763 lines
112 KiB
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763 lines
112 KiB
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<document id="082335BB1FA266F8CCF451FAAE7110F5" ID-CLB-Dataset="281942" ID-DOI="10.5252/geodiversitas2023v45a24" ID-GBIF-Dataset="ccd2ee89-acc4-4fec-97c6-a7895a8ab5ed" ID-ISSN="1638-9395" ID-Zenodo-Dep="10376394" ID-ZooBank="urn:lsid:zoobank.org:pub:73B795BD-23BA-44AA-98CA-0F62DAA3CAD7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702549311693" checkinUser="plazi" docAuthor="Audo, Denis, Kawai, Tadashi, Letenneur, Charlène & Huang, Diying" docDate="2023" docId="7378240DFFC0FFB9FC89FF2FA501CCCD" docLanguage="en" docName="geodiversitas.2023.45.24.pdf" docOrigin="Geodiversitas 45 (24)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a24.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.12:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="11" docTitle="Palaeocambarus licenti" docType="treatment" docVersion="2" lastPageNumber="698" masterDocId="8F415C75FFC7FFB4FFF7FFF6A451CB26" masterDocTitle="Crayfishes from the Jehol biota" masterLastPageNumber="719" masterPageNumber="689" pageNumber="694" updateTime="1702572493601" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="6915ED0EC02D0E66A0A6A611B4E5F925">Crayfishes from the Jehol biota</mods:title>
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<mods:namePart id="BFC2E79639CD9E2ECF1E44A6A874FAD9">Audo, Denis</mods:namePart>
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<mods:affiliation id="01E7E9D12F7BAAE5CE03FB983AC09BB3">Centre de Recherche en Paléontologie - Paris (CR 2 P, UMR 7207), MNHN, CNRS, Sorbonne Université, Muséum national d’Histoire naturelle, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:nameIdentifier id="9FAF5F6F01922E5EB2A5D78CEE84CC05" type="email">denis.audo@mnhn.fr</mods:nameIdentifier>
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<mods:name id="2CF47A5888A6003AFEF282CD5E6A91CB" type="personal">
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<mods:namePart id="099A6FB5F05F7A25C8A307C205B5E347">Kawai, Tadashi</mods:namePart>
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<mods:affiliation id="959957869F31D1BA57C2D774DD3EDB0C">Hokkaido Research Organization, Central Fisheries Research Institute, 238 Hamanaka, Yoichi, 045 - 8555 Hokkaido (Japan)</mods:affiliation>
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<mods:nameIdentifier id="F0C822A7765CE4E6130C2C7F0C3DCFE0" type="email">tadashikawai8@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C3AC4769C7E772C5E50816B98C6D1CBC">Letenneur, Charlène</mods:namePart>
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<mods:affiliation id="D572443E99D8E6EEC327182B74077306">Centre de Recherche en Paléontologie - Paris (CR 2 P, UMR 7207), MNHN, CNRS, Sorbonne Université, Muséum national d’Histoire naturelle, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:nameIdentifier id="D0608437E380B65B4C49B07A20925B3C" type="email">charlene.letenneur@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="10C969C9862DE9F1B9F1EEF744F62215">Huang, Diying</mods:namePart>
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<mods:affiliation id="8A179917FDC49F7497EA75C50FBCCD1C">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008 (China)</mods:affiliation>
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<mods:nameIdentifier id="C3B3591AB4568FD5CC8FB79D10F103CA" type="email">dyhuang@nigpas.ac.cn</mods:nameIdentifier>
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<mods:title id="5EAC5929D8F62B5DD7EA0BC693F807FE">Geodiversitas</mods:title>
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<mods:part id="B5EE02B2D65D2FA08B02ACE240B6B0DC">
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<mods:date id="63653710401B7D268889551F223D429D">2023</mods:date>
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<treatment id="7378240DFFC0FFB9FC89FF2FA501CCCD" ID-DOI="http://doi.org/10.5281/zenodo.10379221" ID-GBIF-Taxon="219863334" ID-Zenodo-Dep="10379221" LSID="urn:lsid:plazi:treatment:7378240DFFC0FFB9FC89FF2FA501CCCD" httpUri="http://treatment.plazi.org/id/7378240DFFC0FFB9FC89FF2FA501CCCD" lastPageId="13" lastPageNumber="698" pageId="7" pageNumber="694">
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<subSubSection id="B3CBC690FFC0FFB3FC89FF2FA10FCBD7" box="[894,1374,215,243]" pageId="7" pageNumber="694" type="nomenclature">
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<paragraph id="FB6E951BFFC0FFB3FC89FF2FA10FCBD7" blockId="7.[894,1374,215,274]" box="[894,1374,215,243]" pageId="7" pageNumber="694">
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<taxonomicName id="3CD1EE98FFC0FFB3FC89FF2FA10FCBD7" authority="(Van Straelen, 1928)" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[894,1374,215,243]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="694" phylum="Arthropoda" rank="species" species="licenti">
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<emphasis id="C9A54909FFC0FFB3FC89FF2FA026CBD5" bold="true" box="[894,1143,217,243]" italics="true" pageId="7" pageNumber="694">Palaeocambarus licenti</emphasis>
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(Van Straelen, 1928)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3CBC690FFC0FFB3FC6BFF01A16ECA37" box="[924,1343,247,274]" pageId="7" pageNumber="694" type="description">
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<paragraph id="FB6E951BFFC0FFB3FC6BFF01A16ECA37" blockId="7.[894,1374,215,274]" box="[924,1343,247,274]" pageId="7" pageNumber="694">
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(
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<figureCitation id="63EA899EFFC0FFB3FC52FF01A7A8CA37" box="[933,1017,247,273]" captionStart="FIG" captionStartId="6.[132,143,1768,1785]" captionTargetBox="[132,1455,215,1725]" captionTargetId="figure-89@6.[129,1459,213,1728]" captionTargetPageId="6" captionText="FIG. 1. — Pleopods of different decapod crustaceans, fossil and extant: A, B, Palaeocambarus licenti (Van Straelen, 1928b) seen in ventral view (A, holotype of Cricoidoscelosus aethus Taylor, 1999, NIGP-126337) and lateral view (B, specimen NIGP-Shen-2), both from the Early Cretaceous of China; C, pleopod 4 of a female Cambaroides dauricus (MNHN-IU-2022-4077, extant, Mandchuria);D, Austropotamobius llopisi (Vía, 1971) (female? – specimen MUPA-LH-14011), from the Early Cretaceous of Spain; E, Palaeobenthesicymus libanensis (Brocchi, 1875) (specimen MNHN.F.A30607) from the Santonian of Lebanon; F, female Polycheles typhlops Heller, 1862 (MNHN-IU-2022-4080, extant, New Caledonia). Photographs: D. Audo. Scale bars: A, C, 1 mm; B, D, E, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10376396" httpUri="https://zenodo.org/record/10376396/files/figure.png" pageId="7" pageNumber="694">Figs 1A</figureCitation>
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;
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<figureCitation id="63EA899EFFC0FFB3FBF2FF01A019CA37" box="[1029,1096,247,274]" captionStart="FIG" captionStartId="8.[132,143,1657,1674]" captionTargetBox="[132,1455,215,1605]" captionTargetId="figure-122@8.[131,1487,214,1605]" captionTargetPageId="8" captionText="FIG. 2. — Comparison of epistomes in fossil and extant crayfishes: A, B, epistome of P. licenti (NIGP-Shen-26), photo (A) and interpretative line-drawing (B); C, epistome of P. licenti (NIGP-Shen-40);D, epistome of a female Cambaroides dauricus (Pallas, 1972) (MNHN-IU-2022-4077, extant, Mandchuria);E, Cambarus bartonii (Fabricius, 1798) (MNHN-IU-2022-4074, extant, Philadelphia, United States); F, Astacus astacus (Linnaeus, 1758) (MNHN-IU-2022-4069, extant, Nissi, Greece). Photos of fossil specimens in cross-polarized light. Abbreviations: a1, antennula; a2.ba, antennal basipodite; a2.cx, antennal coxa; epi, epistome; uo, urinary orifice (nephropore). Photographs: D. Audo. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10376400" httpUri="https://zenodo.org/record/10376400/files/figure.png" pageId="7" pageNumber="694">2B, C</figureCitation>
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;
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||
<figureCitation id="63EA899EFFC0FFB3FBA4FF01A030CA37" box="[1107,1121,247,273]" captionStart="FIG" captionStartId="9.[132,143,1768,1785]" captionTargetBox="[132,1455,215,1732]" captionTargetId="figure-74@9.[-62,1107,214,1736]" captionTargetPageId="9" captionText="FIG. 3. — Ischial hooks in P. licenti: A-D, on specimen NIGP-126337 (holotype of Cricoidoscelosus aethus), ventral view in cross-polarized light (A) and interpretative line-drawing (B); C, D, detailed view of the base of P2-P4 (th5-th7) (C), and interpretative line drawing (D) – note that C and D are aligned horizontally for convenience;E, F, on specimen NIGP-Shen-22,detail of the ischium of P4 (th7). Abbreviations: a1, antennula;a2, antenna; ba, basipod;cx, coxa;epi, epistome; ex, uropodal exopod; go, male gonopod; isc, ischium; md, mandible; me, merus; mxp3, maxilliped 3 (thoracopod 3); P1-P5, pereiopods 1-5 (thoracopod 4-8); pl3, pleopod 3; r, rostrum; s2, pleonite 2; sc, scaphocerite. Photographs: A, D. Audo; C, E, D.-Y. Huang. Scale bars: A, B, 20 mm; C-F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10376402" httpUri="https://zenodo.org/record/10376402/files/figure.png" pageId="7" pageNumber="694">3</figureCitation>
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;
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<figureCitation id="63EA899EFFC0FFB3FB98FF01A0E6CA34" box="[1135,1207,247,274]" captionStart="FIG" captionStartId="10.[132,143,1802,1819]" captionTargetBox="[133,1455,215,1767]" captionTargetId="figure-72@10.[132,1455,215,1772]" captionTargetPageId="10" captionText="FIG. 4. — Sexual dimorphism: A, B, Cambaroides dauricus(AS11,extant,Mandchuria) male (A) and female (B); C, D, possible annulus ventralis in Palaeocambarus licenti (Van Straelen, 1928) specimen NIGP-126342 from the Early Cretaceous of China, photograph (C) and interpretative line-drawing (D); E, annulus ventralis in Cambarus bartonii (Fabricius,1798) (AS90,extant, Philadelphia,United States);Abbreviations: av, annulus ventralis;av?, potential annulus ventralis; gp♀, female gonopore; gp♂, male gonopore; pl1-2♂, pleopods 1-2 (male gonopods). Photographs: D. Audo. Scale bars: A, B, E, 5 mm; C, D, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10376404" httpUri="https://zenodo.org/record/10376404/files/figure.png" pageId="7" pageNumber="694">4C, D</figureCitation>
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;
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<figureCitation id="63EA899EFFC0FFB3FB35FF01A0B6CA37" box="[1218,1255,247,274]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="11.[132,143,1221,1238]" captionStartId-1="12.[132,143,1895,1912]" captionStartId-2="13.[132,143,1542,1559]" captionStartId-3="14.[132,143,1330,1347]" captionTargetBox-0="[132,1452,215,1179]" captionTargetBox-1="[149,1438,215,1877]" captionTargetBox-2="[132,1456,215,1582]" captionTargetBox-3="[185,1397,215,1287]" captionTargetId-0="figure-396@11.[121,815,136,1180]" captionTargetId-1="figure-16@12.[146,1440,214,1877]" captionTargetId-2="figure-221@13.[156,1432,214,1599]" captionTargetId-3="figure-242@14.[185,803,199,1290]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIG. 5. — Neotype of Palaeocambarus licenti (Van Straelen, 1928), specimen NIGP-126342: ventral view in cross-polarized light (A) and interpretative line-drawing (B). Abbreviations: a2, antenna; av?, potential annulus ventralis; dt; digestive tract; epi, epistome; ex, uropodal exopod; mxp3, maxilliped 3 (thoracopod 3); o, eye; P1-P5, pereiopods 1-5 (th4-th8); r, rostrum; s2-s5, pleonites 2-5; sc, scaphocerite. Photograph: D. Audo. Scale bars: 20 mm." captionText-1="FIG. 6. — Palaeocambarus licenti (Van Straelen,1928): three well preserved specimens NIGP-Shen-1,both in left-lateral view,in cross-polarized light (A), interpretative line drawing (B) and close-up of the cephalic area of one of the right crayfish in NIGP-Shen-1 (C), visualisation in Meshlab of the inverted surface volume of NIGP-Shen-GM1 from the photogrammetric model, with Minnaert shader (D). Abbreviations: a, branchiocardiac groove; b, antennal groove; d, gastro-orbital groove; dt, digestive tract; e1e, cervical groove;e1e (os), cervical groove (opposite side); ex, uropodal exopod; o, eye; P1, pereiopod 1; po (d), postorbital carina (dorsal part); po (v), postorbital carina (ventral part); pr, postrostral carina; r1, acumen (distal part) of rostrum; r2, proximal part of rostrum; s1-s6, pleonites 1-6; sc, scaphocerite; su, subdorsal carina; t, telson. Photograph and photogrammetric model: D. Audo. Scale bars: A, B, 20 mm; D, 10 mm." captionText-2="FIG. 7. — Palaeocambarus licenti (Van Straelen,1928),specimen NIGP-175159 with well-preserved ornamentation of first pereiopod (thoracopod 4). Photograph: D.-Y. Huang. Scale bars: 20 mm." captionText-3="FIG. 8. — Syntype of Astacus spinirostrius (synonym of P. licenti), ventral view in cross-polarized light (A) and interpretative line-drawing (B), Abbreviations:a1, antennula:a2, antenna;en, uropodal endopod;epi, epistome;ex, uropodal exopod;go, male gonopod;md, mandible;mxp3, maxilliped 3 (th.3);P1-P5, pereiopods 1-5 (th. 4-8); s2-s6, pleonites 2-6; sc, scaphocerite; t, telson. Photograph: D. Audo. Scale bars: 20 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10376406" figureDoi-1="http://doi.org/10.5281/zenodo.10376408" figureDoi-2="http://doi.org/10.5281/zenodo.10376410" figureDoi-3="http://doi.org/10.5281/zenodo.10376416" httpUri-0="https://zenodo.org/record/10376406/files/figure.png" httpUri-1="https://zenodo.org/record/10376408/files/figure.png" httpUri-2="https://zenodo.org/record/10376410/files/figure.png" httpUri-3="https://zenodo.org/record/10376416/files/figure.png" pageId="7" pageNumber="694">5-8</figureCitation>
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;
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<figureCitation id="63EA899EFFC0FFB3FB02FF01A167CA37" box="[1269,1334,247,274]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="15.[218,228,1628,1645]" captionStartId-1="16.[309,320,854,871]" captionStartId-2="17.[132,143,1391,1408]" captionStartId-3="18.[132,143,1353,1370]" captionTargetBox-0="[216,1367,214,1585]" captionTargetBox-1="[161,1426,214,812]" captionTargetBox-2="[148,1440,214,1349]" captionTargetBox-3="[134,1453,267,1312]" captionTargetId-0="figure-204@15.[210,1377,205,1587]" captionTargetId-1="figure-435@16.[156,1432,209,817]" captionTargetId-2="figure-333@17.[701,1256,726,1264]" captionTargetId-3="figure-326@18.[134,1454,215,1357]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="FIG. 10. — Scientific reconstructions in dorsal view of Palaeastacus licenti (Van Straelen, 1928b). Drawing: C. Letenneur (UMR 7207 CR2P)." captionText-1="FIG. 11. — Scientific reconstructions in lateral view of Palaeastacus licenti. Drawing: C. Letenneur (UMR 7207 CR2P)." captionText-2="FIG. 12. — Eyes and gastroliths: detail of the eye of spec11 (A) and specimen with gastroliths NIGP-126353, whole specimen in dorsal view, in cross-polarized light (B) and close-up of the gastroliths imprint, in cross-polarized light (C). Photographs: D. Audo. Scale bars: A, 1 mm; B, 20 mm; C, 10 mm." captionText-3="FIG. 13. — Two types of preservation: A, B, NIGP-126354,a possible fossil corpse (A) as shown by the presence of an infilled proctodeum (B); C, fossilized moult NIGP-Shen-RT132 (C) as shown by the disconnection of the cephalothorax shield, thinness of the exoskeleton and lack of fossilized soft-parts. Photographs: D. Audo. Scale bars: 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10376420" figureDoi-1="http://doi.org/10.5281/zenodo.10376424" figureDoi-2="http://doi.org/10.5281/zenodo.10376426" figureDoi-3="http://doi.org/10.5281/zenodo.10376436" httpUri-0="https://zenodo.org/record/10376420/files/figure.png" httpUri-1="https://zenodo.org/record/10376424/files/figure.png" httpUri-2="https://zenodo.org/record/10376426/files/figure.png" httpUri-3="https://zenodo.org/record/10376436/files/figure.png" pageId="7" pageNumber="694">10-13</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B3CBC690FFC0FFB3FCDAFEC6A03FC9C6" pageId="7" pageNumber="694" type="reference_group">
|
||
<paragraph id="FB6E951BFFC0FFB3FCDAFEC6A153CA45" blockId="7.[813,1456,304,355]" pageId="7" pageNumber="694">
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<treatmentCitationGroup id="DBC1B235FFC0FFB3FCDAFEC6A153CA45" pageId="7" pageNumber="694">
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<treatmentCitation id="7A70B30AFFC0FFB3FCDAFEC6A08BCA6E" author="VAN STRAELEN V." box="[813,1242,304,328]" page="138" pageId="7" pageNumber="694" year="1928">
|
||
<taxonomicName id="3CD1EE98FFC0FFB3FCDAFEC6A08BCA6E" authority="Van Straelen, 1928 b: 133 - 138" authorityName="Van Straelen" authorityPageNumber="133 - 138" authorityYear="1928" box="[813,1242,304,328]" class="Malacostraca" family="Astacidae" genus="Astacus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="694" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFC0FFB3FCDAFEC6A7E1CA6E" box="[813,944,304,328]" italics="true" pageId="7" pageNumber="694">Astacus licenti</emphasis>
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FC44FEC7A08BCA6E" author="VAN STRAELEN V." box="[947,1242,304,328]" pageId="7" pageNumber="694" pagination="133 - 139" refId="ref14038" refString="VAN STRAELEN V. 1928 b. - On a fossil freshwater crayfish from Eastern Mongolia. Bulletin of the Geological Society of China 7: 133 - 139." type="journal article" year="1928">Van Straelen, 1928b: 133-138</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, pl. 1, figs 1-2. —
|
||
<treatmentCitation id="7A70B30AFFC0FFB3FA7DFEC7A05BCA44" author="IMAIZUMI R." page="173" pageId="7" pageNumber="694" year="1938">
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FA7DFEC7A05BCA44" author="IMAIZUMI R." pageId="7" pageNumber="694" pagination="173 - 178" refId="ref12046" refString="IMAIZUMI R. 1938. - Fossil crayfishes from Jehol. Science reports of the Tohoku Imperial University 2 nd, series, Geology 19 (2): 173 - 178, pls 22 - 23." type="journal article" year="1938">Imaizumi 1938: 173-176</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 23, figs 1, 2, 4-6, 11.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="FB6E951BFFC0FFB3FCDAFE8CA7CBCA8B" blockId="7.[813,1455,378,429]" pageId="7" pageNumber="694">
|
||
<treatmentCitationGroup id="DBC1B235FFC0FFB3FCDAFE8CA7CBCA8B" pageId="7" pageNumber="694">
|
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<treatmentCitation id="7A70B30AFFC0FFB3FCDAFE8CA0A4CAB4" author="IMAIZUMI R." box="[813,1269,378,402]" page="178" pageId="7" pageNumber="694" year="1938">
|
||
<taxonomicName id="3CD1EE98FFC0FFB3FCDAFE8CA0A4CAB4" authority="Imaizumi, 1938: 176 - 178" authorityName=", Imaizumi" authorityPageNumber="176 - 178" authorityYear="1938" box="[813,1269,378,402]" class="Malacostraca" family="Astacidae" genus="Astacus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="694" phylum="Arthropoda" rank="species" species="spinirostrius">
|
||
<emphasis id="C9A54909FFC0FFB3FCDAFE8CA7B8CAB4" box="[813,1001,378,402]" italics="true" pageId="7" pageNumber="694">Astacus spinirostrius</emphasis>
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FC06FE8CA0A4CAB4" author="IMAIZUMI R." box="[1009,1269,378,402]" pageId="7" pageNumber="694" pagination="173 - 178" refId="ref12046" refString="IMAIZUMI R. 1938. - Fossil crayfishes from Jehol. Science reports of the Tohoku Imperial University 2 nd, series, Geology 19 (2): 173 - 178, pls 22 - 23." type="journal article" year="1938">Imaizumi, 1938: 176-178</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, pls 12, 23, figs 9, 10, 12, 13.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="FB6E951BFFC0FFB3FCDAFE32A079C960" blockId="7.[811,1456,452,582]" pageId="7" pageNumber="694">
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||
<treatmentCitationGroup id="DBC1B235FFC0FFB3FCDAFE32A079C960" pageId="7" pageNumber="694">
|
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<treatmentCitation id="7A70B30AFFC0FFB3FCDAFE32A121CAFA" author="TAYLOR R. S. & SCHRAM F. R. & SHEN Y. - B." box="[813,1392,452,476]" page="135" pageId="7" pageNumber="694" year="1999">
|
||
<taxonomicName id="3CD1EE98FFC0FFB3FCDAFE32A121CAFA" authority="Taylor, Schram & Shen, 1999: 130 - 135" authorityName="Taylor, Schram & Shen" authorityPageNumber="130 - 135" authorityYear="1999" box="[813,1392,452,476]" class="Malacostraca" family="Cricoidoscelosidae" genus="Cricoidoscelosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="694" phylum="Arthropoda" rank="species" species="aethus">
|
||
<emphasis id="C9A54909FFC0FFB3FCDAFE32A7A9CAFA" box="[813,1016,452,476]" italics="true" pageId="7" pageNumber="694">Cricoidoscelosus aethus</emphasis>
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FC0CFE32A121CAFA" author="TAYLOR R. S. & SCHRAM F. R. & SHEN Y. - B." box="[1019,1392,452,476]" pageId="7" pageNumber="694" pagination="121 - 136" refId="ref13937" refString="TAYLOR R. S., SCHRAM F. R. & SHEN Y. - B. 1999. - A new crayfish family (Decapoda: Astacida) from the Upper Jurassic of China, with a reinterpretation of other Chinese crayfish taxa. Paleontological Research 3 (2): 121 - 136." type="journal article" year="1999">Taylor, Schram & Shen, 1999: 130-135</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, figs 3, 7, 8. —
|
||
<treatmentCitation id="7A70B30AFFC0FFB3FC8CFE29A0E1CAD1" author="SCHRAM F. R. & SHEN Y. - B." box="[891,1200,479,503]" page="416" pageId="7" pageNumber="694" year="2000">
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FC8CFE29A0E1CAD1" author="SCHRAM F. R. & SHEN Y. - B." box="[891,1200,479,503]" pageId="7" pageNumber="694" pagination="416 - 418" refId="ref13311" refString="SCHRAM F. R. & SHEN Y. - B. 2000. - An unusual specimen of fossil crayfish molt. Acta Palaeontologica Sinica 39 (3): 416 - 418" type="journal article" year="2000">Schram & Shen 2000: 416-418</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 1. —
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FAFAFE29A1F8CAD1" author="SHEN Y. - B. & SHRAM F. R. & TAYLOR R. S." box="[1293,1449,479,503]" pageId="7" pageNumber="694" pagination="26 - 33" refId="ref13672" refString="SHEN Y. - B., SHRAM F. R. & TAYLOR R. S. 2001. - Morphological variation in fossil crayfish of the Jehol biota, Liaoning Province, China and its texonomic discrimination. Chinese Science Bulletin 46 (1): 26 - 33. https: // doi. org / 10.1007 / BF 03183202" type="journal article" year="2001">
|
||
Shen
|
||
<emphasis id="C9A54909FFC0FFB3FAB3FE29A123CAD1" box="[1348,1394,479,503]" italics="true" pageId="7" pageNumber="694">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
: figs 1c, 2a, 3a, 3b. —
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FBFEFE0FA022C937" author="SHEN Y. - B." box="[1033,1139,505,529]" pageId="7" pageNumber="694" pagination="52 - 57" refId="ref13562" refString="SHEN Y. - B. 2008. - Shrimps, in CHANG M. - M., CHEN P. - J., WANG Y. - Q., WANG Y. & MIAO D. - S. The Jehol Fossils, the Emergence of Feathered Dinosaurs, Beaked Birds and Flowering Plants. Academic Press, London, Burlington, San Diego, New York: 52 - 57." type="book chapter" year="2008">Shen 2008</bibRefCitation>
|
||
: figs 62-65. —
|
||
<treatmentCitation id="7A70B30AFFC0FFB3FAE5FE0FA7D9C90A" author="SCHWEITZER C. E. & FELDMANN R. M. & GARASSINO A. & KARASAWA H. & SCHWEIGERT G." page="32" pageId="7" pageNumber="694" year="2010">
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FAE5FE0FA7D9C90A" author="SCHWEITZER C. E. & FELDMANN R. M. & GARASSINO A. & KARASAWA H. & SCHWEIGERT G." pageId="7" pageNumber="694" pagination="1 - 222" refId="ref13425" refString="SCHWEITZER C. E., FELDMANN R. M., GARASSINO A., KARASAWA H. & SCHWEIGERT G. 2010. - Systematic List of Fossil Decapod Crustacean Species. Brill, Leiden: 1 - 222. (Crustaceana Monographs; 10). https: // doi. org / 10.1163 / ej. 9789004178915. i- 222" type="book chapter" year="2010">
|
||
Schweitzer
|
||
<emphasis id="C9A54909FFC0FFB3FA77FE0CA1E1C937" box="[1408,1456,505,529]" italics="true" pageId="7" pageNumber="694">et al.</emphasis>
|
||
2010: 32
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="7A70B30AFFC0FFB3FC44FDE2A0A7C90A" author="CRANDALL K. A. & DE GRAVE S." box="[947,1270,532,556]" page="616" pageId="7" pageNumber="694" year="2017">
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FC44FDE2A0A7C90A" author="CRANDALL K. A. & DE GRAVE S." box="[947,1270,532,556]" pageId="7" pageNumber="694" pagination="615 - 653" refId="ref11085" refString="CRANDALL K. A. & DE GRAVE S. 2017. - An updated classification of the freshwater crayfishes (Decapoda: Astacidea) of the world, with a complete species list. Journal of Crustacean Biology 37: 615 - 653. https: // doi. org / 10.1093 / jcbiol / rux 070" type="journal article" year="2017">Crandall & De Grave 2017: 616</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 636. —
|
||
<treatmentCitation id="7A70B30AFFC0FFB3FAA2FDE2A78DC960" author="BELL P. R. & BICKNELL R. D. C. & SMITH E. T." page="1023" pageId="7" pageNumber="694" year="2020">
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FAA2FDE2A78DC960" author="BELL P. R. & BICKNELL R. D. C. & SMITH E. T." pageId="7" pageNumber="694" pagination="1023 - 1030" refId="ref10286" refString="BELL P. R., BICKNELL R. D. C. & SMITH E. T. 2020. - Crayfish bio-gastroliths from eastern Australia and the middle Cretaceous distribution of Parastacidae. Geological Magazine 157 (7): 1023 - 1030. https: // doi. org / 10.1017 / S 0016756819001092" type="journal article" year="2020">
|
||
Bell
|
||
<emphasis id="C9A54909FFC0FFB3FA76FDE2A1E1C90A" box="[1409,1456,532,556]" italics="true" pageId="7" pageNumber="694">et al.</emphasis>
|
||
2020: 1023-1030
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
,
|
||
<emphasis id="C9A54909FFC0FFB3FC11FDD8A079C960" bold="true" box="[998,1064,558,582]" pageId="7" pageNumber="694">n. syn.</emphasis>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="FB6E951BFFC0FFB3FCDAFDA8A03FC9C6" blockId="7.[811,1455,606,736]" pageId="7" pageNumber="694">
|
||
<treatmentCitationGroup id="DBC1B235FFC0FFB3FCDAFDA8A03FC9C6" pageId="7" pageNumber="694">
|
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<taxonomicName id="3CD1EE98FFC0FFB3FCDAFDA8A053C950" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[813,1026,606,630]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="694" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFC0FFB3FCDAFDA8A053C950" box="[813,1026,606,630]" italics="true" pageId="7" pageNumber="694">Palaeocambarus licenti</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="7A70B30AFFC0FFB3FBE0FDA8A173C950" author="TAYLOR R. S. & SCHRAM F. R. & SHEN Y. - B." box="[1047,1314,606,630]" page="122" pageId="7" pageNumber="694" year="1999">
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FBE0FDA8A173C950" author="TAYLOR R. S. & SCHRAM F. R. & SHEN Y. - B." box="[1047,1314,606,630]" pageId="7" pageNumber="694" pagination="121 - 136" refId="ref13937" refString="TAYLOR R. S., SCHRAM F. R. & SHEN Y. - B. 1999. - A new crayfish family (Decapoda: Astacida) from the Upper Jurassic of China, with a reinterpretation of other Chinese crayfish taxa. Paleontological Research 3 (2): 121 - 136." type="journal article" year="1999">
|
||
Taylor
|
||
<emphasis id="C9A54909FFC0FFB3FBACFDA8A0D9C950" box="[1115,1160,606,630]" italics="true" pageId="7" pageNumber="694">et al.</emphasis>
|
||
1999: 122-130
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs 2, 4-6. —
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FCDCFD8EA780C9B6" author="SHEN Y. - B. & SHRAM F. R. & TAYLOR R. S." box="[811,977,632,656]" pageId="7" pageNumber="694" pagination="26 - 33" refId="ref13672" refString="SHEN Y. - B., SHRAM F. R. & TAYLOR R. S. 2001. - Morphological variation in fossil crayfish of the Jehol biota, Liaoning Province, China and its texonomic discrimination. Chinese Science Bulletin 46 (1): 26 - 33. https: // doi. org / 10.1007 / BF 03183202" type="journal article" year="2001">
|
||
Shen
|
||
<emphasis id="C9A54909FFC0FFB3FC91FD8FA7C6C9B6" box="[870,919,632,656]" italics="true" pageId="7" pageNumber="694">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
: figs 1a, 1b, 2b, 2c, 3c. —
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FB1CFD8EA107C9B6" author="SHEN Y. - B." box="[1259,1366,632,656]" pageId="7" pageNumber="694" pagination="52 - 57" refId="ref13562" refString="SHEN Y. - B. 2008. - Shrimps, in CHANG M. - M., CHEN P. - J., WANG Y. - Q., WANG Y. & MIAO D. - S. The Jehol Fossils, the Emergence of Feathered Dinosaurs, Beaked Birds and Flowering Plants. Academic Press, London, Burlington, San Diego, New York: 52 - 57." type="book chapter" year="2008">Shen 2008</bibRefCitation>
|
||
: figs 66- 67. —
|
||
<treatmentCitation id="7A70B30AFFC0FFB3FC86FD65A03DC98D" author="SCHWEITZER C. E. & FELDMANN R. M. & GARASSINO A. & KARASAWA H. & SCHWEIGERT G." box="[881,1132,659,683]" page="32" pageId="7" pageNumber="694" year="2010">
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FC86FD65A03DC98D" author="SCHWEITZER C. E. & FELDMANN R. M. & GARASSINO A. & KARASAWA H. & SCHWEIGERT G." box="[881,1132,659,683]" pageId="7" pageNumber="694" pagination="1 - 222" refId="ref13425" refString="SCHWEITZER C. E., FELDMANN R. M., GARASSINO A., KARASAWA H. & SCHWEIGERT G. 2010. - Systematic List of Fossil Decapod Crustacean Species. Brill, Leiden: 1 - 222. (Crustaceana Monographs; 10). https: // doi. org / 10.1163 / ej. 9789004178915. i- 222" type="book chapter" year="2010">
|
||
Schweitzer
|
||
<emphasis id="C9A54909FFC0FFB3FC28FD65A05CC98D" box="[991,1037,659,683]" italics="true" pageId="7" pageNumber="694">et al.</emphasis>
|
||
2010: 32
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="7A70B30AFFC0FFB3FB63FD65A707C9E3" author="BRACKEN-GRISSOM H. D. & AHYONG S. T. & WILKINSON R. D. & FELDLMANN R. M. & SCHWEITZER C. E. & BREINHOLT J. W. & BENDALL M. & PALERO F. & CHAN T. - Y. & FELDER D. L. & ROBLES R. & CHU K. - H. & TSANG L. - M. & KIM D. & MARTIN J. W. & CRANDALL K. A." page="465" pageId="7" pageNumber="694" year="2014">
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FB63FD65A707C9E3" author="BRACKEN-GRISSOM H. D. & AHYONG S. T. & WILKINSON R. D. & FELDLMANN R. M. & SCHWEITZER C. E. & BREINHOLT J. W. & BENDALL M. & PALERO F. & CHAN T. - Y. & FELDER D. L. & ROBLES R. & CHU K. - H. & TSANG L. - M. & KIM D. & MARTIN J. W. & CRANDALL K. A." pageId="7" pageNumber="694" pagination="457 - 479" refId="ref10613" refString="BRACKEN-GRISSOM H. D., AHYONG S. T., WILKINSON R. D., FELDLMANN R. M., SCHWEITZER C. E., BREINHOLT J. W., BENDALL M., PALERO F., CHAN T. - Y., FELDER D. L., ROBLES R., CHU K. - H., TSANG L. - M., KIM D., MARTIN J. W. & CRANDALL K. A. 2014. - The emergence of lobsters: phylogenetic relationships, morphological evolution and divergence time comparisons of an ancient group (Decapoda: Achelata, Astacidea, Glypheidea, Polychelida). Systematic Biology 63 (4): 457 - 479. https: // doi. org / 10.1093 / sysbio / syu 008" type="journal article" year="2014">
|
||
Bracken-Grissom
|
||
<emphasis id="C9A54909FFC0FFB3FAB4FD65A120C98D" box="[1347,1393,659,683]" italics="true" pageId="7" pageNumber="694">et al.</emphasis>
|
||
2014: 465
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="7A70B30AFFC0FFB3FC8AFD5BA0EDC9E3" author="CRANDALL K. A. & DE GRAVE S." box="[893,1212,685,709]" page="616" pageId="7" pageNumber="694" year="2017">
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FC8AFD5BA0EDC9E3" author="CRANDALL K. A. & DE GRAVE S." box="[893,1212,685,709]" pageId="7" pageNumber="694" pagination="615 - 653" refId="ref11085" refString="CRANDALL K. A. & DE GRAVE S. 2017. - An updated classification of the freshwater crayfishes (Decapoda: Astacidea) of the world, with a complete species list. Journal of Crustacean Biology 37: 615 - 653. https: // doi. org / 10.1093 / jcbiol / rux 070" type="journal article" year="2017">Crandall & De Grave 2017: 616</bibRefCitation>
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</treatmentCitation>
|
||
, 631. —
|
||
<treatmentCitation id="7A70B30AFFC0FFB3FAE0FD5BA732C9F9" author="BELL P. R. & BICKNELL R. D. C. & SMITH E. T." page="1026" pageId="7" pageNumber="694" year="2020">
|
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<bibRefCitation id="9F40E8EAFFC0FFB3FAE0FD5BA732C9F9" author="BELL P. R. & BICKNELL R. D. C. & SMITH E. T." pageId="7" pageNumber="694" pagination="1023 - 1030" refId="ref10286" refString="BELL P. R., BICKNELL R. D. C. & SMITH E. T. 2020. - Crayfish bio-gastroliths from eastern Australia and the middle Cretaceous distribution of Parastacidae. Geological Magazine 157 (7): 1023 - 1030. https: // doi. org / 10.1017 / S 0016756819001092" type="journal article" year="2020">
|
||
Bell
|
||
<emphasis id="C9A54909FFC0FFB3FAB4FD58A120C9E3" box="[1347,1393,685,709]" italics="true" pageId="7" pageNumber="694">et al.</emphasis>
|
||
2020: 1026
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FC7BFD31A078C9C6" author="XING L. & NIU K. & TAYLOR R. S. & EVANS S. E." box="[908,1065,711,736]" pageId="7" pageNumber="694" pagination="104320" refId="ref14338" refString="XING L., NIU K., TAYLOR R. S. & EVANS S. E. 2020. - Integumentary remains and abdominal contents in the Early Cretaceous Chinese lizard, Yabeinosaurus (Squamata), demonstrate colour banding and a diet including crayfish. Cretaceous Research 108: 104320. https: // doi. org / 10.1016 / j. cretres. 2019.104320" type="journal article" year="2020">
|
||
Xing
|
||
<emphasis id="C9A54909FFC0FFB3FC36FD3EA7A1C9F9" box="[961,1008,711,735]" italics="true" pageId="7" pageNumber="694">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
: fig. 4.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
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|
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|
||
<collectionCode id="9DC00DDEFFC0FFB3FC3EFD0BA7A2C828" box="[969,1011,765,782]" pageId="7" pageNumber="694">AND</collectionCode>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="FD8803C7FFC0FFB3FC0CFD0BA006C828" baselines="778" box="[1019,1111,765,782]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="7" pageNumber="694">
|
||
<collectionCode id="9DC00DDEFFC0FFB3FC0CFD0BA006C828" box="[1019,1111,765,782]" pageId="7" pageNumber="694">HORIZON</collectionCode>
|
||
</smallCapsWord>
|
||
. — Barremian (
|
||
<emphasis id="C9A54909FFC0FFB3FB0AFD0EA15CC829" box="[1277,1293,760,783]" italics="true" pageId="7" pageNumber="694">c.</emphasis>
|
||
125 Ma; Lower Cretaceous) of Yixian Formation, Lingyuan,
|
||
<collectingRegion id="39155BF9FFC0FFB3FB04FCE4A1FDC80C" box="[1267,1452,786,810]" country="China" name="Liaoning" pageId="7" pageNumber="694">Liaoning province</collectingRegion>
|
||
,
|
||
<collectingCountry id="83C6D58BFFC0FFB3FCD9FCDAA73AC862" box="[814,875,812,836]" name="China" pageId="7" pageNumber="694">China</collectingCountry>
|
||
(
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FC8FFCDAA061C862" author="SWISHER C. C. & III & WANG Y. - Q. & WANG X. - L. & XU X. & WANG Y." box="[888,1072,812,836]" pageId="7" pageNumber="694" pagination="58 - 61" refId="ref13818" refString="SWISHER C. C. III, WANG Y. - Q., WANG X. - L., XU X. & WANG Y. 1999. - Cretaceous Age for the feathered dinosaurs of Liaoning, China. Nature 400 (6739): 58 - 61. https: // doi. org / 10.1038 / 21872" type="journal article" year="1999">
|
||
Swisher
|
||
<emphasis id="C9A54909FFC0FFB3FC3EFCDBA7A9C862" box="[969,1016,812,836]" italics="true" pageId="7" pageNumber="694">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FB6E951BFFC0FFB3FCDAFCAAA170C8A8" blockId="7.[813,1455,860,910]" pageId="7" pageNumber="694">
|
||
<materialsCitation id="4BB99F46FFC0FFB3FCDAFCAAA170C8A8" ID-GBIF-Occurrence="4463183302" collectionCode="TYPE, MATERIAL" determinerName="Van Straelen" pageId="7" pageNumber="694" specimenCount="1" typeStatus="holotype">
|
||
<smallCapsWord id="FD8803C7FFC0FFB3FCDAFCAAA70CC857" baselines="877,877" box="[813,861,860,883]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="7" pageNumber="694">TYPE</smallCapsWord>
|
||
<smallCapsWord id="FD8803C7FFC0FFB3FC94FC97A790C854" baselines="878" box="[867,961,865,882]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="7" pageNumber="694">MATERIAL</smallCapsWord>
|
||
. —
|
||
<typeStatus id="246A2BB9FFC0FFB3FC1BFCAAA018C852" box="[1004,1097,860,884]" pageId="7" pageNumber="694" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="3CD1EE98FFC0FFB3FB90FCAAA0BAC852" authorityName="Van Straelen" authorityYear="1928" box="[1127,1259,860,884]" class="Malacostraca" family="Astacidae" genus="Astacus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="694" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFC0FFB3FB90FCAAA0BAC852" box="[1127,1259,860,884]" italics="true" pageId="7" pageNumber="694">Astacus licenti</emphasis>
|
||
</taxonomicName>
|
||
by original designation (
|
||
<determinerName id="0EB279DFFFC0FFB3FC96FC80A7B0C8A8" box="[865,993,886,910]" pageId="7" pageNumber="694">Van Straelen</determinerName>
|
||
, 1928) not found, probably lost.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="FB6E951BFFC0FFB3FCDAFC50A100C898" blockId="7.[813,1361,934,958]" box="[813,1361,934,958]" pageId="7" pageNumber="694">
|
||
<materialsCitation id="4BB99F46FFC0FFB3FCDAFC50A11CC898" ID-GBIF-Occurrence="4463183301" box="[813,1357,934,958]" pageId="7" pageNumber="694">
|
||
<smallCapsWord id="FD8803C7FFC0FFB3FCDAFC50A7D6C89A" baselines="951,952" box="[813,903,934,957]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Neotype" pageId="7" pageNumber="694">
|
||
<typeStatus id="246A2BB9FFC0FFB3FCDAFC50A7D6C89A" box="[813,903,934,957]" pageId="7" pageNumber="694" type="neotype">NEOTYPE</typeStatus>
|
||
</smallCapsWord>
|
||
. — NIGP-126342, herein designated (
|
||
<figureCitation id="63EA899EFFC0FFB3FAFCFC50A112C898" box="[1291,1347,934,958]" captionStart="FIG" captionStartId="11.[132,143,1221,1238]" captionTargetBox="[132,1452,215,1179]" captionTargetId="figure-396@11.[121,815,136,1180]" captionTargetPageId="11" captionText="FIG. 5. — Neotype of Palaeocambarus licenti (Van Straelen, 1928), specimen NIGP-126342: ventral view in cross-polarized light (A) and interpretative line-drawing (B). Abbreviations: a2, antenna; av?, potential annulus ventralis; dt; digestive tract; epi, epistome; ex, uropodal exopod; mxp3, maxilliped 3 (thoracopod 3); o, eye; P1-P5, pereiopods 1-5 (th4-th8); r, rostrum; s2-s5, pleonites 2-5; sc, scaphocerite. Photograph: D. Audo. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10376406" httpUri="https://zenodo.org/record/10376406/files/figure.png" pageId="7" pageNumber="694">Fig. 5</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3CBC690FFC0FFB3FCDAFC20A02AC8CB" box="[813,1147,981,1005]" pageId="7" pageNumber="694" type="description">
|
||
<paragraph id="FB6E951BFFC0FFB3FCDAFC20A02AC8CB" blockId="7.[813,1147,981,1005]" box="[813,1147,981,1005]" pageId="7" pageNumber="694">
|
||
<smallCapsWord id="FD8803C7FFC0FFB3FCDAFC20A7E1C8CA" baselines="999,1000" box="[813,944,982,1005]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Description" pageId="7" pageNumber="694">DESCRIPTION</smallCapsWord>
|
||
. — See
|
||
<figureCitation id="63EA899EFFC0FFB3FBF4FC23A02AC8CB" box="[1027,1147,981,1005]" captionStart="APPENDIX" captionStartId="23.[526,538,219,236]" captionTargetPageId="23" captionText="APPENDIX 1. — Descriptive table of P. licenti (Van Straelen, 1928)." pageId="7" pageNumber="694">Appendix 1.</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3CBC690FFC0FFB3FCDAFBF3A176CE53" pageId="7" pageNumber="694" type="materials_examined">
|
||
<paragraph id="FB6E951BFFC0FFB3FCDAFBF3A176CE53" blockId="7.[809,1457,1029,1397]" pageId="7" pageNumber="694">
|
||
EXAMINED MATERIAL. —
|
||
<specimenCount id="EDD75E92FFC0FFB3FBD5FBF3A0E1CF3B" box="[1058,1200,1029,1053]" count="106" pageId="7" pageNumber="694" type="generic">106 specimens</specimenCount>
|
||
from NIGPAS collections and three from Tohoku University collections:
|
||
<materialsCitation id="4BB99F46FFC0FFB3FAF3FBE9A176CE53" ID-GBIF-Occurrence="4463183303" pageId="7" pageNumber="694">
|
||
NIGP-DYH-3-6, NIGP-Shen-1 (
|
||
<figureCitation id="63EA899EFFC0FFB3FC4BFBCCA7A1CF74" box="[956,1008,1082,1106]" captionStart="FIG" captionStartId="12.[132,143,1895,1912]" captionTargetBox="[149,1438,215,1877]" captionTargetId="figure-16@12.[146,1440,214,1877]" captionTargetPageId="12" captionText="FIG. 6. — Palaeocambarus licenti (Van Straelen,1928): three well preserved specimens NIGP-Shen-1,both in left-lateral view,in cross-polarized light (A), interpretative line drawing (B) and close-up of the cephalic area of one of the right crayfish in NIGP-Shen-1 (C), visualisation in Meshlab of the inverted surface volume of NIGP-Shen-GM1 from the photogrammetric model, with Minnaert shader (D). Abbreviations: a, branchiocardiac groove; b, antennal groove; d, gastro-orbital groove; dt, digestive tract; e1e, cervical groove;e1e (os), cervical groove (opposite side); ex, uropodal exopod; o, eye; P1, pereiopod 1; po (d), postorbital carina (dorsal part); po (v), postorbital carina (ventral part); pr, postrostral carina; r1, acumen (distal part) of rostrum; r2, proximal part of rostrum; s1-s6, pleonites 1-6; sc, scaphocerite; su, subdorsal carina; t, telson. Photograph and photogrammetric model: D. Audo. Scale bars: A, B, 20 mm; D, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10376408" httpUri="https://zenodo.org/record/10376408/files/figure.png" pageId="7" pageNumber="694">Figs 6</figureCitation>
|
||
A-C), Shen-2-3 (
|
||
<figureCitation id="63EA899EFFC0FFB3FB7DFBCCA09FCF74" box="[1162,1230,1082,1106]" captionStart="FIG" captionStartId="6.[132,143,1768,1785]" captionTargetBox="[132,1455,215,1725]" captionTargetId="figure-89@6.[129,1459,213,1728]" captionTargetPageId="6" captionText="FIG. 1. — Pleopods of different decapod crustaceans, fossil and extant: A, B, Palaeocambarus licenti (Van Straelen, 1928b) seen in ventral view (A, holotype of Cricoidoscelosus aethus Taylor, 1999, NIGP-126337) and lateral view (B, specimen NIGP-Shen-2), both from the Early Cretaceous of China; C, pleopod 4 of a female Cambaroides dauricus (MNHN-IU-2022-4077, extant, Mandchuria);D, Austropotamobius llopisi (Vía, 1971) (female? – specimen MUPA-LH-14011), from the Early Cretaceous of Spain; E, Palaeobenthesicymus libanensis (Brocchi, 1875) (specimen MNHN.F.A30607) from the Santonian of Lebanon; F, female Polycheles typhlops Heller, 1862 (MNHN-IU-2022-4080, extant, New Caledonia). Photographs: D. Audo. Scale bars: A, C, 1 mm; B, D, E, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10376396" httpUri="https://zenodo.org/record/10376396/files/figure.png" pageId="7" pageNumber="694">Fig. 1B</figureCitation>
|
||
), Shen-4-6, Shen-11-20, Shen-22-23 (
|
||
<figureCitation id="63EA899EFFC0FFB3FC5FFBA2A054CF4B" box="[936,1029,1108,1133]" captionStart="FIG" captionStartId="9.[132,143,1768,1785]" captionTargetBox="[132,1455,215,1732]" captionTargetId="figure-74@9.[-62,1107,214,1736]" captionTargetPageId="9" captionText="FIG. 3. — Ischial hooks in P. licenti: A-D, on specimen NIGP-126337 (holotype of Cricoidoscelosus aethus), ventral view in cross-polarized light (A) and interpretative line-drawing (B); C, D, detailed view of the base of P2-P4 (th5-th7) (C), and interpretative line drawing (D) – note that C and D are aligned horizontally for convenience;E, F, on specimen NIGP-Shen-22,detail of the ischium of P4 (th7). Abbreviations: a1, antennula;a2, antenna; ba, basipod;cx, coxa;epi, epistome; ex, uropodal exopod; go, male gonopod; isc, ischium; md, mandible; me, merus; mxp3, maxilliped 3 (thoracopod 3); P1-P5, pereiopods 1-5 (thoracopod 4-8); pl3, pleopod 3; r, rostrum; s2, pleonite 2; sc, scaphocerite. Photographs: A, D. Audo; C, E, D.-Y. Huang. Scale bars: A, B, 20 mm; C-F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10376402" httpUri="https://zenodo.org/record/10376402/files/figure.png" pageId="7" pageNumber="694">Fig. 3E, F</figureCitation>
|
||
), Shen-26 (
|
||
<figureCitation id="63EA899EFFC0FFB3FB87FBA2A09ECF4B" box="[1136,1231,1108,1133]" captionStart="FIG" captionStartId="8.[132,143,1657,1674]" captionTargetBox="[132,1455,215,1605]" captionTargetId="figure-122@8.[131,1487,214,1605]" captionTargetPageId="8" captionText="FIG. 2. — Comparison of epistomes in fossil and extant crayfishes: A, B, epistome of P. licenti (NIGP-Shen-26), photo (A) and interpretative line-drawing (B); C, epistome of P. licenti (NIGP-Shen-40);D, epistome of a female Cambaroides dauricus (Pallas, 1972) (MNHN-IU-2022-4077, extant, Mandchuria);E, Cambarus bartonii (Fabricius, 1798) (MNHN-IU-2022-4074, extant, Philadelphia, United States); F, Astacus astacus (Linnaeus, 1758) (MNHN-IU-2022-4069, extant, Nissi, Greece). Photos of fossil specimens in cross-polarized light. Abbreviations: a1, antennula; a2.ba, antennal basipodite; a2.cx, antennal coxa; epi, epistome; uo, urinary orifice (nephropore). Photographs: D. Audo. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10376400" httpUri="https://zenodo.org/record/10376400/files/figure.png" pageId="7" pageNumber="694">Fig. 2A, B</figureCitation>
|
||
), Shen-29, Shen-32-36, Shen-38-39; Shen-40 (
|
||
<figureCitation id="63EA899EFFC0FFB3FC0AFB99A012CFA1" box="[1021,1091,1135,1159]" captionStart="FIG" captionStartId="8.[132,143,1657,1674]" captionTargetBox="[132,1455,215,1605]" captionTargetId="figure-122@8.[131,1487,214,1605]" captionTargetPageId="8" captionText="FIG. 2. — Comparison of epistomes in fossil and extant crayfishes: A, B, epistome of P. licenti (NIGP-Shen-26), photo (A) and interpretative line-drawing (B); C, epistome of P. licenti (NIGP-Shen-40);D, epistome of a female Cambaroides dauricus (Pallas, 1972) (MNHN-IU-2022-4077, extant, Mandchuria);E, Cambarus bartonii (Fabricius, 1798) (MNHN-IU-2022-4074, extant, Philadelphia, United States); F, Astacus astacus (Linnaeus, 1758) (MNHN-IU-2022-4069, extant, Nissi, Greece). Photos of fossil specimens in cross-polarized light. Abbreviations: a1, antennula; a2.ba, antennal basipodite; a2.cx, antennal coxa; epi, epistome; uo, urinary orifice (nephropore). Photographs: D. Audo. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10376400" httpUri="https://zenodo.org/record/10376400/files/figure.png" pageId="7" pageNumber="694">Fig.2C</figureCitation>
|
||
), Shen-42, Shen-47, Shen-50-56, Shen-58, Shen-62, Shen-64, Shen-66-68, Shen-71, Shen-a-b-c, Shen-f-g-h, Shen-GM1 (
|
||
<figureCitation id="63EA899EFFC0FFB3FC51FB52A7A0CF9D" box="[934,1009,1188,1212]" captionStart="FIG" captionStartId="12.[132,143,1895,1912]" captionTargetBox="[149,1438,215,1877]" captionTargetId="figure-16@12.[146,1440,214,1877]" captionTargetPageId="12" captionText="FIG. 6. — Palaeocambarus licenti (Van Straelen,1928): three well preserved specimens NIGP-Shen-1,both in left-lateral view,in cross-polarized light (A), interpretative line drawing (B) and close-up of the cephalic area of one of the right crayfish in NIGP-Shen-1 (C), visualisation in Meshlab of the inverted surface volume of NIGP-Shen-GM1 from the photogrammetric model, with Minnaert shader (D). Abbreviations: a, branchiocardiac groove; b, antennal groove; d, gastro-orbital groove; dt, digestive tract; e1e, cervical groove;e1e (os), cervical groove (opposite side); ex, uropodal exopod; o, eye; P1, pereiopod 1; po (d), postorbital carina (dorsal part); po (v), postorbital carina (ventral part); pr, postrostral carina; r1, acumen (distal part) of rostrum; r2, proximal part of rostrum; s1-s6, pleonites 1-6; sc, scaphocerite; su, subdorsal carina; t, telson. Photograph and photogrammetric model: D. Audo. Scale bars: A, B, 20 mm; D, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10376408" httpUri="https://zenodo.org/record/10376408/files/figure.png" pageId="7" pageNumber="694">Fig. 6D</figureCitation>
|
||
, see also Appendix 3), Shen-RT132 (
|
||
<figureCitation id="63EA899EFFC0FFB3FAB9FB52A1F2CF9A" box="[1358,1443,1188,1212]" captionStart="FIG" captionStartId="18.[132,143,1353,1370]" captionTargetBox="[134,1453,267,1312]" captionTargetId="figure-326@18.[134,1454,215,1357]" captionTargetPageId="18" captionText="FIG. 13. — Two types of preservation: A, B, NIGP-126354,a possible fossil corpse (A) as shown by the presence of an infilled proctodeum (B); C, fossilized moult NIGP-Shen-RT132 (C) as shown by the disconnection of the cephalothorax shield, thinness of the exoskeleton and lack of fossilized soft-parts. Photographs: D. Audo. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10376436" httpUri="https://zenodo.org/record/10376436/files/figure.png" pageId="7" pageNumber="694">Fig. 13C</figureCitation>
|
||
), Shen-RT141, Shen-145,
|
||
<specimenCount id="EDD75E92FFC0FFB3FBEEFB49A0C9CFF0" box="[1049,1176,1214,1238]" count="1" pageId="7" pageNumber="694" type="generic">one specimen</specimenCount>
|
||
from Shen where the number “53” is crossed, NIGP-126337 (
|
||
<typeStatus id="246A2BB9FFC0FFB3FBA6FB2FA0F4CFD7" box="[1105,1189,1241,1265]" pageId="7" pageNumber="694" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="3CD1EE98FFC0FFB3FB49FB2FA141CFD7" authorityName="Taylor, Schram & Shen" authorityYear="1999" box="[1214,1296,1241,1265]" class="Malacostraca" family="Cricoidoscelosidae" genus="Cricoidoscelosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="694" phylum="Arthropoda" rank="species" species="aethus">
|
||
<emphasis id="C9A54909FFC0FFB3FB49FB2FA141CFD7" box="[1214,1296,1241,1265]" italics="true" pageId="7" pageNumber="694">C. aethus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="63EA899EFFC0FFB3FAEBFB2FA132CFD7" box="[1308,1379,1241,1265]" captionStart="FIG" captionStartId="6.[132,143,1768,1785]" captionTargetBox="[132,1455,215,1725]" captionTargetId="figure-89@6.[129,1459,213,1728]" captionTargetPageId="6" captionText="FIG. 1. — Pleopods of different decapod crustaceans, fossil and extant: A, B, Palaeocambarus licenti (Van Straelen, 1928b) seen in ventral view (A, holotype of Cricoidoscelosus aethus Taylor, 1999, NIGP-126337) and lateral view (B, specimen NIGP-Shen-2), both from the Early Cretaceous of China; C, pleopod 4 of a female Cambaroides dauricus (MNHN-IU-2022-4077, extant, Mandchuria);D, Austropotamobius llopisi (Vía, 1971) (female? – specimen MUPA-LH-14011), from the Early Cretaceous of Spain; E, Palaeobenthesicymus libanensis (Brocchi, 1875) (specimen MNHN.F.A30607) from the Santonian of Lebanon; F, female Polycheles typhlops Heller, 1862 (MNHN-IU-2022-4080, extant, New Caledonia). Photographs: D. Audo. Scale bars: A, C, 1 mm; B, D, E, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10376396" httpUri="https://zenodo.org/record/10376396/files/figure.png" pageId="7" pageNumber="694">Figs 1A</figureCitation>
|
||
;
|
||
<figureCitation id="63EA899EFFC0FFB3FA9CFB2FA126CFD7" box="[1387,1399,1241,1265]" captionStart="FIG" captionStartId="9.[132,143,1768,1785]" captionTargetBox="[132,1455,215,1732]" captionTargetId="figure-74@9.[-62,1107,214,1736]" captionTargetPageId="9" captionText="FIG. 3. — Ischial hooks in P. licenti: A-D, on specimen NIGP-126337 (holotype of Cricoidoscelosus aethus), ventral view in cross-polarized light (A) and interpretative line-drawing (B); C, D, detailed view of the base of P2-P4 (th5-th7) (C), and interpretative line drawing (D) – note that C and D are aligned horizontally for convenience;E, F, on specimen NIGP-Shen-22,detail of the ischium of P4 (th7). Abbreviations: a1, antennula;a2, antenna; ba, basipod;cx, coxa;epi, epistome; ex, uropodal exopod; go, male gonopod; isc, ischium; md, mandible; me, merus; mxp3, maxilliped 3 (thoracopod 3); P1-P5, pereiopods 1-5 (thoracopod 4-8); pl3, pleopod 3; r, rostrum; s2, pleonite 2; sc, scaphocerite. Photographs: A, D. Audo; C, E, D.-Y. Huang. Scale bars: A, B, 20 mm; C-F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10376402" httpUri="https://zenodo.org/record/10376402/files/figure.png" pageId="7" pageNumber="694">3</figureCitation>
|
||
A-D), 126338-126347, 126353 (
|
||
<figureCitation id="63EA899EFFC0FFB3FBE8FB05A0D6CE2D" box="[1055,1159,1267,1291]" captionStart="FIG" captionStartId="17.[132,143,1391,1408]" captionTargetBox="[148,1440,214,1349]" captionTargetId="figure-333@17.[701,1256,726,1264]" captionTargetPageId="17" captionText="FIG. 12. — Eyes and gastroliths: detail of the eye of spec11 (A) and specimen with gastroliths NIGP-126353, whole specimen in dorsal view, in cross-polarized light (B) and close-up of the gastroliths imprint, in cross-polarized light (C). Photographs: D. Audo. Scale bars: A, 1 mm; B, 20 mm; C, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10376426" httpUri="https://zenodo.org/record/10376426/files/figure.png" pageId="7" pageNumber="694">Fig.12B, C</figureCitation>
|
||
), 126354 (
|
||
<figureCitation id="63EA899EFFC0FFB3FB1DFB05A102CE2D" box="[1258,1363,1267,1291]" captionStart="FIG" captionStartId="18.[132,143,1353,1370]" captionTargetBox="[134,1453,267,1312]" captionTargetId="figure-326@18.[134,1454,215,1357]" captionTargetPageId="18" captionText="FIG. 13. — Two types of preservation: A, B, NIGP-126354,a possible fossil corpse (A) as shown by the presence of an infilled proctodeum (B); C, fossilized moult NIGP-Shen-RT132 (C) as shown by the disconnection of the cephalothorax shield, thinness of the exoskeleton and lack of fossilized soft-parts. Photographs: D. Audo. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10376436" httpUri="https://zenodo.org/record/10376436/files/figure.png" pageId="7" pageNumber="694">Figs 13A, B</figureCitation>
|
||
), 126366, new unpublished specimens in NIGP collections: NIGP-175159 (
|
||
<figureCitation id="63EA899EFFC0FFB3FCC3FADEA73BCE19" box="[820,874,1320,1344]" captionStart="FIG" captionStartId="13.[132,143,1542,1559]" captionTargetBox="[132,1456,215,1582]" captionTargetId="figure-221@13.[156,1432,214,1599]" captionTargetPageId="13" captionText="FIG. 7. — Palaeocambarus licenti (Van Straelen,1928),specimen NIGP-175159 with well-preserved ornamentation of first pereiopod (thoracopod 4). Photograph: D.-Y. Huang. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10376410" httpUri="https://zenodo.org/record/10376410/files/figure.png" pageId="7" pageNumber="694">Fig. 7</figureCitation>
|
||
), 175159 (
|
||
<figureCitation id="63EA899EFFC0FFB3FC20FADEA05CCE19" box="[983,1037,1320,1344]" captionStart="FIG" captionStartId="13.[132,143,1542,1559]" captionTargetBox="[132,1456,215,1582]" captionTargetId="figure-221@13.[156,1432,214,1599]" captionTargetPageId="13" captionText="FIG. 7. — Palaeocambarus licenti (Van Straelen,1928),specimen NIGP-175159 with well-preserved ornamentation of first pereiopod (thoracopod 4). Photograph: D.-Y. Huang. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10376410" httpUri="https://zenodo.org/record/10376410/files/figure.png" pageId="7" pageNumber="694">Fig. 7</figureCitation>
|
||
), and research specimens without collection number herein numbered sp1-31 (sp11:
|
||
<figureCitation id="63EA899EFFC0FFB3FB68FAB4A0A5CE7D" box="[1183,1268,1346,1371]" captionStart="FIG" captionStartId="17.[132,143,1391,1408]" captionTargetBox="[148,1440,214,1349]" captionTargetId="figure-333@17.[701,1256,726,1264]" captionTargetPageId="17" captionText="FIG. 12. — Eyes and gastroliths: detail of the eye of spec11 (A) and specimen with gastroliths NIGP-126353, whole specimen in dorsal view, in cross-polarized light (B) and close-up of the gastroliths imprint, in cross-polarized light (C). Photographs: D. Audo. Scale bars: A, 1 mm; B, 20 mm; C, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10376426" httpUri="https://zenodo.org/record/10376426/files/figure.png" pageId="7" pageNumber="694">Fig.12A</figureCitation>
|
||
), Tohoku-U-57254, U-57267 and U-57272 (
|
||
<figureCitation id="63EA899EFFC0FFB3FBEEFAABA01FCE53" box="[1049,1102,1373,1397]" captionStart="FIG" captionStartId="14.[132,143,1330,1347]" captionTargetBox="[185,1397,215,1287]" captionTargetId="figure-242@14.[185,803,199,1290]" captionTargetPageId="14" captionText="FIG. 8. — Syntype of Astacus spinirostrius (synonym of P. licenti), ventral view in cross-polarized light (A) and interpretative line-drawing (B), Abbreviations:a1, antennula:a2, antenna;en, uropodal endopod;epi, epistome;ex, uropodal exopod;go, male gonopod;md, mandible;mxp3, maxilliped 3 (th.3);P1-P5, pereiopods 1-5 (th. 4-8); s2-s6, pleonites 2-6; sc, scaphocerite; t, telson. Photograph: D. Audo. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.10376416" httpUri="https://zenodo.org/record/10376416/files/figure.png" pageId="7" pageNumber="694">Fig. 8</figureCitation>
|
||
; see also
|
||
<figureCitation id="63EA899EFFC0FFB3FB53FAABA142CE53" box="[1188,1299,1373,1397]" captionStart="APPENDIX" captionStartId="32.[267,279,219,236]" captionTargetBox="[147,1441,270,1752]" captionTargetId="figure-49@32.[700,1257,825,1383]" captionTargetPageId="32" captionText="APPENDIX 6. — Specimens without registration numbers used in the study, continued from previo us figures. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10380834" httpUri="https://zenodo.org/record/10380834/files/figure.png" pageId="7" pageNumber="694">Appendix 6</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3CBC690FFC0FFB9FCDAFA62A501CCCD" lastPageId="13" lastPageNumber="700" pageId="7" pageNumber="694" type="discussion">
|
||
<paragraph id="FB6E951BFFC0FFB3FCDAFA62A04BCE8A" blockId="7.[811,1457,1428,2028]" box="[813,1050,1428,1454]" pageId="7" pageNumber="694">
|
||
<smallCapsWord id="FD8803C7FFC0FFB3FCDAFA62A7D6CE8A" baselines="1447,1447" box="[813,903,1428,1454]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remark" pageId="7" pageNumber="694">REMARK</smallCapsWord>
|
||
<smallCapsWord id="FD8803C7FFC0FFB3FC79FA6FA7FECE8A" baselines="1447" box="[910,943,1433,1452]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="on" pageId="7" pageNumber="694">ON</smallCapsWord>
|
||
<smallCapsWord id="FD8803C7FFC0FFB3FC41FA6FA7B0CE8A" baselines="1447" box="[950,993,1433,1452]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="the" pageId="7" pageNumber="694">THE</smallCapsWord>
|
||
<smallCapsWord id="FD8803C7FFC0FFB3FC1FFA6FA04BCE8A" baselines="1447" box="[1000,1050,1433,1452]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="type" pageId="7" pageNumber="694">
|
||
<typeStatus id="246A2BB9FFC0FFB3FC1FFA6FA04BCE8A" box="[1000,1050,1433,1452]" pageId="7" pageNumber="694">TYPE</typeStatus>
|
||
</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="FB6E951BFFC0FFBCFCDAFA45A758CCCA" blockId="7.[811,1457,1428,2028]" lastBlockId="8.[131,777,1841,2028]" lastPageId="8" lastPageNumber="695" pageId="7" pageNumber="694">
|
||
The original type material of
|
||
<taxonomicName id="3CD1EE98FFC0FFB3FBA0FA45A16CCEEB" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[1111,1341,1459,1485]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="694" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFC0FFB3FBA0FA45A16CCEEB" box="[1111,1341,1459,1485]" italics="true" pageId="7" pageNumber="694">Palaeocambarus licenti</emphasis>
|
||
</taxonomicName>
|
||
consists of
|
||
<specimenCount id="EDD75E92FFC0FFB3FCDCFA25A786CECB" box="[811,983,1491,1517]" count="3" pageId="7" pageNumber="694" type="generic">three specimens</specimenCount>
|
||
,
|
||
<specimenCount id="EDD75E92FFC0FFB3FC13FA25A020CECB" box="[996,1137,1491,1517]" count="1" pageId="7" pageNumber="694" type="generic" typeStatus="holotype">one holotype</specimenCount>
|
||
(“type”,
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FB22FA25A1F9CECB" author="VAN STRAELEN V." box="[1237,1448,1491,1517]" pageId="7" pageNumber="694" pagination="133 - 139" refId="ref14038" refString="VAN STRAELEN V. 1928 b. - On a fossil freshwater crayfish from Eastern Mongolia. Bulletin of the Geological Society of China 7: 133 - 139." type="journal article" year="1928">Van Straelen 1928b</bibRefCitation>
|
||
:
|
||
<figureCitation id="63EA899EFFC0FFB3FCDAFA05A73DCD2B" box="[813,876,1523,1549]" captionStart="FIG" captionStartId="6.[132,143,1768,1785]" captionTargetBox="[132,1455,215,1725]" captionTargetId="figure-89@6.[129,1459,213,1728]" captionTargetPageId="6" captionText="FIG. 1. — Pleopods of different decapod crustaceans, fossil and extant: A, B, Palaeocambarus licenti (Van Straelen, 1928b) seen in ventral view (A, holotype of Cricoidoscelosus aethus Taylor, 1999, NIGP-126337) and lateral view (B, specimen NIGP-Shen-2), both from the Early Cretaceous of China; C, pleopod 4 of a female Cambaroides dauricus (MNHN-IU-2022-4077, extant, Mandchuria);D, Austropotamobius llopisi (Vía, 1971) (female? – specimen MUPA-LH-14011), from the Early Cretaceous of Spain; E, Palaeobenthesicymus libanensis (Brocchi, 1875) (specimen MNHN.F.A30607) from the Santonian of Lebanon; F, female Polycheles typhlops Heller, 1862 (MNHN-IU-2022-4080, extant, New Caledonia). Photographs: D. Audo. Scale bars: A, C, 1 mm; B, D, E, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10376396" httpUri="https://zenodo.org/record/10376396/files/figure.png" pageId="7" pageNumber="694">Fig. 1</figureCitation>
|
||
), and
|
||
<specimenCount id="EDD75E92FFC0FFB3FC42FA05A01DCD2B" box="[949,1100,1523,1549]" count="2" pageId="7" pageNumber="694" type="generic" typeStatus="paratype">two paratypes</specimenCount>
|
||
(only one originally illustrated). This material was supposedly initially housed in the Huangho Paiho Museum (
|
||
<collectingCountry id="83C6D58BFFC0FFB3FC14F9C4A068CD6A" box="[995,1081,1586,1612]" name="China" pageId="7" pageNumber="694">Tianjin</collectingCountry>
|
||
,
|
||
<collectingCountry id="83C6D58BFFC0FFB3FBB0F9C4A0C1CD6A" box="[1095,1168,1586,1612]" name="China" pageId="7" pageNumber="694">China</collectingCountry>
|
||
). Enquiries were made to try to locate this material again, but were unsuccessful as the specimens do not appear to be in
|
||
<collectingCountry id="83C6D58BFFC0FFB3FB51F984A0A7CDAA" box="[1190,1270,1650,1676]" name="China" pageId="7" pageNumber="694">Tianjin</collectingCountry>
|
||
Natural History Museum (successor to the Huangho Paiho Museum listed by
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FCBAF945A04ECDEA" author="VAN STRAELEN V." box="[845,1055,1714,1741]" pageId="7" pageNumber="694" pagination="133 - 139" refId="ref14038" refString="VAN STRAELEN V. 1928 b. - On a fossil freshwater crayfish from Eastern Mongolia. Bulletin of the Geological Society of China 7: 133 - 139." type="journal article" year="1928">Van Straelen 1928b</bibRefCitation>
|
||
) nor in the Royal Belgian Institute of Natural Sciences (where Van Straelen used to work). For this reason, and in order to preserve nomenclatural stability, we herein designate the specimen NIGP-126342 as the
|
||
<typeStatus id="246A2BB9FFC0FFB3FAABF8E4A1E0CC0A" box="[1372,1457,1810,1836]" pageId="7" pageNumber="694" type="neotype">neotype</typeStatus>
|
||
of
|
||
<taxonomicName id="3CD1EE98FFC0FFB3FCBCF8C4A7F5CC6D" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[843,932,1841,1867]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="694" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFC0FFB3FCBCF8C4A7F5CC6D" box="[843,932,1841,1867]" italics="true" pageId="7" pageNumber="694">P. licenti</emphasis>
|
||
</taxonomicName>
|
||
. This specimen preserves well the rostrum, first pereiopod, epistome, parts of the tailfan and even what is possibly the annulus ventralis, all of which are important for taxonomy. Besides,
|
||
<bibRefCitation id="9F40E8EAFFC0FFB3FBF3F864A0BCCC8D" author="VAN STRAELEN V." box="[1028,1261,1937,1964]" pageId="7" pageNumber="694" pagination="133 - 139" refId="ref14038" refString="VAN STRAELEN V. 1928 b. - On a fossil freshwater crayfish from Eastern Mongolia. Bulletin of the Geological Society of China 7: 133 - 139." type="journal article" year="1928">Van Straelen (1928b)</bibRefCitation>
|
||
indicates that the now lost
|
||
<typeStatus id="246A2BB9FFC0FFB3FC7BF847A7BBCCED" box="[908,1002,1969,1995]" pageId="7" pageNumber="694" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="246A2BB9FFC0FFB3FBEBF844A0D1CCEA" box="[1052,1152,1970,1996]" pageId="7" pageNumber="694" type="paratype">paratypes</typeStatus>
|
||
come from the south-west of Moukden (present day Shenyang). No outcrops of the Yixian Formation are known in the immediate vicinity of Shenyang, suggesting that the type material came from further southeast of the city. Although the exact origin of the type material cannot be determined precisely, it may correspond to Chaoyang. From this point of view, the
|
||
<typeStatus id="246A2BB9FFCFFFBCFE31F844A64DCCEA" box="[454,540,1970,1996]" pageId="8" pageNumber="695" type="neotype">neotype</typeStatus>
|
||
NIGP-126342 from Lingyuan most likely comes from a relatively similar outcrop.
|
||
</paragraph>
|
||
<caption id="AFAEC593FFCFFFBCFF73F98FA6BCCDC3" ID-DOI="http://doi.org/10.5281/zenodo.10376400" ID-Zenodo-Dep="10376400" httpUri="https://zenodo.org/record/10376400/files/figure.png" pageId="8" pageNumber="695" startId="8.[132,143,1657,1674]" targetBox="[132,1455,215,1605]" targetPageId="8" targetType="figure">
|
||
<paragraph id="FB6E951BFFCFFFBCFF73F98FA6BCCDC3" blockId="8.[132,1457,1657,1765]" pageId="8" pageNumber="695">
|
||
<smallCapsWord id="FD8803C7FFCFFFBCFF73F98FA4CCCDAC" baselines="1670,1671" box="[132,157,1657,1674]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="8" pageNumber="695">FIG</smallCapsWord>
|
||
. 2. — Comparison of epistomes in fossil and extant crayfishes:
|
||
<emphasis id="C9A54909FFCFFFBCFD49F98FA69ACDAC" bold="true" box="[702,715,1657,1674]" pageId="8" pageNumber="695">A</emphasis>
|
||
,
|
||
<emphasis id="C9A54909FFCFFFBCFD21F98FA6B2CDAC" bold="true" box="[726,739,1657,1674]" pageId="8" pageNumber="695">B</emphasis>
|
||
, epistome of
|
||
<taxonomicName id="3CD1EE98FFCFFFBCFCAEF98FA7F3CDAC" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[857,930,1657,1674]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="695" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCFFFBCFCAEF98FA7F3CDAC" box="[857,930,1657,1674]" italics="true" pageId="8" pageNumber="695">P. licenti</emphasis>
|
||
</taxonomicName>
|
||
(NIGP-Shen-26), photo (
|
||
<emphasis id="C9A54909FFCFFFBCFB83F98FA0D0CDAC" bold="true" box="[1140,1153,1657,1674]" pageId="8" pageNumber="695">A</emphasis>
|
||
) and interpretative line-drawing (
|
||
<emphasis id="C9A54909FFCFFFBCFA6FF98FA1F4CDAC" bold="true" box="[1432,1445,1657,1674]" pageId="8" pageNumber="695">B</emphasis>
|
||
);
|
||
<emphasis id="C9A54909FFCFFFBCFF73F966A4C3CD87" bold="true" box="[132,146,1680,1697]" pageId="8" pageNumber="695">C</emphasis>
|
||
, epistome of
|
||
<taxonomicName id="3CD1EE98FFCFFFBCFEF5F966A519CD87" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[258,328,1680,1697]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="695" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCFFFBCFEF5F966A519CD87" box="[258,328,1680,1697]" italics="true" pageId="8" pageNumber="695">P. licenti</emphasis>
|
||
</taxonomicName>
|
||
(NIGP-Shen-40);
|
||
<emphasis id="C9A54909FFCFFFBCFE2DF966A5B9CD87" bold="true" box="[474,488,1680,1697]" pageId="8" pageNumber="695">D</emphasis>
|
||
, epistome of a female
|
||
<taxonomicName id="3CD1EE98FFCFFFBCFD55F966A79ECD87" authority="(Pallas, 1972)" baseAuthorityName="Pallas" baseAuthorityYear="1972" box="[674,975,1680,1697]" class="Malacostraca" family="Cambaroididae" genus="Cambaroides" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="695" phylum="Arthropoda" rank="species" species="dauricus">
|
||
<emphasis id="C9A54909FFCFFFBCFD55F966A70DCD87" box="[674,860,1680,1697]" italics="true" pageId="8" pageNumber="695">Cambaroides dauricus</emphasis>
|
||
(Pallas, 1972)
|
||
</taxonomicName>
|
||
(MNHN-IU-2022-4077, extant, Mandchuria);
|
||
<emphasis id="C9A54909FFCFFFBCFAB3F966A101CD87" bold="true" box="[1348,1360,1680,1697]" pageId="8" pageNumber="695">E</emphasis>
|
||
,
|
||
<taxonomicName id="3CD1EE98FFCFFFBCFAADF966A503CD9E" authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="695" phylum="Arthropoda" rank="species" species="bartonii">
|
||
<emphasis id="C9A54909FFCFFFBCFAADF966A493CD9E" italics="true" pageId="8" pageNumber="695">Cambarus bartonii</emphasis>
|
||
(Fabricius, 1798)
|
||
</taxonomicName>
|
||
(MNHN-IU-2022-4074, extant, Philadelphia, United States);
|
||
<emphasis id="C9A54909FFCFFFBCFCBAF951A709CD9E" bold="true" box="[845,856,1703,1720]" pageId="8" pageNumber="695">F</emphasis>
|
||
,
|
||
<taxonomicName id="3CD1EE98FFCFFFBCFC94F951A028CD9E" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[867,1145,1703,1720]" class="Malacostraca" family="Astacidae" genus="Astacus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="695" phylum="Arthropoda" rank="species" species="astacus">
|
||
<emphasis id="C9A54909FFCFFFBCFC94F951A7BACD9E" box="[867,1003,1703,1720]" italics="true" pageId="8" pageNumber="695">Astacus astacus</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
(MNHN-IU-2022-4069, extant, Nissi, Greece). Photos of fossil specimens in cross-polarized light. Abbreviations:
|
||
<emphasis id="C9A54909FFCFFFBCFCFDF94BA74ECDE8" bold="true" box="[778,799,1725,1742]" pageId="8" pageNumber="695">a1</emphasis>
|
||
, antennula;
|
||
<emphasis id="C9A54909FFCFFFBCFC7EF94BA7EBCDE8" bold="true" box="[905,954,1725,1742]" pageId="8" pageNumber="695">a2.ba</emphasis>
|
||
, antennal basipodite;
|
||
<emphasis id="C9A54909FFCFFFBCFB8FF94BA0F6CDE8" bold="true" box="[1144,1191,1725,1742]" pageId="8" pageNumber="695">a2.cx</emphasis>
|
||
, antennal coxa;
|
||
<emphasis id="C9A54909FFCFFFBCFAC2F94BA101CDE8" bold="true" box="[1333,1360,1725,1742]" pageId="8" pageNumber="695">epi</emphasis>
|
||
, epistome;
|
||
<emphasis id="C9A54909FFCFFFBCFF73F922A4CBCDC3" bold="true" box="[132,154,1748,1765]" pageId="8" pageNumber="695">uo</emphasis>
|
||
, urinary orifice (nephropore). Photographs: D. Audo. Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FB6E951BFFCFFFBCFCDAF8C4A7FDCC6C" blockId="8.[813,1455,1842,2028]" box="[813,940,1842,1868]" pageId="8" pageNumber="695">
|
||
<smallCapsWord id="FD8803C7FFCFFFBCFCDAF8C4A7FDCC6C" baselines="1861,1861" box="[813,940,1842,1868]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comments" pageId="8" pageNumber="695">COMMENTS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="FB6E951BFFCFFFBCFCDAF8A7A024CC4D" blockId="8.[813,1455,1842,2028]" box="[813,1141,1873,1900]" pageId="8" pageNumber="695">
|
||
<emphasis id="C9A54909FFCFFFBCFCDAF8A7A024CC4D" box="[813,1141,1873,1900]" italics="true" pageId="8" pageNumber="695">Number of species in Jehol biota</emphasis>
|
||
</paragraph>
|
||
<paragraph id="FB6E951BFFCFFFBFFCDAF884A695CDAA" blockId="8.[813,1455,1842,2028]" lastBlockId="11.[131,777,1363,2028]" lastPageId="11" lastPageNumber="698" pageId="8" pageNumber="695">
|
||
To date, three separate species of crayfishes from the Jehol biota have been described. To justify the second species,
|
||
<taxonomicName id="3CD1EE98FFCFFFBCFC7CF847A159CCED" authority=", Imaizumi (1938)" authorityName=", Imaizumi" authorityYear="1938" box="[907,1288,1969,1996]" class="Malacostraca" family="Astacidae" genus="Astacus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="695" phylum="Arthropoda" rank="species" species="spinirostrius">
|
||
<emphasis id="C9A54909FFCFFFBCFC7CF847A063CCED" box="[907,1074,1969,1995]" italics="true" pageId="8" pageNumber="695">A. spinirostrius</emphasis>
|
||
,
|
||
<bibRefCitation id="9F40E8EAFFCFFFBCFBB4F847A159CCED" author="IMAIZUMI R." box="[1091,1288,1969,1995]" pageId="8" pageNumber="695" pagination="173 - 178" refId="ref12046" refString="IMAIZUMI R. 1938. - Fossil crayfishes from Jehol. Science reports of the Tohoku Imperial University 2 nd, series, Geology 19 (2): 173 - 178, pls 22 - 23." type="journal article" year="1938">Imaizumi (1938)</bibRefCitation>
|
||
</taxonomicName>
|
||
argued that it differed from
|
||
<taxonomicName id="3CD1EE98FFCFFFBCFC26F827A066CCCD" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[977,1079,2001,2027]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="695" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCFFFBCFC26F827A066CCCD" box="[977,1079,2001,2027]" italics="true" pageId="8" pageNumber="695">A. licenti</emphasis>
|
||
</taxonomicName>
|
||
by the presence of dorsal spines on the rostrum (lacking in
|
||
<taxonomicName id="3CD1EE98FFCEFFBDFE4CF864A64ECC8D" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[443,543,1937,1964]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="696" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCEFFBDFE4CF864A64ECC8D" box="[443,543,1937,1964]" italics="true" pageId="9" pageNumber="696">A. licenti</emphasis>
|
||
</taxonomicName>
|
||
), possibly the shape of cervical groove, the absence of spine on gastric region (present in
|
||
<taxonomicName id="3CD1EE98FFCEFFBDFEF1F827A53BCCCD" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[262,362,2001,2027]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="696" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCEFFBDFEF1F827A53BCCCD" box="[262,362,2001,2027]" italics="true" pageId="9" pageNumber="696">A. licenti</emphasis>
|
||
</taxonomicName>
|
||
), trigonal s2 tergopleuron (?) (more rounded in
|
||
<taxonomicName id="3CD1EE98FFCEFFBDFC4CF864A075CC8D" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[955,1060,1937,1964]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="696" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCEFFBDFC4CF864A075CC8D" box="[955,1060,1937,1964]" italics="true" pageId="9" pageNumber="696">A. licenti</emphasis>
|
||
</taxonomicName>
|
||
) and uropodal exopodite shorter than endopodite (inverse in
|
||
<taxonomicName id="3CD1EE98FFCEFFBDFB81F847A08ACCED" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[1142,1243,1969,1995]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="696" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCEFFBDFB81F847A08ACCED" box="[1142,1243,1969,1995]" italics="true" pageId="9" pageNumber="696">A. licenti</emphasis>
|
||
</taxonomicName>
|
||
). These characters were rejected by
|
||
<bibRefCitation id="9F40E8EAFFCEFFBDFC1BF827A099CCCD" author="TAYLOR R. S. & SCHRAM F. R. & SHEN Y. - B." box="[1004,1224,2001,2027]" pageId="9" pageNumber="696" pagination="121 - 136" refId="ref13937" refString="TAYLOR R. S., SCHRAM F. R. & SHEN Y. - B. 1999. - A new crayfish family (Decapoda: Astacida) from the Upper Jurassic of China, with a reinterpretation of other Chinese crayfish taxa. Paleontological Research 3 (2): 121 - 136." type="journal article" year="1999">
|
||
Taylor
|
||
<emphasis id="C9A54909FFCEFFBDFBCBF824A022CCCD" box="[1084,1139,2001,2027]" italics="true" pageId="9" pageNumber="696">et al.</emphasis>
|
||
(1999)
|
||
</bibRefCitation>
|
||
who suggested they mostly represent variations in preservation. We agree with this view, as for instance, the spines dorsal to the rostrum are indeed actually part of the supraorbital carina, on the side of rostrum. Similarly, the relative sizes of the uropodal exopod and endopod appear different in different specimens due to incomplete preservation of the distal part of these structures, or different orientations of the specimens in the sediment. However, we do not agree with
|
||
<bibRefCitation id="9F40E8EAFFCCFFBFFF74FA65A533CE8B" author="TAYLOR R. S. & SCHRAM F. R. & SHEN Y. - B." box="[131,354,1427,1453]" pageId="11" pageNumber="698" pagination="121 - 136" refId="ref13937" refString="TAYLOR R. S., SCHRAM F. R. & SHEN Y. - B. 1999. - A new crayfish family (Decapoda: Astacida) from the Upper Jurassic of China, with a reinterpretation of other Chinese crayfish taxa. Paleontological Research 3 (2): 121 - 136." type="journal article" year="1999">
|
||
Taylor
|
||
<emphasis id="C9A54909FFCCFFBFFF21FA62A55CCE8B" box="[214,269,1427,1453]" italics="true" pageId="11" pageNumber="698">et al.</emphasis>
|
||
(1999)
|
||
</bibRefCitation>
|
||
, or
|
||
<bibRefCitation id="9F40E8EAFFCCFFBFFE63FA65A630CE8B" author="SHEN Y. - B. & SHRAM F. R. & TAYLOR R. S." box="[404,609,1427,1453]" pageId="11" pageNumber="698" pagination="26 - 33" refId="ref13672" refString="SHEN Y. - B., SHRAM F. R. & TAYLOR R. S. 2001. - Morphological variation in fossil crayfish of the Jehol biota, Liaoning Province, China and its texonomic discrimination. Chinese Science Bulletin 46 (1): 26 - 33. https: // doi. org / 10.1007 / BF 03183202" type="journal article" year="2001">
|
||
Shen
|
||
<emphasis id="C9A54909FFCCFFBFFE23FA62A65ACE8B" box="[468,523,1427,1453]" italics="true" pageId="11" pageNumber="698">et al.</emphasis>
|
||
(2001)
|
||
</bibRefCitation>
|
||
that
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFD56FA65A756CE8B" authorityName="Taylor, Schram & Shen" authorityYear="1999" box="[673,775,1427,1453]" class="Malacostraca" family="Cricoidoscelosidae" genus="Cricoidoscelosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="species" species="aethus">
|
||
<emphasis id="C9A54909FFCCFFBFFD56FA65A756CE8B" box="[673,775,1427,1453]" italics="true" pageId="11" pageNumber="698">C. aethus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFE40FA42A647CEEB" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[439,534,1459,1485]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCCFFBFFE40FA42A647CEEB" box="[439,534,1459,1485]" italics="true" pageId="11" pageNumber="698">P. licenti</emphasis>
|
||
</taxonomicName>
|
||
. As explained above, the diagnostic characters of
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFE4CFA25A6C3CECB" authorityName="Taylor, Schram & Shen" authorityYear="1999" box="[443,658,1491,1517]" class="Malacostraca" family="Cricoidoscelosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="family">Cricoidoscelosidae</taxonomicName>
|
||
, which are the same as those distinguishing
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFE0BFA05A630CD2B" authorityName="Taylor, Schram & Shen" authorityYear="1999" box="[508,609,1523,1549]" class="Malacostraca" family="Cricoidoscelosidae" genus="Cricoidoscelosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="species" species="aethus">
|
||
<emphasis id="C9A54909FFCCFFBFFE0BFA05A630CD2B" box="[508,609,1523,1549]" italics="true" pageId="11" pageNumber="698">A. aethus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFD5EFA05A757CD2B" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[681,774,1523,1549]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCCFFBFFD5EFA05A757CD2B" box="[681,774,1523,1549]" italics="true" pageId="11" pageNumber="698">P. licenti</emphasis>
|
||
</taxonomicName>
|
||
are, in our opinion extremely common within crayfishes or misinterpreted. For these reasons, we do not find any compelling characters allowing for the distinction of the three species described from the Jehol biota so far.
|
||
</paragraph>
|
||
<caption id="AFAEC593FFCEFFBDFF73F91EA172CC75" ID-DOI="http://doi.org/10.5281/zenodo.10376402" ID-Zenodo-Dep="10376402" httpUri="https://zenodo.org/record/10376402/files/figure.png" pageId="9" pageNumber="696" startId="9.[132,143,1768,1785]" targetBox="[132,1455,215,1732]" targetPageId="9" targetType="figure">
|
||
<paragraph id="FB6E951BFFCEFFBDFF73F91EA172CC75" blockId="9.[132,1457,1768,1875]" pageId="9" pageNumber="696">
|
||
<smallCapsWord id="FD8803C7FFCEFFBDFF73F91EA4CCCDDE" baselines="1781,1781" box="[132,157,1768,1785]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="9" pageNumber="696">FIG</smallCapsWord>
|
||
. 3. — Ischial hooks in
|
||
<taxonomicName id="3CD1EE98FFCEFFBDFEAEF91EA5F1CDDF" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[345,416,1768,1785]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="696" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCEFFBDFEAEF91EA5F1CDDF" box="[345,416,1768,1785]" italics="true" pageId="9" pageNumber="696">P. licenti</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="C9A54909FFCEFFBDFE5DF91EA59DCDDF" bold="true" box="[426,460,1768,1785]" pageId="9" pageNumber="696">A -D</emphasis>
|
||
, on specimen NIGP-126337 (holotype of
|
||
<taxonomicName id="3CD1EE98FFCEFFBDFCD3F91EA7BECDDF" authorityName="Taylor, Schram & Shen" authorityYear="1999" box="[804,1007,1768,1785]" class="Malacostraca" family="Cricoidoscelosidae" genus="Cricoidoscelosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="696" phylum="Arthropoda" rank="species" species="aethus">
|
||
<emphasis id="C9A54909FFCEFFBDFCD3F91EA7BECDDF" box="[804,1007,1768,1785]" italics="true" pageId="9" pageNumber="696">Cricoidoscelosus aethus</emphasis>
|
||
</taxonomicName>
|
||
), ventral view in cross-polarized light (
|
||
<emphasis id="C9A54909FFCEFFBDFADAF91EA16BCDDF" bold="true" box="[1325,1338,1768,1785]" pageId="9" pageNumber="696">A</emphasis>
|
||
) and interpretative line-drawing (
|
||
<emphasis id="C9A54909FFCEFFBDFEDFF908A564CC29" bold="true" box="[296,309,1790,1807]" pageId="9" pageNumber="696">B</emphasis>
|
||
);
|
||
<emphasis id="C9A54909FFCEFFBDFEB2F908A502CC29" bold="true" box="[325,339,1790,1807]" pageId="9" pageNumber="696">C</emphasis>
|
||
,
|
||
<emphasis id="C9A54909FFCEFFBDFEAAF908A53ACC29" bold="true" box="[349,363,1790,1807]" pageId="9" pageNumber="696">D</emphasis>
|
||
, detailed view of the base of P2-P4 (th5-th7) (
|
||
<emphasis id="C9A54909FFCEFFBDFD1BF908A6ABCC29" bold="true" box="[748,762,1790,1807]" pageId="9" pageNumber="696">C</emphasis>
|
||
), and interpretative line drawing (
|
||
<emphasis id="C9A54909FFCEFFBDFBE7F908A04FCC29" bold="true" box="[1040,1054,1790,1807]" pageId="9" pageNumber="696">D</emphasis>
|
||
) – note that
|
||
<emphasis id="C9A54909FFCEFFBDFB70F908A0C4CC29" bold="true" box="[1159,1173,1790,1807]" pageId="9" pageNumber="696">C</emphasis>
|
||
and
|
||
<emphasis id="C9A54909FFCEFFBDFB48F908A09CCC29" bold="true" box="[1215,1229,1790,1807]" pageId="9" pageNumber="696">D</emphasis>
|
||
are aligned horizontally for convenience;
|
||
<emphasis id="C9A54909FFCEFFBDFF00F8E3A552CC00" bold="true" box="[247,259,1813,1830]" pageId="9" pageNumber="696">E</emphasis>
|
||
,
|
||
<emphasis id="C9A54909FFCEFFBDFEFAF8E3A549CC00" bold="true" box="[269,280,1813,1830]" pageId="9" pageNumber="696">F</emphasis>
|
||
, on specimen NIGP-Shen-22,detail of the ischium of P4 (th7). Abbreviations:
|
||
<emphasis id="C9A54909FFCEFFBDFC60F8E3A7FDCC00" bold="true" box="[919,940,1813,1830]" pageId="9" pageNumber="696">a1</emphasis>
|
||
, antennula;
|
||
<emphasis id="C9A54909FFCEFFBDFBE7F8E3A074CC00" bold="true" box="[1040,1061,1813,1830]" pageId="9" pageNumber="696">a2</emphasis>
|
||
, antenna;
|
||
<emphasis id="C9A54909FFCEFFBDFB8BF8E3A0C3CC00" bold="true" box="[1148,1170,1813,1830]" pageId="9" pageNumber="696">ba</emphasis>
|
||
, basipod;
|
||
<emphasis id="C9A54909FFCEFFBDFB1FF8E3A0ACCC00" bold="true" box="[1256,1277,1813,1830]" pageId="9" pageNumber="696">cx</emphasis>
|
||
, coxa;
|
||
<emphasis id="C9A54909FFCEFFBDFACFF8E3A102CC00" bold="true" box="[1336,1363,1813,1830]" pageId="9" pageNumber="696">epi</emphasis>
|
||
, epistome;
|
||
<emphasis id="C9A54909FFCEFFBDFF73F8DAA4C8CC1B" bold="true" box="[132,153,1836,1853]" pageId="9" pageNumber="696">ex</emphasis>
|
||
, uropodal exopod;
|
||
<emphasis id="C9A54909FFCEFFBDFECAF8DAA505CC1B" bold="true" box="[317,340,1836,1853]" pageId="9" pageNumber="696">go</emphasis>
|
||
, male gonopod;
|
||
<emphasis id="C9A54909FFCEFFBDFE16F8DAA5AACC1B" bold="true" box="[481,507,1836,1853]" pageId="9" pageNumber="696">isc</emphasis>
|
||
, ischium;
|
||
<emphasis id="C9A54909FFCEFFBDFDB8F8DAA63ACC1B" bold="true" box="[591,619,1836,1853]" pageId="9" pageNumber="696">md</emphasis>
|
||
, mandible;
|
||
<emphasis id="C9A54909FFCEFFBDFD3AF8DAA6B8CC1B" bold="true" box="[717,745,1836,1853]" pageId="9" pageNumber="696">me</emphasis>
|
||
, merus;
|
||
<emphasis id="C9A54909FFCEFFBDFCC5F8DAA732CC1B" bold="true" box="[818,867,1836,1853]" pageId="9" pageNumber="696">mxp3</emphasis>
|
||
, maxilliped 3 (thoracopod 3);
|
||
<emphasis id="C9A54909FFCEFFBDFBADF8DAA0DECC1B" bold="true" box="[1114,1167,1836,1853]" pageId="9" pageNumber="696">P1-P5</emphasis>
|
||
, pereiopods 1-5 (thoracopod 4-8);
|
||
<emphasis id="C9A54909FFCEFFBDFF73F8B4A4CECC75" bold="true" box="[132,159,1858,1875]" pageId="9" pageNumber="696">pl3</emphasis>
|
||
, pleopod 3;
|
||
<emphasis id="C9A54909FFCEFFBDFEFFF8B4A55ECC75" bold="true" box="[264,271,1858,1875]" pageId="9" pageNumber="696">r</emphasis>
|
||
, rostrum;
|
||
<emphasis id="C9A54909FFCEFFBDFE92F8B4A528CC75" bold="true" box="[357,377,1858,1875]" pageId="9" pageNumber="696">s2</emphasis>
|
||
, pleonite 2;
|
||
<emphasis id="C9A54909FFCEFFBDFE17F8B4A5A4CC75" bold="true" box="[480,501,1858,1875]" pageId="9" pageNumber="696">sc</emphasis>
|
||
, scaphocerite. Photographs: A, D. Audo; C, E, D.-Y. Huang. Scale bars: A, B, 20 mm; C-F, 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="AFAEC593FFCDFFBEFF73F8FCA0BCCC79" ID-DOI="http://doi.org/10.5281/zenodo.10376404" ID-Zenodo-Dep="10376404" httpUri="https://zenodo.org/record/10376404/files/figure.png" pageId="10" pageNumber="697" startId="10.[132,143,1802,1819]" targetBox="[133,1455,215,1767]" targetPageId="10" targetType="figure">
|
||
<paragraph id="FB6E951BFFCDFFBEFF73F8FCA0BCCC79" blockId="10.[132,1456,1802,1887]" pageId="10" pageNumber="697">
|
||
<smallCapsWord id="FD8803C7FFCDFFBEFF73F8FCA4CCCC3C" baselines="1815,1815" box="[132,157,1802,1819]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="10" pageNumber="697">FIG</smallCapsWord>
|
||
. 4. — Sexual dimorphism:
|
||
<emphasis id="C9A54909FFCDFFBEFE8DF8FCA5D6CC3D" bold="true" box="[378,391,1802,1819]" pageId="10" pageNumber="697">A</emphasis>
|
||
,
|
||
<emphasis id="C9A54909FFCDFFBEFE67F8FCA5CCCC3D" bold="true" box="[400,413,1802,1819]" pageId="10" pageNumber="697">B</emphasis>
|
||
,
|
||
<taxonomicName id="3CD1EE98FFCDFFBEFE51F8FCA631CC3D" baseAuthorityName="Pallas" baseAuthorityYear="1972" box="[422,608,1802,1819]" class="Malacostraca" family="Cambaroididae" genus="Cambaroides" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="697" phylum="Arthropoda" rank="species" species="dauricus">
|
||
<emphasis id="C9A54909FFCDFFBEFE51F8FCA631CC3D" box="[422,608,1802,1819]" italics="true" pageId="10" pageNumber="697">Cambaroides dauricus</emphasis>
|
||
</taxonomicName>
|
||
(AS11,extant,Mandchuria) male (
|
||
<emphasis id="C9A54909FFCDFFBEFC8FF8FCA7D4CC3D" bold="true" box="[888,901,1802,1819]" pageId="10" pageNumber="697">A</emphasis>
|
||
) and female (
|
||
<emphasis id="C9A54909FFCDFFBEFC06F8FCA7AFCC3D" bold="true" box="[1009,1022,1802,1819]" pageId="10" pageNumber="697">B</emphasis>
|
||
);
|
||
<emphasis id="C9A54909FFCDFFBEFBFBF8FCA04BCC3D" bold="true" box="[1036,1050,1802,1819]" pageId="10" pageNumber="697">C</emphasis>
|
||
,
|
||
<emphasis id="C9A54909FFCDFFBEFBD4F8FCA060CC3D" bold="true" box="[1059,1073,1802,1819]" pageId="10" pageNumber="697">D</emphasis>
|
||
, possible annulus ventralis in
|
||
<taxonomicName id="3CD1EE98FFCDFFBEFAD2F8FCA50ECC17" authority="(Van Straelen, 1928)" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="697" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCDFFBEFAD2F8FCA4E4CC17" italics="true" pageId="10" pageNumber="697">Palaeocambarus licenti</emphasis>
|
||
(Van Straelen, 1928)
|
||
</taxonomicName>
|
||
specimen NIGP-126342 from the Early Cretaceous of China, photograph (
|
||
<emphasis id="C9A54909FFCDFFBEFC34F8D6A780CC17" bold="true" box="[963,977,1824,1841]" pageId="10" pageNumber="697">C</emphasis>
|
||
) and interpretative line-drawing (
|
||
<emphasis id="C9A54909FFCDFFBEFB17F8D6A0BFCC17" bold="true" box="[1248,1262,1824,1841]" pageId="10" pageNumber="697">D</emphasis>
|
||
);
|
||
<emphasis id="C9A54909FFCDFFBEFB0BF8D6A159CC17" bold="true" box="[1276,1288,1824,1841]" pageId="10" pageNumber="697">E</emphasis>
|
||
, annulus ventralis in
|
||
<taxonomicName id="3CD1EE98FFCDFFBEFF73F8C1A5F9CC6E" authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[132,424,1847,1864]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="697" phylum="Arthropoda" rank="species" species="bartonii">
|
||
<emphasis id="C9A54909FFCDFFBEFF73F8C1A54DCC6E" box="[132,284,1847,1864]" italics="true" pageId="10" pageNumber="697">Cambarus bartonii</emphasis>
|
||
(Fabricius,1798)
|
||
</taxonomicName>
|
||
(AS90,extant, Philadelphia,United States);Abbreviations:
|
||
<emphasis id="C9A54909FFCDFFBEFC66F8C1A7F4CC6E" bold="true" box="[913,933,1847,1864]" pageId="10" pageNumber="697">av</emphasis>
|
||
, annulus ventralis;
|
||
<emphasis id="C9A54909FFCDFFBEFBB4F8C1A033CC6E" bold="true" box="[1091,1122,1847,1864]" pageId="10" pageNumber="697">av?</emphasis>
|
||
, potential annulus ventralis;
|
||
<emphasis id="C9A54909FFCDFFBEFABAF8C1A135CC6E" bold="true" box="[1357,1380,1847,1864]" pageId="10" pageNumber="697">gp</emphasis>
|
||
♀, female gonopore;
|
||
<emphasis id="C9A54909FFCDFFBEFF28F8B8A4A7CC79" bold="true" box="[223,246,1870,1887]" pageId="10" pageNumber="697">gp</emphasis>
|
||
♂, male gonopore;
|
||
<emphasis id="C9A54909FFCDFFBEFE62F8B8A593CC79" bold="true" box="[405,450,1870,1887]" pageId="10" pageNumber="697">pl1-2</emphasis>
|
||
♂, pleopods 1-2 (male gonopods). Photographs: D. Audo. Scale bars: A, B, E, 5 mm; C, D, 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="AFAEC593FFCCFFBFFF73FB33A089CE25" ID-DOI="http://doi.org/10.5281/zenodo.10376406" ID-Zenodo-Dep="10376406" httpUri="https://zenodo.org/record/10376406/files/figure.png" pageId="11" pageNumber="698" startId="11.[132,143,1221,1238]" targetBox="[132,1452,215,1179]" targetPageId="11" targetType="figure">
|
||
<paragraph id="FB6E951BFFCCFFBFFF73FB33A089CE25" blockId="11.[132,1456,1221,1283]" pageId="11" pageNumber="698">
|
||
<smallCapsWord id="FD8803C7FFCCFFBFFF73FB33A4CCCFF3" baselines="1234,1234" box="[132,157,1221,1238]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="11" pageNumber="698">FIG</smallCapsWord>
|
||
. 5. — Neotype of
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFEC2FB33A6F4CFF0" authority="(Van Straelen, 1928)" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[309,677,1221,1238]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCCFFBFFEC2FB33A5A4CFF0" box="[309,501,1221,1238]" italics="true" pageId="11" pageNumber="698">Palaeocambarus licenti</emphasis>
|
||
(Van Straelen, 1928)
|
||
</taxonomicName>
|
||
, specimen NIGP-126342: ventral view in cross-polarized light (
|
||
<emphasis id="C9A54909FFCCFFBFFB5AFB33A0EBCFF0" bold="true" box="[1197,1210,1221,1238]" pageId="11" pageNumber="698">A</emphasis>
|
||
) and interpretative line-drawing (
|
||
<emphasis id="C9A54909FFCCFFBFFF5FFB2AA4E4CFCB" bold="true" box="[168,181,1244,1261]" pageId="11" pageNumber="698">B</emphasis>
|
||
). Abbreviations:
|
||
<emphasis id="C9A54909FFCCFFBFFEB6FB2AA504CFCB" bold="true" box="[321,341,1244,1261]" pageId="11" pageNumber="698">a2</emphasis>
|
||
, antenna;
|
||
<emphasis id="C9A54909FFCCFFBFFE59FB2AA59CCFCB" bold="true" box="[430,461,1244,1261]" pageId="11" pageNumber="698">av?</emphasis>
|
||
, potential annulus ventralis;
|
||
<emphasis id="C9A54909FFCCFFBFFD4DFB2AA69DCFCB" bold="true" box="[698,716,1244,1261]" pageId="11" pageNumber="698">dt</emphasis>
|
||
; digestive tract;
|
||
<emphasis id="C9A54909FFCCFFBFFCA0FB2AA723CFCB" bold="true" box="[855,882,1244,1261]" pageId="11" pageNumber="698">epi</emphasis>
|
||
, epistome;
|
||
<emphasis id="C9A54909FFCCFFBFFC23FB2AA7B8CFCB" bold="true" box="[980,1001,1244,1261]" pageId="11" pageNumber="698">ex</emphasis>
|
||
, uropodal exopod;
|
||
<emphasis id="C9A54909FFCCFFBFFB7BFB2AA0ECCFCB" bold="true" box="[1164,1213,1244,1261]" pageId="11" pageNumber="698">mxp3</emphasis>
|
||
, maxilliped 3 (thoracopod 3);
|
||
<emphasis id="C9A54909FFCCFFBFFF73FB04A4DECE25" bold="true" box="[132,143,1266,1283]" pageId="11" pageNumber="698">o</emphasis>
|
||
, eye;
|
||
<emphasis id="C9A54909FFCCFFBFFF35FB04A4A6CE25" bold="true" box="[194,247,1266,1283]" pageId="11" pageNumber="698">P1-P5</emphasis>
|
||
, pereiopods 1-5 (th4-th8);
|
||
<emphasis id="C9A54909FFCCFFBFFE21FB04A58CCE25" bold="true" box="[470,477,1266,1283]" pageId="11" pageNumber="698">r</emphasis>
|
||
, rostrum;
|
||
<emphasis id="C9A54909FFCCFFBFFDC4FB04A632CE25" bold="true" box="[563,611,1266,1283]" pageId="11" pageNumber="698">s2-s5</emphasis>
|
||
, pleonites 2-5;
|
||
<emphasis id="C9A54909FFCCFFBFFD12FB04A6ABCE25" bold="true" box="[741,762,1266,1283]" pageId="11" pageNumber="698">sc</emphasis>
|
||
, scaphocerite. Photograph: D. Audo. Scale bars: 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FB6E951BFFCCFFBFFF6CF964A170CC4A" blockId="11.[131,777,1363,2028]" lastBlockId="11.[811,1457,1363,2028]" pageId="11" pageNumber="698">
|
||
Nevertheless, due to the large number of available specimens, the relatively large distribution area where they are found, and the often highly endemic nature of many crayfish species (
|
||
<bibRefCitation id="9F40E8EAFFCCFFBFFE74F904A6C5CC2A" author="CRANDALL K. A. & BUHAY J. E." box="[387,660,1778,1804]" pageId="11" pageNumber="698" pagination="295 - 301" refId="ref11032" refString="CRANDALL K. A. & BUHAY J. E. 2008. - Global diversity of crayfish (Astacidae, Cambaridae, and Parastacidae-Decapoda) in freshwater. Hydrobiologia 595: 295 - 301. https: // doi. org / 10.1007 / s 10750 - 007 - 9120 - 3" type="journal article" year="2008">Crandall & Buhay 2008</bibRefCitation>
|
||
), it would not be impossible to have multiple species in the Jehol biota, especially as species may co-occur (
|
||
<bibRefCitation id="9F40E8EAFFCCFFBFFD94F8C7A493CC4A" author="ALDA & KAWAI T." pageId="11" pageNumber="698" pagination="1 - 4" refId="ref9906" refString="ALDA & KAWAI T. 2022. - Two East Asian crayfish species cohabit in single brook in Liaoning Province, China. Crayfish News 44 (3 - 4): 1 - 4" type="journal article" year="2022">Alda & Kawai 2022</bibRefCitation>
|
||
). We therefore performed a linear morphometric analysis based on measurements of as many specimens as possible, including
|
||
<typeStatus id="246A2BB9FFCCFFBFFE91F864A5C6CC8A" box="[358,407,1938,1964]" pageId="11" pageNumber="698">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFDDCF864A56ECCED" authority="Imaizumi, 1938" authorityName=", Imaizumi" authorityYear="1938" class="Malacostraca" family="Astacidae" genus="Astacus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="species" species="spinirostrius">
|
||
<emphasis id="C9A54909FFCCFFBFFDDCF864A758CC8D" box="[555,777,1938,1964]" italics="true" pageId="11" pageNumber="698">Astacus spinirostrius</emphasis>
|
||
<bibRefCitation id="9F40E8EAFFCCFFBFFF73F847A56ECCED" author="IMAIZUMI R." box="[132,319,1969,1995]" pageId="11" pageNumber="698" pagination="173 - 178" refId="ref12046" refString="IMAIZUMI R. 1938. - Fossil crayfishes from Jehol. Science reports of the Tohoku Imperial University 2 nd, series, Geology 19 (2): 173 - 178, pls 22 - 23." type="journal article" year="1938">Imaizumi, 1938</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFE89F847A533CCCD" authority="Taylor, Schram & Shen, 1999" authorityName="Taylor, Schram & Shen" authorityYear="1999" class="Malacostraca" family="Cricoidoscelosidae" genus="Cricoidoscelosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="species" species="aethus">
|
||
<emphasis id="C9A54909FFCCFFBFFE89F847A626CCED" box="[382,631,1969,1995]" italics="true" pageId="11" pageNumber="698">Cricoidoscelosus aethus</emphasis>
|
||
<bibRefCitation id="9F40E8EAFFCCFFBFFD8AF847A533CCCD" author="TAYLOR R. S. & SCHRAM F. R. & SHEN Y. - B." pageId="11" pageNumber="698" pagination="121 - 136" refId="ref13937" refString="TAYLOR R. S., SCHRAM F. R. & SHEN Y. - B. 1999. - A new crayfish family (Decapoda: Astacida) from the Upper Jurassic of China, with a reinterpretation of other Chinese crayfish taxa. Paleontological Research 3 (2): 121 - 136." type="journal article" year="1999">Taylor, Schram & Shen, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
trying to find variations that could help characterize separate species. Results are given in table 2. As shown out of the eight elements studied, only the relative length of the pleon compared to that of uropodal exopod does not have a normal distribution: the distribution is asymmetrical but not bimodal (
|
||
<figureCitation id="63EA899EFFCCFFBFFB17FA25A171CECB" box="[1248,1312,1491,1517]" captionStart="FIG" captionStartId="14.[838,849,2008,2025]" captionTargetBox="[791,1456,1413,2002]" captionTargetId="graphics-15@14.[826,1454,1427,1968]" captionTargetPageId="14" captionText="FIG. 9. — Distribution of pleon length to uropodal exopod length ratios." figureDoi="http://doi.org/10.5281/zenodo.10380797" httpUri="https://zenodo.org/record/10380797/files/figure.png" pageId="11" pageNumber="698">Fig. 9</figureCitation>
|
||
). A bimodal or multimodal distribution could have hinted at the presence of multiple species in our sample. While the present results cannot exclude the possibility of having multiple species, the parameter distribution is herein attributed to intraspecific variation and variations in preservation. As a result, we cannot at present consider there are more than one species of crayfish in Jehol biota. If more than one species is indeed present, the differences between species are likely mostly obfuscated by fossilisation. We thus herein consider
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFBEDF8E4A12FCC0A" authority="Imaizumi, 1938" authorityName=", Imaizumi" authorityYear="1938" box="[1050,1406,1810,1836]" class="Malacostraca" family="Astacidae" genus="Astacus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="species" species="spinirostrius">
|
||
<emphasis id="C9A54909FFCCFFBFFBEDF8E4A0EECC0D" box="[1050,1215,1810,1836]" italics="true" pageId="11" pageNumber="698">A. spinirostrius</emphasis>
|
||
<bibRefCitation id="9F40E8EAFFCCFFBFFB30F8E4A12FCC0A" author="IMAIZUMI R." box="[1223,1406,1810,1836]" pageId="11" pageNumber="698" pagination="173 - 178" refId="ref12046" refString="IMAIZUMI R. 1938. - Fossil crayfishes from Jehol. Science reports of the Tohoku Imperial University 2 nd, series, Geology 19 (2): 173 - 178, pls 22 - 23." type="journal article" year="1938">Imaizumi, 1938</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFCDBF8C4A0B4CC6A" authority="Taylor, Schram & Shen, 1999" authorityName="Taylor, Schram & Shen" authorityYear="1999" box="[812,1253,1841,1868]" class="Malacostraca" family="Cricoidoscelosidae" genus="Cricoidoscelosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="species" species="aethus">
|
||
<emphasis id="C9A54909FFCCFFBFFCDBF8C4A7C0CC6D" box="[812,913,1841,1868]" italics="true" pageId="11" pageNumber="698">C. aethus</emphasis>
|
||
<bibRefCitation id="9F40E8EAFFCCFFBFFC60F8C7A0B4CC6A" author="TAYLOR R. S. & SCHRAM F. R. & SHEN Y. - B." box="[919,1253,1841,1868]" pageId="11" pageNumber="698" pagination="121 - 136" refId="ref13937" refString="TAYLOR R. S., SCHRAM F. R. & SHEN Y. - B. 1999. - A new crayfish family (Decapoda: Astacida) from the Upper Jurassic of China, with a reinterpretation of other Chinese crayfish taxa. Paleontological Research 3 (2): 121 - 136." type="journal article" year="1999">Taylor, Schram & Shen, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
to be more recent synonyms of
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFC31F8A4A14FCC4A" authority="(Van Straelen, 1928)" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[966,1310,1873,1900]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCCFFBFFC31F8A4A075CC4D" box="[966,1060,1873,1899]" italics="true" pageId="11" pageNumber="698">P. licenti</emphasis>
|
||
(Van Straelen, 1928)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FB6E951BFFCCFFB9FCB4F884A501CCCD" blockId="11.[811,1457,1363,2028]" lastBlockId="13.[131,775,1650,2028]" lastPageId="13" lastPageNumber="700" pageId="11" pageNumber="698">
|
||
Due to the aforementioned diagnostic characters,
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFA74F887A7E8CC8D" authorityName="Taylor, Schram & Shen" authorityYear="1999" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A54909FFCCFFBFFA74F887A7E8CC8D" italics="true" pageId="11" pageNumber="698">Palaeocambarus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFC05F867A0D0CC8D" authorityName="Faxon" authorityYear="1884" box="[1010,1153,1937,1963]" class="Malacostraca" family="Cambaroididae" genus="Cambaroides" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A54909FFCCFFBFFC05F867A0D0CC8D" box="[1010,1153,1937,1963]" italics="true" pageId="11" pageNumber="698">Cambaroides</emphasis>
|
||
</taxonomicName>
|
||
are probably closely allied and forming a monophyletic group. A detailed comparison of
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFCBCF827A7AACCCD" authorityName="Taylor, Schram & Shen" authorityYear="1999" box="[843,1019,2001,2027]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A54909FFCCFFBFFCBCF827A7AACCCD" box="[843,1019,2001,2027]" italics="true" pageId="11" pageNumber="698">Palaeocambarus</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="3CD1EE98FFCCFFBFFBD5F827A0E0CCCD" authorityName="Faxon" authorityYear="1884" box="[1058,1201,2001,2027]" class="Malacostraca" family="Cambaroididae" genus="Cambaroides" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="698" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A54909FFCCFFBFFBD5F827A0E0CCCD" box="[1058,1201,2001,2027]" italics="true" pageId="11" pageNumber="698">Cambaroides</emphasis>
|
||
</taxonomicName>
|
||
is complicated due to the flattened preservation of fossil specimens. It however appears that specimens of
|
||
<taxonomicName id="3CD1EE98FFCAFFB9FE43F964A635CD8A" authorityName="Taylor, Schram & Shen" authorityYear="1999" box="[436,612,1682,1708]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="700" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A54909FFCAFFB9FE43F964A635CD8A" box="[436,612,1682,1708]" italics="true" pageId="13" pageNumber="700">Palaeocambarus</emphasis>
|
||
</taxonomicName>
|
||
differ from of all species of
|
||
<taxonomicName id="3CD1EE98FFCAFFB9FED1F944A5E9CDEA" authorityName="Faxon" authorityYear="1884" box="[294,440,1714,1740]" class="Malacostraca" family="Cambaroididae" genus="Cambaroides" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="700" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A54909FFCAFFB9FED1F944A5E9CDEA" box="[294,440,1714,1740]" italics="true" pageId="13" pageNumber="700">Cambaroides</emphasis>
|
||
</taxonomicName>
|
||
(based on data from
|
||
<bibRefCitation id="9F40E8EAFFCAFFB9FD48F944A551CDCA" author="KAWAI T. & MACHINO Y. & KO H. S." pageId="13" pageNumber="700" pagination="191 - 196" refId="ref12408" refString="KAWAI T., MACHINO Y. & KO H. S. 2003. - Reassessment of Cambaroides dauricus and C. schrenckii (Crustacea: Decapoda: Cambaridae). Korean Journal of Biological Sciences 7 (3): 191 - 196. https: // doi. org / 10.1080 / 12265071.2003.9647704" type="journal article" year="2003">
|
||
Kawai
|
||
<emphasis id="C9A54909FFCAFFB9FF73F925A4EDCDCA" box="[132,188,1746,1772]" italics="true" pageId="13" pageNumber="700">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9F40E8EAFFCAFFB9FEE6F924A51DCDCA" author="KAWAI T. & LABAY V. & FILIPOVA L." box="[273,332,1746,1772]" pageId="13" pageNumber="700" pagination="702 - 717" refId="ref12464" refString="KAWAI T., LABAY V. & FILIPOVA L. 2013. - Taxonomic re-examination of Cambaroides (Decapoda: Cambaridae) with a re-description of C. schrenckii from Sakhalin Island Russia and phylogenetic discussion of the Asian cambarids based on morphological characteristics. Journal of Crustacean Biology 22 (5): 702 - 717. https: // doi. org / 10.1163 / 1937240 X- 00002168" type="journal article" year="2013">2013</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9F40E8EAFFCAFFB9FEAAF924A5C9CDCA" author="KAWAI T. & MIN G. - S. & BARABANSHCHIKOV E. & LABAY V. & KO H. S." box="[349,408,1746,1772]" pageId="13" pageNumber="700" pagination="313 - 368" refId="ref12535" refString="KAWAI T., MIN G. - S., BARABANSHCHIKOV E., LABAY V. & KO H. S. 2016. - Asia, in KAWAI T., FAULKES Z. & SCHOLTZ G. (eds), Freshwater Crayfish, a Global Overview. CRC Press, Boca Raton, London, New York: 313 - 368." type="book chapter" year="2016">2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9F40E8EAFFCAFFB9FE5EF924A6DECDCB" author="KAWAI T. & MIN G. - S." box="[425,655,1746,1773]" pageId="13" pageNumber="700" pagination="777 - 793" refId="ref12322" refString="KAWAI T. & MIN G. - S. 2005. - Re-examination of type material of Cambaroides similis (Koelbel, 1892) (Decapoda: Cambaridae) with a lectotype designation, re-description, and evaluation of geographical variation. Proceedings of the Biological Society of Washington 118 (4): 777 - 793. https: // doi. org / 10.2988 / 0006 - 324 X (2005) 118 [777: ROTMOC] 2.0. CO; 2" type="journal article" year="2005">Kawai & Min 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9F40E8EAFFCAFFB9FD68F924A5DACC2A" author="KAWAI T. & BARABANSHCHIKOV E." pageId="13" pageNumber="700" pagination="27 - 48" refId="ref12138" refString="KAWAI T. & BARABANSHCHIKOV E. 2022. - Re-examination of Cambaroides koshewnikowi and C. wladiwostokiensis with Morphological Observations of the Antennules and Gastric Mill of the Cambaroididae. Freshwater Crayfish 27 (1): 27 - 48. https: // doi. org / 10.5869 / fc. 2022. v 27 - 1.27" type="journal article" year="2022">Kawai & Barabanshchikov 2022</bibRefCitation>
|
||
; and observations of
|
||
<taxonomicName id="3CD1EE98FFCAFFB9FD7FF904A756CC2A" baseAuthorityName="Pallas" baseAuthorityYear="1972" box="[648,775,1778,1804]" class="Malacostraca" family="Cambaroididae" genus="Cambaroides" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="700" phylum="Arthropoda" rank="species" species="dauricus">
|
||
<emphasis id="C9A54909FFCAFFB9FD7FF904A756CC2A" box="[648,775,1778,1804]" italics="true" pageId="13" pageNumber="700">C. dauricus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3CD1EE98FFCAFFB9FF43F8E4A569CC0D" authorityName="De Haan" authorityYear="1841" box="[180,312,1810,1836]" class="Malacostraca" family="Cambaroididae" genus="Cambaroides" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="700" phylum="Arthropoda" rank="species" species="japonicus">
|
||
<emphasis id="C9A54909FFCAFFB9FF43F8E4A569CC0D" box="[180,312,1810,1836]" italics="true" pageId="13" pageNumber="700">C. japonicus</emphasis>
|
||
</taxonomicName>
|
||
from MNHN-IU collections) by a longer acumen of the rostrum, larger spines on postorbital carina and on median line in the cephalic area. For these reasons, and since no detailed phylogeny of the species of
|
||
<taxonomicName id="3CD1EE98FFCAFFB9FF70F867A547CC8D" authorityName="Faxon" authorityYear="1884" box="[135,278,1937,1963]" class="Malacostraca" family="Cambaroididae" genus="Cambaroides" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="700" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9A54909FFCAFFB9FF70F867A547CC8D" box="[135,278,1937,1963]" italics="true" pageId="13" pageNumber="700">Cambaroides</emphasis>
|
||
</taxonomicName>
|
||
including
|
||
<taxonomicName id="3CD1EE98FFCAFFB9FE62F867A6C5CC8D" baseAuthorityName="Van Straelen" baseAuthorityYear="1928" box="[405,660,1937,1963]" class="Malacostraca" family="Cambaridae" genus="Palaeocambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="700" phylum="Arthropoda" rank="species" species="licenti">
|
||
<emphasis id="C9A54909FFCAFFB9FE62F867A6C5CC8D" box="[405,660,1937,1963]" italics="true" pageId="13" pageNumber="700">Palaeocambarus licenti</emphasis>
|
||
</taxonomicName>
|
||
are available, we do not consider the two genera as synonym at present (
|
||
<tableCitation id="B653A0A0FFCAFFB9FF1FF827A56ECCCD" box="[232,319,2001,2027]" captionStart="TABLE" captionStartId="16.[433,444,910,927]" captionTargetBox="[134,1277,985,1235]" captionTargetId="figure-435@16.[156,1432,209,817]" captionTargetPageId="16" captionText="TABLE 1. — Results of linear morphometric analyses on P. licenti (Van Straelen, 1928b)." httpUri="http://table.plazi.org/id/AFAEC593FFD7FFA4FE46FC78A0D5C8B9" pageId="13" pageNumber="700" tableUuid="AFAEC593FFD7FFA4FE46FC78A0D5C8B9">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |