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<document id="853AE44FCE388A565E7A7AF64AEBAC20" ID-CLB-Dataset="293861" ID-DOI="10.11646/zootaxa.5433.2.8" ID-GBIF-Dataset="583e3275-7a32-45ab-9827-ff927f4b3bc1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10954089" ID-ZooBank="AEBF0EF9-61B7-4216-A60D-7A17D926909F" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1712751594845" checkinUser="plazi" docAuthor="Trivedi, Jigneshkumar N., Padate, Vinay P. &amp; Ng, Peter K. L." docDate="2024" docId="038E8784FFACFFE7FF4088D7353AD326" docLanguage="en" docName="zootaxa.5433.2.8.pdf" docOrigin="Zootaxa 5433 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5433.2.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nanosesarma gomantaka Trivedi &amp; Padate &amp; Ng 2024, n. sp." docType="treatment" docVersion="3" lastPageNumber="296" masterDocId="FFB7FFFCFFADFFE0FFD78A2C3123D672" masterDocTitle="A new species of the genus Nanosesarma Tweedie, 1950 (Crustacea: Decapoda: Brachyura: Sesarmidae) from India" masterLastPageNumber="298" masterPageNumber="289" pageNumber="290" updateTime="1713271355224" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BF133E75B27123A077AFE4900B5A65B2">A new species of the genus Nanosesarma Tweedie, 1950 (Crustacea: Decapoda: Brachyura: Sesarmidae) from India</mods:title>
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<mods:namePart id="FB5C1701E283A9404B1D27359DB8B973">Trivedi, Jigneshkumar N.</mods:namePart>
<mods:affiliation id="1029F839B60B4ED8DF3FF4B471D588AA">Animal Taxonomy and Ecology Research Laboratory, Department of Life Sciences, Hemchandracharya North Gujarat University, Patan, Gujarat, India</mods:affiliation>
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<mods:namePart id="CD9C822E1EF23F185E261BDFD05C317A">Padate, Vinay P.</mods:namePart>
<mods:affiliation id="7B7803990A86EC4005EDA9234C93755C">Centre for Marine Living Resources &amp; Ecology, Government of India, Ministry of Earth Sciences, Atal Bhavan, LNG Terminus Road, Puthuvype, Kochi, 682508, Kerala, India</mods:affiliation>
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<mods:namePart id="EE978CD26C3CD6BA000A86D795466AB0">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="C3E743F9C1B02AEBCDAE3660833D38B9">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, 117377 Singapore</mods:affiliation>
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<mods:date id="0BC60BBD589D99FA81B8D2C61F04CC9D">2024</mods:date>
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<taxonomicName id="4C274D11FFACFFE1FF4088D730EFD564" authority="Trivedi &amp; Padate &amp; Ng, 2024" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[151,460,763,790]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="290" phylum="Arthropoda" rank="species" species="gomantaka" status="sp. nov.">
<emphasis id="B953EA80FFACFFE1FF4088D730EFD564" bold="true" box="[151,460,763,790]" italics="true" pageId="1" pageNumber="290">Nanosesarma gomantaka</emphasis>
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<paragraph id="8B983692FFACFFE1FF4089333031D548" blockId="1.[151,534,763,826]" box="[151,274,799,826]" pageId="1" pageNumber="290">
(
<figureCitation id="131C2A17FFACFFE1FF488933302AD548" box="[159,265,799,826]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1375,1399]" captionStartId-1="3.[151,250,1312,1336]" captionStartId-2="4.[151,250,1550,1574]" captionStartId-3="5.[151,250,1214,1238]" captionStartId-4="6.[151,250,1943,1967]" captionTargetBox-0="[151,1436,181,1351]" captionTargetBox-1="[167,1420,184,1286]" captionTargetBox-2="[154,1432,184,1523]" captionTargetBox-3="[155,1431,185,1185]" captionTargetBox-4="[263,1317,189,1905]" captionTargetId-0="figure-21@2.[151,1436,181,1351]" captionTargetId-1="figure-17@3.[164,1423,181,1288]" captionTargetId-2="figure-21@4.[151,1435,181,1526]" captionTargetId-3="figure-17@5.[151,1435,181,1190]" captionTargetId-4="figure-21@6.[256,1332,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live N. gomantaka n. sp., in oyster shell; g, live N. gomantaka n. sp., on rotten wood." captionText-1="FIGURE 2. Nanosesarma gomantaka n. sp., a, c, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. b, d, N. pontianacense (De Man, 1895), male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, b, carapace, dorsal view; c, d, male pleon; e, female pleon." captionText-2="FIGURE 3. Nanosesarma gomantaka n. sp., a, b, d, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, left chela, outer view (setae omitted); b, left chela, dorsal view; d, third maxilliped, dorsal view; e, left vulva." captionText-3="FIGURE 4. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, N. pontianacense (De Man, 1895), male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, c, left G1, dorsal view; b, d, left G1, ventral view." captionText-4="FIGURE 5. Nanosesarma gomantaka n. sp., ad, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, P2, dorsal view; b, P3, dorsal view; c, P4, dorsal view; d, P5, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.10954093" figureDoi-1="http://doi.org/10.5281/zenodo.10954095" figureDoi-2="http://doi.org/10.5281/zenodo.10954097" figureDoi-3="http://doi.org/10.5281/zenodo.10954099" figureDoi-4="http://doi.org/10.5281/zenodo.10954101" httpUri-0="https://zenodo.org/record/10954093/files/figure.png" httpUri-1="https://zenodo.org/record/10954095/files/figure.png" httpUri-2="https://zenodo.org/record/10954097/files/figure.png" httpUri-3="https://zenodo.org/record/10954099/files/figure.png" httpUri-4="https://zenodo.org/record/10954101/files/figure.png" pageId="1" pageNumber="290">Figs. 15</figureCitation>
)
</paragraph>
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<emphasis id="B953EA80FFACFFE1FF40894B3074D5F3" bold="true" box="[151,343,871,897]" pageId="1" pageNumber="290">Type material.</emphasis>
<materialsCitation id="3B4F3CCFFFACFFE1FEA6894B32EAD5D7" collectingDate="2016-05-27" collectionCode="LFSC" collectorName="M. Bhat" country="India" latitude="15.658889" location="India" longLatPrecision="21" longitude="73.79195" pageId="1" pageNumber="290" specimenCount="1" typeStatus="holotype">
<typeStatus id="549C8830FFACFFE1FEA6894B30FFD5F3" box="[369,476,871,897]" pageId="1" pageNumber="290" type="holotype">Holotype</typeStatus>
, male (CL
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, CW
<quantity id="4CDF9B77FFACFFE1FC8A894B32E5D5F3" box="[861,966,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="290" unit="mm" value="4.7">4.7 mm</quantity>
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.ZRC-206), Chapora Estuary (
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;
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), Goa State,
<collectingCountry id="F3307602FFACFFE1FD9E89A733A6D5D7" box="[585,645,907,933]" name="India" pageId="1" pageNumber="290">India</collectingCountry>
, coll.
<collectorName id="26D25344FFACFFE1FD1089A03205D5D7" box="[711,806,907,934]" pageId="1" pageNumber="290">M. Bhat</collectorName>
,
<date id="FF991052FFACFFE1FCE789A732EAD5D7" box="[816,969,907,934]" pageId="1" pageNumber="290" value="2016-05-27">
<collectingDate id="EFDDE9BAFFACFFE1FCE789A732EAD5D7" box="[816,969,907,934]" pageId="1" pageNumber="290" value="2016-05-27">27 May 2016</collectingDate>
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.
<typeStatus id="549C8830FFACFFE1FC0589A03560D5D4" box="[978,1091,908,934]" pageId="1" pageNumber="290" type="paratype">Paratypes</typeStatus>
:
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(CL
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, CW 4.15.0 mm),
<materialsCitation id="3B4F3CCFFFACFFE1FEE28983342AD5BB" box="[309,1289,943,970]" collectingDate="2016-05-27" collectionCode="LFSC" collectorName="M. Bhat" country="India" latitude="15.658889" location="India" longLatPrecision="21" longitude="73.79195" pageId="1" pageNumber="290" specimenCount="8" specimenCount-female="8" typeStatus="paratype">
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(CL
<quantity id="4CDF9B77FFACFFE1FE0C89833347D5B8" box="[475,612,943,970]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.4" metricValueMin="4.4" pageId="1" pageNumber="290" unit="mm" value="4.9" valueMax="5.4" valueMin="4.4">4.45.4 mm</quantity>
, CW
<quantity id="4CDF9B77FFACFFE1FD7489833208D5BB" box="[675,811,943,970]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.1" metricValueMin="4.1" pageId="1" pageNumber="290" unit="mm" value="4.6" valueMax="5.1" valueMin="4.1">4.15.1 mm</quantity>
) (
<collectionCode id="ED36AE57FFACFFE1FC95898332AAD5BB" box="[834,905,943,969]" pageId="1" pageNumber="290">LFSC</collectionCode>
.ZRC-207), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8B983692FFACFFE1FF1089FF3430D3EF" blockId="1.[151,1437,871,2013]" pageId="1" pageNumber="290">
<emphasis id="B953EA80FFACFFE1FF1089FF30F0D59F" bold="true" box="[199,467,979,1005]" pageId="1" pageNumber="290">Comparative material</emphasis>
.
<emphasis id="B953EA80FFACFFE1FE3489FF32DCD59F" bold="true" box="[483,1023,979,1005]" pageId="1" pageNumber="290">
<taxonomicName id="4C274D11FFACFFE1FE3489FF32D9D59F" authority="(De Man, 1895)" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[483,1018,979,1005]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="290" phylum="Arthropoda" rank="species" species="pontianacense">
<emphasis id="B953EA80FFACFFE1FE3489FF3212D59F" bold="true" box="[483,817,979,1005]" italics="true" pageId="1" pageNumber="290">Nanosesarma pontianacense</emphasis>
(
<bibRefCitation id="EFB64B63FFACFFE1FC9489FF32D1D59F" author="De Man, J. G." box="[835,1010,979,1005]" pageId="1" pageNumber="290" pagination="485 - 609" refId="ref4620" refString="De Man, J. G. (1895) Berichte uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4), 485 - 609. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man, 1895</bibRefCitation>
)
</taxonomicName>
.
</emphasis>
<specimenCount id="9D21FD1BFFACFFE1FBDF89FF3575D59F" box="[1032,1110,979,1005]" count="1" pageId="1" pageNumber="290" type="male">1 male</specimenCount>
(CL
<quantity id="4CDF9B77FFACFFE1FB4289FF35CCD59C" box="[1173,1263,979,1006]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="290" unit="mm" value="5.1">5.1 mm</quantity>
, CW
<quantity id="4CDF9B77FFACFFE1FAE489FF34B3D59F" box="[1331,1424,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="290" unit="mm" value="4.7">4.7 mm</quantity>
);
<materialsCitation id="3B4F3CCFFFACFFE1FF4089DB3229D263" box="[151,778,1015,1042]" collectionCode="LFSC" pageId="1" pageNumber="290" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D21FD1BFFACFFE1FF4089DB31D9D263" box="[151,250,1015,1041]" count="1" pageId="1" pageNumber="290" type="female">1 female</specimenCount>
(CL
<quantity id="4CDF9B77FFACFFE1FEEE89DB30B0D260" box="[313,403,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="290" unit="mm" value="4.6">4.6 mm</quantity>
, CW
<quantity id="4CDF9B77FFACFFE1FE0089DB3313D263" box="[471,560,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="1" pageNumber="290" unit="mm" value="4.9">4.9 mm</quantity>
) (
<collectionCode id="ED36AE57FFACFFE1FD9E89DB33B3D263" box="[585,656,1015,1041]" pageId="1" pageNumber="290">LFSC</collectionCode>
.ZRC-208)
</materialsCitation>
,
<materialsCitation id="3B4F3CCFFFACFFE1FCC089DB31F1D20F" collectingDate="2018-07-10" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Chaiya District" location="Project Ban Laem Pho-Pho Sai" municipality="Phum Riang Subdistrict" pageId="1" pageNumber="290" specimenCode="ZRC 2018.0709" specimenCount="18" specimenCount-female="10" specimenCount-male="8" stateProvince="Surat Thani">
<specimenCount id="9D21FD1BFFACFFE1FCC089DB3251D263" box="[791,882,1015,1041]" count="8" pageId="1" pageNumber="290" type="male">8 males</specimenCount>
,
<specimenCount id="9D21FD1BFFACFFE1FC5789DB32DFD263" box="[896,1020,1015,1041]" count="10" pageId="1" pageNumber="290" type="female">10 females</specimenCount>
[3 ovigerous] (
<specimenCode id="DB819EE9FFACFFE1FB7989DB344FD263" box="[1198,1388,1015,1041]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="290">ZRC 2018.0709</specimenCode>
), on floating ropes and algae, mangroves, Forest-in-a-City
<location id="8EF86049FFACFFE1FCDE8E373557D247" LSID="urn:lsid:plazi:treatment:038E8784FFACFFE7FF4088D7353AD326:8EF86049FFACFFE1FCDE8E373557D247" box="[777,1140,1051,1078]" country="Thailand" county="Chaiya District" municipality="Phum Riang Subdistrict" name="Project Ban Laem Pho-Pho Sai" pageId="1" pageNumber="290" stateProvince="Surat Thani">Project Ban Laem Pho-Pho Sai</location>
,
<collectingMunicipality id="6BFCACE8FFACFFE1FB548E3734BBD247" box="[1155,1432,1051,1077]" pageId="1" pageNumber="290">Phum Riang Subdistrict</collectingMunicipality>
,
<collectingCounty id="62F94E1EFFACFFE1FF408E133060D22B" box="[151,323,1087,1113]" pageId="1" pageNumber="290">Chaiya District</collectingCounty>
,
<collectingRegion id="49E3F870FFACFFE1FE9A8E133318D22B" box="[333,571,1087,1113]" country="Thailand" name="Surat Thani" pageId="1" pageNumber="290">Surat Thani Province</collectingRegion>
,
<collectingCountry id="F3307602FFACFFE1FD928E13338FD22B" box="[581,684,1087,1113]" name="Thailand" pageId="1" pageNumber="290">Thailand</collectingCountry>
; 9˚2237.41”N 99˚1624.56”E, lower areas, coll.
<collectorName id="26D25344FFACFFE1FB078E6C3462D228" box="[1232,1345,1088,1114]" pageId="1" pageNumber="290">P.K.L. Ng</collectorName>
,
<date id="FF991052FFACFFE1FA9B8E1331F1D20F" pageId="1" pageNumber="290" value="2018-07-10">
<collectingDate id="EFDDE9BAFFACFFE1FA9B8E1331F1D20F" pageId="1" pageNumber="290" value="2018-07-10">10 July 2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B4F3CCFFFACFFE1FF088E4F34B4D20F" box="[223,1431,1123,1150]" collectingDate="2019-09-24" collectionCode="ZRC" collectorName="Tai O &amp; L. M. Tsang" country="Hong Kong" location="western Lantau Island" pageId="1" pageNumber="290" specimenCode="ZRC 2019.1106" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D21FD1BFFACFFE1FF088E4F3009D20F" box="[223,298,1123,1149]" count="1" pageId="1" pageNumber="290" type="male">1 male</specimenCount>
(
<specimenCode id="DB819EE9FFACFFE1FEED8E4F30D6D20F" box="[314,501,1123,1149]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="290">ZRC 2019.1106</specimenCode>
),
<collectorName id="26D25344FFACFFE1FDDF8E4F336AD20F" box="[520,585,1123,1149]" pageId="1" pageNumber="290">Tai O</collectorName>
,
<location id="8EF86049FFACFFE1FD818E483272D20F" LSID="urn:lsid:plazi:treatment:038E8784FFACFFE7FF4088D7353AD326:8EF86049FFACFFE1FD818E483272D20F" box="[598,849,1123,1150]" country="Hong Kong" name="western Lantau Island" pageId="1" pageNumber="290">western Lantau Island</location>
,
<collectingCountry id="F3307602FFACFFE1FC8A8E4832C7D20C" box="[861,996,1124,1150]" name="Hong Kong" pageId="1" pageNumber="290">Hong Kong</collectingCountry>
, coll.
<collectorName id="26D25344FFACFFE1FBFE8E48358ED20C" box="[1065,1197,1124,1150]" pageId="1" pageNumber="290">L.M. Tsang</collectorName>
,
<date id="FF991052FFACFFE1FB6E8E4F34B4D20F" box="[1209,1431,1123,1149]" pageId="1" pageNumber="290" value="2019-09-24">
<collectingDate id="EFDDE9BAFFACFFE1FB6E8E4F34B4D20F" box="[1209,1431,1123,1149]" pageId="1" pageNumber="290" value="2019-09-24">24 September 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B4F3CCFFFACFFE1FF408EAB3068D2B7" collectingDate="2014-03-13" collectionCode="ZRC" collectorName="K. Wong" country="Hong Kong" location="Tung Chung" pageId="1" pageNumber="290" specimenCode="ZRC 2019.1717" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<specimenCount id="9D21FD1BFFACFFE1FF408EAB31C5D2D3" box="[151,230,1159,1185]" count="1" pageId="1" pageNumber="290" type="male">1 male</specimenCount>
,
<specimenCount id="9D21FD1BFFACFFE1FF248EAB3043D2D3" box="[243,352,1159,1185]" count="3" pageId="1" pageNumber="290" type="female">3 females</specimenCount>
(
<specimenCode id="DB819EE9FFACFFE1FEB88EAB3308D2D3" box="[367,555,1159,1185]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="290">ZRC 2019.1717</specimenCode>
), among oyster clumps, site muddy,
<location id="8EF86049FFACFFE1FC1F8EA43574D2D3" LSID="urn:lsid:plazi:treatment:038E8784FFACFFE7FF4088D7353AD326:8EF86049FFACFFE1FC1F8EA43574D2D3" box="[968,1111,1159,1186]" country="Hong Kong" name="Tung Chung" pageId="1" pageNumber="290">Tung Chung</location>
,
<collectingCountry id="F3307602FFACFFE1FBB38EA435C8D2D0" box="[1124,1259,1160,1186]" name="Hong Kong" pageId="1" pageNumber="290">Hong Kong</collectingCountry>
, coll.
<collectorName id="26D25344FFACFFE1FAE78EA434BBD2D0" box="[1328,1432,1160,1186]" pageId="1" pageNumber="290">K. Wong</collectorName>
,
<date id="FF991052FFACFFE1FF408E873068D2B7" box="[151,331,1195,1221]" pageId="1" pageNumber="290" value="2014-03-13">
<collectingDate id="EFDDE9BAFFACFFE1FF408E873068D2B7" box="[151,331,1195,1221]" pageId="1" pageNumber="290" value="2014-03-13">13 March 2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B4F3CCFFFACFFE1FE8F8E873037D29B" collectingDate="2001-04" collectionCode="ZRC" collectorName="B. C. Jayne" country="Singapore" location="Pandan" pageId="1" pageNumber="290" specimenCode="ZRC 2001.2281" specimenCount="1">
1 ovigerous female (
<specimenCode id="DB819EE9FFACFFE1FD938E873222D2B7" box="[580,769,1195,1221]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="290">ZRC 2001.2281</specimenCode>
),
<location id="8EF86049FFACFFE1FCC08E87324AD2B7" LSID="urn:lsid:plazi:treatment:038E8784FFACFFE7FF4088D7353AD326:8EF86049FFACFFE1FCC08E87324AD2B7" box="[791,873,1195,1221]" country="Singapore" name="Pandan" pageId="1" pageNumber="290">Pandan</location>
mangrove, southern
<collectingCountry id="F3307602FFACFFE1FB8B8E8735F3D2B7" box="[1116,1232,1195,1221]" name="Singapore" pageId="1" pageNumber="290">Singapore</collectingCountry>
, coll.
<collectorName id="26D25344FFACFFE1FACE8E8734BBD2B4" box="[1305,1432,1195,1222]" pageId="1" pageNumber="290">B.C. Jayne</collectorName>
,
<date id="FF991052FFACFFE1FF408EE33037D29B" box="[151,276,1231,1257]" pageId="1" pageNumber="290" value="2001-04">
<collectingDate id="EFDDE9BAFFACFFE1FF408EE33037D29B" box="[151,276,1231,1257]" pageId="1" pageNumber="290" value="2001-04">April 2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B4F3CCFFFACFFE1FEC88EE3338CD37F" collectingDate="2002-06-02" collectionCode="ZRC" collectorName="Y. S. Oei" country="Singapore" location="East Coast" municipality="Victoria Junior College" pageId="1" pageNumber="290" specimenCode="ZRC 2011.0584" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D21FD1BFFACFFE1FEC88EE330A9D29B" box="[287,394,1231,1257]" count="2" pageId="1" pageNumber="290" type="female">2 females</specimenCount>
[broken, poor condition] (
<specimenCode id="DB819EE9FFACFFE1FD788EE3324AD298" box="[687,873,1231,1258]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="290">ZRC 2011.0584</specimenCode>
), canal in
<location id="8EF86049FFACFFE1FC0F8EFC3572D29B" LSID="urn:lsid:plazi:treatment:038E8784FFACFFE7FF4088D7353AD326:8EF86049FFACFFE1FC0F8EFC3572D29B" box="[984,1105,1231,1258]" country="Singapore" municipality="Victoria Junior College" name="East Coast" pageId="1" pageNumber="290">East Coast</location>
, near
<collectingMunicipality id="6BFCACE8FFACFFE1FB468EE334BBD29B" box="[1169,1432,1231,1257]" pageId="1" pageNumber="290">Victoria Junior College</collectingMunicipality>
, eastern
<collectingCountry id="F3307602FFACFFE1FF278EDF3047D37F" box="[240,356,1267,1293]" name="Singapore" pageId="1" pageNumber="290">Singapore</collectingCountry>
, coll.
<collectorName id="26D25344FFACFFE1FE7B8EDF332CD37F" box="[428,527,1267,1293]" pageId="1" pageNumber="290">Y.S. Oei</collectorName>
,
<date id="FF991052FFACFFE1FDC98EDF338CD37F" box="[542,687,1267,1294]" pageId="1" pageNumber="290" value="2002-06-02">
<collectingDate id="EFDDE9BAFFACFFE1FDC98EDF338CD37F" box="[542,687,1267,1294]" pageId="1" pageNumber="290" value="2002-06-02">2 June 2002</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B4F3CCFFFACFFE1FD6B8EDF32A8D343" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; L. W. H. Tan" country="Singapore" location="Lim Chu Kang" pageId="1" pageNumber="290" specimenCode="ZRC 1999.1166" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D21FD1BFFACFFE1FD6B8EDF322ED37F" box="[700,781,1267,1293]" count="1" pageId="1" pageNumber="290" type="male">1 male</specimenCount>
,
<specimenCount id="9D21FD1BFFACFFE1FCCC8EDF325CD37F" box="[795,895,1267,1293]" count="1" pageId="1" pageNumber="290" type="female">1 female</specimenCount>
(
<specimenCode id="DB819EE9FFACFFE1FC468EDF356DD37F" box="[913,1102,1267,1293]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="290">ZRC 1999.1166</specimenCode>
),
<location id="8EF86049FFACFFE1FBB48EDF3432D37C" LSID="urn:lsid:plazi:treatment:038E8784FFACFFE7FF4088D7353AD326:8EF86049FFACFFE1FBB48EDF3432D37C" box="[1123,1297,1267,1294]" country="Singapore" name="Lim Chu Kang" pageId="1" pageNumber="290">Lim Chu Kang</location>
mangroves, northern
<collectingCountry id="F3307602FFACFFE1FF2C8F3B304CD343" box="[251,367,1303,1329]" name="Singapore" pageId="1" pageNumber="290">Singapore</collectingCountry>
, coll.
<collectorName id="26D25344FFACFFE1FE798F34333FD340" box="[430,540,1304,1330]" pageId="1" pageNumber="290">P.K.L. Ng</collectorName>
&amp;
<collectorName id="26D25344FFACFFE1FDEB8F34339DD340" box="[572,702,1304,1330]" pageId="1" pageNumber="290">L.W.H. Tan</collectorName>
, 21
<collectorName id="26D25344FFACFFE1FD3E8F3B3268D343" box="[745,843,1303,1329]" pageId="1" pageNumber="290">Octiober</collectorName>
1987
</materialsCitation>
;
<materialsCitation id="3B4F3CCFFFACFFE1FC438F3B32A2D327" collectingDate="2011-02-24" collectionCode="ZRC" collectorName="B. Y. Lee" country="Singapore" location="Sarimbun" pageId="1" pageNumber="290" specimenCode="ZRC 2012.0299" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D21FD1BFFACFFE1FC438F3B32FED343" box="[916,989,1303,1329]" count="1" pageId="1" pageNumber="290" type="male">1 male</specimenCount>
(
<specimenCode id="DB819EE9FFACFFE1FC3D8F3B3580D343" box="[1002,1187,1303,1329]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="290">ZRC 2012.0299</specimenCode>
),
<location id="8EF86049FFACFFE1FB648F3B3403D343" LSID="urn:lsid:plazi:treatment:038E8784FFACFFE7FF4088D7353AD326:8EF86049FFACFFE1FB648F3B3403D343" box="[1203,1312,1303,1329]" country="Singapore" name="Sarimbun" pageId="1" pageNumber="290">Sarimbun</location>
mangrove, northwestern
<collectingCountry id="F3307602FFACFFE1FEE38F17308BD327" box="[308,424,1339,1365]" name="Singapore" pageId="1" pageNumber="290">Singapore</collectingCountry>
, coll.
<collectorName id="26D25344FFACFFE1FE248F10337AD324" box="[499,601,1340,1366]" pageId="1" pageNumber="290">B.Y. Lee</collectorName>
<emphasis id="B953EA80FFACFFE1FDB48F1033BCD327" box="[611,671,1339,1365]" italics="true" pageId="1" pageNumber="290">et al.</emphasis>
,
<date id="FF991052FFACFFE1FD678F1732A2D327" box="[688,897,1339,1365]" pageId="1" pageNumber="290" value="2011-02-24">
<collectingDate id="EFDDE9BAFFACFFE1FD678F1732A2D327" box="[688,897,1339,1365]" pageId="1" pageNumber="290" value="2011-02-24">24 February 2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B4F3CCFFFACFFE1FC478F1732ECD30B" collectingDate="2011-11-16" collectionCode="ZRC" collectorName="B. Y. Lee" country="Singapore" location="Kranji" pageId="1" pageNumber="290" specimenCode="ZRC 2012.0239" specimenCount="1">
1 ovigerous female (
<specimenCode id="DB819EE9FFACFFE1FB578F17341CD327" box="[1152,1343,1339,1365]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="290">ZRC 2012.0239</specimenCode>
),
<location id="8EF86049FFACFFE1FA828F1734BED327" LSID="urn:lsid:plazi:treatment:038E8784FFACFFE7FF4088D7353AD326:8EF86049FFACFFE1FA828F1734BED327" box="[1365,1437,1339,1365]" country="Singapore" name="Kranji" pageId="1" pageNumber="290">Kranji</location>
mangrove, northern
<collectingCountry id="F3307602FFACFFE1FEA88F7330D0D30B" box="[383,499,1375,1401]" name="Singapore" pageId="1" pageNumber="290">Singapore</collectingCountry>
, coll.
