 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815); D-F, Cardisoma guanhumi Latreille in Latreille, Le Peletier, Serville &amp; Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN-IU-2013-14983 (=MNHN-B12270); G-I, Tuerkayana aff. hirtipes (Dana, 1851) n. comb., ♂ 61.0 ×79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (=MNHN-B24811); J-L, Tuerkayana celeste (Ng &amp; Davie, 2012) n. comb., ♂ 44.4 × 52.2 mm, Australia, Christmas Island, ZRC 2012.0171; M-O, T. magnum (Ng &amp; Shih, 2014) n. comb., ♂ 49.0× 58.7 mm, Indonesia, Java, ZRC 2017.0252. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4336880" httpUri="https://zenodo.org/record/4336880/files/figure.png" pageId="19" pageNumber="564">Fig. 5E, F</figureCitation>
) (
<bibRefCitation id="EFAB4B55FF9C2624FE261D00FD41F8F5" author="NG P. K. L. &amp; WANG C. - H. &amp; HO P. - H. &amp; SHIH H. - T." box="[477,685,1810,1836]" pageId="19" pageNumber="564" pagination="1 - 86" refId="ref59975" refString="NG P. K. L., WANG C. - H., HO P. - H. &amp; SHIH H. - T. 2001. - Anannotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series Taipei 11: 1 - 86." type="journal article" year="2001">Ng &amp; Guinot 2001</bibRefCitation>
: fig. 3A,
<taxonomicName id="4C3A4D27FF9C2624FF791D20FEE0F892" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[130,268,1841,1868]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FF791D20FEE0F892" box="[130,268,1841,1868]" italics="true" pageId="19" pageNumber="564">C. guanhumi</emphasis>
</taxonomicName>
; fig. 3B,
<taxonomicName id="4C3A4D27FF9C2624FE951D20FE33F895" baseAuthorityName="Herbst" baseAuthorityYear="1796" box="[366,479,1842,1868]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FE951D20FE33F895" box="[366,479,1842,1868]" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
</taxonomicName>
; fig. 3C,
<taxonomicName id="4C3A4D27FF9C2624FDB81D20FD28F892" authorityName="Herklots" authorityYear="1861" box="[579,708,1842,1868]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="armatum">
<emphasis id="B94EEAB6FF9C2624FDB81D20FD28F892" box="[579,708,1842,1868]" italics="true" pageId="19" pageNumber="564">C. armatum</emphasis>
</taxonomicName>
); correspondingly, a median septum at level of sternite 6, higher at level of sternite 7, visible after dissection of
<taxonomicName id="4C3A4D27FF9C2624FD701D60FCECF852" baseAuthorityName="Herbst" baseAuthorityYear="1796" box="[651,768,1905,1932]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FD701D60FCECF852" box="[651,768,1905,1932]" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
</taxonomicName>
. Sterno-pleonal cavity very wide. Pleon elongated, may reach sternite 3; somite 6 narrow and conspicuously elongated; telson (
<figureCitation id="13012A21FF9C2624FF711DC3FF0CF835" box="[138,224,2001,2028]" captionStart="FIG" captionStartId="22.[132,143,1894,1912]" captionTargetBox="[237,1384,214,1839]" captionTargetId="figure-233@22.[693,1154,664,1119]" captionTargetPageId="22" captionText="FIG. 5. — Pleon,thoracic sternum and genital region of Gecarcinidae.A, D, G, J, M, thoracic sternum (brushed) with pleon;B, E, H, K, N, thoracic sternum (brushed) without pleon; C, F, I, L, O, genital region with gonopore and penis: A-C, Discoplax longipes A. Milne-Edwards, 1867, ♂ 32.4× 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815); D-F, Cardisoma guanhumi Latreille in Latreille, Le Peletier, Serville &amp; Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN-IU-2013-14983 (=MNHN-B12270); G-I, Tuerkayana aff. hirtipes (Dana, 1851) n. comb., ♂ 61.0 ×79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (=MNHN-B24811); J-L, Tuerkayana celeste (Ng &amp; Davie, 2012) n. comb., ♂ 44.4 × 52.2 mm, Australia, Christmas Island, ZRC 2012.0171; M-O, T. magnum (Ng &amp; Shih, 2014) n. comb., ♂ 49.0× 58.7 mm, Indonesia, Java, ZRC 2017.0252. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4336880" httpUri="https://zenodo.org/record/4336880/files/figure.png" pageId="19" pageNumber="564">Fig. 5D</figureCitation>
) (
<bibRefCitation id="EFAB4B55FF9C2624FF001DC3FE6BF832" author="TURKAY M." box="[251,391,2001,2027]" pageId="19" pageNumber="564" pagination="333 - 354" refId="ref63882" refString="TURKAY M. 1970. - Die Gecarcinidae Amerikas. Mit einem Anhang uber Ucides Rathbun (Crustacea; Decapoda). Senckenbergiana biologica 51: 333 - 354." type="journal article" year="1970">Türkay 1970</bibRefCitation>
: fig. 8b,
<taxonomicName id="4C3A4D27FF9C2624FE121DC3FD9FF832" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[489,627,2001,2027]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FE121DC3FD9FF832" box="[489,627,2001,2027]" italics="true" pageId="19" pageNumber="564">C. guanhumi</emphasis>
</taxonomicName>
; 1973: fig. 8,
<taxonomicName id="4C3A4D27FF9C2624FCD61ACAFC43FF28" authorityName="Herklots" authorityYear="1861" box="[813,943,216,242]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="armatum">
<emphasis id="B94EEAB6FF9C2624FCD61ACAFC43FF28" box="[813,943,216,242]" italics="true" pageId="19" pageNumber="564">C. armatum</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FC451AC5FBB2FF28" author="RATHBUN M. J." box="[958,1118,215,241]" pageId="19" pageNumber="564" pagination="1 - 161" refId="ref61063" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. United States National Museum Bulletin 97: xxii + 461 p., pls 1 - 161. https: // doi. org / 10.5479 / si. 03629236.97. i" type="book chapter" year="1918">Rathbun 1918</bibRefCitation>
: fig. 157, pl. 109,
<taxonomicName id="4C3A4D27FF9C2624FAD51ACAFA4CFF28" authorityName="Smith" authorityYear="1870" box="[1326,1440,216,242]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="crassum">
<emphasis id="B94EEAB6FF9C2624FAD51ACAFA4CFF28" box="[1326,1440,216,242]" italics="true" pageId="19" pageNumber="564">C. crassum</emphasis>
</taxonomicName>
); G1 (
<bibRefCitation id="EFAB4B55FF9C2624FCA61AE5FC09FEC8" author="TURKAY M." box="[861,997,247,273]" pageId="19" pageNumber="564" pagination="333 - 354" refId="ref63882" refString="TURKAY M. 1970. - Die Gecarcinidae Amerikas. Mit einem Anhang uber Ucides Rathbun (Crustacea; Decapoda). Senckenbergiana biologica 51: 333 - 354." type="journal article" year="1970">Türkay 1970</bibRefCitation>
: fig. 8b,
<taxonomicName id="4C3A4D27FF9C2624FBBB1AEAFB2AFEC8" authorityName="Latreille, in Latreille, Le Peletier, Serville &amp; Guerin" authorityYear="1828" box="[1088,1222,247,274]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="guanhumi">
<emphasis id="B94EEAB6FF9C2624FBBB1AEAFB2AFEC8" box="[1088,1222,247,274]" italics="true" pageId="19" pageNumber="564">C. guanhumi</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFAB4B55FF9C2624FB291AE5FAB6FEC8" author="TURKAY M." box="[1234,1370,247,273]" pageId="19" pageNumber="564" pagination="81 - 103" refId="ref63912" refString="TURKAY M. 1973. - Die Gecarcinidae Afrikas (Crustacea: Decapoda). Senckenbergiana Biologica 54 (1 - 3): 81 - 103." type="journal article" year="1973">Türkay 1973</bibRefCitation>
