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<document id="3849C937BCC9D635B9A1EF0D48264C86" ID-DOI="10.5281/zenodo.176487" ID-GBIF-Dataset="815f47b1-6767-416f-a0bf-f925b54845fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="176487" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459834238221" checkinUser="plazi" docAuthor="Shear, William A." docDate="2007" docId="0393879DFFF99621FF7C54DBFE3CC779" docLanguage="en" docName="zt01463p012.pdf" docOrigin="Zootaxa 1463" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Idagona jasperi Shear, 2007, n. sp." docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="FFAAFFE5FFFC9629FFEB514CFFCCC62B" masterDocTitle="Cave millipeds of the United States. V. The genus Idagona Buckett &amp; Gardner (Chordeumatida, Conotylidae, Idagoninae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="6" updateTime="1698223281701" updateUser="plazi">
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<mods:title id="FDCA1347737178EF1BFE1D862C6BAC08">Cave millipeds of the United States. V. The genus Idagona Buckett &amp; Gardner (Chordeumatida, Conotylidae, Idagoninae)</mods:title>
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<mods:namePart id="BE58B0F1B6D757CF44ECE94EF765CDAE">Shear, William A.</mods:namePart>
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<mods:date id="4EA39DCF5D38008D7C1DDC109592455F">2007</mods:date>
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<treatment id="0393879DFFF99621FF7C54DBFE3CC779" ID-DOI="http://doi.org/10.5281/zenodo.6236711" ID-GBIF-Taxon="119358477" ID-Zenodo-Dep="6236711" LSID="urn:lsid:plazi:treatment:0393879DFFF99621FF7C54DBFE3CC779" httpUri="http://treatment.plazi.org/id/0393879DFFF99621FF7C54DBFE3CC779" lastPageId="8" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="C3206500FFF9962CFF7C54DBFE7EC3FF" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8B85368BFFF9962CFF7C54DBFE69C39A" blockId="5.[151,434,1431,1492]" box="[151,421,1431,1457]" pageId="5" pageNumber="6">
<heading id="D0CD81E7FFF9962CFF7C54DBFE69C39A" bold="true" box="[151,421,1431,1457]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="B94EEA99FFF9962CFF7C54DBFE69C39A" bold="true" box="[151,421,1431,1457]" pageId="5" pageNumber="6">
<emphasis id="B94EEA99FFF9962CFF7C54DBFE90C39A" bold="true" box="[151,348,1431,1457]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4D08FFF9962CFF7C54DBFE9AC39A" ID-CoL="3PC3P" box="[151,342,1431,1457]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jasperi" status="sp. nov.">Idagona jasperi</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A27D57E2FFF9962CFE8854DBFE69C39A" box="[355,421,1431,1457]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B85368BFFF9962CFF7C54F6FE7EC3FF" blockId="5.[151,434,1431,1492]" box="[151,434,1466,1492]" pageId="5" pageNumber="6">
<figureCitation id="13012A0EFFF9962CFF7C54F6FF24C3FF" box="[151,232,1466,1492]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1923,1947]" captionTargetBox="[151,1435,729,1899]" captionTargetId="figure@3.[151,1436,729,1900]" captionTargetPageId="3" captionText="FIGURES 1 4. Gonopods of Idagona species. 1, Idagona westcotti, Giant Arch Cave, posteriorventral view. 2. I. westcotti, Ice Capades Cave, posterioventral view. 3. I. jasperi, n. sp. posterioventral view. 4. I. lehmanenesis, n. sp., posterior view." httpUri="https://zenodo.org/record/176488/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
, 7, 11, 17, 19, 20
</paragraph>
</subSubSection>
<subSubSection id="C3206500FFF9962CFF7C5747FB61C05F" pageId="5" pageNumber="6" type="description">
<paragraph id="8B85368BFFF9962CFF7C5747FB61C05F" blockId="5.[151,1436,1546,2012]" pageId="5" pageNumber="6">
<emphasis id="B94EEA99FFF9962CFF7C5747FF1BC00F" box="[151,215,1547,1572]" italics="true" pageId="5" pageNumber="6">
<typeStatus id="54818829FFF9962CFF7C5747FF1BC00F" box="[151,215,1547,1572]" pageId="5" pageNumber="6">Types</typeStatus>
</emphasis>
: Male
<typeStatus id="54818829FFF9962CFED85746FE50C00F" box="[307,412,1546,1572]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
,
<typeStatus id="54818829FFF9962CFE435746FDD4C00F" box="[424,536,1546,1572]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
from
<emphasis id="B94EEA99FFF9962CFDB45746FD6AC00F" bold="true" box="[607,678,1546,1572]" pageId="5" pageNumber="6">Utah:</emphasis>
<emphasis id="B94EEA99FFF9962CFD455747FCFDC00F" box="[686,817,1547,1572]" italics="true" pageId="5" pageNumber="6">Cache Co.,</emphasis>
Nielsens Cave, Tony Grove Recreation Area, 8100 (
<quantity id="4CC29B6EFFF9962CFF74577EFF36C067" box="[159,250,1586,1612]" metricMagnitude="3" metricUnit="m" metricValue="2.469" pageId="5" pageNumber="6" unit="m" value="2469.0">2469 m</quantity>
) elevation, Wasatch-Cache National Forest, J. Jasper,
<date id="FF84104BFFF9962CFC95577EFBDBC067" box="[894,1047,1586,1612]" pageId="5" pageNumber="6" value="2006-08">August 2006</date>
. Additional male
<typeStatus id="54818829FFF9962CFB1C577EFA90C067" box="[1271,1372,1586,1612]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
from same locality, but Main Drain Cave.
<date id="FF84104BFFF9962CFDAD5716FCD2C05F" box="[582,798,1626,1652]" pageId="5" pageNumber="6" value="2006-09-09">9 September 2006</date>
, G.
<collectingCountry id="F32D761BFFF9962CFCA15716FC5CC05F" box="[842,912,1626,1652]" name="United States Pacific Island Wildlife Refuges" pageId="5" pageNumber="6">Baker</collectingCountry>
(all specimens FMNH).
</paragraph>
</subSubSection>
<subSubSection id="C3206500FFF9962CFF2D57CFFB7CC13F" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8B85368BFFF9962CFF2D57CFFB7CC13F" blockId="5.[151,1436,1546,2012]" pageId="5" pageNumber="6">
<emphasis id="B94EEA99FFF9962CFF2D57CFFE8BC0B7" box="[198,327,1667,1692]" italics="true" pageId="5" pageNumber="6">Diagnosis:</emphasis>
Distinct from
<taxonomicName id="4C3A4D08FFF9962CFE1F57CFFDB8C0B7" box="[500,628,1667,1692]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="westcotti">
<emphasis id="B94EEA99FFF9962CFE1F57CFFDB8C0B7" box="[500,628,1667,1692]" italics="true" pageId="5" pageNumber="6">I. westcotti</emphasis>
</taxonomicName>
in the more gracile gonopods (Fig. 7), with many fewer lateral teeth (
<figureCitation id="13012A0EFFF9962CFF7457E6FF2AC0EF" box="[159,230,1706,1732]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1923,1947]" captionTargetBox="[151,1435,729,1899]" captionTargetId="figure@3.[151,1436,729,1900]" captionTargetPageId="3" captionText="FIGURES 1 4. Gonopods of Idagona species. 1, Idagona westcotti, Giant Arch Cave, posteriorventral view. 2. I. westcotti, Ice Capades Cave, posterioventral view. 3. I. jasperi, n. sp. posterioventral view. 4. I. lehmanenesis, n. sp., posterior view." httpUri="https://zenodo.org/record/176488/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
); the posterior gonopod sternum of
<taxonomicName id="4C3A4D08FFF9962CFD6057E7FD3FC0EF" box="[651,755,1707,1732]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="westcotti">
<emphasis id="B94EEA99FFF9962CFD6057E7FD3FC0EF" box="[651,755,1707,1732]" italics="true" pageId="5" pageNumber="6">westcotti</emphasis>
</taxonomicName>
has a median flange with acute corners; this is not seen in
<emphasis id="B94EEA99FFF9962CFF7C579FFF3CC0C7" box="[151,240,1747,1772]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4D08FFF9962CFF7C579FFF21C0C7" box="[151,237,1747,1772]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jasperi">jasperi</taxonomicName>
.