<collectorName id="26D25344FFACFFE1FDED8F4C33BDD308" box="[570,670,1376,1402]" pageId="1" pageNumber="290">B.Y. Lee</collectorName>
<emphasis id="B953EA80FFACFFE1FD708F4C33C2D30B" box="[679,737,1375,1401]" italics="true" pageId="1" pageNumber="290">et al.</emphasis>
,
<date id="FF991052FFACFFE1FD268F7332ECD30B" box="[753,975,1375,1401]" pageId="1" pageNumber="290" value="2011-11-16">
<collectingDate id="EFDDE9BAFFACFFE1FD268F7332ECD30B" box="[753,975,1375,1401]" pageId="1" pageNumber="290" value="2011-11-16">16 November 2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B4F3CCFFFACFFE1FC0A8F73342DD3EF" collectingDate="2011-09-27" collectionCode="ZRC" collectorName="B. Y. Lee" country="Singapore" location="Tuas" pageId="1" pageNumber="290" specimenCode="ZRC 2012.0300" specimenCount="6" specimenCount-female="4" specimenCount-male="2">
<specimenCount id="9D21FD1BFFACFFE1FC0A8F73351BD30B" box="[989,1080,1375,1401]" count="2" pageId="1" pageNumber="290" type="male">2 males</specimenCount>
,
<specimenCount id="9D21FD1BFFACFFE1FB928F733590D30B" box="[1093,1203,1375,1401]" count="4" pageId="1" pageNumber="290" type="female">4 females</specimenCount>
[3 ovigerous] (
<specimenCode id="DB819EE9FFACFFE1FAB38F733030D3EF" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="290">ZRC 2012.0300</specimenCode>
), small patch mangrove,
<location id="8EF86049FFACFFE1FDFD8FA83341D3EC" LSID="urn:lsid:plazi:treatment:038E8784FFACFFE7FF4088D7353AD326:8EF86049FFACFFE1FDFD8FA83341D3EC" box="[554,610,1412,1438]" country="Singapore" name="Tuas" pageId="1" pageNumber="290">Tuas</location>
, western
<collectingCountry id="F3307602FFACFFE1FD1B8FAF3263D3EF" box="[716,832,1411,1437]" name="Singapore" pageId="1" pageNumber="290">Singapore</collectingCountry>
, coll.
<collectorName id="26D25344FFACFFE1FC548FA832C5D3EC" box="[899,998,1412,1438]" pageId="1" pageNumber="290">B.Y. Lee</collectorName>
<emphasis id="B953EA80FFACFFE1FC3A8FA8353DD3EF" box="[1005,1054,1411,1437]" italics="true" pageId="1" pageNumber="290">et al</emphasis>
.,
<date id="FF991052FFACFFE1FBE48FAF342DD3EF" box="[1075,1294,1411,1437]" pageId="1" pageNumber="290" value="2011-09-27">
<collectingDate id="EFDDE9BAFFACFFE1FBE48FAF342DD3EF" box="[1075,1294,1411,1437]" pageId="1" pageNumber="290" value="2011-09-27">27 September 2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6519FFACFFE4FF108F8B32E5D189" lastPageId="4" lastPageNumber="293" pageId="1" pageNumber="290" type="description">
<paragraph id="8B983692FFACFFE1FF108F8B3023D1AF" blockId="1.[151,1437,871,2013]" pageId="1" pageNumber="290">
<emphasis id="B953EA80FFACFFE1FF108F8B307AD3B3" bold="true" box="[199,345,1447,1473]" pageId="1" pageNumber="290">Description.</emphasis>
Carapace longer than wide; dorsal surface thickly setose with conspicuous rugosities demarcating regions; lateral margins almost parallel, curving gently along posterolateral margin to posterior carapace margin (
<figureCitation id="131C2A17FFACFFE1FF488FC33038D078" box="[159,283,1519,1546]" captionStart="FIGURE 1" captionStartId="2.[151,250,1375,1399]" captionTargetBox="[151,1436,181,1351]" captionTargetId="figure-21@2.[151,1436,181,1351]" captionTargetPageId="2" captionText="FIGURE 1. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live N. gomantaka n. sp., in oyster shell; g, live N. gomantaka n. sp., on rotten wood." figureDoi="http://doi.org/10.5281/zenodo.10954093" httpUri="https://zenodo.org/record/10954093/files/figure.png" pageId="1" pageNumber="290">Figs. 1a, c</figureCitation>
;
<figureCitation id="131C2A17FFACFFE1FEFD8FC33064D07B" box="[298,327,1519,1545]" captionStart="FIGURE 2" captionStartId="3.[151,250,1312,1336]" captionTargetBox="[167,1420,184,1286]" captionTargetId="figure-17@3.[164,1423,181,1288]" captionTargetPageId="3" captionText="FIGURE 2. Nanosesarma gomantaka n. sp., a, c, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. b, d, N. pontianacense (De Man, 1895), male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, b, carapace, dorsal view; c, d, male pleon; e, female pleon." figureDoi="http://doi.org/10.5281/zenodo.10954095" httpUri="https://zenodo.org/record/10954095/files/figure.png" pageId="1" pageNumber="290">2a</figureCitation>
). External orbital angle followed by rudimentary epibranchial tooth, tip of external orbital angle not extending beyond tip of rudimentary epibranchial tooth. Frontal margin wide, obliquely directed, distinctly bilobed, clearly in advance of level of orbits, frontal lobes separated by deep V-shaped depression (
<figureCitation id="131C2A17FFACFFE1FACD8C1B34B4D020" box="[1306,1431,1591,1618]" captionStart="FIGURE 1" captionStartId="2.[151,250,1375,1399]" captionTargetBox="[151,1436,181,1351]" captionTargetId="figure-21@2.[151,1436,181,1351]" captionTargetPageId="2" captionText="FIGURE 1. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live N. gomantaka n. sp., in oyster shell; g, live N. gomantaka n. sp., on rotten wood." figureDoi="http://doi.org/10.5281/zenodo.10954093" httpUri="https://zenodo.org/record/10954093/files/figure.png" pageId="1" pageNumber="290">Figs. 1a, c</figureCitation>
;
<figureCitation id="131C2A17FFACFFE1FF408C773196D007" box="[151,181,1627,1653]" captionStart="FIGURE 2" captionStartId="3.[151,250,1312,1336]" captionTargetBox="[167,1420,184,1286]" captionTargetId="figure-17@3.[164,1423,181,1288]" captionTargetPageId="3" captionText="FIGURE 2. Nanosesarma gomantaka n. sp., a, c, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. b, d, N. pontianacense (De Man, 1895), male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, b, carapace, dorsal view; c, d, male pleon; e, female pleon." figureDoi="http://doi.org/10.5281/zenodo.10954095" httpUri="https://zenodo.org/record/10954095/files/figure.png" pageId="1" pageNumber="290">2a</figureCitation>
). Antennular fossae transversely ovate. Antennules with 7 articles, folding transversely, tip flagellate. Antennae long, cylindrical, flagellum located in orbital hiatus, extends beyond ocular peduncle (
<figureCitation id="131C2A17FFACFFE1FBAA8C5335F1D0EB" box="[1149,1234,1663,1689]" captionStart="FIGURE 6" captionStartId="7.[151,250,547,571]" captionTargetBox="[151,1435,181,523]" captionTargetId="figure-17@7.[151,1435,181,523]" captionTargetPageId="7" captionText="FIGURE 6 Nanosesarma gomantaka n. sp., paratype male (CL 5.5 mm, CW 5.1 mm) (LFSC.ZRC-207), Chapora Estuary, Goa State, India. a, frontal view of cephalothorax; b, thoracic sternites 15 and sternopleonal cavity." figureDoi="http://doi.org/10.5281/zenodo.10954105" httpUri="https://zenodo.org/record/10954105/files/figure.png" pageId="1" pageNumber="290">Fig. 6a</figureCitation>
). Orbits rounded, supraorbital margin weakly granular, inner supraorbital angle continuous with frontal margin; infraorbital margin notched under external orbital angle, followed by granular ridge extending below basal antennal segment. Ocular peduncle short, stout, slightly setose, cornea terminal (
<figureCitation id="131C2A17FFACFFE1FCD68CC73277D177" box="[769,852,1771,1797]" captionStart="FIGURE 2" captionStartId="3.[151,250,1312,1336]" captionTargetBox="[167,1420,184,1286]" captionTargetId="figure-17@3.[164,1423,181,1288]" captionTargetPageId="3" captionText="FIGURE 2. Nanosesarma gomantaka n. sp., a, c, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. b, d, N. pontianacense (De Man, 1895), male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, b, carapace, dorsal view; c, d, male pleon; e, female pleon." figureDoi="http://doi.org/10.5281/zenodo.10954095" httpUri="https://zenodo.org/record/10954095/files/figure.png" pageId="1" pageNumber="290">Fig. 2a</figureCitation>
). Buccal cavern wider than long, wider posteriorly; epistome wide, concave, sparsely setose, posterior margin with triangular median lobe and 2 convex lateral lobes, endostomial ridges well-developed. Pterygostomian region covered with reticulate pattern of setae. Third maxillipeds with distinct rhomboidal gap exposing the inner mouth parts when closed; ischium slightly longer than wide, with oblique setose ridge; ischium slightly longer than merus; merus almost as wide as long, with oblique setose ridge; palp slightly shorter than merus, inserted anteroexternally on merus; exopod slender, flattened, with long flagellum (
<figureCitation id="131C2A17FFACFFE1FF488DEF31D0D1AF" box="[159,243,1987,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1550,1574]" captionTargetBox="[154,1432,184,1523]" captionTargetId="figure-21@4.[151,1435,181,1526]" captionTargetPageId="4" captionText="FIGURE 3. Nanosesarma gomantaka n. sp., a, b, d, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, left chela, outer view (setae omitted); b, left chela, dorsal view; d, third maxilliped, dorsal view; e, left vulva." figureDoi="http://doi.org/10.5281/zenodo.10954097" httpUri="https://zenodo.org/record/10954097/files/figure.png" pageId="1" pageNumber="290">Fig. 3d</figureCitation>
).
</paragraph>
<caption id="DF58661AFFAFFFE2FF408F7333B6D391" ID-DOI="http://doi.org/10.5281/zenodo.10954093" ID-Zenodo-Dep="10954093" httpUri="https://zenodo.org/record/10954093/files/figure.png" pageId="2" pageNumber="291" startId="2.[151,250,1375,1399]" targetBox="[151,1436,181,1351]" targetPageId="2" targetType="figure">
<paragraph id="8B983692FFAFFFE2FF408F7333B6D391" blockId="2.[151,1437,1375,1508]" pageId="2" pageNumber="291">
<emphasis id="B953EA80FFAFFFE2FF408F733031D305" bold="true" box="[151,274,1375,1399]" pageId="2" pageNumber="291">FIGURE 1.</emphasis>
<taxonomicName id="4C274D11FFAFFFE2FEC08F4C333AD305" authority="Trivedi &amp; Padate &amp; Ng, 2024" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[279,537,1375,1399]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="291" phylum="Arthropoda" rank="species" species="gomantaka" status="sp. nov.">
<emphasis id="B953EA80FFAFFFE2FEC08F4C333AD305" box="[279,537,1375,1399]" italics="true" pageId="2" pageNumber="291">Nanosesarma gomantaka</emphasis>
</taxonomicName>
<emphasis id="B953EA80FFAFFFE2FDCA8F733376D305" bold="true" box="[541,597,1375,1399]" pageId="2" pageNumber="291">
<taxonomicNameLabel id="A26057FBFFAFFFE2FDCA8F733376D305" box="[541,597,1375,1399]" pageId="2" pageNumber="291" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live
<taxonomicName id="4C274D11FFAFFFE2FBBD8F8435DAD3CD" authority="Trivedi &amp; Padate &amp; Ng, 2024" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[1130,1273,1447,1471]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="291" phylum="Arthropoda" rank="species" species="gomantaka" status="n. sp.">
<emphasis id="B953EA80FFAFFFE2FBBD8F8435DAD3CD" box="[1130,1273,1447,1471]" italics="true" pageId="2" pageNumber="291">N. gomantaka</emphasis>
</taxonomicName>
<emphasis id="B953EA80FFAFFFE2FB298F8B341ED3CD" bold="true" box="[1278,1341,1447,1471]" pageId="2" pageNumber="291">
<taxonomicNameLabel id="A26057FBFFAFFFE2FB298F8B341BD3CD" box="[1278,1336,1447,1471]" pageId="2" pageNumber="291" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
in oyster shell; g, live
<taxonomicName id="4C274D11FFAFFFE2FECD8FE0308AD391" authority="Trivedi &amp; Padate &amp; Ng, 2024" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[282,425,1483,1507]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="291" phylum="Arthropoda" rank="species" species="gomantaka" status="n. sp.">
<emphasis id="B953EA80FFAFFFE2FECD8FE0308AD391" box="[282,425,1483,1507]" italics="true" pageId="2" pageNumber="291">N. gomantaka</emphasis>
</taxonomicName>
<emphasis id="B953EA80FFAFFFE2FE788FE730CBD391" bold="true" box="[431,488,1483,1507]" pageId="2" pageNumber="291">
<taxonomicNameLabel id="A26057FBFFAFFFE2FE788FE730CBD391" box="[431,488,1483,1507]" pageId="2" pageNumber="291" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, on rotten wood.
</paragraph>
</caption>
<paragraph id="8B983692FFAFFFE2FF108C1932DFD1A9" blockId="2.[151,1437,1589,2012]" pageId="2" pageNumber="291">
Male chelipeds subequal, stout, almost as long as carapace (
<figureCitation id="131C2A17FFAFFFE2FC8A8C1932EBD03D" box="[861,968,1589,1615]" captionStart="FIGURE 1" captionStartId="2.[151,250,1375,1399]" captionTargetBox="[151,1436,181,1351]" captionTargetId="figure-21@2.[151,1436,181,1351]" captionTargetPageId="2" captionText="FIGURE 1. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live N. gomantaka n. sp., in oyster shell; g, live N. gomantaka n. sp., on rotten wood." figureDoi="http://doi.org/10.5281/zenodo.10954093" httpUri="https://zenodo.org/record/10954093/files/figure.png" pageId="2" pageNumber="291">Fig. 1a, b</figureCitation>
). Female chelipeds subequal, smaller than male, almost as long as or shorter than carapace (
<figureCitation id="131C2A17FFAFFFE2FD608C753200D001" box="[695,803,1625,1651]" captionStart="FIGURE 1" captionStartId="2.[151,250,1375,1399]" captionTargetBox="[151,1436,181,1351]" captionTargetId="figure-21@2.[151,1436,181,1351]" captionTargetPageId="2" captionText="FIGURE 1. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live N. gomantaka n. sp., in oyster shell; g, live N. gomantaka n. sp., on rotten wood." figureDoi="http://doi.org/10.5281/zenodo.10954093" httpUri="https://zenodo.org/record/10954093/files/figure.png" pageId="2" pageNumber="291">Fig. 1c, d</figureCitation>
). Merus with setose granular striations on dorsal surface. Anterior (inner) surface of merus with 3 minute denticles on lower margin. Carpus with setose granular striations on dorsal surface. Palm short, stout, dorsal surface granular, thickly setose, pectinate crest absent; outer surface with granules in reticulated pattern and 2 rows of granules roughly parallel to ventral granular ridge originating from pollex extending posteriorly across palm; inner surface with 2 rows of large granules. Fingers (dactylus and pollex) stout, curved, with rounded spatulate tips, gaping when closed; dactylus with smooth median longitudinal ridge and proximal granules on dorsal surface, outer surface with strong proximal spine (
<figureCitation id="131C2A17FFAFFFE2FBAC8D1D35EDD139" box="[1147,1230,1841,1867]" captionStart="FIGURE 3" captionStartId="4.[151,250,1550,1574]" captionTargetBox="[154,1432,184,1523]" captionTargetId="figure-21@4.[151,1435,181,1526]" captionTargetPageId="4" captionText="FIGURE 3. Nanosesarma gomantaka n. sp., a, b, d, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, left chela, outer view (setae omitted); b, left chela, dorsal view; d, third maxilliped, dorsal view; e, left vulva." figureDoi="http://doi.org/10.5281/zenodo.10954097" httpUri="https://zenodo.org/record/10954097/files/figure.png" pageId="2" pageNumber="291">Fig. 3b</figureCitation>
). Occlusal surface of dactylus with 3 large blunt teeth; occlusal surface of pollex with 1 sub-distal tooth, 1 large median tooth, and 2 proximal molariform teeth (
<figureCitation id="131C2A17FFAFFFE2FE3C8D55331DD1E1" box="[491,574,1913,1939]" captionStart="FIGURE 3" captionStartId="4.[151,250,1550,1574]" captionTargetBox="[154,1432,184,1523]" captionTargetId="figure-21@4.[151,1435,181,1526]" captionTargetPageId="4" captionText="FIGURE 3. Nanosesarma gomantaka n. sp., a, b, d, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, left chela, outer view (setae omitted); b, left chela, dorsal view; d, third maxilliped, dorsal view; e, left vulva." figureDoi="http://doi.org/10.5281/zenodo.10954097" httpUri="https://zenodo.org/record/10954097/files/figure.png" pageId="2" pageNumber="291">Fig. 3a</figureCitation>
). Female cheliped dactylus short, no proximal spine, dorsal surface without ridges, outer surface with granules in reticulated pattern and 1 row of granules roughly parallel to ventral granular ridge originating from pollex and extending posteriorly across palm (
<figureCitation id="131C2A17FFAFFFE2FC498DED32CDD1A9" box="[926,1006,1985,2011]" captionStart="FIGURE 3" captionStartId="4.[151,250,1550,1574]" captionTargetBox="[154,1432,184,1523]" captionTargetId="figure-21@4.[151,1435,181,1526]" captionTargetPageId="4" captionText="FIGURE 3. Nanosesarma gomantaka n. sp., a, b, d, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, left chela, outer view (setae omitted); b, left chela, dorsal view; d, third maxilliped, dorsal view; e, left vulva." figureDoi="http://doi.org/10.5281/zenodo.10954097" httpUri="https://zenodo.org/record/10954097/files/figure.png" pageId="2" pageNumber="291">Fig. 3c</figureCitation>
).
</paragraph>
<caption id="DF58661AFFAEFFE3FF408F0C3094D3D6" ID-DOI="http://doi.org/10.5281/zenodo.10954095" ID-Zenodo-Dep="10954095" httpUri="https://zenodo.org/record/10954095/files/figure.png" pageId="3" pageNumber="292" startId="3.[151,250,1312,1336]" targetBox="[167,1420,184,1286]" targetPageId="3" targetType="figure">
<paragraph id="8B983692FFAEFFE3FF408F0C3094D3D6" blockId="3.[151,1437,1312,1444]" pageId="3" pageNumber="292">
<emphasis id="B953EA80FFAEFFE3FF408F0C3031D34A" bold="true" box="[151,274,1312,1336]" pageId="3" pageNumber="292">FIGURE 2.</emphasis>
<taxonomicName id="4C274D11FFAEFFE3FEC08F0D333AD34A" authority="Trivedi &amp; Padate &amp; Ng, 2024" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[279,537,1312,1336]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="gomantaka" status="sp. nov.">
<emphasis id="B953EA80FFAEFFE3FEC08F0D333AD34A" box="[279,537,1312,1336]" italics="true" pageId="3" pageNumber="292">Nanosesarma gomantaka</emphasis>
</taxonomicName>
<emphasis id="B953EA80FFAEFFE3FDC98F0C3375D34A" bold="true" box="[542,598,1312,1336]" pageId="3" pageNumber="292">
<taxonomicNameLabel id="A26057FBFFAEFFE3FDC98F0C3375D34A" box="[542,598,1312,1336]" pageId="3" pageNumber="292" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, a, c, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. b, d,
<taxonomicName id="4C274D11FFAEFFE3FA528F6930F7D3F2" authority="(De Man, 1895)" baseAuthorityName="De Man" baseAuthorityYear="1895" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="pontianacense">
<emphasis id="B953EA80FFAEFFE3FA528F69300AD3F2" italics="true" pageId="3" pageNumber="292">N. pontianacense</emphasis>
(
<bibRefCitation id="EFB64B63FFAEFFE3FEE08F4430EFD3F2" author="De Man, J. G." box="[311,460,1384,1408]" pageId="3" pageNumber="292" pagination="485 - 609" refId="ref4620" refString="De Man, J. G. (1895) Berichte uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4), 485 - 609. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man, 1895</bibRefCitation>
)
</taxonomicName>
, male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, b, carapace, dorsal view; c, d, male pleon; e, female pleon.
</paragraph>
</caption>
<paragraph id="8B983692FFAEFFE3FF108FC535E3D01D" blockId="3.[151,1437,1513,2044]" pageId="3" pageNumber="292">
Thoracic sternum wide in both sexes, with smooth, finely pitted ventral surface; male sternites 1 and 2 completely fused, forming broad triangular structure with convex lateral margins; distinct suture present between sternites 3 and 4, lined with granules; sternites 3 and 4 completely fused; sternopleonal cavity extends to suture between sternites 2 and 3, margin of cavity on sternites 3 and 4 cristate, lined with low granules (
<figureCitation id="131C2A17FFAEFFE3FBD98C7935ABD01D" box="[1038,1160,1621,1647]" captionStart="FIGURE 1" captionStartId="2.[151,250,1375,1399]" captionTargetBox="[151,1436,181,1351]" captionTargetId="figure-21@2.[151,1436,181,1351]" captionTargetPageId="2" captionText="FIGURE 1. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live N. gomantaka n. sp., in oyster shell; g, live N. gomantaka n. sp., on rotten wood." figureDoi="http://doi.org/10.5281/zenodo.10954093" httpUri="https://zenodo.org/record/10954093/files/figure.png" pageId="3" pageNumber="292">Figs. 1b, d</figureCitation>
;
<figureCitation id="131C2A17FFAEFFE3FB438C793590D01D" box="[1172,1203,1621,1647]" captionStart="FIGURE 6" captionStartId="7.[151,250,547,571]" captionTargetBox="[151,1435,181,523]" captionTargetId="figure-17@7.[151,1435,181,523]" captionTargetPageId="7" captionText="FIGURE 6 Nanosesarma gomantaka n. sp., paratype male (CL 5.5 mm, CW 5.1 mm) (LFSC.ZRC-207), Chapora Estuary, Goa State, India. a, frontal view of cephalothorax; b, thoracic sternites 15 and sternopleonal cavity." figureDoi="http://doi.org/10.5281/zenodo.10954105" httpUri="https://zenodo.org/record/10954105/files/figure.png" pageId="3" pageNumber="292">6b</figureCitation>
).
</paragraph>
<paragraph id="8B983692FFAEFFE3FF108C5530A3D08D" blockId="3.[151,1437,1513,2044]" pageId="3" pageNumber="292">
Male pleon narrow, triangular, with 6 free somites and telson; surface pitted; lateral margins finely granulated, densely setose; telson longer than wide with anterior surface round; somite 6, 0.6 times wider than long (
<figureCitation id="131C2A17FFAEFFE3FAEE8CB134B4D0C5" box="[1337,1431,1693,1719]" captionStart="FIGURE 1" captionStartId="2.[151,250,1375,1399]" captionTargetBox="[151,1436,181,1351]" captionTargetId="figure-21@2.[151,1436,181,1351]" captionTargetPageId="2" captionText="FIGURE 1. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live N. gomantaka n. sp., in oyster shell; g, live N. gomantaka n. sp., on rotten wood." figureDoi="http://doi.org/10.5281/zenodo.10954093" httpUri="https://zenodo.org/record/10954093/files/figure.png" pageId="3" pageNumber="292">Figs. 1b</figureCitation>
;
<figureCitation id="131C2A17FFAEFFE3FF408CED3196D0A9" box="[151,181,1729,1755]" captionStart="FIGURE 2" captionStartId="3.[151,250,1312,1336]" captionTargetBox="[167,1420,184,1286]" captionTargetId="figure-17@3.[164,1423,181,1288]" captionTargetPageId="3" captionText="FIGURE 2. Nanosesarma gomantaka n. sp., a, c, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. b, d, N. pontianacense (De Man, 1895), male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, b, carapace, dorsal view; c, d, male pleon; e, female pleon." figureDoi="http://doi.org/10.5281/zenodo.10954095" httpUri="https://zenodo.org/record/10954095/files/figure.png" pageId="3" pageNumber="292">2c</figureCitation>
). Female pleon wide, rounded, with 6 free somites and telson; telson rounded, wider than long; margins densely setose (
<figureCitation id="131C2A17FFAEFFE3FF3C8CC9306AD08D" box="[235,329,1765,1791]" captionStart="FIGURE 1" captionStartId="2.[151,250,1375,1399]" captionTargetBox="[151,1436,181,1351]" captionTargetId="figure-21@2.[151,1436,181,1351]" captionTargetPageId="2" captionText="FIGURE 1. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live N. gomantaka n. sp., in oyster shell; g, live N. gomantaka n. sp., on rotten wood." figureDoi="http://doi.org/10.5281/zenodo.10954093" httpUri="https://zenodo.org/record/10954093/files/figure.png" pageId="3" pageNumber="292">Figs. 1d</figureCitation>
;
<figureCitation id="131C2A17FFAEFFE3FE828CC93050D08D" box="[341,371,1765,1791]" captionStart="FIGURE 2" captionStartId="3.[151,250,1312,1336]" captionTargetBox="[167,1420,184,1286]" captionTargetId="figure-17@3.[164,1423,181,1288]" captionTargetPageId="3" captionText="FIGURE 2. Nanosesarma gomantaka n. sp., a, c, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. b, d, N. pontianacense (De Man, 1895), male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, b, carapace, dorsal view; c, d, male pleon; e, female pleon." figureDoi="http://doi.org/10.5281/zenodo.10954095" httpUri="https://zenodo.org/record/10954095/files/figure.png" pageId="3" pageNumber="292">2e</figureCitation>
).
</paragraph>
<paragraph id="8B983692FFAEFFE3FF108D2535DBD18E" blockId="3.[151,1437,1513,2044]" pageId="3" pageNumber="292">
P25, compressed, shorter than chelipeds. P2 short, P3 and P4 longest, subequal in length, P5 short. P45 merus flattened, slightly wider distally; anterolateral margin terminates in blunt angle followed by blunt subdistal spine; dorsal surface with several short, sinuous striations, setose; posterodistal margin with minute granules; P5 merus posterodistal margin dentate, separated from tooth on posterolateral corner by notch. Ventral surfaces of carpus with median longitudinal granular ridge. P2 carpus and propodus densely setose on anterior surface, P3P5 carpus and propodus without dense setae. Propodus 2 times longer than dactylus, posterior margins with scattered long bristles. Dactylus smooth, tips chitinous, slightly curved, with 2 pairs of slender subdistal spines (
<figureCitation id="131C2A17FFAEFFE3FBAA8DCD35C8D18E" box="[1149,1259,2017,2044]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[263,1317,189,1905]" captionTargetId="figure-21@6.[256,1332,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Nanosesarma gomantaka n. sp., ad, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, P2, dorsal view; b, P3, dorsal view; c, P4, dorsal view; d, P5, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10954101" httpUri="https://zenodo.org/record/10954101/files/figure.png" pageId="3" pageNumber="292">Fig. 5ad</figureCitation>
).
</paragraph>
<caption id="DF58661AFFA9FFE4FF408C22343AD01C" ID-DOI="http://doi.org/10.5281/zenodo.10954097" ID-Zenodo-Dep="10954097" httpUri="https://zenodo.org/record/10954097/files/figure.png" pageId="4" pageNumber="293" startId="4.[151,250,1550,1574]" targetBox="[154,1432,184,1523]" targetPageId="4" targetType="figure">
<paragraph id="8B983692FFA9FFE4FF408C22343AD01C" blockId="4.[151,1437,1550,1646]" pageId="4" pageNumber="293">
<emphasis id="B953EA80FFA9FFE4FF408C223036D054" bold="true" box="[151,277,1550,1574]" pageId="4" pageNumber="293">FIGURE 3.</emphasis>
<taxonomicName id="4C274D11FFA9FFE4FEC98C233300D054" authority="Trivedi &amp; Padate &amp; Ng, 2024" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[286,547,1550,1574]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="gomantaka" status="sp. nov.">
<emphasis id="B953EA80FFA9FFE4FEC98C233300D054" box="[286,547,1550,1574]" italics="true" pageId="4" pageNumber="293">Nanosesarma gomantaka</emphasis>
</taxonomicName>
<emphasis id="B953EA80FFA9FFE4FDFC8C223344D054" bold="true" box="[555,615,1550,1574]" pageId="4" pageNumber="293">
<taxonomicNameLabel id="A26057FBFFA9FFE4FDFC8C223344D054" box="[555,615,1550,1574]" pageId="4" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, a, b, d, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, left chela, outer view (setae omitted); b, left chela, dorsal view; d, third maxilliped, dorsal view; e, left vulva.
</paragraph>
</caption>
<paragraph id="8B983692FFA9FFE4FF108CED304CD150" blockId="4.[151,1437,1728,2043]" pageId="4" pageNumber="293">
G1 relatively straight, slender except for distal chitinous part which is gently bent at obtuse angle (
<figureCitation id="131C2A17FFA9FFE4FAED8CEC34ACD0A9" box="[1338,1423,1728,1755]" captionStart="FIGURE 4" captionStartId="5.[151,250,1214,1238]" captionTargetBox="[155,1431,185,1185]" captionTargetId="figure-17@5.[151,1435,181,1190]" captionTargetPageId="5" captionText="FIGURE 4. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, N. pontianacense (De Man, 1895), male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, c, left G1, dorsal view; b, d, left G1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954099" httpUri="https://zenodo.org/record/10954099/files/figure.png" pageId="4" pageNumber="293">Fig. 4a</figureCitation>
); distal part densely covered with long stiff setae which obscures spatuliform chitinous part; main shaft with ventral groove (
<figureCitation id="131C2A17FFA9FFE4FF238D243041D150" box="[244,354,1800,1827]" captionStart="FIGURE 4" captionStartId="5.[151,250,1214,1238]" captionTargetBox="[155,1431,185,1185]" captionTargetId="figure-17@5.[151,1435,181,1190]" captionTargetPageId="5" captionText="FIGURE 4. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, N. pontianacense (De Man, 1895), male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, c, left G1, dorsal view; b, d, left G1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10954099" httpUri="https://zenodo.org/record/10954099/files/figure.png" pageId="4" pageNumber="293">Fig. 4a, b</figureCitation>
).
</paragraph>
<paragraph id="8B983692FFA9FFE4FF108D0032FDD119" blockId="4.[151,1437,1728,2043]" pageId="4" pageNumber="293">
Vulva on distal half of thoracic sternite 6, not touching suture with sternite 5, with semicircular operculum on inner side; lateral parts with low rim but no obvious vulvar cover (
<figureCitation id="131C2A17FFA9FFE4FCA88D7C32F2D119" box="[895,977,1872,1899]" captionStart="FIGURE 3" captionStartId="4.[151,250,1550,1574]" captionTargetBox="[154,1432,184,1523]" captionTargetId="figure-21@4.[151,1435,181,1526]" captionTargetPageId="4" captionText="FIGURE 3. Nanosesarma gomantaka n. sp., a, b, d, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, e, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, left chela, outer view (setae omitted); b, left chela, dorsal view; d, third maxilliped, dorsal view; e, left vulva." figureDoi="http://doi.org/10.5281/zenodo.10954097" httpUri="https://zenodo.org/record/10954097/files/figure.png" pageId="4" pageNumber="293">Fig. 3e</figureCitation>
).
</paragraph>
<paragraph id="8B983692FFA9FFE4FF108D58303DD1C0" blockId="4.[151,1437,1728,2043]" pageId="4" pageNumber="293">
<emphasis id="B953EA80FFA9FFE4FF108D58332BD1FC" bold="true" box="[199,520,1908,1934]" pageId="4" pageNumber="293">Geographical distribution.</emphasis>
<materialsCitation id="3B4F3CCFFFA9FFE4FDC58D583039D1C0" country="India" location="Chapora Estuary" municipality="Goa State" pageId="4" pageNumber="293" specimenCount="1">
The species is so far known only from its
<typeStatus id="549C8830FFA9FFE4FBD58D593511D1FD" box="[1026,1074,1909,1935]" pageId="4" pageNumber="293">type</typeStatus>
locality,
<location id="8EF86049FFA9FFE4FB778D583443D1FD" LSID="urn:lsid:plazi:treatment:038E8784FFACFFE7FF4088D7353AD326:8EF86049FFA9FFE4FB778D583443D1FD" box="[1184,1376,1908,1935]" country="India" municipality="Goa State" name="Chapora Estuary" pageId="4" pageNumber="293">Chapora Estuary</location>
,
<collectingMunicipality id="6BFCACE8FFA9FFE4FAB98D5931F1D1C1" pageId="4" pageNumber="293">Goa State</collectingMunicipality>
,
<collectingCountry id="F3307602FFA9FFE4FF0A8DB43039D1C0" box="[221,282,1944,1970]" name="India" pageId="4" pageNumber="293">India</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8B983692FFA9FFE4FF108D9032E5D189" blockId="4.[151,1437,1728,2043]" pageId="4" pageNumber="293">
<emphasis id="B953EA80FFA9FFE4FF108D903009D1A4" bold="true" box="[199,298,1980,2006]" pageId="4" pageNumber="293">Habitat.</emphasis>
This species inhabits mangrove forest habitat where it lives among the aerial roots of mangroves, under decaying wood, and under large dead shells of oysters (
<figureCitation id="131C2A17FFA9FFE4FC9D8DCC329AD189" box="[842,953,2016,2043]" captionStart="FIGURE 1" captionStartId="2.[151,250,1375,1399]" captionTargetBox="[151,1436,181,1351]" captionTargetId="figure-21@2.[151,1436,181,1351]" captionTargetPageId="2" captionText="FIGURE 1. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live N. gomantaka n. sp., in oyster shell; g, live N. gomantaka n. sp., on rotten wood." figureDoi="http://doi.org/10.5281/zenodo.10954093" httpUri="https://zenodo.org/record/10954093/files/figure.png" pageId="4" pageNumber="293">Fig. 1eg</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF58661AFFA8FFE5FF408E923373D36D" ID-DOI="http://doi.org/10.5281/zenodo.10954099" ID-Zenodo-Dep="10954099" httpUri="https://zenodo.org/record/10954099/files/figure.png" pageId="5" pageNumber="294" startId="5.[151,250,1214,1238]" targetBox="[155,1431,185,1185]" targetPageId="5" targetType="figure">
<paragraph id="8B983692FFA8FFE5FF408E923373D36D" blockId="5.[151,1436,1214,1311]" pageId="5" pageNumber="294">
<emphasis id="B953EA80FFA8FFE5FF408E923031D2A4" bold="true" box="[151,274,1214,1238]" pageId="5" pageNumber="294">FIGURE 4.</emphasis>
<taxonomicName id="4C274D11FFA8FFE5FEC08E93333AD2A5" authority="Trivedi &amp; Padate &amp; Ng, 2024" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[279,537,1215,1239]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="gomantaka" status="sp. nov.">
<emphasis id="B953EA80FFA8FFE5FEC08E93333AD2A5" box="[279,537,1215,1239]" italics="true" pageId="5" pageNumber="294">Nanosesarma gomantaka</emphasis>
</taxonomicName>
<emphasis id="B953EA80FFA8FFE5FDCA8E933376D2A5" bold="true" box="[541,597,1215,1239]" pageId="5" pageNumber="294">
<taxonomicNameLabel id="A26057FBFFA8FFE5FDCA8E933376D2A5" box="[541,597,1215,1239]" pageId="5" pageNumber="294" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d,
<taxonomicName id="4C274D11FFA8FFE5FEA28ECF33EED288" authority="(De Man, 1895)" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[373,717,1250,1275]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="pontianacense">
<emphasis id="B953EA80FFA8FFE5FEA28ECF3307D288" box="[373,548,1251,1274]" italics="true" pageId="5" pageNumber="294">N. pontianacense</emphasis>
(
<bibRefCitation id="EFB64B63FFA8FFE5FDE68ECF33E6D288" author="De Man, J. G." box="[561,709,1250,1275]" pageId="5" pageNumber="294" pagination="485 - 609" refId="ref4620" refString="De Man, J. G. (1895) Berichte uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4), 485 - 609. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man, 1895</bibRefCitation>
)
</taxonomicName>
, male (CL 6.0 mm, CW 5.3 mm) (ZRC 2012.0299), Singapore. a, c, left G1, dorsal view; b, d, left G1, ventral view.
</paragraph>
</caption>
<subSubSection id="C33D6519FFA8FFE5FF108F5D34B9D385" pageId="5" pageNumber="294" type="etymology">
<paragraph id="8B983692FFA8FFE5FF108F5D3544D3DD" blockId="5.[151,1437,1393,2032]" pageId="5" pageNumber="294">
<emphasis id="B953EA80FFA8FFE5FF108F5D306DD3F9" bold="true" box="[199,334,1393,1419]" pageId="5" pageNumber="294">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C274D11FFA8FFE5FDE98F5D33EBD3F9" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[574,712,1393,1419]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="gomantaka">“gomantaka</taxonomicName>
” is derived from the ancient Sanskrit name of Goa State,
<collectingCountry id="F3307602FFA8FFE5FA8B8F5D34BBD3F9" box="[1372,1432,1393,1419]" name="India" pageId="5" pageNumber="294">India</collectingCountry>
, which is the
<typeStatus id="549C8830FFA8FFE5FEF18FB93075D3DD" box="[294,342,1429,1455]" pageId="5" pageNumber="294">type</typeStatus>
locality of the new species. The name is used as a noun in apposition.
</paragraph>
<paragraph id="8B983692FFA8FFE5FF108F9534B9D385" blockId="5.[151,1437,1393,2032]" pageId="5" pageNumber="294">
<emphasis id="B953EA80FFA8FFE5FF108F95303ED3A1" bold="true" box="[199,285,1465,1491]" pageId="5" pageNumber="294">Colour.</emphasis>
Carapace of live specimens dark brown over anterior half, darker posteriorly. Chelipeds, pereopods and pleon light brown, pereopods with brown transverse bands. Dark brown setae on chelipeds and pereopods (
<figureCitation id="131C2A17FFA8FFE5FAF38FF134AFD385" box="[1316,1420,1501,1527]" captionStart="FIGURE 1" captionStartId="2.[151,250,1375,1399]" captionTargetBox="[151,1436,181,1351]" captionTargetId="figure-21@2.[151,1436,181,1351]" captionTargetPageId="2" captionText="FIGURE 1. Nanosesarma gomantaka n. sp., a, b, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India; c, d, paratype female (CL 5.4 mm, CW 5.1 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, c, dorsal view; b, d, ventral view; e, mangrove habitat of Chapora Estuary, Goa State, India; f, live N. gomantaka n. sp., in oyster shell; g, live N. gomantaka n. sp., on rotten wood." figureDoi="http://doi.org/10.5281/zenodo.10954093" httpUri="https://zenodo.org/record/10954093/files/figure.png" pageId="5" pageNumber="294">Fig. 1ad</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33D6519FFA8FFE7FF108C2D353AD326" lastPageId="7" lastPageNumber="296" pageId="5" pageNumber="294" type="discussion">
<paragraph id="8B983692FFA8FFE7FF108C2D3287D5F2" blockId="5.[151,1437,1393,2032]" lastBlockId="7.[151,1437,690,1364]" lastPageId="7" lastPageNumber="296" pageId="5" pageNumber="294">
<emphasis id="B953EA80FFA8FFE5FF108C2D3018D069" bold="true" box="[199,315,1537,1563]" pageId="5" pageNumber="294">Remarks.</emphasis>
<bibRefCitation id="EFB64B63FFA8FFE5FE948C2D3305D069" author="De Man, J. G." box="[323,550,1537,1564]" pageId="5" pageNumber="294" pagination="485 - 609" refId="ref4620" refString="De Man, J. G. (1895) Berichte uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4), 485 - 609. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man (1895: 178</bibRefCitation>
, pl. 30 fig. 33) described
<taxonomicName id="4C274D11FFA8FFE5FC9A8C2D35DED069" authorityName="De Man" authorityYear="1895" box="[845,1277,1537,1563]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="pontianacensis" subGenus="Episesarma">
<emphasis id="B953EA80FFA8FFE5FC9A8C2D328ED069" box="[845,941,1537,1563]" italics="true" pageId="5" pageNumber="294">Sesarma</emphasis>
(
<emphasis id="B953EA80FFA8FFE5FC688C2E3560D069" box="[959,1091,1538,1563]" italics="true" pageId="5" pageNumber="294">Episesarma</emphasis>
)
<emphasis id="B953EA80FFA8FFE5FB828C2E35DED069" box="[1109,1277,1538,1563]" italics="true" pageId="5" pageNumber="294">pontianacensis</emphasis>
</taxonomicName>
from a single female specimen (CL
<quantity id="4CDF9B77FFA8FFE5FE418C0930DCD04D" box="[406,511,1573,1599]" metricMagnitude="-3" metricUnit="m" metricValue="8.17" pageId="5" pageNumber="294" unit="mm" value="8.17">8.17 mm</quantity>
, CW
<quantity id="4CDF9B77FFA8FFE5FD938C09338FD04D" box="[580,684,1573,1600]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" pageId="5" pageNumber="294" unit="mm" value="7.75">7.75 mm</quantity>
) from “Pontianak, west coast of Borneo”, with the author noting its general resemblance to much larger S.
<taxonomicName id="4C274D11FFA8FFE5FD568C65351DD011" authority="(H. Milne Edwards, 1837)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[641,1086,1609,1635]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="trapezoidea">
<emphasis id="B953EA80FFA8FFE5FD568C653226D011" box="[641,773,1609,1635]" italics="true" pageId="5" pageNumber="294">trapezoidea</emphasis>
(H.
<bibRefCitation id="EFB64B63FFA8FFE5FCEA8C653515D011" author="Milne Edwards, H. &amp; Histoire Naturelle des Crustaces, Comprenant" box="[829,1078,1609,1635]" pageId="5" pageNumber="294" refId="ref5004" refString="Milne Edwards, H. (1834 - 1840) Histoire Naturelle des Crustaces, Comprenant l´Anatomie, la Physiologie et la Classification de ces Animaux. Vols. I - III. Encyclopedique Roret, Paris, xxxv + 468 pp. (1834), 532 pp. (1837), 638 pp., 42 pls. (1840). https: // doi. org / 10.5962 / bhl. title. 16170" type="book" year="1837">Milne Edwards, 1837</bibRefCitation>
)
</taxonomicName>
(now in
<taxonomicName id="4C274D11FFA8FFE5FB7B8C653026D0F5" authority="Naruse &amp; Ng, 2020" authorityName="Naruse &amp; Ng" authorityYear="2020" class="Malacostraca" family="Sesarmidae" genus="Shinobium" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="B953EA80FFA8FFE5FB7B8C653407D011" box="[1196,1316,1609,1635]" italics="true" pageId="5" pageNumber="294">Shinobium</emphasis>
<bibRefCitation id="EFB64B63FFA8FFE5FAFA8C653026D0F5" author="Naruse, T. &amp; Ng, P. K. L." pageId="5" pageNumber="294" pagination="445 - 532" refId="ref5233" refString="Naruse, T. &amp; Ng, P. K. L. (2020) Revision of the sesarmid crab genera Labuanium Serene and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003, and Namlacium Sere`ne and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species. Journal of Natural History, 54 (7 - 8), 445 - 532. https: // doi. org / 10.1080 / 00222933.2020.1763491" type="journal article" year="2020">Naruse &amp; Ng, 2020</bibRefCitation>
</taxonomicName>
). Of the species he placed in the subgenus
<taxonomicName id="4C274D11FFA8FFE5FD3E8C423539D0FA" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[745,1050,1645,1672]" pageId="5" pageNumber="294" rank="subGenus" subGenus="Episesarma">
<emphasis id="B953EA80FFA8FFE5FD3E8C42324ED0F5" box="[745,877,1646,1671]" italics="true" pageId="5" pageNumber="294">Episesarma</emphasis>
<bibRefCitation id="EFB64B63FFA8FFE5FCA28C413539D0FA" author="De Man, J. G." box="[885,1050,1645,1672]" pageId="5" pageNumber="294" pagination="485 - 609" refId="ref4620" refString="De Man, J. G. (1895) Berichte uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4), 485 - 609. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man, 1895</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C274D11FFA8FFE5FBF28C4135C9D0F5" authorityName="De Man" authorityYear="1895" box="[1061,1258,1645,1671]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="pontianacensis">
<emphasis id="B953EA80FFA8FFE5FBF28C4135C9D0F5" box="[1061,1258,1645,1671]" italics="true" pageId="5" pageNumber="294">S. pontianacensis</emphasis>
</taxonomicName>
is most distinct in its carapace being longer than wide, with a setose body and ambulatory legs which resemble some majoid crabs. Unfortunately, the type female specimen is almost certainly lost. The type was collected during the Hugo Storm expedition to Asia, and extant
<typeStatus id="549C8830FFA8FFE5FD908CF5338BD081" box="[583,680,1753,1779]" pageId="5" pageNumber="294" type="syntype">syntypes</typeStatus>
are in the Rijksmuseum van Natuurlijke Historie in Leiden (now Naturalis Biodiversity Center); with some supposed material in the Munich Museum (
<bibRefCitation id="EFB64B63FFA8FFE5FB828CD13412D165" author="Fransen, C. H. J. M. &amp; Holthuis, L. B. &amp; Adema, J. P. H. M." box="[1109,1329,1789,1815]" pageId="5" pageNumber="294" pagination="1 - 344" refId="ref4695" refString="Fransen, C. H. J. M., Holthuis, L. B. &amp; Adema, J. P. H. M. (1997) Type-catalogue of the Decapod Crustacea in the collections of the Nationaal Naturhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologische Verhandelingen, 311, i - xvi + 1 - 344." type="journal article" year="1997">
Fransen
<emphasis id="B953EA80FFA8FFE5FB618CD235CDD165" box="[1206,1262,1789,1815]" italics="true" pageId="5" pageNumber="294">et al.</emphasis>
1997
</bibRefCitation>
). The last author has checked for the type in these institutions as well as Muséum national dHistoire naturelle in Paris and Forschungsinstitut Senckenberg where De Mans material is known to be kept; the type of
<taxonomicName id="4C274D11FFA8FFE5FBB18D69344FD12D" authorityName="De Man" authorityYear="1895" box="[1126,1388,1861,1887]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="pontianacensis">
<emphasis id="B953EA80FFA8FFE5FBB18D69344FD12D" box="[1126,1388,1861,1887]" italics="true" pageId="5" pageNumber="294">Sesarma pontianacensis</emphasis>
</taxonomicName>
was not found in these locations, with the late Michael Türkay (personal communication) advising that it was probably lost. A
<typeStatus id="549C8830FFA8FFE5FF668DA13024D1D5" box="[177,263,1933,1959]" pageId="5" pageNumber="294" type="neotype">neotype</typeStatus>
for
<taxonomicName id="4C274D11FFA8FFE5FEEF8DA13363D1D5" authorityName="De Man" authorityYear="1895" box="[312,576,1933,1959]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="pontianacensis">
<emphasis id="B953EA80FFA8FFE5FEEF8DA13363D1D5" box="[312,576,1933,1959]" italics="true" pageId="5" pageNumber="294">Sesarma pontianacensis</emphasis>
</taxonomicName>
should be selected when the revision of the genus is done.
<bibRefCitation id="EFB64B63FFA8FFE5FB038DA134BFD1D5" author="Tesch, J. J." box="[1236,1436,1933,1959]" pageId="5" pageNumber="294" pagination="127 - 260" refId="ref5953" refString="Tesch, J. J. (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the Indo-Pacific species. Zoologische Mededelingen, 3 (2 - 3), 127 - 260. https: // doi. org / 10.5962 / bhl. title. 10640" type="journal article" year="1917">Tesch (1917: 190)</bibRefCitation>
grouped
<taxonomicName id="4C274D11FFA8FFE5FF208D9D3097D1B9" authorityName="De Man" authorityYear="1895" box="[247,436,1969,1995]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="pontianacensis">
<emphasis id="B953EA80FFA8FFE5FF208D9D3097D1B9" box="[247,436,1969,1995]" italics="true" pageId="5" pageNumber="294">S. pontianacensis</emphasis>
</taxonomicName>
with
<taxonomicName id="4C274D11FFA8FFE5FE258D9D33CED1B9" authority="De Man, 1888" authorityName="De Man" authorityYear="1888" box="[498,749,1969,1995]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="polita">
<emphasis id="B953EA80FFA8FFE5FE258D9D3368D1B9" box="[498,587,1969,1995]" italics="true" pageId="5" pageNumber="294">S. polita</emphasis>
<bibRefCitation id="EFB64B63FFA8FFE5FD858D9D33CED1B9" author="De Man, J. G." box="[594,749,1969,1995]" pageId="5" pageNumber="294" pagination="129 - 176" refId="ref4533" refString="De Man, J. G. (1888) Report on the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F. R. S., Superintendent of the Museum. Part III. Journal of the Linnean Society of London, Zoology, 22, 129 - 176. https: // doi. org / 10.1111 / j. 1096 - 3642.1887. tb 00027. x" type="journal article" year="1888">De Man, 1888</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C274D11FFA8FFE5FD2C8D9D354AD1BE" authority="H. Milne Edwards, 1853" authorityName="H. Milne Edwards" authorityYear="1853" box="[763,1129,1969,1996]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="B953EA80FFA8FFE5FD2C8D9D327BD1B9" box="[763,856,1969,1995]" italics="true" pageId="5" pageNumber="294">S. smithi</emphasis>
H.
<bibRefCitation id="EFB64B63FFA8FFE5FC578D9D354AD1BE" author="Milne Edwards, H." box="[896,1129,1969,1996]" pageId="5" pageNumber="294" pagination="163 - 228" refId="ref5083" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodiens. Annales des Sciences Naturelles, 3 e Serie, 20, 163 - 228, pls. 6 - 11." type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C274D11FFA8FFE5FBA08D9D3459D1BE" authority="Hess, 1865" authorityName="Hess" authorityYear="1865" box="[1143,1402,1969,1996]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="rotundata">
<emphasis id="B953EA80FFA8FFE5FBA08D9D35DFD1B9" box="[1143,1276,1969,1995]" italics="true" pageId="5" pageNumber="294">S. rotundata</emphasis>
<bibRefCitation id="EFB64B63FFA8FFE5FAD58D9D3459D1BE" author="Hess, W." box="[1282,1402,1969,1996]" pageId="5" pageNumber="294" pagination="127 - 173" refId="ref4812" refString="Hess, W. (1865) Beitragezur Kenntnis der Decapoden-Krebse Ost-Australiens. Archiv fur Naturgeschicte, 31 (1), 127 - 173, pls. 6 - 8. https: // doi. org / 10.5962 / bhl. part. 15862" type="journal article" year="1865">Hess, 1865</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C274D11FFA8FFE5FA5F8D9D30A5D19D" authority="Burger, 1893" authorityName="Burger" authorityYear="1893" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="cruciata">
<emphasis id="B953EA80FFA8FFE5FA5F8D9D31D1D19D" italics="true" pageId="5" pageNumber="294">S. cruciata</emphasis>
<bibRefCitation id="EFB64B63FFA8FFE5FF2F8DF930A5D19D" author="Burger, O." box="[248,390,2005,2031]" pageId="5" pageNumber="294" pagination="613 - 632" refId="ref4089" refString="Burger, O. (1893) Beitrage zur Kenntniss der Gattung Sesarma. Zoologischen Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7 (4), 613 - 632, pl. 21." type="journal article" year="1893">Bürger, 1893</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C274D11FFA8FFE5FE438DF933B6D19D" authority="Burger, 1893" authorityName="Burger" authorityYear="1893" box="[404,661,2005,2031]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="B953EA80FFA8FFE5FE438DF930DCD19D" box="[404,511,2005,2031]" italics="true" pageId="5" pageNumber="294">S. demani</emphasis>
<bibRefCitation id="EFB64B63FFA8FFE5FDD18DF933B6D19D" author="Burger, O." box="[518,661,2005,2031]" pageId="5" pageNumber="294" pagination="613 - 632" refId="ref4089" refString="Burger, O. (1893) Beitrage zur Kenntniss der Gattung Sesarma. Zoologischen Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7 (4), 613 - 632, pl. 21." type="journal article" year="1893">Bürger, 1893</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C274D11FFA8FFE5FD748DF93281D19D" authority="Ihle, 1912" authorityName="Ihle" authorityYear="1912" box="[675,930,2005,2031]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B953EA80FFA8FFE5FD748DF93208D19D" box="[675,811,2005,2031]" italics="true" pageId="5" pageNumber="294">S. jacobsoni</emphasis>
<bibRefCitation id="EFB64B63FFA8FFE5FCE58DF93281D19D" author="Ihle, J. E. W." box="[818,930,2005,2031]" pageId="5" pageNumber="294" pagination="17 - 182" refId="ref4896" refString="Ihle, J. E. W. (1912) Ueber eine kleine Brachyuren-Sammlung aus unterirdischen Flussen von Java. Notes from the Leyden Museum, 34 (3 - 4), 17 - 182, pl. 9." type="journal article" year="1912">Ihle, 1912</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C274D11FFA8FFE5FC678DF935E5D182" authority="Hess, 1865" authorityName="Hess" authorityYear="1865" box="[944,1222,2005,2032]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="atrorubens">
<emphasis id="B953EA80FFA8FFE5FC678DF93566D19D" box="[944,1093,2005,2031]" italics="true" pageId="5" pageNumber="294">S. atrorubens</emphasis>
<bibRefCitation id="EFB64B63FFA8FFE5FB9C8DF935E5D182" author="Hess, W." box="[1099,1222,2005,2032]" pageId="5" pageNumber="294" pagination="127 - 173" refId="ref4812" refString="Hess, W. (1865) Beitragezur Kenntnis der Decapoden-Krebse Ost-Australiens. Archiv fur Naturgeschicte, 31 (1), 127 - 173, pls. 6 - 8. https: // doi. org / 10.5962 / bhl. part. 15862" type="journal article" year="1865">Hess, 1865</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C274D11FFA8FFE7FB2B8DF9308FD4BE" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" lastPageId="7" lastPageNumber="296" order="Decapoda" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="trapezoidea">
<emphasis id="B953EA80FFA8FFE5FB2B8DF934B8D19D" box="[1276,1435,2005,2031]" italics="true" pageId="5" pageNumber="294">S. trapezoidea</emphasis>
H.
<bibRefCitation id="EFB64B63FFAAFFE7FF6E889E308FD4BE" author="Milne Edwards, H. &amp; Histoire Naturelle des Crustaces, Comprenant" box="[185,428,690,716]" pageId="7" pageNumber="296" refId="ref5004" refString="Milne Edwards, H. (1834 - 1840) Histoire Naturelle des Crustaces, Comprenant l´Anatomie, la Physiologie et la Classification de ces Animaux. Vols. I - III. Encyclopedique Roret, Paris, xxxv + 468 pp. (1834), 532 pp. (1837), 638 pp., 42 pls. (1840). https: // doi. org / 10.5962 / bhl. title. 16170" type="book" year="1837">Milne Edwards, 1837</bibRefCitation>
</taxonomicName>
, because of the “distance between external orbital angles distinctly less than the length of the carapace in the median line.” These species are now known to be unrelated and are placed in other genera (
<bibRefCitation id="EFB64B63FFAAFFE7FA9888FA3001D566" author="Serene, R. &amp; Soh, C. L." pageId="7" pageNumber="296" pagination="387 - 416" refId="ref5912" refString="Serene, R. &amp; Soh, C. L. (1970) New Indo-Pacific genera allied to Sesarma Say 1817 (Brachyura, Decapoda, Crustacea). Treubia, 27 (4), 387 - 416." type="journal article" year="1970">Serène &amp; Soh 1970</bibRefCitation>
;
<bibRefCitation id="EFB64B63FFAAFFE7FEFA88D630DBD566" author="Davie, P. J. F. &amp; Ng, P. K. L." box="[301,504,762,788]" pageId="7" pageNumber="296" pagination="227 - 231" refId="ref4410" refString="Davie, P. J. F. &amp; Ng, P. K. L. (2007) A new genus for cave-dwelling crabs previously assigned to Sesarmoides (Crustacea: Decapoda: Brachyura: Sesarmidae). Raffles Bulletin of Zoology, Supplement 16, 227 - 231." type="journal article" year="2007">Davie &amp; Ng 2007</bibRefCitation>
;
<bibRefCitation id="EFB64B63FFAAFFE7FDD488D633F9D566" author="Naruse, T. &amp; Ng, P. K. L." box="[515,730,762,788]" pageId="7" pageNumber="296" pagination="445 - 532" refId="ref5233" refString="Naruse, T. &amp; Ng, P. K. L. (2020) Revision of the sesarmid crab genera Labuanium Serene and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003, and Namlacium Sere`ne and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species. Journal of Natural History, 54 (7 - 8), 445 - 532. https: // doi. org / 10.1080 / 00222933.2020.1763491" type="journal article" year="2020">Naruse &amp; Ng 2020</bibRefCitation>
;
<bibRefCitation id="EFB64B63FFAAFFE7FD3188D632A9D566" author="Ng, P. K. L. &amp; Li, J. - J. &amp; Shih, H. - T." box="[742,906,762,788]" pageId="7" pageNumber="296" pagination="1 - 39" refId="ref5487" refString="Ng, P. K. L., Li, J. - J. &amp; Shih, H. - T. (2020) What is Sesarmops impressus (H. Milne Edwards, 1837) (Crustacea: Brachyura: Sesarmidae)? Zoological Studies, 59 (27), 1 - 39." type="journal article" year="2020">
Ng
<emphasis id="B953EA80FFAAFFE7FCD888D63264D566" box="[783,839,762,788]" italics="true" pageId="7" pageNumber="296">et al.</emphasis>
2020
</bibRefCitation>
). In establishing
<taxonomicName id="4C274D11FFAAFFE7FB9088D634BFD566" authority=", Tweedie (1950)" authorityName=", Tweedie" authorityYear="1950" box="[1095,1436,761,788]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B953EA80FFAAFFE7FB9088D635FCD561" box="[1095,1247,762,787]" italics="true" pageId="7" pageNumber="296">Nanosesarma</emphasis>
,
<bibRefCitation id="EFB64B63FFAAFFE7FB3A88D634BFD566" author="Tweedie, M. W. F." box="[1261,1436,761,788]" pageId="7" pageNumber="296" pagination="310 - 316" refId="ref6123" refString="Tweedie, M. W. F. (1950) Notes on grapsoid crabs from the Raffles Museum. Bulletin of the Raffles Museum, 23, 310 - 316." type="journal article" year="1950">Tweedie (1950)</bibRefCitation>
</taxonomicName>
did not treat
<taxonomicName id="4C274D11FFAAFFE7FEF4893230C5D545" authorityName="De Man" authorityYear="1895" box="[291,486,798,824]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="pontianacensis">
<emphasis id="B953EA80FFAAFFE7FEF4893230C5D545" box="[291,486,798,824]" italics="true" pageId="7" pageNumber="296">S. pontianacensis</emphasis>
</taxonomicName>
, and it was
<bibRefCitation id="EFB64B63FFAAFFE7FDBD893232AED54A" author="Serene, R. &amp; Soh, C. L." box="[618,909,798,824]" pageId="7" pageNumber="296" pagination="387 - 416" refId="ref5912" refString="Serene, R. &amp; Soh, C. L. (1970) New Indo-Pacific genera allied to Sesarma Say 1817 (Brachyura, Decapoda, Crustacea). Treubia, 27 (4), 387 - 416." type="journal article" year="1970">Serène &amp; Soh (1970: 393)</bibRefCitation>
who formally referred the species to the genus.
<taxonomicName id="4C274D11FFAAFFE7FF40896E30F9D529" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[151,474,834,859]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="pontianacense">
<emphasis id="B953EA80FFAAFFE7FF40896E30F9D529" box="[151,474,834,859]" italics="true" pageId="7" pageNumber="296">Nanosesarma pontianacense</emphasis>
</taxonomicName>
has since been reported from many locations in Southeast and
<collectingCountry id="F3307602FFAAFFE7FB41896E35DDD52E" box="[1174,1278,834,860]" name="China" pageId="7" pageNumber="296">East Asia</collectingCountry>
(
<bibRefCitation id="EFB64B63FFAAFFE7FADA896E34B4D52E" author="Nobili, G." box="[1293,1431,834,860]" pageId="7" pageNumber="296" pagination="1 - 32" refId="ref5562" refString="Nobili, G. (1903) Contributo alla fauna carcinologica di Borneo. Bolletino dei Musei Zoologia ed Anatamia comprata della R. Universita di Torino, 18 (447), 1 - 32." type="journal article" year="1903">Nobili 1903</bibRefCitation>
;
<bibRefCitation id="EFB64B63FFAAFFE7FF40894A301BD5F2" author="Tweedie, M. W. F." box="[151,312,870,896]" pageId="7" pageNumber="296" pagination="88 - 113" refId="ref6086" refString="Tweedie, M. W. F. (1940) New and interesting Malaysian species of Sesarma and Utica (Crustacea Brachyura). Bulletin of the Raffles Museum, 16, 88 - 113." type="journal article" year="1940">Tweedie 1940</bibRefCitation>
;
<bibRefCitation id="EFB64B63FFAAFFE7FE94894A30CED50D" author="Dai, A. - Y. &amp; Yang, S. - L. &amp; Song, Y. - Z. &amp; Chen, G. - X." box="[323,493,869,896]" pageId="7" pageNumber="296" refId="ref4176" refString="Dai, A. - Y., Yang, S. - L., Song, Y. - Z. &amp; Chen, G. - X. (1986) Crabs of the China Seas. China Ocean Press, Beijing, 11 + 642 pp. [in Chinese]" type="book" year="1986">
Dai
<emphasis id="B953EA80FFAAFFE7FEA4894A3088D5F2" box="[371,427,870,896]" italics="true" pageId="7" pageNumber="296">et al.</emphasis>
1986
</bibRefCitation>
;
<bibRefCitation id="EFB64B63FFAAFFE7FE2E894A33E3D5F2" author="Dai, A. - Y. &amp; Yang, S. - L." box="[505,704,870,896]" pageId="7" pageNumber="296" refId="ref4129" refString="Dai, A. - Y. &amp; Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing and Springer-Verlag, Berlin, 21 + 608 pp., 295 figs., 74 pls," type="book" year="1991">Dai &amp; Yang 1991</bibRefCitation>
;
<bibRefCitation id="EFB64B63FFAAFFE7FD1C894A32BBD5F2" author="Ng, P. K. L. &amp; Davie, P. J. F." box="[715,920,870,896]" pageId="7" pageNumber="296" pagination="369 - 384" refId="ref5327" refString="Ng, P. K. L. &amp; Davie, P. J. F. (2002) A checklist of the brachyuran crabs of Phuket and western Thailand. - Biodiversity of Crustacea of the Andaman Sea. In: Bruce, N. L., Berggren, M. &amp; Bussawarit, S. (Eds.), Proceedings of the International Workshop on the Biodiversity of Crustacea in the Andaman Sea. Phuket Marine Biological Center, 29 November to 20 December 1998. Phuket Marine Biological Center Special Publication, 23 (2), 369 - 384." type="journal article" year="2002">Ng &amp; Davie 2002</bibRefCitation>
).
</paragraph>
<caption id="DF58661AFFABFFE6FF408DBB356FD1A1" ID-DOI="http://doi.org/10.5281/zenodo.10954101" ID-Zenodo-Dep="10954101" httpUri="https://zenodo.org/record/10954101/files/figure.png" pageId="6" pageNumber="295" startId="6.[151,250,1943,1967]" targetBox="[263,1317,189,1905]" targetPageId="6" targetType="figure">
<paragraph id="8B983692FFABFFE6FF408DBB356FD1A1" blockId="6.[151,1437,1943,2003]" pageId="6" pageNumber="295">
<emphasis id="B953EA80FFABFFE6FF408DBB3031D1DD" bold="true" box="[151,274,1943,1967]" pageId="6" pageNumber="295">FIGURE 5.</emphasis>
<taxonomicName id="4C274D11FFABFFE6FEC08DB4333BD1DD" authority="Trivedi &amp; Padate &amp; Ng, 2024" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[279,536,1943,1967]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="gomantaka" status="sp. nov.">
<emphasis id="B953EA80FFABFFE6FEC08DB4333BD1DD" box="[279,536,1943,1967]" italics="true" pageId="6" pageNumber="295">Nanosesarma gomantaka</emphasis>
</taxonomicName>
<emphasis id="B953EA80FFABFFE6FDCA8DBB3376D1DD" bold="true" box="[541,597,1943,1967]" pageId="6" pageNumber="295">
<taxonomicNameLabel id="A26057FBFFABFFE6FDCA8DBB3376D1DD" box="[541,597,1943,1967]" pageId="6" pageNumber="295" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, ad, holotype male (CL 5.1 mm, CW 4.7 mm) (LFSC.ZRC-206), Chapora Estuary, Goa State, India. a, P2, dorsal view; b, P3, dorsal view; c, P4, dorsal view; d, P5, dorsal view.
</paragraph>
</caption>
<caption id="DF58661AFFAAFFE7FF40880F357DD42D" ID-DOI="http://doi.org/10.5281/zenodo.10954105" ID-Zenodo-Dep="10954105" httpUri="https://zenodo.org/record/10954105/files/figure.png" pageId="7" pageNumber="296" startId="7.[151,250,547,571]" targetBox="[151,1435,181,523]" targetPageId="7" targetType="figure">
<paragraph id="8B983692FFAAFFE7FF40880F357DD42D" blockId="7.[151,1436,547,608]" pageId="7" pageNumber="296">
<emphasis id="B953EA80FFAAFFE7FF40880F302FD449" bold="true" box="[151,268,547,571]" pageId="7" pageNumber="296">FIGURE 6</emphasis>
<taxonomicName id="4C274D11FFAAFFE7FEC588083337D449" authority="Trivedi &amp; Padate &amp; Ng, 2024" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[274,532,547,571]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="gomantaka" status="sp. nov.">
<emphasis id="B953EA80FFAAFFE7FEC588083337D449" box="[274,532,547,571]" italics="true" pageId="7" pageNumber="296">Nanosesarma gomantaka</emphasis>
</taxonomicName>
<emphasis id="B953EA80FFAAFFE7FDCE88083372D44E" bold="true" box="[537,593,548,572]" pageId="7" pageNumber="296">
<taxonomicNameLabel id="A26057FBFFAAFFE7FDCE88083372D44E" box="[537,593,548,572]" pageId="7" pageNumber="296" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype male (CL 5.5 mm, CW 5.1 mm) (LFSC.ZRC-207), Chapora Estuary, Goa State, India. a, frontal view of cephalothorax; b, thoracic sternites 15 and sternopleonal cavity.
</paragraph>
</caption>
<paragraph id="8B983692FFAAFFE7FF1089A63434D2D2" blockId="7.[151,1437,690,1364]" pageId="7" pageNumber="296">
<bibRefCitation id="EFB64B63FFAAFFE7FF1089A63046D5D1" author="Serene, R." box="[199,357,905,932]" pageId="7" pageNumber="296" pagination="817 - 827" refId="ref5863" refString="Serene, R. (1967) Sur deux especes nouvelles de Brachyoures (Crustaces Decapodes) et sur une troisieme peu connue recoltees dans la region Malaise. Bulletin du Museum National d'histoire Naturelle, Serie 2 e, 38 (6), 817 - 827." type="journal article" year="1967">Serène (1967)</bibRefCitation>
described
<taxonomicName id="4C274D11FFAAFFE7FE0889A633FED5D6" authorityName="SERENE" authorityYear="1967" box="[479,733,906,932]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="tweediei">
<emphasis id="B953EA80FFAAFFE7FE0889A633FED5D6" box="[479,733,906,932]" italics="true" pageId="7" pageNumber="296">Nanosesarma tweediei</emphasis>
</taxonomicName>
on the basis of specimens collected from
<collectingCountry id="F3307602FFAAFFE7FB6189A6343CD5D6" box="[1206,1311,906,932]" name="Malaysia" pageId="7" pageNumber="296">Malaysia</collectingCountry>
,
<collectingCountry id="F3307602FFAAFFE7FAFC89A634BED5D6" box="[1323,1437,906,932]" name="Singapore" pageId="7" pageNumber="296">Singapore</collectingCountry>
and
<collectingCountry id="F3307602FFAAFFE7FF1F89823009D5BA" box="[200,298,942,968]" name="Vietnam" pageId="7" pageNumber="296">Vietnam</collectingCountry>
; the
<typeStatus id="549C8830FFAAFFE7FEB4898230E6D5BA" box="[355,453,942,968]" pageId="7" pageNumber="296" type="holotype">holotype</typeStatus>
being a male specimen (CL
<quantity id="4CDF9B77FFAAFFE7FCC68981324FD5BA" box="[785,876,941,968]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="7" pageNumber="296" unit="mm" value="5.8">5.8 mm</quantity>
; CW 5.0 mm) from Mersing, southern
<collectingCountry id="F3307602FFAAFFE7FAF8898234BAD5BA" box="[1327,1433,942,968]" name="Malaysia" pageId="7" pageNumber="296">Malaysia</collectingCountry>
.
<bibRefCitation id="EFB64B63FFAAFFE7FF4089FE3014D599" author="Serene, R." box="[151,311,977,1004]" pageId="7" pageNumber="296" pagination="817 - 827" refId="ref5863" refString="Serene, R. (1967) Sur deux especes nouvelles de Brachyoures (Crustaces Decapodes) et sur une troisieme peu connue recoltees dans la region Malaise. Bulletin du Museum National d'histoire Naturelle, Serie 2 e, 38 (6), 817 - 827." type="journal article" year="1967">Serène (1967)</bibRefCitation>
compared
<taxonomicName id="4C274D11FFAAFFE7FE6189FE3315D59E" authorityName="SERENE" authorityYear="1967" box="[438,566,978,1004]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="tweediei">
<emphasis id="B953EA80FFAAFFE7FE6189FE3315D59E" box="[438,566,978,1004]" italics="true" pageId="7" pageNumber="296">N. tweediei</emphasis>
</taxonomicName>
with
<taxonomicName id="4C274D11FFAAFFE7FDAE89FE329ED59E" authority="(De Man, 1887)" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[633,957,978,1004]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="minutum">
<emphasis id="B953EA80FFAAFFE7FDAE89FE33DED599" box="[633,765,978,1003]" italics="true" pageId="7" pageNumber="296">N. minutum</emphasis>
(
<bibRefCitation id="EFB64B63FFAAFFE7FCD989FE3296D59E" author="De Man, J. G." box="[782,949,978,1004]" pageId="7" pageNumber="296" pagination="639 - 722" refId="ref4462" refString="De Man, J. G. (1887) Uebersicht der indo-pacifischen Arten der Gattung Sesarma Say, nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschriebenen Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2, 639 - 722. [https: // biostor. org / reference / 4033]" type="journal article" year="1887">De Man, 1887</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C274D11FFAAFFE7FC2089FE3460D59E" authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1015,1347,977,1004]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="vestitum">
<emphasis id="B953EA80FFAAFFE7FC2089FE3529D599" box="[1015,1034,978,1003]" italics="true" pageId="7" pageNumber="296">N</emphasis>
.
<emphasis id="B953EA80FFAAFFE7FBCE89FE3555D599" box="[1049,1142,978,1003]" italics="true" pageId="7" pageNumber="296">vestitum</emphasis>
(Stimpson, 1858)
</taxonomicName>
but not with
<taxonomicName id="4C274D11FFAAFFE7FF0489DA30BBD27D" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[211,408,1014,1039]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="pontianacense">
<emphasis id="B953EA80FFAAFFE7FF0489DA30BBD27D" box="[211,408,1014,1039]" italics="true" pageId="7" pageNumber="296">N. pontianacense</emphasis>
</taxonomicName>
, and appears to have not considered this species. In his later work (
<bibRefCitation id="EFB64B63FFAAFFE7FB6789DA31E6D246" author="Serene, R. &amp; Soh, C. L." pageId="7" pageNumber="296" pagination="387 - 416" refId="ref5912" refString="Serene, R. &amp; Soh, C. L. (1970) New Indo-Pacific genera allied to Sesarma Say 1817 (Brachyura, Decapoda, Crustacea). Treubia, 27 (4), 387 - 416." type="journal article" year="1970">Serène &amp; Soh 1970: 394</bibRefCitation>
), he corrected himself when he noted that “
<taxonomicName id="4C274D11FFAAFFE7FD628E363545D241" authority="SERENE 1967" authorityName="SERENE" authorityYear="1967" box="[693,1126,1049,1076]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="tweediei">
<emphasis id="B953EA80FFAAFFE7FD628E363297D246" box="[693,948,1050,1076]" italics="true" pageId="7" pageNumber="296">Nanosesarma tweediei</emphasis>
<bibRefCitation id="EFB64B63FFAAFFE7FC6A8E363545D241" author="Serene, R." box="[957,1126,1049,1076]" pageId="7" pageNumber="296" pagination="817 - 827" refId="ref5863" refString="Serene, R. (1967) Sur deux especes nouvelles de Brachyoures (Crustaces Decapodes) et sur une troisieme peu connue recoltees dans la region Malaise. Bulletin du Museum National d'histoire Naturelle, Serie 2 e, 38 (6), 817 - 827." type="journal article" year="1967">SERENE 1967</bibRefCitation>
</taxonomicName>
corresponds to the male of
<taxonomicName id="4C274D11FFAAFFE7FF408E123041D225" authorityName="De Man" authorityYear="1895" box="[151,354,1086,1112]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="pontianacensis">
<emphasis id="B953EA80FFAAFFE7FF408E123189D225" box="[151,170,1086,1111]" italics="true" pageId="7" pageNumber="296">N</emphasis>
.
<emphasis id="B953EA80FFAAFFE7FF6D8E123041D225" box="[186,354,1086,1111]" italics="true" pageId="7" pageNumber="296">pontianacensis</emphasis>
</taxonomicName>
and is its synonym. The two species are marine and have the same habitat.”
<bibRefCitation id="EFB64B63FFAAFFE7FB0B8E1234BBD22A" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[1244,1432,1086,1112]" pageId="7" pageNumber="296" pagination="1 - 286" refId="ref5434" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="B953EA80FFAAFFE7FAD08E123461D22A" box="[1287,1346,1086,1112]" italics="true" pageId="7" pageNumber="296">et al.</emphasis>
(2008)
</bibRefCitation>
, however, inadvertently left
<taxonomicName id="4C274D11FFAAFFE7FE1C8E4E336BD20E" authorityName="SERENE" authorityYear="1967" box="[459,584,1122,1148]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="tweediei">
<emphasis id="B953EA80FFAAFFE7FE1C8E4E336BD20E" box="[459,584,1122,1148]" italics="true" pageId="7" pageNumber="296">N. tweediei</emphasis>
</taxonomicName>
as a valid species of
<taxonomicName id="4C274D11FFAAFFE7FCE28E4E32ECD209" authorityName=", Tweedie" authorityYear="1950" box="[821,975,1122,1147]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B953EA80FFAAFFE7FCE28E4E32ECD209" box="[821,975,1122,1147]" italics="true" pageId="7" pageNumber="296">Nanosesarma</emphasis>
</taxonomicName>
. We have compared the material on hand and we agree with
<bibRefCitation id="EFB64B63FFAAFFE7FEBC8EAA3349D2D2" author="Serene, R. &amp; Soh, C. L." box="[363,618,1158,1184]" pageId="7" pageNumber="296" pagination="387 - 416" refId="ref5912" refString="Serene, R. &amp; Soh, C. L. (1970) New Indo-Pacific genera allied to Sesarma Say 1817 (Brachyura, Decapoda, Crustacea). Treubia, 27 (4), 387 - 416." type="journal article" year="1970">Serène &amp; Sohs (1970)</bibRefCitation>
decision - there are no major characters distinguishing them.
</paragraph>
<paragraph id="8B983692FFAAFFE7FF108E86353AD326" blockId="7.[151,1437,690,1364]" pageId="7" pageNumber="296">
The specimens of
<taxonomicName id="4C274D11FFAAFFE7FE4B8E863342D2B1" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[412,609,1194,1219]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="pontianacense">
<emphasis id="B953EA80FFAAFFE7FE4B8E863342D2B1" box="[412,609,1194,1219]" italics="true" pageId="7" pageNumber="296">N. pontianacense</emphasis>
</taxonomicName>
from Southeast Asia examined in the present study agree well with the descriptions and figures by (
<bibRefCitation id="EFB64B63FFAAFFE7FE368EE233A6D29A" author="De Man, J. G." box="[481,645,1229,1256]" pageId="7" pageNumber="296" pagination="485 - 609" refId="ref4620" refString="De Man, J. G. (1895) Berichte uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4), 485 - 609. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man 1895</bibRefCitation>
;
<bibRefCitation id="EFB64B63FFAAFFE7FD438EE23214D29A" author="Tweedie, M. W. F." box="[660,823,1230,1256]" pageId="7" pageNumber="296" pagination="88 - 113" refId="ref6086" refString="Tweedie, M. W. F. (1940) New and interesting Malaysian species of Sesarma and Utica (Crustacea Brachyura). Bulletin of the Raffles Museum, 16, 88 - 113." type="journal article" year="1940">Tweedie 1940</bibRefCitation>
;
<bibRefCitation id="EFB64B63FFAAFFE7FC918EE232FBD295" author="Serene, R." box="[838,984,1229,1256]" pageId="7" pageNumber="296" pagination="817 - 827" refId="ref5863" refString="Serene, R. (1967) Sur deux especes nouvelles de Brachyoures (Crustaces Decapodes) et sur une troisieme peu connue recoltees dans la region Malaise. Bulletin du Museum National d'histoire Naturelle, Serie 2 e, 38 (6), 817 - 827." type="journal article" year="1967">Serène 1967</bibRefCitation>
).
<taxonomicName id="4C274D11FFAAFFE7FC388EE2342CD29A" authority="Trivedi &amp; Padate &amp; Ng, 2024" authorityName="Trivedi &amp; Padate &amp; Ng" authorityYear="2024" box="[1007,1295,1230,1256]" class="Malacostraca" family="Sesarmidae" genus="Nanosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="gomantaka" status="sp. nov.">
<emphasis id="B953EA80FFAAFFE7FC388EE2342CD29A" box="[1007,1295,1230,1256]" italics="true" pageId="7" pageNumber="296">Nanosesarma gomantaka</emphasis>
</taxonomicName>
<emphasis id="B953EA80FFAAFFE7FACD8EE23478D29A" bold="true" box="[1306,1371,1230,1256]" pageId="7" pageNumber="296">
<taxonomicNameLabel id="A26057FBFFAAFFE7FACD8EE23478D29A" box="[1306,1371,1230,1256]" pageId="7" pageNumber="296" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
most closely resembles
<taxonomicName id="4C274D11FFAAFFE7FEB28EDE330DD379" authorityName="De Man" authorityYear="1895" box="[357,558,1266,1291]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="pontianacensis">
<emphasis id="B953EA80FFAAFFE7FEB28EDE330DD379" box="[357,558,1266,1291]" italics="true" pageId="7" pageNumber="296">N. pontianacensis</emphasis>
</taxonomicName>
but can easily be distinguished in the form of the frontal lobes and strength of the median notch, shape of the carapace, notably the lateral margins, structure of the external orbital angle, as well as proportions of the male pleon (notably that of somite 6) and the G1 (
<tableCitation id="C6A50329FFAAFFE7FC628F16352BD326" box="[949,1032,1338,1364]" captionStart="TABLE 1" captionStartId="7.[152,244,1411,1435]" captionTargetBox="[163,1418,1487,1859]" captionTargetPageId="7" captionText="TABLE 1. Morphological comparison between Nanosesarma gomantaka n. sp. and Nanosesarma pontianacense (De Man, 1895)." httpUri="http://table.plazi.org/id/DF58661AFFAAFFE7FF4F8FAF3038D3CD" pageId="7" pageNumber="296" tableUuid="DF58661AFFAAFFE7FF4F8FAF3038D3CD">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>