: fig. 4b,
<taxonomicName id="4C3A4D27FF9C2624FCD01B05FC4DFEE8" baseAuthorityName="Herbst" baseAuthorityYear="1796" box="[811,929,279,305]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FCD01B05FC4DFEE8" box="[811,929,279,305]" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
</taxonomicName>
; 1974a: fig.11,
<taxonomicName id="4C3A4D27FF9C2624FBAC1B05FB22FEE8" baseAuthorityName="Herbst" baseAuthorityYear="1796" box="[1111,1230,279,305]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FBAC1B05FB22FEE8" box="[1111,1230,279,305]" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
</taxonomicName>
). Locking structure absent: no press-button on smooth sternite 5, covered with setae (
<figureCitation id="13012A21FF9C2624FC971B45FC56FEAB" box="[876,954,343,370]" captionStart="FIG" captionStartId="22.[132,143,1894,1912]" captionTargetBox="[237,1384,214,1839]" captionTargetId="figure-233@22.[693,1154,664,1119]" captionTargetPageId="22" captionText="FIG. 5. — Pleon,thoracic sternum and genital region of Gecarcinidae.A, D, G, J, M, thoracic sternum (brushed) with pleon;B, E, H, K, N, thoracic sternum (brushed) without pleon; C, F, I, L, O, genital region with gonopore and penis: A-C, Discoplax longipes A. Milne-Edwards, 1867, ♂ 32.4× 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815); D-F, Cardisoma guanhumi Latreille in Latreille, Le Peletier, Serville &amp; Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN-IU-2013-14983 (=MNHN-B12270); G-I, Tuerkayana aff. hirtipes (Dana, 1851) n. comb., ♂ 61.0 ×79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (=MNHN-B24811); J-L, Tuerkayana celeste (Ng &amp; Davie, 2012) n. comb., ♂ 44.4 × 52.2 mm, Australia, Christmas Island, ZRC 2012.0171; M-O, T. magnum (Ng &amp; Shih, 2014) n. comb., ♂ 49.0× 58.7 mm, Indonesia, Java, ZRC 2017.0252. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4336880" httpUri="https://zenodo.org/record/4336880/files/figure.png" pageId="19" pageNumber="564">Fig. 5E</figureCitation>
) (
<bibRefCitation id="EFAB4B55FF9C2624FC2A1B45FBB1FEA8" author="GUINOT D." box="[977,1117,343,369]" pageId="19" pageNumber="564" pagination="1 - 354" refId="ref53121" refString="GUINOT D. 1979. - Morphologie et phylogenese des Brachyoures. Museum national d'Histoire naturelle, Paris (Memoires du Museum national d'Histoire naturelle, Ser. A - Zoologie; 112): 1 - 354, pls 1 - 25." type="journal article" year="1979">Guinot 1979</bibRefCitation>
;
<bibRefCitation id="EFAB4B55FF9C2624FB931B45FA92FEA8" author="GUINOT D. &amp; BOUCHARD J. - M." box="[1128,1406,343,369]" pageId="19" pageNumber="564" pagination="613 - 694" refId="ref53418" refString="GUINOT D. &amp; BOUCHARD J. - M. 1998. - Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema 20 (4): 613 - 694." type="journal article" year="1998">Guinot &amp; Bouchard 1998</bibRefCitation>
); no socket on pleonal somite 6, only some remants of a pleonal socket may be discernible in
<taxonomicName id="4C3A4D27FF9C2624FB981B85FB39FE68" baseAuthorityName="Herbst" baseAuthorityYear="1796" box="[1123,1237,407,433]" class="Malacostraca" family="Gecarcinidae" genus="Cardisoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="564" phylum="Arthropoda" rank="species" species="carnifex">
<emphasis id="B94EEAB6FF9C2624FB981B85FB39FE68" box="[1123,1237,407,433]" italics="true" pageId="19" pageNumber="564">C. carnifex</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFAB4B55FF9C2624FB1E1B84FA44FE68" author="KOHNK S. &amp; GORB S. &amp; BRANDIS D." box="[1253,1448,406,433]" pageId="19" pageNumber="564" pagination="2087 - 2132" refId="ref56245" refString="KOHNK S., GORB S. &amp; BRANDIS D. 2017. - The morphological and functional variability of pleon-holding mechanisms in selected Eubrachyura (Crustacea: Decapoda). Journal of Natural History 51 (35 - 36): 2087 - 2132. https: // doi. org / 10.1080 / 00222933.2017.1355076" type="journal article" year="2017">
Köhnk
<emphasis id="B94EEAB6FF9C2624FACD1B85FA86FE69" box="[1334,1386,406,432]" italics="true" pageId="19" pageNumber="564">et al.</emphasis>
2017
</bibRefCitation>
: fig. 20a, b). Male gonopore close to P5 coxo-sternal condyle; penis proximally narrow, then more expanded (
<figureCitation id="13012A21FF9C2624FAE21BC4FA8AFE28" box="[1305,1382,470,497]" captionStart="FIG" captionStartId="22.[132,143,1894,1912]" captionTargetBox="[237,1384,214,1839]" captionTargetId="figure-233@22.[693,1154,664,1119]" captionTargetPageId="22" captionText="FIG. 5. — Pleon,thoracic sternum and genital region of Gecarcinidae.A, D, G, J, M, thoracic sternum (brushed) with pleon;B, E, H, K, N, thoracic sternum (brushed) without pleon; C, F, I, L, O, genital region with gonopore and penis: A-C, Discoplax longipes A. Milne-Edwards, 1867, ♂ 32.4× 37.2 mm, Loyalty Islands, Lifou, Inegoj Cave, MNHN-IU-2008-11402 (= MNHN-B24815); D-F, Cardisoma guanhumi Latreille in Latreille, Le Peletier, Serville &amp; Guérin, 1828, ♂ 56.5 × 68.0 mm, Antilles, MNHN-IU-2013-14983 (=MNHN-B12270); G-I, Tuerkayana aff. hirtipes (Dana, 1851) n. comb., ♂ 61.0 ×79.0 mm, Loyalty Islands, Lifou Island, We cave, MNHN-IU-2017-8397 (=MNHN-B24811); J-L, Tuerkayana celeste (Ng &amp; Davie, 2012) n. comb., ♂ 44.4 × 52.2 mm, Australia, Christmas Island, ZRC 2012.0171; M-O, T. magnum (Ng &amp; Shih, 2014) n. comb., ♂ 49.0× 58.7 mm, Indonesia, Java, ZRC 2017.0252. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4336880" httpUri="https://zenodo.org/record/4336880/files/figure.png" pageId="19" pageNumber="564">Fig. 5F</figureCitation>
) (
<bibRefCitation id="EFAB4B55FF9C2624FA871BC4FC7DFDC9" author="GUINOT D." pageId="19" pageNumber="564" pagination="1 - 354" refId="ref53121" refString="GUINOT D. 1979. - Morphologie et phylogenese des Brachyoures. Museum national d'Histoire naturelle, Paris (Memoires du Museum national d'Histoire naturelle, Ser. A - Zoologie; 112): 1 - 354, pls 1 - 25." type="journal article" year="1979">Guinot 1979</bibRefCitation>
: fig. 54A-C;
<bibRefCitation id="EFAB4B55FF9C2624FBE41BE4FB0BFDC9" author="GUINOT D. &amp; TAVARES M. &amp; CASTRO P." box="[1055,1255,502,528]" pageId="19" pageNumber="564" pagination="1 - 414" refId="ref53507" refString="GUINOT D., TAVARES M. &amp; CASTRO P. 2013. - Significance of sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zootaxa 3665 (1): 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
Guinot
<emphasis id="B94EEAB6FF9C2624FB8D1BE5FB45FDC9" box="[1142,1193,502,528]" italics="true" pageId="19" pageNumber="564">et al.</emphasis>
2013
</bibRefCitation>
: fig. 23A).
</paragraph>
</subSubSection>
</treatment>
</document>