156 lines
20 KiB
XML
156 lines
20 KiB
XML
<document ID-DOI="10.5281/zenodo.163451" ID-GBIF-Dataset="f9566778-1df4-44d6-9a4c-849de39a4b6c" ID-GBIF-Taxon="124955169" ID-Zenodo-Dep="163451" checkinTime="1480975261095" checkinUser="plazi" docAuthor="Vincent J. Kalkman" docDate="2016" docId="03938796FFD6FFE227BA8DC8FEB1FE6E" docLanguage="en" docName="10.5281zenodo.163451_Kalkman.pdf" docOrigin="Odonatologica 45 (3)" docStyle="DocumentStyle{}" docTitle="Celebargiolestes cinctus Selys 1886" docType="treatment" docVersion="11" lastPageId="12" lastPageNumber="247" masterDocId="FFAAFFEEFFDEFFEE27258934FFCBFF99" masterDocTitle="Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae)" masterLastPageNumber="269" masterPageNumber="235" pageId="8" pageNumber="243" updateTime="1643502553511" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vincent J. Kalkman</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Odonatologica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-12-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>45</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>235</mods:start>
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<mods:end>269</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.163451</mods:identifier>
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<mods:identifier type="GBIF-Dataset">f9566778-1df4-44d6-9a4c-849de39a4b6c</mods:identifier>
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<mods:identifier type="Zenodo-Dep">163451</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="124955169" LSID="urn:lsid:plazi:treatment:03938796FFD6FFE227BA8DC8FEB1FE6E" httpUri="http://treatment.plazi.org/id/03938796FFD6FFE227BA8DC8FEB1FE6E" lastPageId="12" lastPageNumber="247" pageId="8" pageNumber="243">
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<subSubSection box="[159,779,1276,1303]" pageId="8" pageNumber="243" type="nomenclature">
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<paragraph blockId="8.[159,1036,1276,1554]" box="[159,779,1276,1303]" pageId="8" pageNumber="243">
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<heading bold="true" box="[159,779,1276,1303]" fontSize="11" level="3" pageId="8" pageNumber="243" reason="0">
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<emphasis bold="true" box="[159,779,1276,1303]" pageId="8" pageNumber="243">
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<taxonomicName authority="(Selys, 1886)" authorityName="Selys" authorityYear="1886" baseAuthorityName="Selys" baseAuthorityYear="1886" box="[159,603,1276,1303]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="243" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis bold="true" box="[159,441,1276,1303]" italics="true" pageId="8" pageNumber="243">Celebargiolestes cinctus</emphasis>
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(Selys, 1886)
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</taxonomicName>
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, ‘red leg’ form
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="9" lastPageNumber="244" pageId="8" pageNumber="243" type="materials_examined">
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<paragraph blockId="8.[159,1036,1276,1554]" box="[159,360,1313,1339]" pageId="8" pageNumber="243">
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<heading bold="true" box="[159,360,1313,1339]" fontSize="11" level="3" pageId="8" pageNumber="243" reason="0">
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<emphasis bold="true" box="[159,360,1313,1339]" pageId="8" pageNumber="243">Material studied</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="8.[159,1036,1276,1554]" box="[159,341,1346,1370]" pageId="8" pageNumber="243">
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<heading bold="true" box="[159,341,1346,1370]" fontSize="10" level="6" pageId="8" pageNumber="243" reason="6">
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<emphasis bold="true" box="[159,341,1346,1370]" pageId="8" pageNumber="243">Sulawesi Tengah</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="8.[159,1036,1276,1554]" lastBlockId="9.[159,1036,253,369]" lastPageId="9" lastPageNumber="244" pageId="8" pageNumber="243">2♂, 15 km NNE of Luwuk, Sg. Biak, clear running water with rapids and pools, 150m, 27-i-1989, J. van Tol leg.; 8♂ 2♀, 25 km NE of Luwuk, tributary of Sg. Bantajan, clear running water, shaded by disturbed primary forest, 100 m a.s.l., 30-i-1989, J. van Tol leg.; 1♂, 15 km NNE Luwuk, Sg. Biak, near road Biak to Poh, disturbed rainforest, 06-x-1989, J.P. Duffels leg.; 2♂ 1♀, 15 km NNE of Luwuk, Sg. Biak at road from Biak to Poh, with ponded sites and currents, semi-cultivated areas, nearly unshaded, some emergent vegetation, 06- and 21-x-1989, J. van Tol leg.; 7♂ 1♀, 20 km ENE of Biak: Sg. Takilalang, stream 2–3 m wide in somewhat disturbed lowland evergreen rainforest, clear water, bottom sand and uplifted coral reef, mostly shaded, 08-x-1989, J. van Tol leg.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="246" pageId="9" pageNumber="244" type="description">
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<paragraph blockId="9.[159,1036,424,673]" box="[159,220,424,450]" pageId="9" pageNumber="244">
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<heading bold="true" box="[159,220,424,450]" fontSize="11" level="3" pageId="9" pageNumber="244" reason="0">
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<emphasis bold="true" box="[159,220,424,450]" pageId="9" pageNumber="244">Male</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="9.[159,1036,424,673]" pageId="9" pageNumber="244">Description of male specimen from Ranotongkor (JvT6476; collected by L. Coomans de Ruiter in 1940)</paragraph>
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<paragraph blockId="9.[159,1036,424,673]" pageId="9" pageNumber="244">
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Head − Anterior ⅔ of labium blackish, posterior third yellow-brown. Labrum shining metallic black. Front of head, including mandibles, clypeus and sockets of antennae ochre (
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<figureCitation box="[537,609,610,637]" captionStart="Figures 7 – 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 – 14. Head of Celebargiolestes males. 7 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 – C. askewi sp. nov., holotype; 11 – C. toli sp. nov., holotype; 12 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 – C. orri sp. nov., holotype; 14 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="9" pageNumber="244">Fig. 7</figureCitation>
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). Antennae brown, dorsum of head black.