</emphasis>
The adenostyle of femur three (
<figureCitation id="13012A0EFFF9962CFD9F579EFD0BC0C7" box="[628,711,1746,1772]" captionStart="FIGURES 9 14" captionStartId="6.[151,269,1722,1746]" captionTargetBox="[151,1436,357,1706]" captionTargetId="figure@6.[151,1436,357,1707]" captionTargetPageId="6" captionText="FIGURES 9 14. Pregonopodal leg modifications of Idagona species. 9 12, third legs, anterior views. 9. Idagona westcotti, Giant Arch Cave. 10.. I. westcotti, Ice Capades Cave. 11. I. jasperi n. sp. 12. I. lehmanensis n. sp. 13. Leg 4 of I. lehmanensis n. sp., anterior view. 14. Denticles near adenostyle of leg 3 femur of I. lehmanensis n. sp." httpUri="https://zenodo.org/record/176489/files/figure.png" pageId="5" pageNumber="6">Fig. 11</figureCitation>
) is more strongly constricted at the base in
<emphasis id="B94EEA99FFF9962CFB26579FFAEAC0C7" box="[1229,1318,1747,1772]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4D08FFF9962CFB26579FFAEEC0C7" box="[1229,1314,1747,1772]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jasperi">jasperi</taxonomicName>
,
</emphasis>
and has a very obvious apical depression and large pore; in
<taxonomicName id="4C3A4D08FFF9962CFD3657B7FC89C13F" box="[733,837,1787,1812]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="westcotti">
<emphasis id="B94EEA99FFF9962CFD3657B7FC89C13F" box="[733,837,1787,1812]" italics="true" pageId="5" pageNumber="6">westcotti</emphasis>
</taxonomicName>
there are multiple small pores.
</paragraph>
</subSubSection>
<subSubSection id="C3206500FFF9962CFF2D566FFC51C117" box="[198,925,1826,1852]" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8B85368BFFF9962CFF2D566FFC51C117" blockId="5.[151,1436,1546,2012]" box="[198,925,1826,1852]" pageId="5" pageNumber="6">
<emphasis id="B94EEA99FFF9962CFF2D566FFE81C117" box="[198,333,1827,1852]" italics="true" pageId="5" pageNumber="6">Etymology:</emphasis>
The species is named for the collector, Jon Jasper.
</paragraph>
</subSubSection>
<subSubSection id="C3206500FFF9962FFF2D5607FD43C729" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="8B85368BFFF9962FFF2D5607FD43C729" blockId="5.[151,1436,1546,2012]" lastBlockId="6.[151,1437,152,258]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B94EEA99FFF9962CFF2D5607FE9DC14F" box="[198,337,1867,1892]" italics="true" pageId="5" pageNumber="6">Description</emphasis>
: Male: 13.5 mm long, 1.0 mm wide. Fourteen ocelli in oval patch, ocelli round, black, compactly arranged. Legpair 3 (
<figureCitation id="13012A0EFFF9962CFE09563EFDF9C1A7" box="[482,565,1906,1932]" captionStart="FIGURES 9 14" captionStartId="6.[151,269,1722,1746]" captionTargetBox="[151,1436,357,1706]" captionTargetId="figure@6.[151,1436,357,1707]" captionTargetPageId="6" captionText="FIGURES 9 14. Pregonopodal leg modifications of Idagona species. 9 12, third legs, anterior views. 9. Idagona westcotti, Giant Arch Cave. 10.. I. westcotti, Ice Capades Cave. 11. I. jasperi n. sp. 12. I. lehmanensis n. sp. 13. Leg 4 of I. lehmanensis n. sp., anterior view. 14. Denticles near adenostyle of leg 3 femur of I. lehmanensis n. sp." httpUri="https://zenodo.org/record/176489/files/figure.png" pageId="5" pageNumber="6">Fig. 11</figureCitation>
) with femur enlarged, bearing prominent median adenostyle strongly constricted at base. Legpair 4 enlarged but with unmodified femora. Gonopods (
<figureCitation id="13012A0EFFF9962CFBAC56D6FB57C19F" box="[1095,1179,1946,1972]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1923,1947]" captionTargetBox="[151,1435,729,1899]" captionTargetId="figure@3.[151,1436,729,1900]" captionTargetPageId="3" captionText="FIGURES 1 4. Gonopods of Idagona species. 1, Idagona westcotti, Giant Arch Cave, posteriorventral view. 2. I. westcotti, Ice Capades Cave, posterioventral view. 3. I. jasperi, n. sp. posterioventral view. 4. I. lehmanenesis, n. sp., posterior view." httpUri="https://zenodo.org/record/176488/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
, 7, 19, 20) similar to those of
<emphasis id="B94EEA99FFF9962CFEE8568FFE43C1F7" box="[259,399,1987,2012]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4D08FFF9962CFEE8568FFE46C1F7" box="[259,394,1987,2012]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="westcotti">I. westcotti</taxonomicName>
,
</emphasis>
but markedly more slender, more abruptly tapered distally; lateral apical teeth much reduced, median extension of posterior gonopod sternum low. Coxae 10 (
<figureCitation id="13012A0EFFFA962FFBE851D4FB95C699" box="[1027,1113,152,178]" captionStart="FIGURES 15 18" captionStartId="7.[151,269,1493,1517]" captionTargetBox="[151,1435,357,1466]" captionTargetId="figure@7.[151,1436,357,1467]" captionTargetPageId="7" captionText="FIGURES 15 18. Leg 10 coxae of Idagona species, posterior views. 15.. Idagona westcotti, Giant Arch Cave. 16. I. westcotti, Ice Capades Cave. 17. I. jasperi n. sp. 18. I. lehmanensis n. sp., anterior view." httpUri="https://zenodo.org/record/176490/files/figure.png" pageId="6" pageNumber="7">Fig. 17</figureCitation>
) enlarged but not extended distal to trochanteral joint, distal knob strongly acute, gland opening ventral. Coloration medium tan, extensively mottled with darker purplish-brown.
</paragraph>
</subSubSection>
<caption id="DF456603FFFA962FFF7C57F6FB35C13C" httpUri="https://zenodo.org/record/176489/files/figure.png" pageId="6" pageNumber="7" targetBox="[151,1436,357,1706]" targetPageId="6">
<paragraph id="8B85368BFFFA962FFF7C57F6FB35C13C" blockId="6.[151,1436,1722,1815]" pageId="6" pageNumber="7">
<emphasis id="B94EEA99FFFA962FFF7C57F6FE83C0F9" bold="true" box="[151,335,1722,1746]" pageId="6" pageNumber="7">FIGURES 914.</emphasis>
Pregonopodal leg modifications of
<taxonomicName id="4C3A4D08FFFA962FFD2057F6FCE8C0FA" box="[715,804,1722,1745]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA99FFFA962FFD2057F6FCE8C0FA" box="[715,804,1722,1745]" italics="true" pageId="6" pageNumber="7">Idagona</emphasis>
</taxonomicName>
species. 912, third legs, anterior views. 9.
<taxonomicName id="4C3A4D08FFFA962FFAEE57F6FE5CC0DF" authority="Giant Arch Cave." authorityName="Giant Arch Cave." class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="westcotti">
<emphasis id="B94EEA99FFFA962FFAEE57F6FF00C0DF" italics="true" pageId="6" pageNumber="7">Idagona westcotti,</emphasis>
Giant Arch Cave.
</taxonomicName>
10..
<taxonomicName id="4C3A4D08FFFA962FFE3F5791FCD1C0DF" authority="Ice Capades Cave." authorityName="Ice Capades Cave." box="[468,797,1757,1780]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="westcotti">
<emphasis id="B94EEA99FFFA962FFE3F5791FD9CC0DF" box="[468,592,1757,1780]" italics="true" pageId="6" pageNumber="7">I. westcotti,</emphasis>
Ice Capades Cave.
</taxonomicName>
11.
<taxonomicName id="4C3A4D08FFFA962FFCB85791FC7FC0DF" box="[851,947,1757,1780]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="jasperi" status="sp. nov.">
<emphasis id="B94EEA99FFFA962FFCB85791FC7FC0DF" box="[851,947,1757,1780]" italics="true" pageId="6" pageNumber="7">I. jasperi</emphasis>
</taxonomicName>
<emphasis id="B94EEA99FFFA962FFC515790FC3AC0DF" bold="true" box="[954,1014,1756,1780]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A27D57E2FFFA962FFC515790FC3AC0DF" box="[954,1014,1756,1780]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
12.
<taxonomicName id="4C3A4D08FFFA962FFBC65791FB0BC0DF" box="[1069,1223,1757,1780]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="lehmanensis" status="sp. nov.">
<emphasis id="B94EEA99FFFA962FFBC65791FB0BC0DF" box="[1069,1223,1757,1780]" italics="true" pageId="6" pageNumber="7">I. lehmanensis</emphasis>
</taxonomicName>
<emphasis id="B94EEA99FFFA962FFB255790FAC7C0DF" bold="true" box="[1230,1291,1756,1780]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A27D57E2FFFA962FFB255790FAC7C0DF" box="[1230,1291,1756,1780]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
13. Leg 4 of
<taxonomicName id="4C3A4D08FFFA962FFF7C564CFEFCC13C" box="[151,304,1792,1815]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="lehmanensis" status="sp. nov.">
<emphasis id="B94EEA99FFFA962FFF7C564CFEFCC13C" box="[151,304,1792,1815]" italics="true" pageId="6" pageNumber="7">I. lehmanensis</emphasis>
</taxonomicName>
<emphasis id="B94EEA99FFFA962FFEDC57B3FEBFC13C" bold="true" box="[311,371,1791,1815]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A27D57E2FFFA962FFEDC57B3FEBFC13C" box="[311,371,1791,1815]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, anterior view. 14. Denticles near adenostyle of leg 3 femur of
<taxonomicName id="4C3A4D08FFFA962FFBF7564CFB7AC13C" box="[1052,1206,1792,1815]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="lehmanensis" status="sp. nov.">
<emphasis id="B94EEA99FFFA962FFBF7564CFB7AC13C" box="[1052,1206,1792,1815]" italics="true" pageId="6" pageNumber="7">I. lehmanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57E2FFFA962FFB5657B3FB35C13C" box="[1213,1273,1791,1815]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="C3206500FFFA9621FF2D561CFE3CC779" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="distribution">
<paragraph id="8B85368BFFFA962FFF2D561CFA4BC1BA" blockId="6.[151,1436,1871,2017]" pageId="6" pageNumber="7">
<emphasis id="B94EEA99FFFA962FFF2D561CFEACC142" box="[198,352,1872,1897]" italics="true" pageId="6" pageNumber="7">Distribution:</emphasis>
Known only from the
<typeStatus id="54818829FFFA962FFD9F5603FD6AC142" box="[628,678,1871,1897]" pageId="6" pageNumber="7">type</typeStatus>
localities. The caves with
<taxonomicName id="4C3A4D08FFFA962FFC03561CFB98C142" box="[1000,1108,1872,1897]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="jasperi">
<emphasis id="B94EEA99FFFA962FFC03561CFB98C142" box="[1000,1108,1872,1897]" italics="true" pageId="6" pageNumber="7">I. jasperi</emphasis>
</taxonomicName>
are in the upper reaches of Logan Canyon, about
<quantity id="4CC29B6EFFFA962FFE70563BFDEBC1BA" box="[411,551,1911,1937]" metricMagnitude="4" metricUnit="m" metricValue="3.621024" metricValueMax="4.02336" metricValueMin="3.218688" pageId="6" pageNumber="7" unit="mi" value="22.5" valueMax="25.0" valueMin="20.0">20-25 miles</quantity>
northeast of Logan, Utah, in the southeast flank of the Bear River Range.
