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<document id="0B9573DC5ADA43C1E8C8703ADA6DF02E" ID-CLB-Dataset="26024" ID-DOI="10.11646/zootaxa.4656.2.11" ID-GBIF-Dataset="a796f578-1291-4e1e-9a39-8edf97e99954" ID-ISSN="1175-5326" ID-Zenodo-Dep="3368127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565768028779" checkinUser="plazi" docAuthor="Xu, Hao, Zhang, Chuntian &amp; Yao, Zhiyuan" docDate="2019" docId="E97E87DAE70D047E8DA23481EB0FFCC8" docLanguage="en" docName="zootaxa.4656.2.11.pdf" docOrigin="Zootaxa 4656 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pholcus mixiaoqii Xu &amp; Zhang &amp; Yao 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="388" masterDocId="1547FFA2E70C047A8D353152E82AFFF2" masterDocTitle="Pholcus mixiaoqii sp. nov., one new spider species from Tibet, China (Araneae: Pholcidae)" masterLastPageNumber="388" masterPageNumber="384" pageNumber="385" updateTime="1698750901774" updateUser="ExternalLinkService">
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<mods:title id="2BC05CF6C3D8BE49366FEE78939DABEC">Pholcus mixiaoqii sp. nov., one new spider species from Tibet, China (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="5A03EA8D8C4BF25C5BF1942FC7ABABA8">Xu, Hao</mods:namePart>
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<mods:namePart id="2D48C31DA21A3151D2769625C24F5A89">Zhang, Chuntian</mods:namePart>
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<mods:namePart id="1AA4F0700AA62C4B0F2D3775087ED65C">Yao, Zhiyuan</mods:namePart>
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<mods:title id="5C4FE015516FC20037B3B6682C7C6D22">Zootaxa</mods:title>
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<mods:date id="9FA4E0260C727C061E83EA28265F9D6B">2019</mods:date>
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<mods:number id="5E35FCF6173EAF9F7E4BAD20E5A33A11">2019-08-14</mods:number>
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<treatment id="E97E87DAE70D047E8DA23481EB0FFCC8" ID-DOI="http://doi.org/10.5281/zenodo.5941433" ID-GBIF-Taxon="159133481" ID-Zenodo-Dep="5941433" LSID="urn:lsid:plazi:treatment:E97E87DAE70D047E8DA23481EB0FFCC8" httpUri="http://treatment.plazi.org/id/E97E87DAE70D047E8DA23481EB0FFCC8" lastPageId="4" lastPageNumber="388" pageId="1" pageNumber="385">
<subSubSection id="29CD6547E70D047B8DA23481EA73FA1C" box="[151,601,1491,1518]" pageId="1" pageNumber="385" type="nomenclature">
<paragraph id="616836CCE70D047B8DA23481EA73FA1C" blockId="1.[151,1437,1491,2022]" box="[151,601,1491,1518]" pageId="1" pageNumber="385">
<heading id="3A2081A0E70D047B8DA23481EA73FA1C" bold="true" box="[151,601,1491,1518]" fontSize="11" level="1" pageId="1" pageNumber="385" reason="1">
<emphasis id="53A3EADEE70D047B8DA23481EA73FA1C" bold="true" box="[151,601,1491,1518]" pageId="1" pageNumber="385">
<taxonomicName id="A6D74D4FE70D047B8DA23481E958FA1C" ID-CoL="95GG5" authority="Xu &amp; Zhang &amp; Yao, 2019" authorityName="Xu &amp; Zhang &amp; Yao" authorityYear="2019" box="[151,370,1491,1518]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="mixiaoqii" status="sp. nov.">
<emphasis id="53A3EADEE70D047B8DA23481E958FA1C" bold="true" box="[151,370,1491,1518]" italics="true" pageId="1" pageNumber="385">Pholcus mixiaoqii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="489057A5E70D047B8C4F3486E9FDFA1C" box="[378,471,1492,1518]" pageId="1" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="F9EC2A49E70D047B8CD23486EA78FA1C" box="[487,594,1492,1518]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="2.[151,244,1940,1962]" captionStartId-1="3.[151,244,1899,1921]" captionTargetBox-0="[204,1383,192,1912]" captionTargetBox-1="[212,1375,167,1879]" captionTargetId-0="figure@2.[198,1389,167,1919]" captionTargetId-1="figure@3.[212,1375,167,1879]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 2. Pholcus mixiaoqii sp. nov., holotype male.A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis); C–D. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow points at two prolatero-dorsal spines). a = appendix, b = bulb, e = embolus, pr = procursus, u = uncus. Scale bars: 0.20 (A–B), 0.05 (C–D)." captionText-1="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri-0="https://zenodo.org/record/3368131/files/figure.png" httpUri-1="https://zenodo.org/record/3368133/files/figure.png" pageId="1" pageNumber="385">Figs 2–3</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="29CD6547E70D047E8DA234A8EB0FFCC8" lastPageId="4" lastPageNumber="388" pageId="1" pageNumber="385" type="materials_examined">
<paragraph id="616836CCE70D047B8DA234A8EAC3F9AB" blockId="1.[151,1437,1491,2022]" pageId="1" pageNumber="385">
<emphasis id="53A3EADEE70D047B8DA234A8E914F9E3" bold="true" box="[151,318,1529,1554]" pageId="1" pageNumber="385">Type material.</emphasis>
<materialsCitation id="D1BF3C91E70D047B8C7334A8EB8CF9C7" ID-GBIF-Occurrence="2332527315" collectingDate="2015-06-15" collectorName="X. Mi &amp; K. Guo" country="China" county="Zanda County" elevation="2987" latitude="31.782984" location="Diya Village" longLatPrecision="1" longitude="78.87252" municipality="Diya Town" pageId="1" pageNumber="385" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="53A3EADEE70D047B8C7334A8E99DF9E0" bold="true" box="[326,439,1530,1554]" pageId="1" pageNumber="385">
<typeStatus id="BE6C886EE70D047B8C7334A8E998F9E0" box="[326,434,1530,1554]" pageId="1" pageNumber="385" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (SYNU-Ar00001),
<location id="64086017E70D047B8FFE34ABEB7EF9E3" LSID="urn:lsid:plazi:treatment:E97E87DAE70D047E8DA23481EB0FFCC8:64086017E70D047B8FFE34ABEB7EF9E3" box="[715,852,1529,1553]" country="China" county="Zanda County" latitude="31.782984" longLatPrecision="1" longitude="78.87252" municipality="Diya Town" name="Diya Village" pageId="1" pageNumber="385" stateProvince="Xizang">Diya Village</location>
(
<geoCoordinate id="04E3500BE70D047B8E5134ABEBD8F9E3" box="[868,1010,1529,1553]" degrees="31" direction="north" minutes="46.979" orientation="latitude" pageId="1" pageNumber="385" precision="1" value="31.782984">31°46.979’N</geoCoordinate>
,
<geoCoordinate id="04E3500BE70D047B8ECB34ABECA3F9E3" box="[1022,1161,1529,1553]" degrees="78" direction="east" minutes="52.351" orientation="longitude" pageId="1" pageNumber="385" precision="1" value="78.87252">78°52.351’E</geoCoordinate>
,
<elevation id="EAFAD1FFE70D047B89A034ABED7EF9E0" box="[1173,1364,1529,1554]" metricMagnitude="3" metricUnit="m" metricValue="2.9869999999999997" pageId="1" pageNumber="385" unit="m" value="2987.0">
elevation
<quantity id="A62F9B29E70D047B89CA34A8ED7EF9E0" box="[1279,1364,1530,1554]" metricMagnitude="3" metricUnit="m" metricValue="2.9869999999999997" pageId="1" pageNumber="385" unit="m" value="2987.0">2987 m</quantity>
</elevation>
),
<collectingMunicipality id="810CACB6E70D047B885D34ABE8FCF9C7" pageId="1" pageNumber="385">Diya Town</collectingMunicipality>
,
<collectingCounty id="88094E40E70D047B8DD7374FE956F9C4" box="[226,380,1565,1590]" pageId="1" pageNumber="385">Zanda County</collectingCounty>
,
<emphasis id="53A3EADEE70D047B8CB2374FE9E9F9C7" bold="true" box="[391,451,1565,1589]" pageId="1" pageNumber="385">
<collectingRegion id="A313F82EE70D047B8CB2374FE9E9F9C7" box="[391,451,1565,1589]" country="China" name="Xizang" pageId="1" pageNumber="385">Tibet</collectingRegion>
</emphasis>
,
<emphasis id="53A3EADEE70D047B8CE4374FEA3DF9C7" bold="true" box="[465,535,1565,1589]" pageId="1" pageNumber="385">
<collectingCountry id="19C0765CE70D047B8CE4374FEA3DF9C7" box="[465,535,1565,1589]" name="China" pageId="1" pageNumber="385">China</collectingCountry>
</emphasis>
,
<date id="1569100CE70D047B8F10374FEA92F9C7" box="[549,696,1565,1589]" pageId="1" pageNumber="385" value="2015-06-15">
<collectingDate id="052DE9E4E70D047B8F10374FEA92F9C7" box="[549,696,1565,1589]" pageId="1" pageNumber="385" value="2015-06-15">15 June 2015</collectingDate>
</date>
,
<collectorName id="CC22531AE70D047B8FF6374CEB2EF9C7" box="[707,772,1565,1589]" pageId="1" pageNumber="385">X. Mi</collectorName>
&amp;
<collectorName id="CC22531AE70D047B8E1D374CEB5DF9C4" box="[808,887,1566,1590]" pageId="1" pageNumber="385">K. Guo</collectorName>
leg.
</materialsCitation>
<emphasis id="53A3EADEE70D047B8E98374CEC02F9C4" bold="true" box="[941,1064,1566,1590]" pageId="1" pageNumber="385">
<typeStatus id="BE6C886EE70D047B8E98374CEC09F9C4" box="[941,1059,1566,1590]" pageId="1" pageNumber="385" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="77D1FD45E70D047B8905374CECA8F9C7" box="[1072,1154,1565,1590]" pageId="1" pageNumber="385" type="male">2 males</specimenCount>
(SYNU-Ar00002–00003) and
<materialsCitation id="D1BF3C91E70D047B8DF13710EACFF9AB" ID-GBIF-Occurrence="2332527318" box="[196,741,1601,1626]" collectingDate="2015-06-15" collectorName="X. Mi &amp; K. Guo" country="China" county="Zanda County" elevation="2987" latitude="31.782984" location="Diya Village" longLatPrecision="1" longitude="78.87252" municipality="Diya Town" pageId="1" pageNumber="385" specimenCount="1" specimenCount-female="1" stateProvince="Xizang" typeStatus="paratype">
<specimenCount id="77D1FD45E70D047B8DF13710E90BF9AB" box="[196,289,1601,1626]" pageId="1" pageNumber="385" type="female">1 female</specimenCount>
(SYNU-Ar00004), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="616836CCE70D047B8DF23734E9BDF953" blockId="1.[151,1437,1491,2022]" pageId="1" pageNumber="385">
<emphasis id="53A3EADEE70D047B8DF23734E962F98C" bold="true" box="[199,328,1638,1662]" pageId="1" pageNumber="385">Etymology.</emphasis>
The specific name is a patronym in honor of the collector Xiaoqi Mi (Tongren University), and is a noun (name) in genitive case.
</paragraph>
<paragraph id="616836CCE70D047B8DF237FFEA69F888" blockId="1.[151,1437,1491,2022]" pageId="1" pageNumber="385">
<emphasis id="53A3EADEE70D047B8DF237FFE916F937" bold="true" box="[199,316,1709,1733]" pageId="1" pageNumber="385">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="A6D74D4FE70D047B8F0A37FFEBF0F934" authority="Yao &amp; Li, 2017" authorityName="Yao &amp; Li" authorityYear="2017" box="[575,986,1709,1734]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="shuguanensis">
<emphasis id="53A3EADEE70D047B8F0A37FFEB06F937" box="[575,812,1709,1733]" italics="true" pageId="1" pageNumber="385">Pholcus shuguanensis</emphasis>
Yao &amp; Li, 2017
</taxonomicName>
(see
<bibRefCitation id="05464B3DE70D047B892137FCECE6F934" author="Dong, T. &amp; Zheng, G. &amp; Davlatov, A. &amp; Yao, Z. &amp; Li, S." box="[1044,1228,1709,1734]" pageId="1" pageNumber="385" pagination="1 - 14" refId="ref2149" refString="Dong, T., Zheng, G., Davlatov, A., Yao, Z. &amp; Li, S. (2017) Two new species of the spider genus Pholcus Walckenaer, 1805 (Araneae, Pholcidae) from Tajikistan, with the first description of female Pholcus sidorenkoi Dunin, 1994. European Journal of Taxonomy, 301, 1 - 14. https: // doi. org / 10.5852 / ejt. 2017.30 1" type="journal article" year="2017">
Dong
<emphasis id="53A3EADEE70D047B896337FCECA6F937" box="[1110,1164,1709,1733]" italics="true" pageId="1" pageNumber="385">et al.</emphasis>
2017
</bibRefCitation>
: figs 3A–D, 4A–E) and
<taxonomicName id="A6D74D4FE70D047B8DF63783EA3DF918" authority="Dunin, 1994" authorityName="Dunin" authorityYear="1994" box="[195,535,1745,1770]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="sidorenkoi">
<emphasis id="53A3EADEE70D047B8DF63783E9A5F91B" box="[195,399,1745,1769]" italics="true" pageId="1" pageNumber="385">Pholcus sidorenkoi</emphasis>
<bibRefCitation id="05464B3DE70D047B8CA13783EA3DF918" author="Dunin, P. M." box="[404,535,1745,1770]" pageId="1" pageNumber="385" pagination="136 - 138" refId="ref2231" refString="Dunin, P. M. (1994) Pholcus sidorenkoi sp. n., a new species of pholcid spiders (Aranei, Haplogynae, Pholcidae) from the Volga region. Zoologicheskii Zhurnal, 73 (3), 136 - 138." type="journal article" year="1994">Dunin, 1994</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="05464B3DE70D047B8F7B3783EAE5F918" author="Dunin, P. M." box="[590,719,1745,1770]" pageId="1" pageNumber="385" pagination="136 - 138" refId="ref2231" refString="Dunin, P. M. (1994) Pholcus sidorenkoi sp. n., a new species of pholcid spiders (Aranei, Haplogynae, Pholcidae) from the Volga region. Zoologicheskii Zhurnal, 73 (3), 136 - 138." type="journal article" year="1994">Dunin 1994</bibRefCitation>
: figs 1–7;
<bibRefCitation id="05464B3DE70D047B8E0F3783EB91F918" author="Huber, B. A." box="[826,955,1745,1770]" pageId="1" pageNumber="385" pagination="1 - 509" refId="ref2394" refString="Huber, B. A. (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner Zoologische Monographien, 58, 1 - 509." type="journal article" year="2011">Huber 2011</bibRefCitation>
: figs 1538,
<date id="1569100CE70D047B89063783EC84F91B" box="[1075,1198,1745,1769]" pageId="1" pageNumber="385" value="1642" valueMax="1644">1642–1644</date>
;
<bibRefCitation id="05464B3DE70D047B898D3780ED41F918" author="Dong, T. &amp; Zheng, G. &amp; Davlatov, A. &amp; Yao, Z. &amp; Li, S." box="[1208,1387,1745,1770]" pageId="1" pageNumber="385" pagination="1 - 14" refId="ref2149" refString="Dong, T., Zheng, G., Davlatov, A., Yao, Z. &amp; Li, S. (2017) Two new species of the spider genus Pholcus Walckenaer, 1805 (Araneae, Pholcidae) from Tajikistan, with the first description of female Pholcus sidorenkoi Dunin, 1994. European Journal of Taxonomy, 301, 1 - 14. https: // doi. org / 10.5852 / ejt. 2017.30 1" type="journal article" year="2017">
Dong
<emphasis id="53A3EADEE70D047B89CD3780ED07F91B" box="[1272,1325,1745,1769]" italics="true" pageId="1" pageNumber="385">et al.</emphasis>
2017
</bibRefCitation>
: figs 5A–D, 6A–H) with similar male chelicerae (
<figureCitation id="F9EC2A49E70D047B8F4437A7EAEEF8FC" box="[625,708,1781,1806]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="1" pageNumber="385">Fig. 3D</figureCitation>
) but can be distinguished by procursus prolatero-distally sclerotized (
<figureCitation id="F9EC2A49E70D047B8DAA364BE93DF8C0" box="[159,279,1817,1842]" captionStart="FIGURE 2" captionStartId="2.[151,244,1940,1962]" captionTargetBox="[204,1383,192,1912]" captionTargetId="figure@2.[198,1389,167,1919]" captionTargetPageId="2" captionText="FIGURE 2. Pholcus mixiaoqii sp. nov., holotype male.A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis); C–D. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow points at two prolatero-dorsal spines). a = appendix, b = bulb, e = embolus, pr = procursus, u = uncus. Scale bars: 0.20 (A–B), 0.05 (C–D)." httpUri="https://zenodo.org/record/3368131/files/figure.png" pageId="1" pageNumber="385">Figs 2A, C</figureCitation>
) and by uncus distally wide (
<figureCitation id="F9EC2A49E70D047B8F60364BEA83F8C0" box="[597,681,1817,1842]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="1" pageNumber="385">Fig. 3C</figureCitation>
); also distinguished from
<taxonomicName id="A6D74D4FE70D047B8EF43648EC46F8C3" authorityName="Yao &amp; Li" authorityYear="2017" box="[961,1132,1817,1841]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="shuguanensis">
<emphasis id="53A3EADEE70D047B8EF43648EC46F8C3" box="[961,1132,1817,1841]" italics="true" pageId="1" pageNumber="385">P. shuguanensis</emphasis>
</taxonomicName>
by appendix L-shaped (
<figureCitation id="F9EC2A49E70D047B8844364BE893F8A4" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="1" pageNumber="385">Fig. 3C</figureCitation>
); also distinguished from
<taxonomicName id="A6D74D4FE70D047B8CFB366CEA72F8A7" authorityName="Dunin" authorityYear="1994" box="[462,600,1853,1877]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="sidorenkoi">
<emphasis id="53A3EADEE70D047B8CFB366CEA72F8A7" box="[462,600,1853,1877]" italics="true" pageId="1" pageNumber="385">P. sidorenkoi</emphasis>
</taxonomicName>
by epigynum with pair of lateral apophyses anteriorly (arrows in
<figureCitation id="F9EC2A49E70D047B8820366FED4DF8A4" box="[1301,1383,1853,1878]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="1" pageNumber="385">Fig. 3A</figureCitation>
) and by pore plates nearly rounded (
<figureCitation id="F9EC2A49E70D047B8CD63633EA1CF888" box="[483,566,1889,1914]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="1" pageNumber="385">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="616836CCE70D047E8DF236D7EBC1FDE8" blockId="1.[151,1437,1491,2022]" lastBlockId="4.[151,1437,153,826]" lastPageId="4" lastPageNumber="388" pageId="1" pageNumber="385">
<emphasis id="53A3EADEE70D047B8DF236D7EA32F86F" bold="true" box="[199,536,1925,1950]" pageId="1" pageNumber="385">
Description. Male (
<typeStatus id="BE6C886EE70D047B8C9636D7EA21F86F" box="[419,523,1925,1949]" pageId="1" pageNumber="385" type="holotype">holotype</typeStatus>
):
</emphasis>
Total length 5.56 (5.75 with clypeus), carapace 1.41 long, 1.63 wide, opisthosoma 4.15 long, 1.66 wide. Leg I: 33.13 (8.63 + 0.73 + 8.84 + 12.88 + 2.05), leg II: 23.89 (6.56 + 0.70 + 5.94 + 9.38 + 1.31), leg III: 17.08 (5.36 + 0.66 + 4.47 + 5.54 + 1.05), leg IV: 23.26 (6.68 + 0.68 + 5.79 + 8.81 + 1.30); tibia I L/d: 49. Distance PME-PME 0.29, diameter PME 0.14, distance PME-ALE 0.04, distance AME-AME 0.05, diameter AME 0.07. Sternum wider than long (1.11/0.86). Habitus as in
<figureCitation id="F9EC2A49E708047E8FA031ECEAFFFF24" box="[661,725,190,214]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="4" pageNumber="388">Figs 3</figureCitation>
E–F. Carapace yellowish, with brown marks medially; ocular area and clypeus yellowish; sternum yellowish, with brownish marks. Legs yellowish, but dark brown on patellae, without darker rings. Opisthosoma yellowish, without spots. Ocular area elevated, each eye triad on top of short laterally directed eye-stalk. Thoracic furrow absent. Chelicerae as in
<figureCitation id="F9EC2A49E708047E8FF63078EB32FEB0" box="[707,792,298,322]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="4" pageNumber="388">Fig. 3D</figureCitation>
, with pair of large proximo-lateral apophyses, pair of distal apophyses provided with two teeth each, and pair of frontal apophyses. Pedipalps as in
<figureCitation id="F9EC2A49E708047E8975301CECA8FE94" box="[1088,1154,334,358]" captionStart="FIGURE 2" captionStartId="2.[151,244,1940,1962]" captionTargetBox="[204,1383,192,1912]" captionTargetId="figure@2.[198,1389,167,1919]" captionTargetPageId="2" captionText="FIGURE 2. Pholcus mixiaoqii sp. nov., holotype male.A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis); C–D. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow points at two prolatero-dorsal spines). a = appendix, b = bulb, e = embolus, pr = procursus, u = uncus. Scale bars: 0.20 (A–B), 0.05 (C–D)." httpUri="https://zenodo.org/record/3368131/files/figure.png" pageId="4" pageNumber="388">Figs 2</figureCitation>
A–B; trochanter with long ventral apophysis; femur with small dorsal apophysis proximally (arrow in
<figureCitation id="F9EC2A49E708047E8EFB3020EC0BFE78" box="[974,1057,370,394]" captionStart="FIGURE 2" captionStartId="2.[151,244,1940,1962]" captionTargetBox="[204,1383,192,1912]" captionTargetId="figure@2.[198,1389,167,1919]" captionTargetPageId="2" captionText="FIGURE 2. Pholcus mixiaoqii sp. nov., holotype male.A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis); C–D. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow points at two prolatero-dorsal spines). a = appendix, b = bulb, e = embolus, pr = procursus, u = uncus. Scale bars: 0.20 (A–B), 0.05 (C–D)." httpUri="https://zenodo.org/record/3368131/files/figure.png" pageId="4" pageNumber="388">Fig. 2B</figureCitation>
) and distinct ventral protuberance; procursus simple, with two prolatero-dorsal spines (arrow in
<figureCitation id="F9EC2A49E708047E8E0D30C4EBBAFE5C" box="[824,912,406,430]" captionStart="FIGURE 2" captionStartId="2.[151,244,1940,1962]" captionTargetBox="[204,1383,192,1912]" captionTargetId="figure@2.[198,1389,167,1919]" captionTargetPageId="2" captionText="FIGURE 2. Pholcus mixiaoqii sp. nov., holotype male.A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis); C–D. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow points at two prolatero-dorsal spines). a = appendix, b = bulb, e = embolus, pr = procursus, u = uncus. Scale bars: 0.20 (A–B), 0.05 (C–D)." httpUri="https://zenodo.org/record/3368131/files/figure.png" pageId="4" pageNumber="388">Fig. 2D</figureCitation>
); uncus with scaly edge (
<figureCitation id="F9EC2A49E708047E899F30C4ED2BFE5C" box="[1194,1281,406,430]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="4" pageNumber="388">Fig. 3C</figureCitation>
); appendix Lshaped, flat and large distally (
<figureCitation id="F9EC2A49E708047E8CD130E8EA02FE20" box="[484,552,442,466]" captionStart="FIGURE 2" captionStartId="2.[151,244,1940,1962]" captionTargetBox="[204,1383,192,1912]" captionTargetId="figure@2.[198,1389,167,1919]" captionTargetPageId="2" captionText="FIGURE 2. Pholcus mixiaoqii sp. nov., holotype male.A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis); C–D. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow points at two prolatero-dorsal spines). a = appendix, b = bulb, e = embolus, pr = procursus, u = uncus. Scale bars: 0.20 (A–B), 0.05 (C–D)." httpUri="https://zenodo.org/record/3368131/files/figure.png" pageId="4" pageNumber="388">Figs 2</figureCitation>
A–B, 3C); embolus weakly sclerotized, with some transparent projections distally (
<figureCitation id="F9EC2A49E708047E8DAA308CE8DEFE04" box="[159,244,478,502]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="4" pageNumber="388">Fig. 3C</figureCitation>
). Retrolateral trichobothrium of tibia I at 6% proximally; legs with short vertical setae on tibiae, metatarsi and tarsi, without spines and curved setae; tarsus I with 28 distinct pseudosegments.
</paragraph>
<caption id="35A86644E70E04788DA236C6EC1FF801" httpUri="https://zenodo.org/record/3368131/files/figure.png" pageId="2" pageNumber="386" startId="2.[151,244,1940,1962]" targetBox="[204,1383,192,1912]" targetPageId="2">
<paragraph id="616836CCE70E04788DA236C6EC1FF801" blockId="2.[151,1437,1940,2035]" pageId="2" pageNumber="386">
<emphasis id="53A3EADEE70E04788DA236C6E926F85B" bold="true" box="[151,268,1940,1962]" pageId="2" pageNumber="386">FIGURE 2.</emphasis>
<taxonomicName id="A6D74D4FE70E04788C2436C6E995F858" authority="Xu &amp; Zhang &amp; Yao, 2019" authorityName="Xu &amp; Zhang &amp; Yao" authorityYear="2019" box="[273,447,1940,1962]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="mixiaoqii" status="sp. nov.">
<emphasis id="53A3EADEE70E04788C2436C6E995F858" box="[273,447,1940,1962]" italics="true" pageId="2" pageNumber="386">Pholcus mixiaoqii</emphasis>
</taxonomicName>
<emphasis id="53A3EADEE70E04788CF136C7EA3AF858" bold="true" box="[452,528,1941,1962]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="489057A5E70E04788CF136C7EA3AF858" box="[452,528,1941,1962]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis); C–D. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow points at two prolatero-dorsal spines). a = appendix, b = bulb, e = embolus, pr = procursus, u = uncus. Scale bars: 0.20 (A–B), 0.05 (C–D).
</paragraph>
</caption>
<caption id="35A86644E70F04798DA23639EB4CF81C" httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="3" pageNumber="387" startId="3.[151,244,1899,1921]" targetBox="[212,1375,167,1879]" targetPageId="3">
<paragraph id="616836CCE70F04798DA23639EB4CF81C" blockId="3.[151,1436,1899,2030]" pageId="3" pageNumber="387">
<emphasis id="53A3EADEE70F04798DA23639E924F873" bold="true" box="[151,270,1899,1921]" pageId="3" pageNumber="387">FIGURE 3.</emphasis>
<taxonomicName id="A6D74D4FE70F04798C20363EE9EEF873" authority="Xu &amp; Zhang &amp; Yao, 2019" authorityName="Xu &amp; Zhang &amp; Yao" authorityYear="2019" box="[277,452,1900,1921]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="mixiaoqii" status="sp. nov.">
<emphasis id="53A3EADEE70F04798C20363EE9EEF873" box="[277,452,1900,1921]" italics="true" pageId="3" pageNumber="387">Pholcus mixiaoqii</emphasis>
</taxonomicName>
<emphasis id="53A3EADEE70F04798CFE363EEA32F873" bold="true" box="[459,536,1900,1921]" pageId="3" pageNumber="387">
<taxonomicNameLabel id="489057A5E70F04798CFE363EEA32F873" box="[459,536,1900,1921]" pageId="3" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H).
</paragraph>
</caption>
<paragraph id="616836CCE708047E8DF23374EA43FD3C" blockId="4.[151,1437,153,826]" pageId="4" pageNumber="388">
<emphasis id="53A3EADEE708047E8DF23374E909FDCC" bold="true" box="[199,291,550,574]" pageId="4" pageNumber="388">Female:</emphasis>
Similar to male, habitus as in
<figureCitation id="F9EC2A49E708047E8F453374EA98FDCC" box="[624,690,550,574]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="4" pageNumber="388">Figs 3</figureCitation>
G–H. Total length 5.77 (5.91 with clypeus), carapace 1.48 long, 1.55 wide, opisthosoma 4.29 long, 1.94 wide; tibia I: 7.76; tibia I L/d: 49. Distance PME-PME 0.26, diameter PME 0.13, distance PME-ALE 0.04, distance AME-AME 0.05, diameter AME 0.07. Sternum wider than long (1.05/0.82). Epigynum (
<figureCitation id="F9EC2A49E708047E8DAA33C0E8DBFD58" box="[159,241,658,682]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="4" pageNumber="388">Fig. 3A</figureCitation>
) with knob and pair of lateral apophyses anteriorly (arrows in
<figureCitation id="F9EC2A49E708047E8EB233C0EBF1FD58" box="[903,987,658,682]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="4" pageNumber="388">Fig. 3A</figureCitation>
). Vulva (
<figureCitation id="F9EC2A49E708047E890E33C0ECA6FD58" box="[1083,1164,658,682]" captionStart="FIGURE 3" captionStartId="3.[151,244,1899,1921]" captionTargetBox="[212,1375,167,1879]" captionTargetId="figure@3.[212,1375,167,1879]" captionTargetPageId="3" captionText="FIGURE 3. Pholcus mixiaoqii sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum, ventral view, arrows point at lateral apophyses; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). a = appendix, da = distal apophysis, e = embolus, fa = frontal apophysis, pa = proximo-lateral apophysis, pp = pore plate, u = uncus. Scale bars: 0.10 (A–D), 1.00 (E–H)." httpUri="https://zenodo.org/record/3368133/files/figure.png" pageId="4" pageNumber="388">Fig. 3B</figureCitation>
) with sclerotized anterior arch and pair of nearly rounded pore plates.
</paragraph>
<paragraph id="616836CCE708047E8DF2338BEC49FD00" blockId="4.[151,1437,153,826]" box="[199,1123,729,754]" pageId="4" pageNumber="388">
<emphasis id="53A3EADEE708047E8DF2338BE916FD03" bold="true" box="[199,316,729,753]" pageId="4" pageNumber="388">Variation:</emphasis>
Tibia I in one male
<typeStatus id="BE6C886EE708047E8F273388EA44FD00" box="[530,622,730,754]" pageId="4" pageNumber="388" type="paratype">paratype</typeStatus>
(leg I missing in another male
<typeStatus id="BE6C886EE708047E8E8F3388EC36FD00" box="[954,1052,730,754]" pageId="4" pageNumber="388" type="paratype">paratype</typeStatus>
): 9.03.
</paragraph>
<paragraph id="616836CCE708047E8DF233AFEAFEFCE4" blockId="4.[151,1437,153,826]" box="[199,724,765,790]" pageId="4" pageNumber="388">
<emphasis id="53A3EADEE708047E8DF233AFE970FCE7" bold="true" box="[199,346,765,789]" pageId="4" pageNumber="388">Distribution.</emphasis>
<collectingCountry id="19C0765CE708047E8C5533ACE98AFCE4" box="[352,416,766,790]" name="China" pageId="4" pageNumber="388">China</collectingCountry>
(
<collectingRegion id="A313F82EE708047E8C9833ACE9C3FCE4" box="[429,489,766,790]" country="China" name="Xizang" pageId="4" pageNumber="388">Tibet</collectingRegion>
,
<typeStatus id="BE6C886EE708047E8CC133ACEA08FCE4" box="[500,546,766,790]" pageId="4" pageNumber="388">type</typeStatus>
locality;
<figureCitation id="F9EC2A49E708047E8FB333ACEAEEFCE4" box="[646,708,766,790]" captionStart="FIGURE 1" captionStartId="1.[151,244,1171,1193]" captionTargetBox="[151,1436,346,1150]" captionTargetId="figure@1.[151,1436,346,1150]" captionTargetPageId="1" captionText="FIGURE 1. Distribution records of the Pholcus nenjukovi species-group. 1. Pholcus afghanus Senglet, 2008 (Afghanistan); 2. Pholcus arsacius Senglet, 2008 (Iran); 3. Pholcus cophenius Senglet, 2008 (Afghanistan); 4. Pholcus mixiaoqii sp. nov. (Tibet, China); 5. Pholcus nenjukovi Spassky, 1936 (Tajikistan); 6. Pholcus parthicus Senglet, 2008 (Iran); 7. Pholcus saidovi Yao &amp; Li, 2017 (Tajikistan); 8. Pholcus shuguanensis Yao &amp; Li, 2017 (Tajikistan); 9. Pholcus sidorenkoi Dunin, 1994 (Tajikistan; for the single dubious record from Russia, see Huber 2011: 339, fig. 1636). Locality coordinates for P. parthicus, P. afghanus, P. arsacius, P. cophenius, P. saidovi, and P. shuguanensis were recovered from the original descriptions (see Senglet 2008, Dong et al. 2017), and for P.nenjukovi and P. sidorenkoi from other revisions (see Huber 2011, Dong et al. 2017)." httpUri="https://zenodo.org/record/3368129/files/figure.png" pageId="4" pageNumber="388">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="616836CCE708047E8DF23270EB0FFCC8" blockId="4.[151,1437,153,826]" box="[199,805,801,826]" pageId="4" pageNumber="388">
<emphasis id="53A3EADEE708047E8DF23270E956FCCB" bold="true" box="[199,380,801,826]" pageId="4" pageNumber="388">Natural history.</emphasis>
The species was found in an old house.
</paragraph>
</subSubSection>
</treatment>
</document>
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