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</paragraph>
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<paragraph blockId="9.[159,1036,727,941]" pageId="9" pageNumber="244">
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Thorax − Sides of prothorax dull yellow; anterior lobe of pronotum dull yellow with outer rim brown-black, median part of middle and posterior lobe blackish, remainder dull yellow. Posterior lobe of pronotum broad, seen from lateral gently sloping and with the outer border steeply raised, lateral corners rounded. Pattern of thorax as in
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<figureCitation box="[636,749,877,904]" captionStart="Figures 15 – 18" captionStartId="14.[159,238,1441,1466]" captionTargetBox="[203,991,248,1401]" captionTargetId="figure@14.[203,991,248,1401]" captionTargetPageId="14" captionText="Figures 15 – 18. Body of Celebargiolestes males. 15 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 16 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 17 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 18 – C. askewi sp. nov., holotype." httpUri="https://zenodo.org/record/192856/files/figure.png" pageId="9" pageNumber="244">Figure 15</figureCitation>
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; darker areas black, pale areas dull yellow.
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</paragraph>
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<paragraph blockId="9.[159,1036,994,1207]" pageId="9" pageNumber="244">Legs − Coxae and trochanters dull yellow, femora yellow with two broad but poorly defined black rings, knees black, tibiae and tarsi yellow-brown. Spines pale orange-brown; femora with respectively 9–11, 8–9 and 12–13 spines laterally; tibiae of fore legs with 8–9 smaller and 5–6 larger spines laterally; tibiae of mid and hind legs with 11–12 and 11–13 spines laterally respectively.</paragraph>
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<paragraph blockId="9.[159,1035,1261,1399]" pageId="9" pageNumber="244">Wings − Venation black, membrane hyaline, Fw and Hw with 2 Ax, Fw with 35–38 Px and Hw with 33 Px. Ac at about a tenth the distance between Ax1 and Ax2. Brace vein absent. Pterostigma brown. Three, rarely four rows of cells between anal vein and border of Hw.</paragraph>
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<paragraph blockId="9.[159,1036,1453,1554]" lastBlockId="10.[159,1036,253,542]" lastPageId="10" lastPageNumber="245" pageId="9" pageNumber="244">
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Abdomen − Pattern as in
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<figureCitation box="[469,583,1453,1480]" captionStart="Figures 15 – 18" captionStartId="14.[159,238,1441,1466]" captionTargetBox="[203,991,248,1401]" captionTargetId="figure@14.[203,991,248,1401]" captionTargetPageId="14" captionText="Figures 15 – 18. Body of Celebargiolestes males. 15 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 16 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 17 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 18 – C. askewi sp. nov., holotype." httpUri="https://zenodo.org/record/192856/files/figure.png" pageId="9" pageNumber="244">Figure 15</figureCitation>
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; dark pattern elements dark brown to black, pale pattern elements yellow-brown. Anterior half of S1 yellow, posterior half brown to black, S2 with dorsum black and anterior section laterally yellow, S3–8 dorsally largely dark, ventrally and lower lateral section yellow, anteriorly of S3–8 the yellow reaches to the dorsum forming a yellow ring. S9–10 black, the latter strongly depressed in the middle, its hind bor- der without modifications. Appendages as shown in
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<figureCitation box="[775,942,365,391]" captionStart="Figures 34 – 39" captionStartId="27.[159,238,1413,1438]" captionTargetBox="[156,1038,480,1372]" captionTargetId="figure@27.[156,1038,480,1372]" captionTargetPageId="27" captionText="Figures 34 – 39. Appendages of Celebargiolestes males. 34 – C. cinctus, Ranotongkor, 02 - viii- 1940, JvT 6476, dorsal view; 35 – C. cinctus, Ranotongkor, 02 - viii- 1940, JvT 6476, lateral view; 36 – Ranotongkor, 02 - viii- 1940, JvT 6476, ventral view; 37 – C. askewi sp. nov., holotype, dorsal view; 38 – C. askewi sp. nov., holotype, lateral view; 39 – C. askewi sp. nov., holotype, ventral view." httpUri="https://zenodo.org/record/192860/files/figure.png" pageId="10" pageNumber="245">Figures 34–36</figureCitation>
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; base of lower appendages yellow, remainder dark brown. Lower appendages long, in lateral view slightly longer than ½ of upper appendages and, when seen from lateral, with tip reaching level of the upper appendages. Genital ligula as in
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<figureCitation box="[220,386,515,542]" captionStart="Figures 23 – 27" captionStartId="25.[159,237,1412,1437]" captionTargetBox="[208,986,249,1370]" captionTargetId="figure@25.[208,986,249,1370]" captionTargetPageId="25" captionText="Figures 23 – 27. Genital ligula of Celebargiolestes males. 23 – C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, lateral view; 24 – C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, ventral view; 25 – C. cinctus ‘ red leg’, Banggai Peninsula, 08 - x- 1989, latero-ventral view; 26 – C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, lateral view; 27 – C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, ventral view." httpUri="https://zenodo.org/record/192858/files/figure.png" pageId="10" pageNumber="245">Figures 23–25</figureCitation>
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, with a deep apical cleft and with large lateral lobes.
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</paragraph>
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<caption httpUri="https://zenodo.org/record/192854/files/figure.png" pageId="10" pageNumber="245" targetBox="[173,1024,597,1348]" targetPageId="10">
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<paragraph blockId="10.[159,1037,1412,1554]" pageId="10" pageNumber="245">
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<emphasis bold="true" box="[159,296,1412,1437]" pageId="10" pageNumber="245">Figures 1–6.</emphasis>
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Details of
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<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[424,685,1412,1436]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis box="[424,685,1412,1436]" italics="true" pageId="10" pageNumber="245">Celebargiolestes cinctus</emphasis>
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</taxonomicName>
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. 1 – Prothorax and pronotum, male; 2 – Venation, male, Ranotongkor, 02-viii-1940, JvT6476; 3 – Pterostigma of fore and hind wing, male, Mapanget, 23-vi-1940, JvT6638; 4 – Pterostigma of fore and hind wing, Mapanget, 23-vi-1940, female, JvT6639; 5 – Body, female, Ton- dano, 03-ix-1940, JvT6494; 6 – Ovipositor, female, Tondano, 03-ix-1940, JvT6494.
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</paragraph>
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</caption>
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<paragraph blockId="11.[159,1036,253,877]" box="[159,289,253,279]" pageId="11" pageNumber="246">
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<heading bold="true" box="[159,289,253,279]" fontSize="11" level="3" pageId="11" pageNumber="246" reason="0">
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<emphasis bold="true" box="[159,289,253,279]" pageId="11" pageNumber="246">Variability</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="11.[159,1036,253,877]" pageId="11" pageNumber="246">
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In ventral view the inferior appendages shows minute differences in shape between localities. The inferior appendages are however often slightly distorted making it difficult to judge these differences. In part of the material from the Banggai Penisula, listed here as
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<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[647,761,403,429]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="246" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis box="[647,761,403,429]" italics="true" pageId="11" pageNumber="246">C. cinctus</emphasis>
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</taxonomicName>
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‘red leg’ form, the pale coloration of head and thorax has a more reddish tinge (
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<figureCitation box="[832,917,440,466]" captionStart="Figures 15 – 18" captionStartId="14.[159,238,1441,1466]" captionTargetBox="[203,991,248,1401]" captionTargetId="figure@14.[203,991,248,1401]" captionTargetPageId="14" captionText="Figures 15 – 18. Body of Celebargiolestes males. 15 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 16 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 17 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 18 – C. askewi sp. nov., holotype." httpUri="https://zenodo.org/record/192856/files/figure.png" pageId="11" pageNumber="246">Fig. 17</figureCitation>
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). In addition to this the black rings on the femora are less pronounced and often hardly detectable (
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<figureCitation box="[381,467,515,541]" captionStart="Figures 15 – 18" captionStartId="14.[159,238,1441,1466]" captionTargetBox="[203,991,248,1401]" captionTargetId="figure@14.[203,991,248,1401]" captionTargetPageId="14" captionText="Figures 15 – 18. Body of Celebargiolestes males. 15 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 16 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 17 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 18 – C. askewi sp. nov., holotype." httpUri="https://zenodo.org/record/192856/files/figure.png" pageId="11" pageNumber="246">Fig. 17</figureCitation>
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). I decided to give no taxonomic status to this form as these specimens do not differ in morphology from specimens from other localities and as there are only differences in tint of the colouration but not in the pattern. The specimens from Sangihe Islands have a relatively open venation with a low number of Px. Despite the variability found within
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<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[159,271,702,728]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="246" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis box="[159,271,702,728]" italics="true" pageId="11" pageNumber="246">C.cinctus</emphasis>
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</taxonomicName>
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it is likely to be monophyletic.
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</paragraph>
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<paragraph blockId="11.[159,1036,253,877]" pageId="11" pageNumber="246">Wings − Fw and Hw with 2 Ax; Fw with 35–38 Px (29–30 in material from Sangihe) and Hw with 32–33 Px (26–28 in material from Sangihe). Measurements [mm] − Total length 44–48, abdomen 32–38, Fw 28–34; Hw 29–34.</paragraph>
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<paragraph blockId="11.[159,569,947,1011]" box="[159,247,947,973]" pageId="11" pageNumber="246">
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<heading bold="true" box="[159,247,947,973]" fontSize="11" level="3" pageId="11" pageNumber="246" reason="0">
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<emphasis bold="true" box="[159,247,947,973]" pageId="11" pageNumber="246">Female</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="11.[159,569,947,1011]" box="[159,569,984,1011]" pageId="11" pageNumber="246">
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<heading box="[159,569,984,1011]" fontSize="11" level="5" pageId="11" pageNumber="246" reason="1">Head − Pattern on head as in male.</heading>
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</paragraph>
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<paragraph blockId="11.[159,1036,1059,1197]" pageId="11" pageNumber="246">Thorax − Pattern on prothorax, synthorax and legs as in male. Fw and Hw with 2 Ax; Fw with 27–32 Px (25 in material from Sangihe) and Hw with 25–31 Px (23 in material from Sangihe). Pt white with a slight yellow tinge, outer border often mottled greyish-black.</paragraph>
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<paragraph blockId="11.[159,1036,1237,1450]" pageId="11" pageNumber="246">
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Abdomen − Pattern as in
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<figureCitation box="[457,554,1237,1264]" captionStart="Figures 7 – 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 – 14. Head of Celebargiolestes males. 7 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 – C. askewi sp. nov., holotype; 11 – C. toli sp. nov., holotype; 12 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 – C. orri sp. nov., holotype; 14 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="11" pageNumber="246">Figure 7</figureCitation>
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a; dark sections dark brown to black, pale sections yellow-brown. S3–9 dorsally largely dark, ventrally and lower lateral area yellow, anteriorly on each segment the yellow reaches to the dorsum forming a yellow ring covering approximately ¹/₅ to ¹/₁0 of the segment. S10 black, cerci brown, base of valvifer yellow becoming brown to black towards the tip.
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</paragraph>
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<paragraph blockId="11.[159,1036,1490,1554]" pageId="11" pageNumber="246">Measurements [mm] − Total length 36–40; abdomen 28–31, Fw 27–31; Hw 27–31.</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="247" type="diagnosis">
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<paragraph blockId="12.[159,1036,253,504]" box="[159,424,253,279]" pageId="12" pageNumber="247">
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<heading bold="true" box="[159,424,253,279]" fontSize="11" level="3" pageId="12" pageNumber="247" reason="0">
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<emphasis bold="true" box="[159,424,253,279]" pageId="12" pageNumber="247">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="12.[159,1036,253,504]" pageId="12" pageNumber="247">
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Males are distinguished from other species of
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<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[705,884,291,317]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="247" phylum="Arthropoda" rank="genus">
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<emphasis box="[705,884,291,317]" italics="true" pageId="12" pageNumber="247">Celebargiolestes</emphasis>
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</taxonomicName>
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by the combination of the following characters: (i) labrum dark brown to black; (ii)postclypeus and frons yellow-brown; (iii) genital ligula with a deep apical cleft; (iv) lower appendages long, about half the length of the upper appendages and, when seen from lateral, with the tip reaching the level of the upper appendages.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |