treatments-xml/data/C5/12/87/C51287DE5B65FFE8FDFB4C36FE58FD1E.xml

<document id="E4D889825F44CC6E14E0B0AA8391A259" ID-CLB-Dataset="21972" ID-DOI="10.5852/ejt.2016.188" ID-GBIF-Dataset="94987553-aa5d-4444-a739-c8ff9bf54d30" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837557" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590066436525" checkinUser="valdenar" docAuthor="Horká, Ivona, Fransen, Charles H. J. M. &amp; Ďuriš, Zdeněk" docDate="2016" docId="C51287DE5B65FFE8FDFB4C36FE58FD1E" docLanguage="en" docName="ejt-2016-188.pdf.imf" docOrigin="European Journal of Taxonomy 188 (188)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hamodactylus paraqabai Horká, Fransen &amp; Ďuriš, 2016, sp. nov." docType="treatment" docUuid="D50ED903-8CDF-4B68-BE54-8E4F2177FB2B" docUuidSource="ZooBank" docVersion="7" lastPageNumber="12" masterDocId="392BFFA65B61FFE3FF8C4D07FF90FFFF" masterDocTitle="Two new species of shrimp of the Indo-West Pacific genus Hamodactylus Holthuis, 1952 (Crustacea: Decapoda: Palaemonidae)" masterLastPageNumber="26" masterPageNumber="1" pageNumber="5" updateTime="1698840598216" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C211AAD14962A9EBBFDEA7BE6885E96F">Two new species of shrimp of the Indo-West Pacific genus Hamodactylus Holthuis, 1952 (Crustacea: Decapoda: Palaemonidae)</mods:title>
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<mods:namePart id="E3463459E36BA64F75E944269FEF0B74">Horká, Ivona</mods:namePart>
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<mods:affiliation id="F35C786A85B05B82A9C8FE5DC5E75AF7">Department of Biology and Ecology, and Institute of Environmental Technologies, University of Ostrava, Chittussiho 10, CZ- 71000 Ostrava, Czech Republic. &amp; ivona. horka @ osu. cz &amp; urn: lsid: zoobank. org: author: CCA 47494 - EA 9 C- 46 D 9 - B 579 - 90772 B 584 F 35 &amp; Department of Ecology, Charles University, Viničná 7, CZ- 12844 Prague, Czech Republic.</mods:affiliation>
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<mods:namePart id="DE54C41C18C6B7019EE9D7448074C793">Fransen, Charles H. J. M.</mods:namePart>
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<mods:affiliation id="2D4ADC518FBBB8A7EB9BAAD119E411BA">Department of Taxonomy &amp; Systematics, Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands. &amp; charles. fransen @ naturalis. nl &amp; urn: lsid: zoobank. org: author: 08 C 8 BF 56 - A 737 - 4 B 4 F-BC 80 - 56333 AE 6 AB 3 A</mods:affiliation>
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<mods:namePart id="3D51B18E31D17A297BE880FCCADDFCDC">Ďuriš, Zdeněk</mods:namePart>
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<mods:affiliation id="FE8AB67980AF6AFE9D09A9C2B2B534D7">Department of Biology and Ecology, and Institute of Environmental Technologies, University of Ostrava, Chittussiho 10, CZ- 71000 Ostrava, Czech Republic. &amp; urn: lsid: zoobank. org: author: 0 CF 5 D 3 F 9 - 9663 - 4 B 76 - BF 91 - 713 D 9 BE 50 BC 3 &amp; Corresponding author: zdenek. duris @ osu. cz</mods:affiliation>
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<heading id="164C81A45B65FFE7FDFB4C36FB99FEB3" box="[631,1033,305,332]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
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<emphasis id="7FCFEADA5B65FFE7FDFB4C36FC36FEB3" bold="true" box="[631,934,305,332]" italics="true" pageId="4" pageNumber="5">Hamodactylus paraqabai</emphasis>
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<taxonomicNameLabel id="64FC57A15B65FFE7FC224C35FB99FEB3" box="[942,1033,306,332]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="4D0436C85B65FFE7FD504C70FC34FE6D" blockId="4.[401,1262,305,402]" box="[732,932,375,402]" pageId="4" pageNumber="5">
<figureCitation id="D5802A4D5B65FFE7FD504C70FCD4FE6D" box="[732,836,375,402]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="4.[227,270,2001,2027]" captionStartId-1="5.[227,270,1812,1838]" captionStartId-2="6.[227,270,1810,1836]" captionStartId-3="8.[227,270,1967,1993]" captionStartId-4="9.[225,268,1967,1993]" captionTargetBox-0="[254,1423,1063,1924]" captionTargetBox-1="[289,1391,403,1760]" captionTargetBox-2="[251,1405,438,1742]" captionTargetBox-3="[284,1396,760,1911]" captionTargetBox-4="[280,1360,1049,1913]" captionTargetId-0="figure@4.[227,1436,1051,1930]" captionTargetId-1="figure@5.[230,1439,386,1767]" captionTargetId-2="figure@6.[227,1436,408,1758]" captionTargetId-3="figure@8.[227,1436,735,1926]" captionTargetId-4="figure@9.[227,1436,1032,1926]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Fig. 1. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New" captionText-1="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." captionText-2="Fig. 3. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Right first pereiopod. B. Same as previous, chela and carpus. C. Same as previous, fingers. D. Same as previous, detail of denticulations. E. Second pereiopod, lateral. F. Same as previous, dactylus and distal propodus. G. Third pereiopod. H. Same as previous, dactylus and distal propodus. I. Fifth pereiopod, dactylus and distal propodus." captionText-3="Fig. 4. Hamodactylus paraqabai sp. nov., ♀, paratype (PoCL 1.6 mm, Madang, Papua New Guinea, MNHN-IU-2013-11090), mouthparts. A. Mandible. B. Same as previous, turned. C. Same as previous," captionText-4="Fig. 5. Hamodactylus paraqabai sp. nov., Ƌ (PoCL 1.1 mm, Lizard Is., Australia, MTQ-33218). A. Carapace and rostrum, lateral. B. First pleopod. C. Same as previous, endopod. D. Second pleopod. E. Same as previous, appendices interna and masculina." figureDoi-0="http://doi.org/10.5281/zenodo.3837559" figureDoi-1="http://doi.org/10.5281/zenodo.3837561" figureDoi-2="http://doi.org/10.5281/zenodo.3837563" figureDoi-3="http://doi.org/10.5281/zenodo.3837565" figureDoi-4="http://doi.org/10.5281/zenodo.3837567" httpUri-0="https://zenodo.org/record/3837559/files/figure.png" httpUri-1="https://zenodo.org/record/3837561/files/figure.png" httpUri-2="https://zenodo.org/record/3837563/files/figure.png" httpUri-3="https://zenodo.org/record/3837565/files/figure.png" httpUri-4="https://zenodo.org/record/3837567/files/figure.png" pageId="4" pageNumber="5">Figs 1–5</figureCitation>
, 
<figureCitation id="D5802A4D5B65FFE7FCDC4C7FFCFBFE6D" box="[848,875,376,402]" captionStart="Fig" captionStartId="16.[227,270,1862,1888]" captionTargetBox="[227,1436,711,1820]" captionTargetId="figure@16.[227,1436,711,1820]" captionTargetPageId="16" captionText="Fig. 10. Colour pattern of three relatives of the genus Hamodactylus. — A. H. aqabai Bruce &amp; Svoboda, 1983, ovigerous ♀ (UO Aq09-55) on alcyonacean coral Nephthea sp., Aqaba, Red Sea. — B–C. H. paraqabai sp. nov., ovigerous ♀♀, Madang, Papua New Guinea, on Nephthea sp. B. Holotype (MNHN-IU-2013-11092). C. Paratype (MNHN-IU-2013-11090). — D. H. pseudaqabai sp. nov., ovigerous ♀, holotype, N. Sulawesi, Indonesia (RMNH.CRUS.D.57195). Photos: Z. Ďuriš (A), T.-" figureDoi="http://doi.org/10.5281/zenodo.3837577" httpUri="https://zenodo.org/record/3837577/files/figure.png" pageId="4" pageNumber="5">10</figureCitation>
B–C
</paragraph>
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<subSubSection id="05A165435B65FFE7FF6F4CBEFAC8FDDA" pageId="4" pageNumber="5" type="etymology">
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<emphasis id="7FCFEADA5B65FFE7FF6F4CBEFEFAFE2C" bold="true" box="[227,362,441,467]" pageId="4" pageNumber="5">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="4D0436C85B65FFE7FF6F4CE0FAC8FDDA" blockId="4.[227,1436,487,549]" pageId="4" pageNumber="5">
The specific name 
<taxonomicName id="8ABB4D4B5B65FFE7FE444CE0FDD2FDFE" authorityName="Horka, Fransen &amp; Duris" authorityYear="2016" box="[456,578,487,513]" class="Malacostraca" family="Palaemonidae" genus="Hamodactylus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="paraqabai">
<emphasis id="7FCFEADA5B65FFE7FE444CE0FDD2FDFE" box="[456,578,487,513]" italics="true" pageId="4" pageNumber="5">paraqabai</emphasis>
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is a noun composed of the prefix 
<emphasis id="7FCFEADA5B65FFE7FC6B4CEFFB8EFDFE" box="[999,1054,488,513]" italics="true" pageId="4" pageNumber="5">para</emphasis>
- (from Greek: besides) and the adjective - 
<taxonomicName id="8ABB4D4B5B65FFE7FED24F0CFE3FFDDA" authorityName="Bruce &amp; Svoboda" authorityYear="1983" box="[350,431,523,549]" class="Malacostraca" family="Palaemonidae" genus="Hamodactylus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aqabai">
<emphasis id="7FCFEADA5B65FFE7FED24F0CFE3FFDDA" box="[350,431,523,549]" italics="true" pageId="4" pageNumber="5">aqabai</emphasis>
</taxonomicName>
, a species name used for 
<taxonomicName id="8ABB4D4B5B65FFE7FD504F0BFCC1FDDA" authorityName="Bruce &amp; Svoboda" authorityYear="1983" box="[732,849,523,549]" class="Malacostraca" family="Palaemonidae" genus="Hamodactylus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aqabai">
<emphasis id="7FCFEADA5B65FFE7FD504F0BFCC1FDDA" box="[732,849,523,549]" italics="true" pageId="4" pageNumber="5">H. aqabai</emphasis>
</taxonomicName>
, a closely similar species from the Red Sea.
</paragraph>
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<paragraph id="4D0436C85B65FFE7FF6F4F4BFE40FD99" blockId="4.[227,464,588,614]" box="[227,464,588,614]" pageId="4" pageNumber="5">
<heading id="164C81A45B65FFE7FF6F4F4BFE40FD99" bold="true" box="[227,464,588,614]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="7FCFEADA5B65FFE7FF6F4F4BFE40FD99" bold="true" box="[227,464,588,614]" pageId="4" pageNumber="5">Material examined</emphasis>
</heading>
</paragraph>
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<paragraph id="4D0436C85B65FFE7FE9C4F7CFE12FD6A" blockId="4.[227,1436,635,767]" box="[272,386,635,661]" pageId="4" pageNumber="5">
<emphasis id="7FCFEADA5B65FFE7FE9C4F7CFE12FD6A" bold="true" box="[272,386,635,661]" pageId="4" pageNumber="5">
<typeStatus id="9200886A5B65FFE7FE9C4F7CFE12FD6A" box="[272,386,635,661]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="4D0436C85B65FFE7FF6F4F99FC94FD01" blockId="4.[227,1436,635,767]" pageId="4" pageNumber="5">
<collectingCountry id="35AC76585B65FFE7FF6F4F99FD98FD47" box="[227,520,670,696]" name="Papua New Guinea" pageId="4" pageNumber="5">PAPUA NEW GUINEA</collectingCountry>
: ovigerous 
<specimenCount id="5BBDFD415B65FFE7FD104F99FD22FD47" box="[668,690,670,696]" pageId="4" pageNumber="5" type="female">♀</specimenCount>
(PoCL 
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), 
<collectingRegion id="8F7FF82A5B65FFE7FC1D4F99FC64FD47" box="[913,1012,670,696]" country="Papua New Guinea" name="Madang" pageId="4" pageNumber="5">Madang</collectingRegion>
, 
<collectorName id="E04E531E5B65FFE7FB884F99FB0BFD47" box="[1028,1179,670,696]" pageId="4" pageNumber="5">Krangket Is.</collectorName>
, stn PR3, 
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<geoCoordinate id="288F500F5B65FFE7FF6F4FC6FEFDFD23" box="[227,365,705,732]" degrees="145" direction="east" minutes="49.1" orientation="longitude" pageId="4" pageNumber="5" precision="92" value="145.81833">145°49.1' E</geoCoordinate>
, lagoon, depth 
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<elevation id="C696D1FB5B65FFE7FD924FC6FDCAFD23" box="[542,602,705,732]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="5" unit="m" value="13.0">13 m</elevation>
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, 
<collectedFrom id="FEA775355B65FFE7FDEA4FC5FD69FD24" box="[614,761,705,732]" pageId="4" pageNumber="5">
on 
<taxonomicName id="8ABB4D4B5B65FFE7FD054FC6FD69FD24" box="[649,761,705,731]" class="Anthozoa" family="Nephtheidae" genus="Nephthea" kingdom="Animalia" order="Alcyonacea" pageId="4" pageNumber="5" phylum="Cnidaria" rank="genus">
<emphasis id="7FCFEADA5B65FFE7FD054FC6FD69FD24" box="[649,761,705,731]" italics="true" pageId="4" pageNumber="5">Nephthea</emphasis>
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</collectedFrom>
sp., 
<date id="390510085B65FFE7FCA34FC6FC50FD24" box="[815,960,705,732]" pageId="4" pageNumber="5" value="2012-11-06">
<collectingDate id="2941E9E05B65FFE7FCA34FC6FC50FD24" box="[815,960,705,732]" pageId="4" pageNumber="5" value="2012-11-06">6 Nov. 2012</collectingDate>
</date>
, coll. 
<collectorName id="E04E531E5B65FFE7FB884FC5FBF8FD24" box="[1028,1128,703,732]" pageId="4" pageNumber="5">Z. Ďuriš</collectorName>
, fcn PNG12-17 (GenBank acc. nr. 
<accessionNumber id="52E8AB2B5B65FFE7FEB24FE3FE51FD00" box="[318,449,740,767]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KR088751" pageId="4" pageNumber="5">KR088751</accessionNumber>
) (MNHN-IU-2013-11092).
</paragraph>
</materialsCitation>
<paragraph id="4D0436C85B65FFE7FE9C4E21FE1DFCBF" blockId="4.[227,1437,806,973]" box="[272,397,806,832]" pageId="4" pageNumber="5">
<emphasis id="7FCFEADA5B65FFE7FE9C4E21FE1DFCBF" bold="true" box="[272,397,806,832]" pageId="4" pageNumber="5">
<typeStatus id="9200886A5B65FFE7FE9C4E21FE1DFCBF" box="[272,397,806,832]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="4D0436C85B65FFE7FF6F4E4EFD39FC32" blockId="4.[227,1437,806,973]" pageId="4" pageNumber="5">
<collectingCountry id="35AC76585B65FFE7FF6F4E4EFE69FC9C" box="[227,505,841,867]" name="Papua New Guinea" pageId="4" pageNumber="5">PAPUA NEW GUINEA</collectingCountry>
: 1 ovigerous 
<specimenCount id="5BBDFD415B65FFE7FD1D4E4EFD37FC9C" box="[657,679,841,867]" pageId="4" pageNumber="5" type="female">♀</specimenCount>
(PoCL 
<quantity id="8A439B2D5B65FFE7FD714E4EFCC8FC9C" box="[765,856,841,867]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="5" unit="mm" value="1.9">1.9 mm</quantity>
), same data as 
<typeStatus id="9200886A5B65FFE7FC704E4EFBF3FC9C" box="[1020,1123,841,867]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
(MNHN-IU-2013-10551); 
<specimenCount id="5BBDFD415B65FFE7FF6F4E6BFE83FC78" box="[227,275,876,903]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
(PoCL 
<quantity id="8A439B2D5B65FFE7FEFA4E6BFE47FC78" box="[374,471,876,903]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="5" unit="mm" value="1.6">1.6 mm</quantity>
, mouthparts dissected), 
<collectingRegion id="8F7FF82A5B65FFE7FD714E6BFCCDFC79" box="[765,861,876,902]" country="Papua New Guinea" name="Madang" pageId="4" pageNumber="5">Madang</collectingRegion>
Resort, 
<geoCoordinate id="288F500F5B65FFE7FC4B4E6BFBD4FC79" box="[967,1092,876,903]" degrees="05" direction="south" minutes="12.5" orientation="latitude" pageId="4" pageNumber="5" precision="92" value="-5.2083335">05°12.5' S</geoCoordinate>
<geoCoordinate id="288F500F5B65FFE7FBC34E6BFB4FFC78" box="[1103,1247,876,903]" degrees="145" direction="east" minutes="48.5" orientation="longitude" pageId="4" pageNumber="5" precision="92" value="145.80833">145°48.5' E</geoCoordinate>
, stn PR227, on 
<taxonomicName id="8ABB4D4B5B65FFE7FF6F4E88FEC3FC56" box="[227,339,911,937]" class="Anthozoa" family="Nephtheidae" genus="Nephthea" kingdom="Animalia" order="Alcyonacea" pageId="4" pageNumber="5" phylum="Cnidaria" rank="genus">
<emphasis id="7FCFEADA5B65FFE7FF6F4E88FEC3FC56" box="[227,339,911,937]" italics="true" pageId="4" pageNumber="5">Nephthea</emphasis>
</taxonomicName>
sp., 
<date id="390510085B65FFE7FE1E4E88FDAAFC56" box="[402,570,911,938]" pageId="4" pageNumber="5" value="2012-12-10">10 Dec. 2012</date>
, coll. A.F. Berberian, leg. Z. Ďuriš, fcn 
<collectingCountry id="35AC76585B65FFE7FBA44E88FBF7FC56" box="[1064,1127,911,937]" name="Papua New Guinea" pageId="4" pageNumber="5">PNG</collectingCountry>
12-638 (GenBank acc. nr. 
<accessionNumber id="52E8AB2B5B65FFE7FF6F4EB5FEF6FC32" box="[227,358,946,973]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KR088752" pageId="4" pageNumber="5">KR088752</accessionNumber>
) (MNHN-IU-2013-11090).
</paragraph>
<caption id="19C466405B65FFE7FF6F4AD6FA0CF813" ID-DOI="http://doi.org/10.5281/zenodo.3837559" ID-Zenodo-Dep="3837559" box="[227,1436,1997,2028]" httpUri="https://zenodo.org/record/3837559/files/figure.png" pageId="4" pageNumber="5" startId="4.[227,270,2001,2027]" targetBox="[254,1423,1063,1924]" targetPageId="4">
<paragraph id="4D0436C85B65FFE7FF6F4AD6FA0CF813" blockId="4.[227,1436,1997,2028]" box="[227,1436,1997,2028]" pageId="4" pageNumber="5">
<emphasis id="7FCFEADA5B65FFE7FF6F4AD6FEA1F814" bold="true" box="[227,305,2001,2027]" pageId="4" pageNumber="5">Fig. 1.</emphasis>
<taxonomicName id="8ABB4D4B5B65FFE7FEB54AD6FDF4F814" authorityName="Horka, Fransen &amp; Duris" authorityYear="2016" box="[313,612,2001,2027]" class="Malacostraca" family="Palaemonidae" genus="Hamodactylus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="paraqabai" status="sp. nov.">
<emphasis id="7FCFEADA5B65FFE7FEB54AD6FDF4F814" box="[313,612,2001,2027]" italics="true" pageId="4" pageNumber="5">Hamodactylus paraqabai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64FC57A15B65FFE7FDE04AD5FD5AF813" box="[620,714,2002,2028]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New
</paragraph>
</caption>
<paragraph id="4D0436C85B65FFE4FF6F4AF3FD97FEB5" blockId="4.[227,312,2036,2062]" lastBlockId="7.[227,1436,304,507]" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
<collectingCountry id="35AC76585B65FFE7FF6F4AF3FEA8F7F1" box="[227,312,2036,2062]" name="Guinea-Bissau" pageId="4" pageNumber="5">Guinea</collectingCountry>
), habitus. Scale bar = 
<quantity id="8A439B2D5B65FFE7FDB34AF2FD16F7F0" box="[575,646,2037,2063]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="mm" value="1.0">1 mm</quantity>
. 
<emphasis id="7FCFEADA5B66FFE4FE9C4C37FD97FEB5" bold="true" box="[272,519,304,330]" pageId="7" pageNumber="8">Additional material</emphasis>
</paragraph>
<caption id="19C466405B64FFE6FF6F4A13FC80F844" ID-DOI="http://doi.org/10.5281/zenodo.3837561" ID-Zenodo-Dep="3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="5" pageNumber="6" startId="5.[227,270,1812,1838]" targetBox="[289,1391,403,1760]" targetPageId="5">
<paragraph id="4D0436C85B64FFE6FF6F4A13FC80F844" blockId="5.[227,1438,1808,1979]" pageId="5" pageNumber="6">
<emphasis id="7FCFEADA5B64FFE6FF6F4A13FEA1F8D1" bold="true" box="[227,305,1812,1838]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
<taxonomicName id="8ABB4D4B5B64FFE6FEB54A13FDF5F8D1" authorityName="Horka, Fransen &amp; Duris" authorityYear="2016" box="[313,613,1812,1838]" class="Malacostraca" family="Palaemonidae" genus="Hamodactylus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="paraqabai" status="sp. nov.">
<emphasis id="7FCFEADA5B64FFE6FEB54A13FDF5F8D1" box="[313,613,1812,1838]" italics="true" pageId="5" pageNumber="6">Hamodactylus paraqabai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64FC57A15B64FFE6FDE14A12FD5BF8D0" box="[621,715,1813,1839]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). 
<emphasis id="7FCFEADA5B64FFE6FEDD4A30FEF6F8AE" bold="true" box="[337,358,1847,1873]" pageId="5" pageNumber="6">A</emphasis>
. Carapace and rostrum, lateral. 
<emphasis id="7FCFEADA5B64FFE6FD534A3FFD63F8AD" bold="true" box="[735,755,1848,1874]" pageId="5" pageNumber="6">B</emphasis>
. Anterior cephalothorax and appendages, dorsal. 
<emphasis id="7FCFEADA5B64FFE6FAB74A30FAC0F8AE" bold="true" box="[1339,1360,1847,1873]" pageId="5" pageNumber="6">C</emphasis>
. Sixth abdominal segment, telson and uropod, dorsal. 
<emphasis id="7FCFEADA5B64FFE6FC844A5CFC8DF88A" bold="true" box="[776,797,1883,1909]" pageId="5" pageNumber="6">D</emphasis>
. Posterior part of telson, dorsal. 
<emphasis id="7FCFEADA5B64FFE6FB1A4A5CFB3AF88A" bold="true" box="[1174,1194,1883,1909]" pageId="5" pageNumber="6">E</emphasis>
. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. 
<emphasis id="7FCFEADA5B64FFE6FB224A79FB50F867" bold="true" box="[1198,1216,1918,1944]" pageId="5" pageNumber="6">F</emphasis>
. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="19C466405B67FFE5FF6F4A15FD2DF846" ID-DOI="http://doi.org/10.5281/zenodo.3837563" ID-Zenodo-Dep="3837563" httpUri="https://zenodo.org/record/3837563/files/figure.png" pageId="6" pageNumber="7" startId="6.[227,270,1810,1836]" targetBox="[251,1405,438,1742]" targetPageId="6">
<paragraph id="4D0436C85B67FFE5FF6F4A15FD2DF846" blockId="6.[227,1437,1806,1977]" pageId="6" pageNumber="7">
<emphasis id="7FCFEADA5B67FFE5FF6F4A15FEA1F8D3" bold="true" box="[227,305,1810,1836]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
<taxonomicName id="8ABB4D4B5B67FFE5FEB54A15FDF4F8D3" authorityName="Horka, Fransen &amp; Duris" authorityYear="2016" box="[313,612,1810,1836]" class="Malacostraca" family="Palaemonidae" genus="Hamodactylus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="paraqabai" status="sp. nov.">
<emphasis id="7FCFEADA5B67FFE5FEB54A15FDF4F8D3" box="[313,612,1810,1836]" italics="true" pageId="6" pageNumber="7">Hamodactylus paraqabai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64FC57A15B67FFE5FDE04A14FD5AF8D2" box="[620,714,1811,1837]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). 
<emphasis id="7FCFEADA5B67FFE5FEDC4A32FEF5F8B0" bold="true" box="[336,357,1845,1871]" pageId="6" pageNumber="7">A</emphasis>
. Right first pereiopod. 
<emphasis id="7FCFEADA5B67FFE5FDF94A31FD19F8AF" bold="true" box="[629,649,1846,1872]" pageId="6" pageNumber="7">B</emphasis>
. Same as previous, chela and carpus. 
<emphasis id="7FCFEADA5B67FFE5FBC94A32FBCAF8B0" bold="true" box="[1093,1114,1845,1871]" pageId="6" pageNumber="7">C</emphasis>
. Same as previous, fingers. 
<emphasis id="7FCFEADA5B67FFE5FF6F4A5EFE90F88C" bold="true" box="[227,256,1881,1907]" pageId="6" pageNumber="7">D.</emphasis>
Same as previous, detail of denticulations. 
<emphasis id="7FCFEADA5B67FFE5FD7D4A5EFC95F88C" bold="true" box="[753,773,1881,1907]" pageId="6" pageNumber="7">E</emphasis>
. Second pereiopod, lateral. 
<emphasis id="7FCFEADA5B67FFE5FBC94A5EFBC7F88C" bold="true" box="[1093,1111,1881,1907]" pageId="6" pageNumber="7">F</emphasis>
. Same as previous, dactylus and distal propodus. 
<emphasis id="7FCFEADA5B67FFE5FE524A7BFE65F869" bold="true" box="[478,501,1916,1942]" pageId="6" pageNumber="7">G</emphasis>
. Third pereiopod. 
<emphasis id="7FCFEADA5B67FFE5FD5A4A7BFD7DF869" bold="true" box="[726,749,1916,1942]" pageId="6" pageNumber="7">H</emphasis>
. Same as previous, dactylus and distal propodus. 
<emphasis id="7FCFEADA5B67FFE5FACB4A7BFAC2F869" bold="true" box="[1351,1362,1916,1942]" pageId="6" pageNumber="7">I</emphasis>
. Fifth pereiopod, dactylus and distal propodus.
</paragraph>
</caption>
<paragraph id="4D0436C85B66FFE4FF6F4C53FC73FE04" blockId="7.[227,1436,304,507]" pageId="7" pageNumber="8">
<collectingCountry id="35AC76585B66FFE4FF6F4C53FE19FE91" box="[227,393,340,366]" name="Australia" pageId="7" pageNumber="8">AUSTRALIA</collectingCountry>
, Great Barrier Reef, Lizard Island, from soft branchy corals (
<taxonomicName id="8ABB4D4B5B66FFE4FBD94C53FB76FE90" box="[1109,1254,340,367]" class="Anthozoa" kingdom="Animalia" order="Alcyonacea" pageId="7" pageNumber="8" phylum="Cnidaria" rank="order">Alcyonacea</taxonomicName>
), coll. &amp; leg. Z. Ďuriš: 
<specimenCount id="5BBDFD415B66FFE4FEB84C70FEF0FE6D" box="[308,352,375,402]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
(PoCL 
<quantity id="8A439B2D5B66FFE4FE304C70FD8AFE6D" box="[444,538,375,402]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="8" unit="mm" value="1.3">1.3 mm</quantity>
), stn LI10-47, 
<geoCoordinate id="288F500F5B66FFE4FD4A4C70FCAFFE6E" box="[710,831,375,401]" degrees="14" direction="south" minutes="41.8" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="-14.696667">14°41.8' S</geoCoordinate>
<geoCoordinate id="288F500F5B66FFE4FCCB4C70FC43FE6D" box="[839,979,375,402]" degrees="145" direction="east" minutes="27.0" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="145.45">145°27.0' E</geoCoordinate>
, North Point, South of Mermaid Cove, depth 
<quantity id="8A439B2D5B66FFE4FEA04C9DFECBFE4A" box="[300,347,410,437]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="m" value="9.0">9 m</quantity>
, on 
<taxonomicName id="8ABB4D4B5B66FFE4FE004C9DFE6CFE4B" box="[396,508,410,436]" class="Anthozoa" family="Nephtheidae" genus="Nephthea" kingdom="Animalia" order="Alcyonacea" pageId="7" pageNumber="8" phylum="Cnidaria" rank="genus">
<emphasis id="7FCFEADA5B66FFE4FE004C9DFE6CFE4B" box="[396,508,410,436]" italics="true" pageId="7" pageNumber="8">Nephthea</emphasis>
</taxonomicName>
sp., 
<date id="390510085B66FFE4FDB94C9DFD54FE4B" box="[565,708,410,436]" pageId="7" pageNumber="8" value="2010-09-01">1 Sep. 2010</date>
, fcn Au10-78B (GenBank acc. nr. 
<accessionNumber id="52E8AB2B5B66FFE4FBD64C9DFB4DFE4A" box="[1114,1245,410,437]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KR088753" pageId="7" pageNumber="8">KR088753</accessionNumber>
) (MTQ-33208); 
<specimenCount id="5BBDFD415B66FFE4FF6F4CB9FE9FFE27" box="[227,271,446,472]" pageId="7" pageNumber="8" type="male">1 ♂</specimenCount>
(PoCL 
<quantity id="8A439B2D5B66FFE4FEE74CB9FE59FE27" box="[363,457,446,472]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="8" unit="mm" value="1.1">1.1 mm</quantity>
), stn LI10-59, 
<geoCoordinate id="288F500F5B66FFE4FDF94CB9FD7EFE27" box="[629,750,446,472]" degrees="14" direction="south" minutes="42.0" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="-14.7">14°42.0' S</geoCoordinate>
<geoCoordinate id="288F500F5B66FFE4FD7A4CBAFC11FE27" box="[758,897,445,472]" degrees="145" direction="east" minutes="26.9" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="145.44833">145°26.9' E</geoCoordinate>
, lagoon, depth 
<quantity id="8A439B2D5B66FFE4FBB94CBAFBEAFE27" box="[1077,1146,445,472]" metricMagnitude="0" metricUnit="m" metricValue="8.6" pageId="7" pageNumber="8" unit="m" value="8.6">8.6 m</quantity>
, on 
<taxonomicName id="8ABB4D4B5B66FFE4FB274CBAFA8BFE28" box="[1195,1307,445,471]" class="Anthozoa" family="Nephtheidae" genus="Nephthea" kingdom="Animalia" order="Alcyonacea" pageId="7" pageNumber="8" phylum="Cnidaria" rank="genus">
<emphasis id="7FCFEADA5B66FFE4FB274CBAFA8BFE28" box="[1195,1307,445,471]" italics="true" pageId="7" pageNumber="8">Nephthea</emphasis>
</taxonomicName>
sp., 
<date id="390510085B66FFE4FADF4CB9FEB1FE04" pageId="7" pageNumber="8" value="2010-09-02">2 Sep. 2010</date>
, fcn Au10-85 (GenBank acc. nr. 
<accessionNumber id="52E8AB2B5B66FFE4FD2E4CE7FCB5FE04" box="[674,805,480,507]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KR088754" pageId="7" pageNumber="8">KR088754</accessionNumber>
) (MTQ-33218).
</paragraph>
</subSubSection>
<subSubSection id="05A165435B66FFE8FF6F4F25FDEEFE91" lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8" type="description">
<paragraph id="4D0436C85B66FFE4FF6F4F25FD97FDC3" blockId="7.[227,519,546,572]" box="[227,519,546,572]" pageId="7" pageNumber="8">
<heading id="164C81A45B66FFE4FF6F4F25FD97FDC3" bold="true" box="[227,519,546,572]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="7FCFEADA5B66FFE4FF6F4F25FD97FDC3" bold="true" box="[227,519,546,572]" pageId="7" pageNumber="8">
Description of 
<typeStatus id="9200886A5B66FFE4FE174F25FD97FDC3" box="[411,519,546,572]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="4D0436C85B66FFE4FF6F4F56FB7CFD94" blockId="7.[227,1260,593,619]" box="[227,1260,593,619]" pageId="7" pageNumber="8">
Small sized shrimp, ovigerous female; specimen complete, with all appendages (
<figureCitation id="D5802A4D5B66FFE4FB1A4F56FB4BFD94" box="[1174,1243,593,619]" captionStart="Fig" captionStartId="4.[227,270,2001,2027]" captionTargetBox="[254,1423,1063,1924]" captionTargetId="figure@4.[227,1436,1051,1930]" captionTargetPageId="4" captionText="Fig. 1. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New" figureDoi="http://doi.org/10.5281/zenodo.3837559" httpUri="https://zenodo.org/record/3837559/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="4D0436C85B66FFE4FF6F4F95FCD5FC5D" blockId="7.[227,1437,658,930]" pageId="7" pageNumber="8">
Carapace smooth. Rostrum (
<figureCitation id="D5802A4D5B66FFE4FDB64F95FD0AFD53" box="[570,666,658,684]" captionStart="Fig" captionStartId="5.[227,270,1812,1838]" captionTargetBox="[289,1391,403,1760]" captionTargetId="figure@5.[230,1439,386,1767]" captionTargetPageId="5" captionText="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="7" pageNumber="8">Fig. 2A</figureCitation>
) well developed, exceeding intermediate segment of antennular peduncle (
<figureCitation id="D5802A4D5B66FFE4FED14FB2FE29FD30" box="[349,441,693,719]" captionStart="Fig" captionStartId="5.[227,270,1812,1838]" captionTargetBox="[289,1391,403,1760]" captionTargetId="figure@5.[230,1439,386,1767]" captionTargetPageId="5" captionText="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="7" pageNumber="8">Fig. 2B</figureCitation>
); lamina deep, lateral carina situated slightly above straight, setose, toothless ventral margin; dorsal margin convex, elevated, strongly compressed, with six subequal teeth, of which first situated above level of posterior orbital margin. Supra-orbital and epigastric spines absent. Orbit obsolescent, inferior orbital angle acutely produced in lateral view. Antennal spine small and slender, marginal, situated closely below inferior orbital angle. Hepatic spine larger and more robust than antennal, situated just behind level of posterior orbital margin and well below level of antennal spine. Antero-lateral angle moderately produced, rounded.
</paragraph>
<paragraph id="4D0436C85B66FFE4FF6F4ECDFB6EFBD4" blockId="7.[227,1437,970,1067]" pageId="7" pageNumber="8">
Abdominal segments smooth (
<figureCitation id="D5802A4D5B66FFE4FDC74ECDFD01FC1B" box="[587,657,970,996]" captionStart="Fig" captionStartId="4.[227,270,2001,2027]" captionTargetBox="[254,1423,1063,1924]" captionTargetId="figure@4.[227,1436,1051,1930]" captionTargetPageId="4" captionText="Fig. 1. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New" figureDoi="http://doi.org/10.5281/zenodo.3837559" httpUri="https://zenodo.org/record/3837559/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
). First segment pleuron highly produced anteriorly, third segment not produced posterodorsally, sixth segment twice as long as deep. Pleura all broadly rounded. Posteroventral angle of sixth segment feebly produced, postero-lateral angle feebly acute.
</paragraph>
<paragraph id="4D0436C85B66FFE4FF6F4955FD5DFAE3" blockId="7.[227,1437,1106,1308]" pageId="7" pageNumber="8">
Telson 0.8 of length of sixth abdominal segment and 3.2 times as long as maximum width (
<figureCitation id="D5802A4D5B66FFE4FABC4955FA1FFB93" box="[1328,1423,1106,1132]" captionStart="Fig" captionStartId="5.[227,270,1812,1838]" captionTargetBox="[289,1391,403,1760]" captionTargetId="figure@5.[230,1439,386,1767]" captionTargetPageId="5" captionText="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="7" pageNumber="8">Fig. 2C</figureCitation>
). Lateral margins convex in anterior half, posteriorly converging. Two pairs of very small marginal dorsal spines, about 0.05 of telson length, present at 0.55 and 0.8 of telson length. Posterior margin (
<figureCitation id="D5802A4D5B66FFE4FABB499FFA03FB4D" box="[1335,1427,1176,1202]" captionStart="Fig" captionStartId="5.[227,270,1812,1838]" captionTargetBox="[289,1391,403,1760]" captionTargetId="figure@5.[230,1439,386,1767]" captionTargetPageId="5" captionText="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="7" pageNumber="8">Fig. 2D</figureCitation>
) rounded, about half of anterior width, with three pairs of spines; lateral spines short but distinctly twice as long as dorsal spines. Intermediate spines largest, about 0.2 of telson length and 1.5 times length of more slender, setulose, submedian spines.
</paragraph>
<paragraph id="4D0436C85B66FFE4FF6F4843FC97FA7F" blockId="7.[227,1436,1347,1409]" pageId="7" pageNumber="8">
Eyes (
<figureCitation id="D5802A4D5B66FFE4FEA04844FE16FAA2" box="[300,390,1347,1373]" captionStart="Fig" captionStartId="5.[227,270,1812,1838]" captionTargetBox="[289,1391,403,1760]" captionTargetId="figure@5.[230,1439,386,1767]" captionTargetPageId="5" captionText="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="7" pageNumber="8">Fig. 2B</figureCitation>
) well developed. Cornea ellipsoid, obliquely set on stalks; stalk swollen proximally, with distinct accessory pigment spot anterodorsally.
</paragraph>
<paragraph id="4D0436C85B66FFE4FF6F48AFFABEF969" blockId="7.[227,1436,1448,1686]" pageId="7" pageNumber="8">
Antennular peduncle (
<figureCitation id="D5802A4D5B66FFE4FE6948AFFDADFA3D" box="[485,573,1448,1474]" captionStart="Fig" captionStartId="5.[227,270,1812,1838]" captionTargetBox="[289,1391,403,1760]" captionTargetId="figure@5.[230,1439,386,1767]" captionTargetPageId="5" captionText="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="7" pageNumber="8">Fig. 2B</figureCitation>
) normal, exceeding tip of rostrum by distal segment. Basal segment broad, 2 times as long as wide, with stylocerite slender, acute, reaching almost to middle of segment; lateral border convex, anterolateral margin slightly acutely produced mesially, with 2 acute distolateral teeth (
<figureCitation id="D5802A4D5B66FFE4FF674B16FED8F9D3" box="[235,328,1553,1580]" captionStart="Fig" captionStartId="5.[227,270,1812,1838]" captionTargetBox="[289,1391,403,1760]" captionTargetId="figure@5.[230,1439,386,1767]" captionTargetPageId="5" captionText="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="7" pageNumber="8">Fig. 2E</figureCitation>
); medioventral margin with tooth in front of statocyst level. Intermediate and distal segments short, together equal to 0.3 of proximal segment length. Upper flagellum biramous, with first 7 segments fused, shorter free ramus 3-segmented and longer ramus consisting of about 10 slender segments. About 6 groups of aesthetascs present. Lower flagellum consisting of about 11–13 slender segments.
</paragraph>
<paragraph id="4D0436C85B66FFE4FF6F4BBAFB48F8BF" blockId="7.[227,1437,1724,1857]" pageId="7" pageNumber="8">
Antenna (
<figureCitation id="D5802A4D5B66FFE4FED74BBBFE2AF928" box="[347,442,1724,1751]" captionStart="Fig" captionStartId="5.[227,270,1812,1838]" captionTargetBox="[289,1391,403,1760]" captionTargetId="figure@5.[230,1439,386,1767]" captionTargetPageId="5" captionText="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="7" pageNumber="8">Fig. 2B</figureCitation>
) with robust basicerite without lateral tooth. Carpocerite slender, short, reaching almost to middle of scaphocerite. Scaphocerite large, extending well beyond antennular peduncle, about 2.5 times as long as broad, with greatest width in midlength, distomedially angulated; lateral border almost straight, with acute distal tooth; lamina extending far beyond distolateral tooth.
</paragraph>
<paragraph id="4D0436C85B66FFE4FF6F4A60FDFAF87E" blockId="7.[227,618,1895,1921]" box="[227,618,1895,1921]" pageId="7" pageNumber="8">Fourth thoracic sternite unarmed.</paragraph>
<paragraph id="4D0436C85B66FFEBFF6F4AAEFB71FE4A" blockId="7.[227,1436,1961,2058]" lastBlockId="8.[227,1436,305,437]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
First pereiopod (
<figureCitation id="D5802A4D5B66FFE4FE244AAEFE7DF83C" box="[424,493,1961,1987]" captionStart="Fig" captionStartId="6.[227,270,1810,1836]" captionTargetBox="[251,1405,438,1742]" captionTargetId="figure@6.[227,1436,408,1758]" captionTargetPageId="6" captionText="Fig. 3. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Right first pereiopod. B. Same as previous, chela and carpus. C. Same as previous, fingers. D. Same as previous, detail of denticulations. E. Second pereiopod, lateral. F. Same as previous, dactylus and distal propodus. G. Third pereiopod. H. Same as previous, dactylus and distal propodus. I. Fifth pereiopod, dactylus and distal propodus." figureDoi="http://doi.org/10.5281/zenodo.3837563" httpUri="https://zenodo.org/record/3837563/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
A–D) slender, reaching to end of scaphocerite. Chela with palm subcylindrical, slightly bowed and compressed, about 4 times as long as wide. Fingers about 0.75 of palm length; cutting edges gaping distally, bearing series of 5 small blunt teeth at distal third on dactylus, and 3 such teeth on fixed finger; very thin denticulate lamella present laterally along latter series of teeth on both fingers, tip of fingers slender, hooked. Cleaning setae present proximally on palm and on disto-ventral end of carpus, latter being subequal to chela length and 6.5 times as long as wide distally. Merus about 1.3 times as long as carpus and almost twice length of ischium. Basis and coxa normal.
</paragraph>
<paragraph id="4D0436C85B69FFEBFF6F4CDBFA0DFD7C" blockId="8.[227,1437,476,644]" pageId="8" pageNumber="9">
Second pereiopods (
<figureCitation id="D5802A4D5B69FFEBFE5E4CDBFD86FE09" box="[466,534,476,502]" captionStart="Fig" captionStartId="6.[227,270,1810,1836]" captionTargetBox="[251,1405,438,1742]" captionTargetId="figure@6.[227,1436,408,1758]" captionTargetPageId="6" captionText="Fig. 3. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Right first pereiopod. B. Same as previous, chela and carpus. C. Same as previous, fingers. D. Same as previous, detail of denticulations. E. Second pereiopod, lateral. F. Same as previous, dactylus and distal propodus. G. Third pereiopod. H. Same as previous, dactylus and distal propodus. I. Fifth pereiopod, dactylus and distal propodus." figureDoi="http://doi.org/10.5281/zenodo.3837563" httpUri="https://zenodo.org/record/3837563/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
E–F) greatly reduced, small, reaching almost to end of carpocerite, subequal and similar, resembling ambulatory pereiopod. Chela without any trace of fixed finger; dactylus resembles that of walking legs. Unguis distinct, occupying distal 0.25 of dactylus length, corpus slender, 4.2 times as long as basal width, slightly curved, without accessory spines or teeth, long setae arising from distal and subdistal part of corpus. Propodus subcylindrical, unarmed, about 6.5 times as long as wide and 2
</paragraph>
<caption id="19C466405B69FFEBFF6F4AA8FA0CF813" ID-DOI="http://doi.org/10.5281/zenodo.3837565" ID-Zenodo-Dep="3837565" httpUri="https://zenodo.org/record/3837565/files/figure.png" pageId="8" pageNumber="9" startId="8.[227,270,1967,1993]" targetBox="[284,1396,760,1911]" targetPageId="8">
<paragraph id="4D0436C85B69FFEBFF6F4AA8FA0CF813" blockId="8.[227,1436,1963,2029]" pageId="8" pageNumber="9">
<emphasis id="7FCFEADA5B69FFEBFF6F4AA8FEA3F836" bold="true" box="[227,307,1967,1993]" pageId="8" pageNumber="9">Fig. 4.</emphasis>
<taxonomicName id="8ABB4D4B5B69FFEBFEB14AA8FDFAF836" authorityName="Horka, Fransen &amp; Duris" authorityYear="2016" box="[317,618,1967,1993]" class="Malacostraca" family="Palaemonidae" genus="Hamodactylus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="paraqabai" status="sp. nov.">
<emphasis id="7FCFEADA5B69FFEBFEB14AA8FDFAF836" box="[317,618,1967,1993]" italics="true" pageId="8" pageNumber="9">Hamodactylus paraqabai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64FC57A15B69FFEBFDF84AB7FD44F835" box="[628,724,1968,1994]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, ♀, paratype (PoCL 1.6 mm, Madang, Papua New Guinea, MNHN-IU-2013-11090), mouthparts. 
<emphasis id="7FCFEADA5B69FFEBFD2B4AD5FD2CF813" bold="true" box="[679,700,2002,2028]" pageId="8" pageNumber="9">A</emphasis>
. Mandible. 
<emphasis id="7FCFEADA5B69FFEBFCC64AD4FCCEF812" bold="true" box="[842,862,2003,2029]" pageId="8" pageNumber="9">B</emphasis>
. Same as previous, turned. 
<emphasis id="7FCFEADA5B69FFEBFB2F4AD5FB28F813" bold="true" box="[1187,1208,2002,2028]" pageId="8" pageNumber="9">C</emphasis>
. Same as previous,
</paragraph>
</caption>
<paragraph id="4D0436C85B69FFEBFF6F4AF1FA0DF7EF" blockId="8.[227,295,2038,2064]" box="[227,1437,2038,2064]" lastBlockId="8.[1303,1437,2038,2064]" pageId="8" pageNumber="9">molar maxilliped.</paragraph>
<paragraph id="4D0436C85B69FFEAFEA14AF1FC22FE90" blockId="8.[301,1297,2037,2064]" lastBlockId="9.[227,1436,306,368]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
process. 
<emphasis id="7FCFEADA5B69FFEBFE1E4AF1FE37F7EF" bold="true" box="[402,423,2038,2064]" pageId="8" pageNumber="9">D</emphasis>
. Maxillula. 
<emphasis id="7FCFEADA5B69FFEBFDB84AF1FDD8F7EF" bold="true" box="[564,584,2038,2064]" pageId="8" pageNumber="9">E</emphasis>
. Maxilla. 
<emphasis id="7FCFEADA5B69FFEBFD314AF1FD5FF7EF" bold="true" box="[701,719,2038,2064]" pageId="8" pageNumber="9">F</emphasis>
. First maxilliped. 
<emphasis id="7FCFEADA5B69FFEBFC2E4AF2FC29F7F0" bold="true" box="[930,953,2037,2063]" pageId="8" pageNumber="9">G</emphasis>
. Second maxilliped. 
<emphasis id="7FCFEADA5B69FFEBFB214AF1FB54F7EF" bold="true" box="[1197,1220,2038,2064]" pageId="8" pageNumber="9">H</emphasis>
. Third times length of dactylus; dactylus unarmed. Carpus, merus and ischium unarmed, lengths in ratio of 1.2, 1.0 and 1.6 times length of propodus. Basis and coxa normal.
</paragraph>
<paragraph id="4D0436C85B68FFEAFF6F4C91FA16FD9F" blockId="9.[227,1437,406,609]" pageId="9" pageNumber="10">
Ambulatory pereiopods slender, distinctly longer and more robust than second pereiopods. Third pereiopod (
<figureCitation id="D5802A4D5B68FFEAFEE04CBDFE5BFE2B" box="[364,459,442,468]" captionStart="Fig" captionStartId="6.[227,270,1810,1836]" captionTargetBox="[251,1405,438,1742]" captionTargetId="figure@6.[227,1436,408,1758]" captionTargetPageId="6" captionText="Fig. 3. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Right first pereiopod. B. Same as previous, chela and carpus. C. Same as previous, fingers. D. Same as previous, detail of denticulations. E. Second pereiopod, lateral. F. Same as previous, dactylus and distal propodus. G. Third pereiopod. H. Same as previous, dactylus and distal propodus. I. Fifth pereiopod, dactylus and distal propodus." figureDoi="http://doi.org/10.5281/zenodo.3837563" httpUri="https://zenodo.org/record/3837563/files/figure.png" pageId="9" pageNumber="10">Fig. 3G</figureCitation>
) exceeds tip of scaphocerite by dactylus. Dactylus (
<figureCitation id="D5802A4D5B68FFEAFBC14CBDFB3CFE2B" box="[1101,1196,442,468]" captionStart="Fig" captionStartId="6.[227,270,1810,1836]" captionTargetBox="[251,1405,438,1742]" captionTargetId="figure@6.[227,1436,408,1758]" captionTargetPageId="6" captionText="Fig. 3. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Right first pereiopod. B. Same as previous, chela and carpus. C. Same as previous, fingers. D. Same as previous, detail of denticulations. E. Second pereiopod, lateral. F. Same as previous, dactylus and distal propodus. G. Third pereiopod. H. Same as previous, dactylus and distal propodus. I. Fifth pereiopod, dactylus and distal propodus." figureDoi="http://doi.org/10.5281/zenodo.3837563" httpUri="https://zenodo.org/record/3837563/files/figure.png" pageId="9" pageNumber="10">Fig. 3H</figureCitation>
) slender, uniformly tapering, unarmed and with a distinct unguis on distal 0.45 of length, total length about 5 times width near base. Propodus about 12 times as long as wide, 3 times length of dactyl, and devoid of spines. Carpus, merus and ischium 0.5, 0.8 and 0.5 of propodus length, unarmed. Fourth and fifth pereiopods similar, dactylus of fifth pereiopod (
<figureCitation id="D5802A4D5B68FFEAFD044F41FD48FD9E" box="[648,728,582,609]" captionStart="Fig" captionStartId="6.[227,270,1810,1836]" captionTargetBox="[251,1405,438,1742]" captionTargetId="figure@6.[227,1436,408,1758]" captionTargetPageId="6" captionText="Fig. 3. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Right first pereiopod. B. Same as previous, chela and carpus. C. Same as previous, fingers. D. Same as previous, detail of denticulations. E. Second pereiopod, lateral. F. Same as previous, dactylus and distal propodus. G. Third pereiopod. H. Same as previous, dactylus and distal propodus. I. Fifth pereiopod, dactylus and distal propodus." figureDoi="http://doi.org/10.5281/zenodo.3837563" httpUri="https://zenodo.org/record/3837563/files/figure.png" pageId="9" pageNumber="10">Fig. 3I</figureCitation>
) slightly more slender and straighter than that of third leg.
</paragraph>
<paragraph id="4D0436C85B68FFEAFF6F4F8FFB4DFD17" blockId="9.[227,1436,648,745]" pageId="9" pageNumber="10">
Uropods (
<figureCitation id="D5802A4D5B68FFEAFED44F8FFE22FD5D" box="[344,434,648,674]" captionStart="Fig" captionStartId="5.[227,270,1812,1838]" captionTargetBox="[289,1391,403,1760]" captionTargetId="figure@5.[230,1439,386,1767]" captionTargetPageId="5" captionText="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="9" pageNumber="10">Fig. 2C</figureCitation>
) slender, extending well beyond tip of telson. Protopodite unarmed laterally. Exopod about 3 times as long as wide, extending well beyond endopod, with lateral border almost straight, entire, terminating in small mobile spine (
<figureCitation id="D5802A4D5B68FFEAFD434FC9FCB8FD17" box="[719,808,718,744]" captionStart="Fig" captionStartId="5.[227,270,1812,1838]" captionTargetBox="[289,1391,403,1760]" captionTargetId="figure@5.[230,1439,386,1767]" captionTargetPageId="5" captionText="Fig. 2. Hamodactylus paraqabai sp. nov., ovigerous ♀, holotype (PoCL 1.5 mm; Madang, Papua New Guinea). A. Carapace and rostrum, lateral. B. Anterior cephalothorax and appendages, dorsal. C. Sixth abdominal segment, telson and uropod, dorsal. D. Posterior part of telson, dorsal. E. Antennule, detail of distolateral angle of proximal segment and lateral border of intermediate segment. F. Distolateral angle of uropodal exopod, dorsal. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837561" httpUri="https://zenodo.org/record/3837561/files/figure.png" pageId="9" pageNumber="10">Fig. 2F</figureCitation>
). Endopod 3.3 times as long as wide.
</paragraph>
<paragraph id="4D0436C85B68FFEAFF6F4E17FEA6FC6C" blockId="9.[227,1436,784,915]" pageId="9" pageNumber="10">
Mouthparts (
<figureCitation id="D5802A4D5B68FFEAFEF14E17FE59FCD5" box="[381,457,784,810]" captionStart="Fig" captionStartId="8.[227,270,1967,1993]" captionTargetBox="[284,1396,760,1911]" captionTargetId="figure@8.[227,1436,735,1926]" captionTargetPageId="8" captionText="Fig. 4. Hamodactylus paraqabai sp. nov., ♀, paratype (PoCL 1.6 mm, Madang, Papua New Guinea, MNHN-IU-2013-11090), mouthparts. A. Mandible. B. Same as previous, turned. C. Same as previous," figureDoi="http://doi.org/10.5281/zenodo.3837565" httpUri="https://zenodo.org/record/3837565/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
; dissected from female 
<typeStatus id="9200886A5B68FFEAFD674E17FCC0FCD5" box="[747,848,784,810]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
MNHN-IU-2013-11090). Mandible (
<figureCitation id="D5802A4D5B68FFEAFA9F4E17FACAFCD5" box="[1299,1370,784,810]" captionStart="Fig" captionStartId="8.[227,270,1967,1993]" captionTargetBox="[284,1396,760,1911]" captionTargetId="figure@8.[227,1436,735,1926]" captionTargetPageId="8" captionText="Fig. 4. Hamodactylus paraqabai sp. nov., ♀, paratype (PoCL 1.6 mm, Madang, Papua New Guinea, MNHN-IU-2013-11090), mouthparts. A. Mandible. B. Same as previous, turned. C. Same as previous," figureDoi="http://doi.org/10.5281/zenodo.3837565" httpUri="https://zenodo.org/record/3837565/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
A–C) with reduced cylindrical molar process bearing a few brushes of setae distally (
<figureCitation id="D5802A4D5B68FFEAFBF04E34FB48FCB2" box="[1148,1240,819,845]" captionStart="Fig" captionStartId="8.[227,270,1967,1993]" captionTargetBox="[284,1396,760,1911]" captionTargetId="figure@8.[227,1436,735,1926]" captionTargetPageId="8" captionText="Fig. 4. Hamodactylus paraqabai sp. nov., ♀, paratype (PoCL 1.6 mm, Madang, Papua New Guinea, MNHN-IU-2013-11090), mouthparts. A. Mandible. B. Same as previous, turned. C. Same as previous," figureDoi="http://doi.org/10.5281/zenodo.3837565" httpUri="https://zenodo.org/record/3837565/files/figure.png" pageId="9" pageNumber="10">Fig. 4C</figureCitation>
). Incisor process slightly expanded distally, with four large teeth on truncated tip, of which lateral most enlarged. Palp absent.
</paragraph>
<caption id="19C466405B68FFEAFF6D4AA8FCE2F7EF" ID-DOI="http://doi.org/10.5281/zenodo.3837567" ID-Zenodo-Dep="3837567" httpUri="https://zenodo.org/record/3837567/files/figure.png" pageId="9" pageNumber="10" startId="9.[225,268,1967,1993]" targetBox="[280,1360,1049,1913]" targetPageId="9">
<paragraph id="4D0436C85B68FFEAFF6D4AA8FCE2F7EF" blockId="9.[225,1437,1966,2065]" pageId="9" pageNumber="10">
<emphasis id="7FCFEADA5B68FFEAFF6D4AA8FEA5F835" bold="true" box="[225,309,1967,1994]" pageId="9" pageNumber="10">Fig. 5.</emphasis>
<taxonomicName id="8ABB4D4B5B68FFEAFECF4AB7FDE4F835" authorityName="Horka, Fransen &amp; Duris" authorityYear="2016" box="[323,628,1968,1994]" class="Malacostraca" family="Palaemonidae" genus="Hamodactylus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="paraqabai" status="sp. nov.">
<emphasis id="7FCFEADA5B68FFEAFECF4AB7FDE4F835" box="[323,628,1968,1994]" italics="true" pageId="9" pageNumber="10">Hamodactylus paraqabai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64FC57A15B68FFEAFD0E4AB7FD75F835" box="[642,741,1968,1994]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, Ƌ (PoCL 1.1 mm, Lizard Is., Australia, MTQ-33218). 
<emphasis id="7FCFEADA5B68FFEAFF6D4AD4FF66F812" bold="true" box="[225,246,2003,2029]" pageId="9" pageNumber="10">A</emphasis>
. Carapace and rostrum, lateral. 
<emphasis id="7FCFEADA5B68FFEAFDFC4AD4FD14F812" bold="true" box="[624,644,2003,2029]" pageId="9" pageNumber="10">B</emphasis>
. First pleopod. 
<emphasis id="7FCFEADA5B68FFEAFCB24AD4FCC3F812" bold="true" box="[830,851,2003,2029]" pageId="9" pageNumber="10">C</emphasis>
. Same as previous, endopod. 
<emphasis id="7FCFEADA5B68FFEAFB384AD4FB59F812" bold="true" box="[1204,1225,2003,2029]" pageId="9" pageNumber="10">D</emphasis>
. Second pleopod. 
<emphasis id="7FCFEADA5B68FFEAFF6D4AF1FF65F7EF" bold="true" box="[225,245,2038,2064]" pageId="9" pageNumber="10">E</emphasis>
. Same as previous, appendices interna and masculina.
</paragraph>
</caption>
<paragraph id="4D0436C85B6BFFE9FF6F4C36FD28FE6E" blockId="10.[227,1436,305,402]" pageId="10" pageNumber="11">
Maxillula (
<figureCitation id="D5802A4D5B6BFFE9FEEB4C36FE55FEB4" box="[359,453,305,331]" captionStart="Fig" captionStartId="8.[227,270,1967,1993]" captionTargetBox="[284,1396,760,1911]" captionTargetId="figure@8.[227,1436,735,1926]" captionTargetPageId="8" captionText="Fig. 4. Hamodactylus paraqabai sp. nov., ♀, paratype (PoCL 1.6 mm, Madang, Papua New Guinea, MNHN-IU-2013-11090), mouthparts. A. Mandible. B. Same as previous, turned. C. Same as previous," figureDoi="http://doi.org/10.5281/zenodo.3837565" httpUri="https://zenodo.org/record/3837565/files/figure.png" pageId="10" pageNumber="11">Fig. 4D</figureCitation>
) with slender, feebly bilobed palp, lower lobe bearing a small seta distally. Upper lacinia short and stout, truncate distally, with about 10 robust spines distally. Lower lacinia more slender, with group of long setae on truncate tip.
</paragraph>
<paragraph id="4D0436C85B6BFFE9FF6F4CBEFE4FFDC3" blockId="10.[227,1436,441,573]" pageId="10" pageNumber="11">
Maxilla (
<figureCitation id="D5802A4D5B6BFFE9FEDD4CBEFE3CFE2C" box="[337,428,441,467]" captionStart="Fig" captionStartId="8.[227,270,1967,1993]" captionTargetBox="[284,1396,760,1911]" captionTargetId="figure@8.[227,1436,735,1926]" captionTargetPageId="8" captionText="Fig. 4. Hamodactylus paraqabai sp. nov., ♀, paratype (PoCL 1.6 mm, Madang, Papua New Guinea, MNHN-IU-2013-11090), mouthparts. A. Mandible. B. Same as previous, turned. C. Same as previous," figureDoi="http://doi.org/10.5281/zenodo.3837565" httpUri="https://zenodo.org/record/3837565/files/figure.png" pageId="10" pageNumber="11">Fig. 4E</figureCitation>
) with distally tapering, non-setose palp. Basal endite simple, short, stout and blunt, with simple seta distally. Coxal endite obsolete, medial region slightly convex. Scaphognathite normal, widest centrally, about 2 times as long as broad, with more slender posterior lobe. Marginal plumose setae well developed.
</paragraph>
<paragraph id="4D0436C85B6BFFE9FF6F4F63FAC5FD17" blockId="10.[227,1437,612,744]" pageId="10" pageNumber="11">
First maxilliped (
<figureCitation id="D5802A4D5B6BFFE9FE3F4F63FD9CFD81" box="[435,524,612,638]" captionStart="Fig" captionStartId="8.[227,270,1967,1993]" captionTargetBox="[284,1396,760,1911]" captionTargetId="figure@8.[227,1436,735,1926]" captionTargetPageId="8" captionText="Fig. 4. Hamodactylus paraqabai sp. nov., ♀, paratype (PoCL 1.6 mm, Madang, Papua New Guinea, MNHN-IU-2013-11090), mouthparts. A. Mandible. B. Same as previous, turned. C. Same as previous," figureDoi="http://doi.org/10.5281/zenodo.3837565" httpUri="https://zenodo.org/record/3837565/files/figure.png" pageId="10" pageNumber="11">Fig. 4F</figureCitation>
) with slender, tapering, non-setose palp. Basal region broad and round, not distinctly separated from coxal region. Median margin straight, sparsely provided with slender setulose setae. Caridean lobe of exopod distinct, with plumose marginal setae; flagellum greatly reduced, devoid of setae, slightly overreaching caridean lobe. Epipod small, somewhat elongate, rounded distally.
</paragraph>
<paragraph id="4D0436C85B6BFFE9FF6F4E08FC90FC26" blockId="10.[227,1437,783,985]" pageId="10" pageNumber="11">
Second maxilliped (
<figureCitation id="D5802A4D5B6BFFE9FE594E08FDA5FCD6" box="[469,565,783,809]" captionStart="Fig" captionStartId="8.[227,270,1967,1993]" captionTargetBox="[284,1396,760,1911]" captionTargetId="figure@8.[227,1436,735,1926]" captionTargetPageId="8" captionText="Fig. 4. Hamodactylus paraqabai sp. nov., ♀, paratype (PoCL 1.6 mm, Madang, Papua New Guinea, MNHN-IU-2013-11090), mouthparts. A. Mandible. B. Same as previous, turned. C. Same as previous," figureDoi="http://doi.org/10.5281/zenodo.3837565" httpUri="https://zenodo.org/record/3837565/files/figure.png" pageId="10" pageNumber="11">Fig. 4G</figureCitation>
) with small dactylar segment, about 1.6 times as long as wide, convex medially, bearing row of stout, serrulate spines. Propodal segment large, twice as long as dactylar segment, twice as long as wide, with subrectangular distomedial angle. Medial margin broadly convex, with row of coarsely serrulate setae. Carpus short. Merus partly fused to ischium, ischium partly fused to basis, basis without exopod; combined segment stout, about twice as long as wide. Coxa produced medially, with small, suboval epipod laterally.
</paragraph>
<paragraph id="4D0436C85B6BFFE9FF6F4907FB85FB11" blockId="10.[227,1437,1024,1262]" pageId="10" pageNumber="11">
Third maxilliped (
<figureCitation id="D5802A4D5B6BFFE9FE304907FD89FBE4" box="[444,537,1024,1051]" captionStart="Fig" captionStartId="8.[227,270,1967,1993]" captionTargetBox="[284,1396,760,1911]" captionTargetId="figure@8.[227,1436,735,1926]" captionTargetPageId="8" captionText="Fig. 4. Hamodactylus paraqabai sp. nov., ♀, paratype (PoCL 1.6 mm, Madang, Papua New Guinea, MNHN-IU-2013-11090), mouthparts. A. Mandible. B. Same as previous, turned. C. Same as previous," figureDoi="http://doi.org/10.5281/zenodo.3837565" httpUri="https://zenodo.org/record/3837565/files/figure.png" pageId="10" pageNumber="11">Fig. 4H</figureCitation>
) with broad antepenultimate segment, about 4 times as long as wide. Basis completely fused with ischiomerus, junction indicated medially by small knob; median margin sparsely provided with serrulate setae over proximal two thirds. Penultimate segment slender, about 3 times as long as wide, 0.33 of length of antepenultimate segment and with 2 slender, serrulate setae on medial border. Terminal segment 3 times as long as wide, slightly shorter than penultimate segment, with groups of serrulate setae medially and slender terminal setae. Exopod absent. Coxa not produced medially, with broadly rounded lateral lobe and small reduced arthrobranch laterally.
</paragraph>
<paragraph id="4D0436C85B6BFFE9FE9C4812FE71FAD0" blockId="10.[227,1437,1301,1609]" box="[272,481,1301,1327]" pageId="10" pageNumber="11">
<emphasis id="7FCFEADA5B6BFFE9FE9C4812FE71FAD0" bold="true" box="[272,481,1301,1327]" pageId="10" pageNumber="11">Other specimens</emphasis>
</paragraph>
<paragraph id="4D0436C85B6BFFE9FF6F483FFADFF9B6" blockId="10.[227,1437,1301,1609]" pageId="10" pageNumber="11">
Females similar to 
<typeStatus id="9200886A5B6BFFE9FE47483FFDA6FAAD" box="[459,566,1336,1362]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
. Dorsal rostrum teeth varying from 4 (smaller specimen, PoCL 
<quantity id="8A439B2D5B6BFFE9FAB5483FFA08FAAC" box="[1337,1432,1336,1363]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="11" unit="mm" value="1.2">1.2 mm</quantity>
, damaged, lost) to 5, lamina shallower in smallest specimens, first pereiopod fingers with 3–4 distal cutting teeth. Single male specimen available (
<figureCitation id="D5802A4D5B6BFFE9FC904878FCF5FA65" box="[796,869,1407,1434]" captionStart="Fig" captionStartId="9.[225,268,1967,1993]" captionTargetBox="[280,1360,1049,1913]" captionTargetId="figure@9.[227,1436,1032,1926]" captionTargetPageId="9" captionText="Fig. 5. Hamodactylus paraqabai sp. nov., Ƌ (PoCL 1.1 mm, Lizard Is., Australia, MTQ-33218). A. Carapace and rostrum, lateral. B. First pleopod. C. Same as previous, endopod. D. Second pleopod. E. Same as previous, appendices interna and masculina." figureDoi="http://doi.org/10.5281/zenodo.3837567" httpUri="https://zenodo.org/record/3837567/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
) comparatively smaller and more slender than adult females; rostrum (
<figureCitation id="D5802A4D5B6BFFE9FE7348A5FDCCFA42" box="[511,604,1442,1469]" captionStart="Fig" captionStartId="9.[225,268,1967,1993]" captionTargetBox="[280,1360,1049,1913]" captionTargetId="figure@9.[227,1436,1032,1926]" captionTargetPageId="9" captionText="Fig. 5. Hamodactylus paraqabai sp. nov., Ƌ (PoCL 1.1 mm, Lizard Is., Australia, MTQ-33218). A. Carapace and rostrum, lateral. B. First pleopod. C. Same as previous, endopod. D. Second pleopod. E. Same as previous, appendices interna and masculina." figureDoi="http://doi.org/10.5281/zenodo.3837567" httpUri="https://zenodo.org/record/3837567/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
) more shallow, bearing 4 dorsal teeth; first male pleopod (
<figureCitation id="D5802A4D5B6BFFE9FA9A48A5FACBFA42" box="[1302,1371,1442,1469]" captionStart="Fig" captionStartId="9.[225,268,1967,1993]" captionTargetBox="[280,1360,1049,1913]" captionTargetId="figure@9.[227,1436,1032,1926]" captionTargetPageId="9" captionText="Fig. 5. Hamodactylus paraqabai sp. nov., Ƌ (PoCL 1.1 mm, Lizard Is., Australia, MTQ-33218). A. Carapace and rostrum, lateral. B. First pleopod. C. Same as previous, endopod. D. Second pleopod. E. Same as previous, appendices interna and masculina." figureDoi="http://doi.org/10.5281/zenodo.3837567" httpUri="https://zenodo.org/record/3837567/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
B–C) with endopod almost half length of exopod, enlarged distally, with apex rounded, with 3 short simple setae proximally on medial border and 2 short setae more distally on lateral border; second pleopod (
<figureCitation id="D5802A4D5B6BFFE9FAE148EFFF60F9D9" captionStart="Fig" captionStartId="9.[225,268,1967,1993]" captionTargetBox="[280,1360,1049,1913]" captionTargetId="figure@9.[227,1436,1032,1926]" captionTargetPageId="9" captionText="Fig. 5. Hamodactylus paraqabai sp. nov., Ƌ (PoCL 1.1 mm, Lizard Is., Australia, MTQ-33218). A. Carapace and rostrum, lateral. B. First pleopod. C. Same as previous, endopod. D. Second pleopod. E. Same as previous, appendices interna and masculina." figureDoi="http://doi.org/10.5281/zenodo.3837567" httpUri="https://zenodo.org/record/3837567/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
D–E) with endopod slightly shorter than exopod, with appendix masculina about as long as appendix interna, with two unequal distal serrulate setae and one simple seta subdistally on lateral margin.
</paragraph>
<paragraph id="4D0436C85B6BFFE9FF6F4B77FDFBF974" blockId="10.[227,619,1648,1675]" box="[227,619,1648,1675]" pageId="10" pageNumber="11">
<emphasis id="7FCFEADA5B6BFFE9FF6F4B77FE0BF975" bold="true" box="[227,411,1648,1674]" pageId="10" pageNumber="11">Measurements</emphasis>
(PoCL, RL; mm)
</paragraph>
<paragraph id="4D0436C85B6BFFE9FF6F4B98FBC0F922" blockId="10.[227,1436,1695,1757]" pageId="10" pageNumber="11">
Ovigerous females: 1.9, 1.8; 
<typeStatus id="9200886A5B6BFFE9FDBB4B98FD32F946" box="[567,674,1695,1721]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
: 1.5, 1.5. Other females 1.3–1.6, 1.2–1.4 (in female of PoCL 1.6, TL 
<quantity id="8A439B2D5B6BFFE9FE814BC5FEFCF923" box="[269,364,1730,1757]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="10" pageNumber="11" unit="mm" value="9.5">9.5 mm</quantity>
). Male: 1.1, 1.0. Eggs (with eye spots): 0.54–0.65 × 0.40–0.45.
</paragraph>
<paragraph id="4D0436C85B6BFFE9FF6F4A04FD8EF8E2" blockId="10.[227,542,1795,1821]" box="[227,542,1795,1821]" pageId="10" pageNumber="11">
<emphasis id="7FCFEADA5B6BFFE9FF6F4A04FEEBF8E2" bold="true" box="[227,379,1795,1821]" pageId="10" pageNumber="11">Colouration</emphasis>
(
<figureCitation id="D5802A4D5B6BFFE9FE064A04FE4DF8E2" box="[394,477,1795,1821]" captionStart="Fig" captionStartId="16.[227,270,1862,1888]" captionTargetBox="[227,1436,711,1820]" captionTargetId="figure@16.[227,1436,711,1820]" captionTargetPageId="16" captionText="Fig. 10. Colour pattern of three relatives of the genus Hamodactylus. — A. H. aqabai Bruce &amp; Svoboda, 1983, ovigerous ♀ (UO Aq09-55) on alcyonacean coral Nephthea sp., Aqaba, Red Sea. — B–C. H. paraqabai sp. nov., ovigerous ♀♀, Madang, Papua New Guinea, on Nephthea sp. B. Holotype (MNHN-IU-2013-11092). C. Paratype (MNHN-IU-2013-11090). — D. H. pseudaqabai sp. nov., ovigerous ♀, holotype, N. Sulawesi, Indonesia (RMNH.CRUS.D.57195). Photos: Z. Ďuriš (A), T.-" figureDoi="http://doi.org/10.5281/zenodo.3837577" httpUri="https://zenodo.org/record/3837577/files/figure.png" pageId="10" pageNumber="11">Fig. 10</figureCitation>
B–C)
</paragraph>
<paragraph id="4D0436C85B6BFFE8FF6F4A35FDEEFE91" blockId="10.[227,1437,1842,2044]" lastBlockId="11.[227,1436,305,366]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Males largely transparent, with diffuse yellow and white speckles, more dense on cephalothorax. Adult females more heavily coloured. Body generally transparent, faintly speckled with yellow and white colour, with yellow dots on base of the rostrum and on eyestalks, yellow colour more condensed to transverse bands on sides of each basis of antennae, two bands on carapace, and one wide band on each of 1 
<superScript id="BACE9B805B6BFFE9FE984ABAFE8FF834" attach="left" box="[276,287,1981,1995]" fontSize="6" pageId="10" pageNumber="11">st</superScript>
to 5 
<superScript id="BACE9B805B6BFFE9FED84ABAFEF1F834" attach="left" box="[340,353,1981,1995]" fontSize="6" pageId="10" pageNumber="11">th</superScript>
abdominal segments, all interspaced by faint white dots. Whitish path transversely on the tailfan articulations to the last abdominal segment. Anterior rostrum, antennal flagella, pereiopods, the sixth abdominal segment, and tailfan, predominantly transparent. One female from 
<collectingCountry id="35AC76585B6AFFE8FB354C35FA0CFEB4" box="[1209,1436,305,332]" name="Papua New Guinea" pageId="11" pageNumber="12">Papua New Guinea</collectingCountry>
reddish coloured instead of yellow.
</paragraph>
</subSubSection>
<subSubSection id="05A165435B6AFFE8FF6F4C91FDB7FDFD" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="4D0436C85B6AFFE8FF6F4C91FE8EFE4F" blockId="11.[227,286,406,432]" box="[227,286,406,432]" pageId="11" pageNumber="12">
<heading id="164C81A45B6AFFE8FF6F4C91FE8EFE4F" bold="true" box="[227,286,406,432]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="7FCFEADA5B6AFFE8FF6F4C91FE8EFE4F" bold="true" box="[227,286,406,432]" pageId="11" pageNumber="12">Host</emphasis>
</heading>
</paragraph>
<paragraph id="4D0436C85B6AFFE8FF6F4CC3FDB7FDFD" blockId="11.[227,1436,452,514]" pageId="11" pageNumber="12">
Associated with branching soft corals of the genus 
<taxonomicName id="8ABB4D4B5B6AFFE8FCCD4CC3FC21FE21" box="[833,945,452,478]" class="Anthozoa" family="Nephtheidae" genus="Nephthea" kingdom="Animalia" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="genus">
<emphasis id="7FCFEADA5B6AFFE8FCCD4CC3FC21FE21" box="[833,945,452,478]" italics="true" pageId="11" pageNumber="12">Nephthea</emphasis>
</taxonomicName>
sp. (
<figureCitation id="D5802A4D5B6AFFE8FC604CC3FBC6FE21" box="[1004,1110,452,478]" captionStart="Fig" captionStartId="16.[227,270,1862,1888]" captionTargetBox="[227,1436,711,1820]" captionTargetId="figure@16.[227,1436,711,1820]" captionTargetPageId="16" captionText="Fig. 10. Colour pattern of three relatives of the genus Hamodactylus. — A. H. aqabai Bruce &amp; Svoboda, 1983, ovigerous ♀ (UO Aq09-55) on alcyonacean coral Nephthea sp., Aqaba, Red Sea. — B–C. H. paraqabai sp. nov., ovigerous ♀♀, Madang, Papua New Guinea, on Nephthea sp. B. Holotype (MNHN-IU-2013-11092). C. Paratype (MNHN-IU-2013-11090). — D. H. pseudaqabai sp. nov., ovigerous ♀, holotype, N. Sulawesi, Indonesia (RMNH.CRUS.D.57195). Photos: Z. Ďuriš (A), T.-" figureDoi="http://doi.org/10.5281/zenodo.3837577" httpUri="https://zenodo.org/record/3837577/files/figure.png" pageId="11" pageNumber="12">Fig. 10C</figureCitation>
) of the family 
<taxonomicName id="8ABB4D4B5B6AFFE8FA844CC3FA0CFE21" box="[1288,1436,452,478]" class="Anthozoa" family="Nephtheidae" kingdom="Animalia" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="family">Nephtheidae</taxonomicName>
(Octocorallia: 
<taxonomicName id="8ABB4D4B5B6AFFE8FE074CE0FD8BFDFD" box="[395,539,487,514]" class="Anthozoa" kingdom="Animalia" order="Alcyonacea" pageId="11" pageNumber="12" phylum="Cnidaria" rank="order">Alcyonacea</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="05A165435B6AFFE8FF6F4F2EFA0DFD8E" box="[227,1437,553,626]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="4D0436C85B6AFFE8FF6F4F2EFEECFDBC" blockId="11.[227,380,553,579]" box="[227,380,553,579]" pageId="11" pageNumber="12">
<heading id="164C81A45B6AFFE8FF6F4F2EFEECFDBC" bold="true" box="[227,380,553,579]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="7FCFEADA5B6AFFE8FF6F4F2EFEECFDBC" bold="true" box="[227,380,553,579]" pageId="11" pageNumber="12">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="4D0436C85B6AFFE8FF6F4F5FFA0DFD8E" blockId="11.[227,1437,599,626]" box="[227,1437,599,626]" pageId="11" pageNumber="12">
Up to date known from the 
<collectingRegion id="8F7FF82A5B6AFFE8FDAA4F50FD16FD8E" box="[550,646,599,625]" country="Papua New Guinea" name="Madang" pageId="11" pageNumber="12">Madang</collectingRegion>
Lagoon, 
<collectingCountry id="35AC76585B6AFFE8FD794F5FFC4BFD8E" box="[757,987,599,626]" name="Papua New Guinea" pageId="11" pageNumber="12">Papua New Guinea</collectingCountry>
, and the Great Barrier Reef, 
<collectingCountry id="35AC76585B6AFFE8FAA54F50FA08FD8E" box="[1321,1432,599,625]" name="Australia" pageId="11" pageNumber="12">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="05A165435B6AFFE8FF6F4F9EFE58FD1E" box="[227,456,665,738]" pageId="11" pageNumber="12" type="discussion">
<paragraph id="4D0436C85B6AFFE8FF6F4F9EFEC5FD4C" blockId="11.[227,341,665,691]" box="[227,341,665,691]" pageId="11" pageNumber="12">
<heading id="164C81A45B6AFFE8FF6F4F9EFEC5FD4C" bold="true" box="[227,341,665,691]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="7FCFEADA5B6AFFE8FF6F4F9EFEC5FD4C" bold="true" box="[227,341,665,691]" pageId="11" pageNumber="12">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="4D0436C85B6AFFE8FF6F4FCFFE58FD1E" blockId="11.[227,456,711,738]" box="[227,456,711,738]" pageId="11" pageNumber="12">See the Discussion.</paragraph>
</subSubSection>
</treatment>
</document>