</paragraph>
<paragraph id="8B85368BFFFA962EFF2D56ECFC8FC701" blockId="6.[151,1436,1871,2017]" lastBlockId="7.[151,1436,152,298]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B94EEA99FFFA962FFF2D56ECFEDFC192" box="[198,275,1952,1977]" italics="true" pageId="6" pageNumber="7">Notes:</emphasis>
There is extensive karst development in Logan Canyon (
<bibRefCitation id="EFAB4B7AFFFA962FFC5A56D3FBACC192" author="Spangler" box="[945,1120,1951,1977]" pageId="6" pageNumber="12" refString="Spangler, L. E. (2001) Delineation of recharge areas for karst springs in Logan Canyon, Bear River Range, northern Utah. United States Geological Survey Karst Interest Group Proceedings, Water-Resources Investigations Report 01 - 4011, 186 - 193." type="book chapter" year="2001">Spangler 2001</bibRefCitation>
;
<bibRefCitation id="EFAB4B7AFFFA962FFB8756D3FB35C192" author="Jasper" box="[1132,1273,1951,1977]" pageId="6" pageNumber="12" refString="Jasper, J. &amp; Kowallis, B. (2006) Exploring the deep caves of Tony Grove. Utah Caver Annual, 2005, 17." type="journal article" year="2006">Jasper 2006</bibRefCitation>
) and many of the caves are at high altitudes, above 8000. Nielsons Cave, among others, harbors permanent ice or snowpack, as do some of the Snake River Plain lava tubes.
<taxonomicName id="4C3A4D08FFFB962EFCDE51D4FC3EC69A" box="[821,1010,152,177]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="jasperi">
<emphasis id="B94EEA99FFFB962EFCDE51D4FC3EC69A" box="[821,1010,152,177]" italics="true" pageId="7" pageNumber="8">Idagona jasperi</emphasis>
</taxonomicName>
specimens are recorded as actively walking over ice and snow, and like other elements of the faunas of these caves, are well-adapted to nearfreezing temperatures. The millipeds are described by collectors as being very abundant on wood and other organic debris in the caves (J. Jasper, pers. comm..to WS).
</paragraph>
<caption id="DF456603FFFB962EFF7C5499FB9FC024" httpUri="https://zenodo.org/record/176490/files/figure.png" pageId="7" pageNumber="8" targetBox="[151,1435,357,1466]" targetPageId="7">
<paragraph id="8B85368BFFFB962EFF7C5499FB9FC024" blockId="7.[151,1436,1493,1552]" pageId="7" pageNumber="8">
<emphasis id="B94EEA99FFFB962EFF7C5499FE92C3C6" bold="true" box="[151,350,1493,1517]" pageId="7" pageNumber="8">FIGURES 1518.</emphasis>
Leg 10 coxae of
<taxonomicName id="4C3A4D08FFFB962EFDC9549AFDB7C3C6" box="[546,635,1494,1517]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA99FFFB962EFDC9549AFDB7C3C6" box="[546,635,1494,1517]" italics="true" pageId="7" pageNumber="8">Idagona</emphasis>
</taxonomicName>
species, posterior views. 15..
<taxonomicName id="4C3A4D08FFFB962EFC27549AFA9AC3C7" authority="Giant Arch Cave." authorityName="Giant Arch Cave." box="[972,1366,1493,1517]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="westcotti">
<emphasis id="B94EEA99FFFB962EFC27549AFB5EC3C6" box="[972,1170,1494,1517]" italics="true" pageId="7" pageNumber="8">Idagona westcotti,</emphasis>
Giant Arch Cave.
</taxonomicName>
16.
<taxonomicName id="4C3A4D08FFFB962EFA67549AFE05C024" authority="Ice Capades Cave." authorityName="Ice Capades Cave." class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="westcotti">
<emphasis id="B94EEA99FFFB962EFA67549AFF30C024" italics="true" pageId="7" pageNumber="8">I. westcotti,</emphasis>
Ice Capades Cave.
</taxonomicName>
17.
<taxonomicName id="4C3A4D08FFFB962EFE1554B4FD92C024" box="[510,606,1528,1551]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="jasperi" status="sp. nov.">
<emphasis id="B94EEA99FFFB962EFE1554B4FD92C024" box="[510,606,1528,1551]" italics="true" pageId="7" pageNumber="8">I. jasperi</emphasis>
</taxonomicName>
<emphasis id="B94EEA99FFFB962EFD8E54B4FD6DC03B" bold="true" box="[613,673,1528,1552]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A27D57E2FFFB962EFD8E54B4FD6DC03B" box="[613,673,1528,1552]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
18.
<taxonomicName id="4C3A4D08FFFB962EFD3D54B4FCBCC024" box="[726,880,1528,1551]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lehmanensis" status="sp. nov.">
<emphasis id="B94EEA99FFFB962EFD3D54B4FCBCC024" box="[726,880,1528,1551]" italics="true" pageId="7" pageNumber="8">I. lehmanensis</emphasis>
</taxonomicName>
<emphasis id="B94EEA99FFFB962EFC9C54B4FC7FC03B" bold="true" box="[887,947,1528,1552]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A27D57E2FFFB962EFC9C54B4FC7FC03B" box="[887,947,1528,1552]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, anterior view.
</paragraph>
</caption>
<paragraph id="8B85368BFFFB962EFF2D570FFDB5C0FE" blockId="7.[151,1437,1603,2029]" pageId="7" pageNumber="8">
In contrast to the lava tubes of the Snake River Plain, the fauna of the caves of the Bear River Range is little known, and this report is the first for troglobitic or troglophilic terrestrial animals there. The compact, wellformed eyepatch and relatively dark pigmentation (when compared to
<taxonomicName id="4C3A4D08FFFB962EFC3157DFFB90C087" box="[986,1116,1683,1708]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="westcotti">
<emphasis id="B94EEA99FFFB962EFC3157DFFB90C087" box="[986,1116,1683,1708]" italics="true" pageId="7" pageNumber="8">I. westcotti</emphasis>
</taxonomicName>
) would indicate a minimal degree of cave-adaptation in this species.
</paragraph>
<paragraph id="8B85368BFFFB9621FF2D57AFFE3CC779" blockId="7.[151,1437,1603,2029]" lastBlockId="8.[151,1436,152,338]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Earlier,
<bibRefCitation id="EFAB4B7AFFFB962EFECD57AFFE7BC0D6" author="Peck" box="[294,439,1763,1789]" pageId="7" pageNumber="12" refString="Peck, S. B. (1981) The invertebrate fauna of the caves of the Uinta Mountains, northeastern Utah. Great Basin Naturalist, 41, 207 - 212." type="journal article" year="1981">Peck (1981)</bibRefCitation>
described invertebrate collections from the Uinta Mountains, to the southeast, and found that of 38 species collected in caves there, only three could be regarded as cave-limited.
<bibRefCitation id="EFAB4B7AFFFB962EFAE75647FA50C10E" author="Peck" box="[1292,1436,1803,1829]" pageId="7" pageNumber="12" refString="Peck, S. B. (1981) The invertebrate fauna of the caves of the Uinta Mountains, northeastern Utah. Great Basin Naturalist, 41, 207 - 212." type="journal article" year="1981">Peck (1981)</bibRefCitation>
attributed the relatively low number of troglobionts to the short time available since colonization. Up to about 12,000 years ago, severe periglacial conditions would have obtained, and as the glaciers melted, caves would have been scoured by powerful streams. No doubt this same analysis can also be applied to the high altitude caves of Logan Canyon, but with even less time available since colonization, as the montane glaciers in the more northerly Bear River Range would have retreated more slowly. Moist conferous forests surround the caves where
<taxonomicName id="4C3A4D08FFF49621FEB051D4FE09C69A" box="[347,453,152,177]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="jasperi">
<emphasis id="B94EEA99FFF49621FEB051D4FE09C69A" box="[347,453,152,177]" italics="true" pageId="8" pageNumber="9">I. jasperi</emphasis>
</taxonomicName>
has been found, and surface-dwelling populations may well exist there, but no collecting has been done. The caves may in fact only be refuges for the millipeds from the severe winter conditions that obtain in the region, and the animals may move freely between caves and forest litter. I strongly suspect that other, as yet uncollected, species of
<taxonomicName id="4C3A4D08FFF49621FD24505CFCFDC702" box="[719,817,272,297]" class="Diplopoda" family="Conotylidae" genus="Idagona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA99FFF49621FD24505CFCFDC702" box="[719,817,272,297]" italics="true" pageId="8" pageNumber="9">Idagona</emphasis>
</taxonomicName>
exist in caves and high altitude forests in other Utah (and Idaho) mountain ranges.
</paragraph>
</subSubSection>
</treatment>
</document>