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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.4039.2.3" ID-GBIF-Dataset="e8810e56-3876-475f-9354-d78b6eef32de" ID-ISSN="1175-5326" ID-Zenodo-Dep="236226" ID-ZooBank="7DD7CB5E-E4E5-4424-A3CE-0D822B440512" checkinTime="1461331489926" checkinUser="plazi" docAuthor="Lukić, Marko, Porco, David, Bedos, Anne &amp; Deharveng, Louis" docDate="2015" docId="504DC64DFFEC9413FF25FE84FB65FB9A" docLanguage="en" docName="zt04039p275.pdf" docOrigin="Zootaxa 4039 (2)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@4e3283f6" docStyleName="zootaxa.2013.journal_article" docTitle="Heteromurus (Verhoeffiella) absoloni Kseneman 1938" docType="treatment" docVersion="8" lastPageNumber="266" masterDocId="AC74BE35FFE99402FFB2FFACFFF9FFE7" masterDocTitle="The puzzling distribution of Heteromurus (Verhoeffiella) absoloni Kseneman, 1938 (Collembola: Entomobryidae: Heteromurinae) resolved: detailed redescription of the nominal species and description of a new species from Catalonia (Spain)" masterLastPageNumber="275" masterPageNumber="249" pageNumber="254" updateTime="1635476460527" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The puzzling distribution of Heteromurus (Verhoeffiella) absoloni Kseneman, 1938 (Collembola: Entomobryidae: Heteromurinae) resolved: detailed redescription of the nominal species and description of a new species from Catalonia (Spain)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lukić, Marko</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Porco, David</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bedos, Anne</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deharveng, Louis</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>4039</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>249</mods:start>
<mods:end>275</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.4039.2.3</mods:identifier>
<mods:identifier type="GBIF-Dataset">e8810e56-3876-475f-9354-d78b6eef32de</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">236226</mods:identifier>
<mods:identifier type="ZooBank">7DD7CB5E-E4E5-4424-A3CE-0D822B440512</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5611549" ID-Zenodo-Dep="5611549" LSID="urn:lsid:plazi:treatment:504DC64DFFEC9413FF25FE84FB65FB9A" httpUri="http://treatment.plazi.org/id/504DC64DFFEC9413FF25FE84FB65FB9A" lastPageId="17" lastPageNumber="266" pageId="5" pageNumber="254">
<subSubSection pageId="5" pageNumber="254" type="description">
<paragraph blockId="5.[151,1031,295,322]" box="[151,1031,295,322]" pageId="5" pageNumber="254">
<heading bold="true" box="[151,1031,295,322]" fontSize="11" level="1" pageId="5" pageNumber="254" reason="1">
<emphasis bold="true" box="[151,1031,295,322]" pageId="5" pageNumber="254">
Redescription of
<taxonomicName authority="Kseneman, 1938" authorityName="Kseneman" authorityYear="1938" box="[364,1031,295,322]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis bold="true" box="[364,526,295,321]" italics="true" pageId="5" pageNumber="254">Heteromurus</emphasis>
(
<emphasis bold="true" box="[542,698,295,321]" italics="true" pageId="5" pageNumber="254">Verhoeffiella</emphasis>
)
<emphasis bold="true" box="[715,818,295,321]" italics="true" pageId="5" pageNumber="254">absoloni</emphasis>
<bibRefCitation author="Kseneman" box="[825,1031,296,322]" pageId="5" pageNumber="254" refString="Kseneman, M. (1938) Vorlaufige Mitteilung uber eine neue Collembolenart aus den Hohlen des Dinarischen Karstes und uber die Gattung Typhlopodura Absolon. Folia entomologica, 1, 69 - 70." type="journal article" year="1938">
Kseneman,
<number box="[972,1031,296,322]" pageId="5" pageNumber="254" value="1938.0">1938</number>
</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[151,1437,366,1111]" pageId="5" pageNumber="254">
<emphasis bold="true" box="[151,430,366,391]" pageId="5" pageNumber="254">
Designation of
<typeStatus box="[330,425,366,391]" pageId="5" pageNumber="254">neotype</typeStatus>
.
</emphasis>
The
<typeStatus box="[488,536,367,391]" pageId="5" pageNumber="254">type</typeStatus>
material of
<taxonomicName box="[673,1092,366,391]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[673,820,367,390]" italics="true" pageId="5" pageNumber="254">Heteromurus</emphasis>
(
<emphasis box="[837,979,366,391]" italics="true" pageId="5" pageNumber="254">Verhoeffiella</emphasis>
)
<emphasis box="[996,1092,366,391]" italics="true" pageId="5" pageNumber="254">absoloni</emphasis>
</taxonomicName>
has not been retrieved in spite of thorough investigations in regional Museums and in
<collectingCountry box="[801,976,402,427]" name="Czech Republic" pageId="5" pageNumber="254">Czech republic</collectingCountry>
, including: Natural History Museum of
<collectingCountry box="[151,291,439,463]" name="Serbia" pageId="5" pageNumber="254">Montenegro</collectingCountry>
, Podgorica, National Museum of
<collectingCountry box="[708,1002,438,463]" name="Bosnia and Herzegovina" pageId="5" pageNumber="254">Bosnia and Herzegovina</collectingCountry>
, Sarajevo, Croatian Natural History Museum, Zagreb, The Moravian Museum, Brno, Mendel University in Brno - Department of Zoology, Fisheries, Hydrobiology and Apiculture (formally Zoologicky ustav vysoke školy zemedelske v Brne where Kseneman was working, pers. com. Jan Bezdek). It was also not possible to locate Ksenemans collection (Rusek and Weiner, pers. com.).
<typeStatus box="[232,287,583,607]" pageId="5" pageNumber="254">Type</typeStatus>
material can be therefore considered as lost. We propose here as
<typeStatus box="[1046,1135,583,607]" pageId="5" pageNumber="254">neotype</typeStatus>
a specimen from the
<typeStatus box="[1388,1436,583,607]" pageId="5" pageNumber="254">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="254" type="materials_examined">
<paragraph blockId="5.[151,1437,366,1111]" pageId="5" pageNumber="254">
<emphasis bold="true" box="[199,446,654,679]" pageId="5" pageNumber="254">Material examined.</emphasis>
<collectingCountry box="[466,606,655,679]" name="Serbia" pageId="5" pageNumber="254">Montenegro</collectingCountry>
, Boka kotorska, Risan, Golodražnica (cave) (coordinates WGS
<date pageId="5" pageNumber="254" value="1984-10">
<number box="[1409,1436,655,679]" pageId="5" pageNumber="254" value="84.0">84</number>
X
</date>
=
<number box="[187,297,690,715]" pageId="5" pageNumber="254" value="18.69962">18.69962</number>
°,
<collectingCountry box="[318,338,690,715]" name="Italy" pageId="5" pageNumber="254">Y</collectingCountry>
=
<number box="[353,464,690,715]" pageId="5" pageNumber="254" value="42.50282">42.50282</number>
°, alt.
<specimenCount box="[518,600,690,715]" pageId="5" pageNumber="254" type="generic">
<quantity box="[518,596,690,715]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="254" unit="m" value="110.0">110 m</quantity>
.
</specimenCount>
).
<typeStatus box="[621,716,691,715]" pageId="5" pageNumber="254">Neotype</typeStatus>
adult female on slide,
<date box="[975,1105,690,715]" pageId="5" pageNumber="254" value="2014-12-04">
04.xii.
<number box="[1044,1105,690,715]" pageId="5" pageNumber="254" value="2014.0">2014</number>
</date>
, M. Lukić leg.(CLL
<number box="[1341,1402,690,715]" pageId="5" pageNumber="254" value="4557.0">4557</number>
);
<number box="[1422,1436,690,715]" pageId="5" pageNumber="254" value="5.0">5</number>
ex. on slides and
<number box="[349,377,727,751]" pageId="5" pageNumber="254" value="19.0">19</number>
ex. in
<number box="[456,485,726,751]" pageId="5" pageNumber="254" value="96.0">96</number>
% alcohol, same data (CLL
<number box="[803,864,726,751]" pageId="5" pageNumber="254" value="4555.0">4555</number>
). Three ex. directly mounted on slides and one ex. mounted on slide after
<collectionCode LSID="urn:lsid:biocol.org:col:15536" box="[426,487,763,787]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="5" pageNumber="254">DNA</collectionCode>
extraction (
<collectionCode box="[629,707,763,787]" pageId="5" pageNumber="254">BOLD</collectionCode>
Sample ID:
<number box="[847,905,763,787]" pageId="5" pageNumber="254" value="4028.0">4028</number>
C01_MNECLL
<number box="[1086,1143,763,787]" pageId="5" pageNumber="254" value="2278.0">2278</number>
),
<number box="[1167,1195,763,787]" pageId="5" pageNumber="254" value="24.0">24</number>
ex. in
<number box="[1284,1313,762,787]" pageId="5" pageNumber="254" value="96.0">96</number>
% alcohol,
<date box="[151,271,798,823]" pageId="5" pageNumber="254" value="2010-04-24">
<number box="[151,181,798,823]" pageId="5" pageNumber="254" value="24.0">24</number>
.iv.
<number box="[210,271,798,823]" pageId="5" pageNumber="254" value="2010.0">2010</number>
</date>
, J. Bedek leg.(CLL
<number box="[497,557,799,823]" pageId="5" pageNumber="254" value="2278.0">2278</number>
);
<number box="[578,592,799,823]" pageId="5" pageNumber="254" value="3.0">3</number>
ex. on slides,
<number box="[752,766,799,823]" pageId="5" pageNumber="254" value="8.0">8</number>
ex. in
<number box="[843,872,798,823]" pageId="5" pageNumber="254" value="96.0">96</number>
% alcohol,
<date box="[998,1119,798,823]" pageId="5" pageNumber="254" value="2010-04-24">
<number box="[998,1028,798,823]" pageId="5" pageNumber="254" value="24.0">24</number>
.iv.
<number box="[1058,1119,798,823]" pageId="5" pageNumber="254" value="2010.0">2010</number>
</date>
, A. Kirin leg.(CLL
<number box="[1342,1403,799,823]" pageId="5" pageNumber="254" value="2279.0">2279</number>
);
<number box="[1422,1436,799,823]" pageId="5" pageNumber="254" value="2.0">2</number>
ex. on slides,
<number box="[315,329,835,859]" pageId="5" pageNumber="254" value="4.0">4</number>
ex. in
<number box="[416,445,834,859]" pageId="5" pageNumber="254" value="96.0">96</number>
% alcohol,
<date box="[579,699,834,859]" pageId="5" pageNumber="254" value="2010-04-24">
<number box="[579,609,834,859]" pageId="5" pageNumber="254" value="24.0">24</number>
.iv.
<number box="[638,699,834,859]" pageId="5" pageNumber="254" value="2010.0">2010</number>
</date>
, A. Komerički leg.(CLL
<number box="[997,1058,835,859]" pageId="5" pageNumber="254" value="2280.0">2280</number>
);
<number box="[1080,1094,835,859]" pageId="5" pageNumber="254" value="3.0">3</number>
ex. on slides,
<number box="[1270,1298,835,859]" pageId="5" pageNumber="254" value="34.0">34</number>
ex. in
<number box="[1385,1414,834,859]" pageId="5" pageNumber="254" value="96.0">96</number>
% alcohol,
<date box="[249,373,870,895]" pageId="5" pageNumber="254" value="2012-03-31">
<number box="[249,277,870,895]" pageId="5" pageNumber="254" value="31.0">31</number>
.iii.
<number box="[310,373,870,895]" pageId="5" pageNumber="254" value="2012.0">2012</number>
</date>
, J. Bedek leg.(CLL
<number box="[607,668,871,895]" pageId="5" pageNumber="254" value="4000.0">4000</number>
);
<number box="[689,703,871,895]" pageId="5" pageNumber="254" value="2.0">2</number>
ex. on slides,
<number box="[872,900,871,894]" pageId="5" pageNumber="254" value="77.0">77</number>
ex. in
<number box="[983,1012,870,895]" pageId="5" pageNumber="254" value="96.0">96</number>
% alcohol,
<date box="[1142,1266,870,895]" pageId="5" pageNumber="254" value="2012-03-31">
<number box="[1142,1170,870,895]" pageId="5" pageNumber="254" value="31.0">31</number>
.iii.
<number box="[1203,1266,870,895]" pageId="5" pageNumber="254" value="2012.0">2012</number>
</date>
, M. Lukić leg. (CLL
<number box="[220,281,907,931]" pageId="5" pageNumber="254" value="4001.0">4001</number>
);
<number box="[300,328,907,931]" pageId="5" pageNumber="254" value="17.0">17</number>
ex. in
<number box="[404,433,906,931]" pageId="5" pageNumber="254" value="96.0">96</number>
% alcohol,
<date box="[559,683,906,931]" pageId="5" pageNumber="254" value="2012-03-31">
<number box="[559,587,906,931]" pageId="5" pageNumber="254" value="31.0">31</number>
.iii.
<number box="[620,683,906,931]" pageId="5" pageNumber="254" value="2012.0">2012</number>
</date>
, A. Komerički leg. (CLL
<number box="[973,1034,906,931]" pageId="5" pageNumber="254" value="4185.0">4185</number>
).
</paragraph>
<paragraph blockId="5.[151,1437,366,1111]" pageId="5" pageNumber="254">
<emphasis bold="true" box="[199,440,942,967]" pageId="5" pageNumber="254">Material deposition.</emphasis>
All material in
<number box="[618,647,942,967]" pageId="5" pageNumber="254" value="96.0">96</number>
% alcohol and following ex. on slides, including
<typeStatus box="[1197,1289,943,967]" pageId="5" pageNumber="254">neotype</typeStatus>
, deposited in collection of the Croatian Biospeleological Society, Zagreb,
<collectingCountry box="[843,930,978,1003]" name="Croatia" pageId="5" pageNumber="254">Croatia</collectingCountry>
:
<number box="[942,956,979,1003]" pageId="5" pageNumber="254" value="2.0">2</number>
ex. (CLL
<number box="[1080,1140,979,1003]" pageId="5" pageNumber="254" value="2278.0">2278</number>
),
<number box="[1161,1175,979,1003]" pageId="5" pageNumber="254" value="3.0">3</number>
ex. (CLL
<number box="[1298,1358,979,1003]" pageId="5" pageNumber="254" value="2279.0">2279</number>
),
<number box="[1379,1393,979,1003]" pageId="5" pageNumber="254" value="1.0">1</number>
ex. (CLL
<number box="[223,283,1015,1039]" pageId="5" pageNumber="254" value="2280.0">2280</number>
),
<number box="[305,319,1015,1039]" pageId="5" pageNumber="254" value="1.0">1</number>
ex. (CLL
<number box="[445,505,1015,1039]" pageId="5" pageNumber="254" value="4000.0">4000</number>
),
<number box="[528,542,1015,1039]" pageId="5" pageNumber="254" value="1.0">1</number>
ex. (CLL
<number box="[668,728,1015,1039]" pageId="5" pageNumber="254" value="4001.0">4001</number>
),
<number box="[750,764,1014,1039]" pageId="5" pageNumber="254" value="6.0">6</number>
ex. (CLL
<number box="[890,951,1014,1039]" pageId="5" pageNumber="254" value="4555.0">4555</number>
). Following ex. on slides deposited in the collection of the Muséum National dHistoire Naturelle de Paris:
<number box="[888,902,1051,1075]" pageId="5" pageNumber="254" value="1.0">1</number>
ex. (CLL
<number box="[1020,1080,1051,1075]" pageId="5" pageNumber="254" value="2278.0">2278</number>
),
<number box="[1099,1113,1051,1075]" pageId="5" pageNumber="254" value="1.0">1</number>
ex. (CLL
<number box="[1232,1292,1051,1075]" pageId="5" pageNumber="254" value="2280.0">2280</number>
),
<number box="[1311,1325,1051,1075]" pageId="5" pageNumber="254" value="2.0">2</number>
ex. (CLL
<number box="[151,211,1087,1111]" pageId="5" pageNumber="254" value="4000.0">4000</number>
),
<number box="[230,244,1087,1111]" pageId="5" pageNumber="254" value="1.0">1</number>
ex. (CLL
<number box="[361,422,1087,1111]" pageId="5" pageNumber="254" value="4001.0">4001</number>
).
</paragraph>
</subSubSection>
<caption box="[151,911,1799,1821]" httpUri="https://zenodo.org/record/236227/files/figure.png" pageId="5" pageNumber="254" targetBox="[151,1436,1164,1777]" targetPageId="5">
<paragraph blockId="5.[151,911,1799,1821]" box="[151,911,1799,1821]" pageId="5" pageNumber="254">
<emphasis bold="true" box="[151,275,1799,1821]" pageId="5" pageNumber="254">FIGURE 1.</emphasis>
<taxonomicName box="[281,659,1799,1821]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[281,414,1800,1821]" italics="true" pageId="5" pageNumber="254">Heteromurus</emphasis>
(
<emphasis box="[428,557,1799,1821]" italics="true" pageId="5" pageNumber="254">Verhoeffiella</emphasis>
)
<emphasis box="[572,659,1799,1821]" italics="true" pageId="5" pageNumber="254">absoloni</emphasis>
</taxonomicName>
: habitus (scale 500 µm).
</paragraph>
</caption>
<subSubSection lastPageId="6" lastPageNumber="255" pageId="5" pageNumber="254" type="diagnosis">
<paragraph blockId="5.[151,1436,1860,2018]" pageId="5" pageNumber="254">
<emphasis bold="true" box="[199,321,1860,1885]" pageId="5" pageNumber="254">Diagnosis.</emphasis>
Color pale orange, pigment as orange dots. Scales present on dorsal and ventral side of head and body, on Ant. Ib and on Ant. II, on legs, ventrally and dorsally on manubrium, ventrally on dens. Antennae
<number pageId="5" pageNumber="254" value="0.6" valueMax="0.7" valueMin="0.5">0.5 to 0.7</number>
as long as (head + body) length, Ant. III and IV annulated, often fused. Ant. segment ratio as Ia:Ib:II:III:IV =
<number box="[151,208,1960,1985]" pageId="5" pageNumber="254" value="0.15">0.15</number>
:
<number box="[212,268,1960,1985]" pageId="5" pageNumber="254" value="0.61">0.61</number>
:
<number box="[271,281,1960,1985]" pageId="5" pageNumber="254" value="1.0">1</number>
:
<number box="[285,322,1960,1985]" pageId="5" pageNumber="254" value="1.5">1.5</number>
:2.0. Tita of leg III longer than those of legs I and II; length
<number box="[1011,1063,1961,1985]" pageId="5" pageNumber="254" value="0.4">0.40</number>
,
<number box="[1075,1127,1961,1985]" pageId="5" pageNumber="254" value="0.49">0.49</number>
,
<quantity box="[1140,1241,1960,1985]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="5" pageNumber="254" unit="mm" value="0.58">0.58 mm</quantity>
for Tita I, II, III. Labial triangle with
<number box="[379,435,1994,2018]" pageId="5" pageNumber="254" value="10.5" valueMax="12.0" valueMin="9.0">912</number>
smooth chaetae, and no scales. Dorsal macrochaetotaxy as (
<number box="[1113,1126,1994,2018]" pageId="5" pageNumber="254" value="3.0">3</number>
+
<number box="[1143,1158,1994,2018]" pageId="5" pageNumber="254" value="4.0">4</number>
),(
<number box="[1179,1192,1994,2018]" pageId="5" pageNumber="254" value="3.0">3</number>
+
<number box="[1209,1222,1994,2018]" pageId="5" pageNumber="254" value="3.0">3</number>
)/0,2,3,(
<number box="[1316,1329,1994,2018]" pageId="5" pageNumber="254" value="3.0">3</number>
+0+
<number box="[1377,1392,1994,2018]" pageId="5" pageNumber="254" value="4.0">4</number>
+
<number box="[1409,1424,1994,2018]" pageId="5" pageNumber="254" value="4.0">4</number>
).
</paragraph>
<paragraph blockId="6.[151,1437,151,810]" pageId="6" pageNumber="255">
S-chaetotaxy: ms formula
<number box="[446,455,152,176]" pageId="6" pageNumber="255" value="1.0">1</number>
,?
<number box="[471,480,152,176]" pageId="6" pageNumber="255" value="1.0">1</number>
/1,0,1,0,0; sens formula, 2,2/1,3,3,3,3, with S-like chaetae at least
<number box="[1229,1257,152,176]" pageId="6" pageNumber="255" value="10.0">10</number>
+
<number box="[1273,1301,152,176]" pageId="6" pageNumber="255" value="10.0">10</number>
on Abd. IV. Unguis thin and elongated, with one pair of inner basal teeth at
<number box="[871,900,185,210]" pageId="6" pageNumber="255" value="25.0">25</number>
% from its base and one unpaired inner tooth at
<number box="[151,180,218,243]" pageId="6" pageNumber="255" value="50.0">50</number>
% from its base; unguiculus pointed,
<number box="[623,652,218,243]" pageId="6" pageNumber="255" value="75.0">75</number>
% as long as inner edge of unguis; tenent hair acuminate, smooth. Manubrium with about
<number box="[418,446,252,277]" pageId="6" pageNumber="255" value="15.0">15</number>
smooth pointed mac dorsally, unpaired ventro-basal smooth mac and
<number box="[1237,1251,252,276]" pageId="6" pageNumber="255" value="2.0">2</number>
+
<number box="[1267,1281,252,276]" pageId="6" pageNumber="255" value="2.0">2</number>
ventro-apical serrated mes; dens with one short internal basal spine, mucro bidentate with one basal spine.
</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="266" pageId="6" pageNumber="255" type="description">
<paragraph blockId="6.[151,1437,151,810]" pageId="6" pageNumber="255">
<emphasis bold="true" box="[199,372,318,343]" pageId="6" pageNumber="255">Redescription.</emphasis>
Head and body length combined without antennae:
<quantity box="[961,1119,319,343]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="6" pageNumber="255" unit="mm" value="2.95" valueMax="3.2" valueMin="2.7">2.7 to 3.2 mm</quantity>
(N=
<number box="[1176,1189,319,343]" pageId="6" pageNumber="255" value="8.0">8</number>
). Habitus as in
<figureCitation box="[1364,1429,318,343]" captionStart="FIGURE 1" captionStartId="5.[151,250,1799,1821]" captionTargetBox="[151,1436,1164,1777]" captionTargetId="figure@5.[151,1436,1164,1778]" captionTargetPageId="5" captionText="FIGURE 1. Heteromurus (Verhoeffiella) absoloni: habitus (scale 500 µm)." httpUri="https://zenodo.org/record/236227/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[1415,1429,319,343]" pageId="6" pageNumber="255" value="1.0">1</number>
</figureCitation>
. Abd. IV
<number box="[256,354,351,376]" pageId="6" pageNumber="255" value="1.875" valueMax="2.0" valueMin="1.75">1.75 2</number>
times as long as Abd. III. Color pale orange, pigment scattered as orange dots ventrally and dorsally on body, manubrium, coxa, paler on trochanter and femur. Intersegment area, mouth parts, antennae, other leg segments, and dens unpigmented. Body dorsally and laterally with white regular patches on an orange background (
<figureCitation box="[300,366,452,477]" captionStart="FIGURE 1" captionStartId="5.[151,250,1799,1821]" captionTargetBox="[151,1436,1164,1777]" captionTargetId="figure@5.[151,1436,1164,1778]" captionTargetPageId="5" captionText="FIGURE 1. Heteromurus (Verhoeffiella) absoloni: habitus (scale 500 µm)." httpUri="https://zenodo.org/record/236227/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[353,366,452,476]" pageId="6" pageNumber="255" value="1.0">1</number>
</figureCitation>
). Eyes absent, no pigment patch at their place. Body, mouthparts and appendages with a large diversity of phaneres. Most can be assigned to the traditional four
<typeStatus box="[915,974,486,510]" pageId="6" pageNumber="255">types</typeStatus>
of chaetae: ordinary chaetae, S-chaetae, trichobothria and scales; several other kinds of specialized chaetae occur on mouth and antennae. Pseudopores of same size as mac socket, observed on antennae, tergites, legs and manubrium.
</paragraph>
<paragraph blockId="6.[151,1437,151,810]" pageId="6" pageNumber="255">
<emphasis bold="true" box="[199,314,585,610]" pageId="6" pageNumber="255">Antennae</emphasis>
(
<figureCitation box="[329,446,585,610]" captionStart-0="FIGURES 2 8" captionStart-1="FIGURES 9 14" captionStartId-0="7.[151,264,1704,1727]" captionStartId-1="8.[151,264,1597,1620]" captionTargetBox-0="[235,1327,204,1639]" captionTargetBox-1="[169,1417,186,1574]" captionTargetId-0="figure@7.[223,1332,193,1651]" captionTargetId-1="figure@8.[151,1436,171,1589]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." captionText-1="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri-0="https://zenodo.org/record/236228/files/figure.png" httpUri-1="https://zenodo.org/record/236229/files/figure.png" pageId="6" pageNumber="255">
Figs
<number box="[386,446,585,609]" pageId="6" pageNumber="255" value="8.0" valueMax="14.0" valueMin="2.0">214</number>
</figureCitation>
). Antennae
<number box="[579,713,585,610]" pageId="6" pageNumber="255" value="0.6100000000000001" valueMax="0.68" valueMin="0.54">0.54 to 0.68</number>
as long as (head + body) length (N=
<number box="[1129,1142,585,609]" pageId="6" pageNumber="255" value="8.0">8</number>
). Antennae
<number box="[1276,1290,585,609]" pageId="6" pageNumber="255" value="3.0">3</number>
times longer than head diagonal. Ant. segment ratio as Ia:Ib:II:III:IV =
<number box="[808,864,618,643]" pageId="6" pageNumber="255" value="0.15">0.15</number>
:
<number box="[869,924,618,643]" pageId="6" pageNumber="255" value="0.61">0.61</number>
:
<number box="[927,937,618,643]" pageId="6" pageNumber="255" value="1.0">1</number>
:
<number box="[941,978,618,643]" pageId="6" pageNumber="255" value="1.5">1.5</number>
:2.0 (mean values, N=
<number box="[1233,1247,619,643]" pageId="6" pageNumber="255" value="9.0">9</number>
). Ant. III and IV annulated, except the proximal part of Ant. III and the distal part of Ant. IV. Antennae often asymmetrical, because of the fusion of Ant. III and IV. Pseudopores present ventro-apically on Ant. Ib (
<number box="[1068,1114,685,710]" pageId="6" pageNumber="255" value="3.5" valueMax="5.0" valueMin="2.0">25</number>
), Ant. II (
<number box="[1228,1285,686,710]" pageId="6" pageNumber="255" value="11.0" valueMax="14.0" valueMin="8.0">814</number>
) and Ant. III (
<number box="[160,175,718,743]" pageId="6" pageNumber="255" value="5.0">5</number>
), located in the intersegment just above sensorial areas.
</paragraph>
<paragraph blockId="6.[151,1437,151,810]" pageId="6" pageNumber="255">
Antennal phaneres very diversified (ordinary chaetae, S-chaetae, scales and papillae). The following categories have been recognized (
<figureCitation box="[409,478,785,810]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[461,478,786,810]" pageId="6" pageNumber="255" value="2.0">2</number>
</figureCitation>
):
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,1128,852,877]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,853,877]" pageId="6" pageNumber="255">Type</typeStatus>
a—serrated mac, dorsally on Ant. Ib and II, shorter than those of head and body.
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,861,885,910]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,886,910]" pageId="6" pageNumber="255">Type</typeStatus>
b—thick serrated mes, on all antennal segments (
<figureCitation box="[779,847,885,910]" captionStart="FIGURES 9 14" captionStartId="8.[151,264,1597,1620]" captionTargetBox="[169,1417,186,1574]" captionTargetId="figure@8.[151,1436,171,1589]" captionTargetPageId="8" captionText="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[831,847,886,910]" pageId="6" pageNumber="255" value="9.0">9</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,1051,918,943]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,919,943]" pageId="6" pageNumber="255">Type</typeStatus>
c—thin serrated mes, less ciliated than
<typeStatus box="[667,715,919,943]" pageId="6" pageNumber="255">type</typeStatus>
b, on Ant. III and IV (
<figureCitation box="[968,1038,918,943]" captionStart="FIGURES 9 14" captionStartId="8.[151,264,1597,1620]" captionTargetBox="[169,1417,186,1574]" captionTargetId="figure@8.[151,1436,171,1589]" captionTargetPageId="8" captionText="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[1021,1038,919,943]" pageId="6" pageNumber="255" value="9.0">9</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,812,951,976]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,952,976]" pageId="6" pageNumber="255">Type</typeStatus>
d—pointed smooth mac, ventrally on Ant. Ib and II.
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,844,985,1010]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,986,1010]" pageId="6" pageNumber="255">Type</typeStatus>
e—pointed smooth mes, ventrally on Ant. Ia, Ib and II.
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,1019,1043]" pageId="6" pageNumber="255">Type</typeStatus>
f—acuminate smooth mes, thin, basally thinner than e-type mes, straight and inserted perpendicular to integument, on all antennal segments but only ventrally on Ant. Ia, Ib (
<figureCitation box="[992,1060,1052,1077]" captionStart="FIGURES 9 14" captionStartId="8.[151,264,1597,1620]" captionTargetBox="[169,1417,186,1574]" captionTargetId="figure@8.[151,1436,171,1589]" captionTargetPageId="8" captionText="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[1044,1060,1052,1076]" pageId="6" pageNumber="255" value="9.0">9</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,1359,1085,1110]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,1086,1110]" pageId="6" pageNumber="255">Type</typeStatus>
g—pointed smooth mic similar in shape to e-type mes but shorter, basally on Ant. Ia and II (
<figureCitation box="[1262,1346,1085,1110]" captionStart="FIGURES 9 14" captionStartId="8.[151,264,1597,1620]" captionTargetBox="[169,1417,186,1574]" captionTargetId="figure@8.[151,1436,171,1589]" captionTargetPageId="8" captionText="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[1315,1346,1086,1110]" pageId="6" pageNumber="255" value="10.0">10</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,1385,1118,1143]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,1119,1143]" pageId="6" pageNumber="255">Type</typeStatus>
h—swollen S-chaetae (
<figureCitation box="[488,569,1118,1143]" captionStart="FIGURES 9 14" captionStartId="8.[151,264,1597,1620]" captionTargetBox="[169,1417,186,1574]" captionTargetId="figure@8.[151,1436,171,1589]" captionTargetPageId="8" captionText="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[540,569,1119,1143]" pageId="6" pageNumber="255" value="11.0">11</number>
</figureCitation>
), dorso-externo-apically on Ant. Ib (
<number box="[981,997,1119,1143]" pageId="6" pageNumber="255" value="2.0">2</number>
), II (
<number box="[1053,1097,1118,1143]" pageId="6" pageNumber="255" value="5.0" valueMax="6.0" valueMin="4.0">46</number>
) and III (
<number box="[1203,1218,1119,1143]" pageId="6" pageNumber="255" value="2.0">2</number>
) (
<figureCitation box="[1242,1372,1118,1143]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="6" pageNumber="255">
Figs 4, 5,
<number box="[1355,1372,1118,1143]" pageId="6" pageNumber="255" value="6.0">6</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,1152,1176]" pageId="6" pageNumber="255">Type</typeStatus>
i—thick subcylindrical S-chaetae, on all antennal segments (
<figureCitation box="[936,1024,1151,1176]" captionStart="FIGURES 9 14" captionStartId="8.[151,264,1597,1620]" captionTargetBox="[169,1417,186,1574]" captionTargetId="figure@8.[151,1436,171,1589]" captionTargetPageId="8" captionText="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[992,1024,1152,1176]" pageId="6" pageNumber="255" value="12.0">12</number>
</figureCitation>
):
<number box="[1048,1089,1152,1176]" pageId="6" pageNumber="255" value="3.5" valueMax="4.0" valueMin="3.0">34</number>
ventrally on Ant. Ia (
<figureCitation box="[1350,1421,1151,1176]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[1406,1421,1152,1176]" pageId="6" pageNumber="255" value="3.0">3</number>
</figureCitation>
), several grouped in ventro-externo-distal sensorial fields on Ant. Ib, II and III (
<figureCitation box="[1075,1144,1185,1210]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[1128,1144,1186,1210]" pageId="6" pageNumber="255" value="4.0">4</number>
</figureCitation>
), and
<number box="[1213,1255,1186,1210]" pageId="6" pageNumber="255" value="4.5" valueMax="7.0" valueMin="2.0">27</number>
distally on Ant. IV (
<figureCitation box="[242,310,1218,1243]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[294,310,1219,1243]" pageId="6" pageNumber="255" value="7.0">7</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,1253,1277]" pageId="6" pageNumber="255">Type</typeStatus>
j—short, pointed, bent S-chaetae (
<figureCitation box="[609,691,1252,1277]" captionStart="FIGURES 9 14" captionStartId="8.[151,264,1597,1620]" captionTargetBox="[169,1417,186,1574]" captionTargetId="figure@8.[151,1436,171,1589]" captionTargetPageId="8" captionText="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[661,691,1252,1276]" pageId="6" pageNumber="255" value="13.0">13</number>
</figureCitation>
),
<number box="[712,754,1252,1276]" pageId="6" pageNumber="255" value="5.0" valueMax="8.0" valueMin="2.0">28</number>
distally on Ant. II and
<number box="[1013,1055,1252,1276]" pageId="6" pageNumber="255" value="3.5" valueMax="4.0" valueMin="3.0">34</number>
on Ant. III, located near sensorial area; not seen on Ant. Ib.
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,1319,1343]" pageId="6" pageNumber="255">Type</typeStatus>
k—thin, short, hyaline S-chaetae (
<figureCitation box="[620,704,1318,1343]" captionStart="FIGURES 9 14" captionStartId="8.[151,264,1597,1620]" captionTargetBox="[169,1417,186,1574]" captionTargetId="figure@8.[151,1436,171,1589]" captionTargetPageId="8" captionText="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[674,704,1319,1343]" pageId="6" pageNumber="255" value="14.0">14</number>
</figureCitation>
), on the whole length of Ant. III and IV and a few ventrally on Ant. Ib and II, some larger and intermediate with
<typeStatus box="[752,800,1352,1376]" pageId="6" pageNumber="255">type</typeStatus>
i, especially on Ant. IV.
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" pageId="6" pageNumber="255">
<typeStatus box="[165,221,1386,1410]" pageId="6" pageNumber="255">Type</typeStatus>
l—thin, long S-chaetae (
<figureCitation box="[520,608,1385,1410]" captionStart="FIGURES 9 14" captionStartId="8.[151,264,1597,1620]" captionTargetBox="[169,1417,186,1574]" captionTargetId="figure@8.[151,1436,171,1589]" captionTargetPageId="8" captionText="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[577,608,1386,1410]" pageId="6" pageNumber="255" value="10.0">10</number>
</figureCitation>
), similar in shape and size to f-type mes, but blunt at the tip, thinner proximally and not straight, present ventrally on Ant. Ia and Ib.
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[199,1436,1452,1477]" pageId="6" pageNumber="255">Scales rounded, of medium size, their surface covered with numerous short spicules arranged longitudinally;</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" pageId="6" pageNumber="255">
numerous dorsally on Ant. Ib and II, few ventrally on Ant. Ib; absent on Ant. Ia, III and IV. Subapical organite of Ant. IV long, thick and slightly bifurcate (
<figureCitation box="[918,986,1518,1543]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[970,986,1519,1543]" pageId="6" pageNumber="255" value="7.0">7</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[199,846,1551,1576]" pageId="6" pageNumber="255">
Subapical papilla of Ant. IV, devoid of &quot;pin seta&quot; (
<figureCitation box="[764,832,1551,1576]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[816,832,1552,1576]" pageId="6" pageNumber="255" value="7.0">7</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" pageId="6" pageNumber="255">
The different categories of phaneres are distributed as follows on each antennal segment: Ant. Ia dorsally with
<number box="[440,454,1619,1643]" pageId="6" pageNumber="255" value="2.0">2</number>
basal mic (
<typeStatus box="[585,635,1619,1643]" pageId="6" pageNumber="255">type</typeStatus>
g) and thick serrated mes (
<typeStatus box="[943,993,1619,1643]" pageId="6" pageNumber="255">type</typeStatus>
b); ventrally with
<number box="[1204,1218,1619,1643]" pageId="6" pageNumber="255" value="3.0">3</number>
basal mic (
<typeStatus box="[1348,1398,1619,1643]" pageId="6" pageNumber="255">type</typeStatus>
g),
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,1436,1652,1677]" pageId="6" pageNumber="255">
pointed smooth mes (
<typeStatus box="[394,444,1652,1676]" pageId="6" pageNumber="255">type</typeStatus>
e), acuminate smooth mes (
<typeStatus box="[762,812,1652,1676]" pageId="6" pageNumber="255">type</typeStatus>
f) and sensorial area constituted of
<number box="[1218,1288,1652,1676]" pageId="6" pageNumber="255" value="11.0" valueMax="12.0" valueMin="10.0">1012</number>
S-chaetae of
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,747,1685,1710]" pageId="6" pageNumber="255">
<typeStatus box="[151,199,1686,1710]" pageId="6" pageNumber="255">type</typeStatus>
l and
<number box="[268,310,1686,1710]" pageId="6" pageNumber="255" value="3.5" valueMax="4.0" valueMin="3.0">34</number>
S-chaetae of
<typeStatus box="[463,511,1686,1710]" pageId="6" pageNumber="255">type</typeStatus>
i (
<figureCitation box="[542,610,1685,1710]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="6" pageNumber="255">
Fig.
<number box="[594,610,1686,1710]" pageId="6" pageNumber="255" value="3.0">3</number>
</figureCitation>
). No scales.
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[199,1436,1718,1743]" pageId="6" pageNumber="255">
Ant. Ib dorsally with dense cover of scales, without chaetae except apical row of serrated mac (
<typeStatus box="[1285,1335,1719,1743]" pageId="6" pageNumber="255">type</typeStatus>
a), thick
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,1436,1751,1776]" pageId="6" pageNumber="255">
serrated mes (
<typeStatus box="[312,362,1752,1776]" pageId="6" pageNumber="255">type</typeStatus>
b), and two externo-apical small swollen S-chaetae (
<typeStatus box="[980,1030,1752,1776]" pageId="6" pageNumber="255">type</typeStatus>
h); ventrally with very few scales,
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,1436,1785,1810]" pageId="6" pageNumber="255">
large number of pointed smooth mes (
<typeStatus box="[594,644,1786,1810]" pageId="6" pageNumber="255">type</typeStatus>
e), thick serrated mes (
<typeStatus box="[919,969,1786,1810]" pageId="6" pageNumber="255">type</typeStatus>
b), few acuminate smooth mes (
<typeStatus box="[1351,1401,1786,1810]" pageId="6" pageNumber="255">type</typeStatus>
f),
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,1436,1818,1843]" pageId="6" pageNumber="255">
distal row of pointed smooth mac (
<typeStatus box="[558,608,1819,1843]" pageId="6" pageNumber="255">type</typeStatus>
d) or large pointed smooth mes (
<typeStatus box="[997,1047,1819,1843]" pageId="6" pageNumber="255">type</typeStatus>
e), ventro-external sensorial field
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,1436,1852,1877]" pageId="6" pageNumber="255">
constituted of several S-chaetae of
<typeStatus box="[558,606,1853,1877]" pageId="6" pageNumber="255">type</typeStatus>
i, few short S-chaetae of
<typeStatus box="[912,960,1853,1877]" pageId="6" pageNumber="255">type</typeStatus>
k externo-apically to sensorial field, and
</paragraph>
<paragraph blockId="6.[151,1437,852,1910]" box="[151,934,1885,1910]" pageId="6" pageNumber="255">
several S-chaetae of
<typeStatus box="[383,442,1886,1910]" pageId="6" pageNumber="255">types</typeStatus>
i, k and l proximally to field. No basal mic.
</paragraph>
<caption httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="7" pageNumber="256" targetBox="[235,1327,204,1639]" targetPageId="7">
<paragraph blockId="7.[151,1436,1704,2034]" pageId="7" pageNumber="256">
<emphasis bold="true" box="[151,317,1704,1727]" pageId="7" pageNumber="256">FIGURES 28.</emphasis>
<taxonomicName box="[325,711,1705,1727]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="256" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[325,458,1706,1727]" italics="true" pageId="7" pageNumber="256">Heteromurus</emphasis>
(
<emphasis box="[475,604,1705,1727]" italics="true" pageId="7" pageNumber="256">Verhoeffiella</emphasis>
)
<emphasis box="[621,711,1705,1727]" italics="true" pageId="7" pageNumber="256">absoloni</emphasis>
</taxonomicName>
, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm.
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="8" pageNumber="257" targetBox="[169,1417,186,1574]" targetPageId="8">
<paragraph blockId="8.[151,1437,1597,1712]" pageId="8" pageNumber="257">
<emphasis bold="true" box="[151,326,1597,1620]" pageId="8" pageNumber="257">FIGURES 914.</emphasis>
<taxonomicName box="[332,709,1598,1620]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="257" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[332,465,1599,1620]" italics="true" pageId="8" pageNumber="257">Heteromurus</emphasis>
(
<emphasis box="[479,608,1598,1620]" italics="true" pageId="8" pageNumber="257">Verhoeffiella</emphasis>
)
<emphasis box="[622,709,1598,1620]" italics="true" pageId="8" pageNumber="257">absoloni</emphasis>
</taxonomicName>
, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm.
</paragraph>
</caption>
<paragraph blockId="8.[151,1437,1757,2034]" pageId="8" pageNumber="257">
Ant. II dorsally with dense cover of scales mixed with sparse serrated mac and mes (
<typeStatus box="[1158,1219,1758,1782]" pageId="8" pageNumber="257">types</typeStatus>
a and b), distally a few acuminate smooth mes (
<typeStatus box="[478,528,1793,1817]" pageId="8" pageNumber="257">type</typeStatus>
f), apical row of small mac (
<typeStatus box="[864,914,1793,1817]" pageId="8" pageNumber="257">type</typeStatus>
d) or large straight mes (
<typeStatus box="[1205,1255,1793,1817]" pageId="8" pageNumber="257">type</typeStatus>
e) and externoapical group of
<number box="[335,377,1830,1854]" pageId="8" pageNumber="257" value="5.5" valueMax="7.0" valueMin="4.0">47</number>
swollen S-chaetae of
<typeStatus box="[635,683,1830,1854]" pageId="8" pageNumber="257">type</typeStatus>
h (
<collectionCode box="[724,787,1830,1854]" pageId="8" pageNumber="257">AIIO</collectionCode>
,
<figureCitation box="[801,871,1829,1854]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="8" pageNumber="257">
Fig.
<number box="[855,871,1829,1854]" pageId="8" pageNumber="257" value="5.0">5</number>
</figureCitation>
); ventrally without scales, with many short thick serrated mes (
<typeStatus box="[309,359,1865,1889]" pageId="8" pageNumber="257">type</typeStatus>
b), pointed smooth mes (
<typeStatus box="[651,701,1865,1889]" pageId="8" pageNumber="257">type</typeStatus>
e, less numerous), acuminate smooth mes (
<typeStatus box="[1199,1249,1865,1889]" pageId="8" pageNumber="257">type</typeStatus>
f), distal row of small mac (
<typeStatus box="[282,332,1902,1926]" pageId="8" pageNumber="257">type</typeStatus>
d) or large smooth mes (
<typeStatus box="[616,666,1902,1926]" pageId="8" pageNumber="257">type</typeStatus>
e) and externo-apical sensorial field constituted of
<number box="[1249,1319,1901,1926]" pageId="8" pageNumber="257" value="27.5" valueMax="30.0" valueMin="25.0">2530</number>
S-chaetae of
<typeStatus box="[181,229,1938,1962]" pageId="8" pageNumber="257">type</typeStatus>
i, similar to that of Ant. Ib, and
<number box="[591,633,1937,1961]" pageId="8" pageNumber="257" value="5.0" valueMax="8.0" valueMin="2.0">28</number>
chaetae of
<typeStatus box="[761,809,1938,1962]" pageId="8" pageNumber="257">type</typeStatus>
j;
<number box="[839,880,1937,1961]" pageId="8" pageNumber="257" value="2.5" valueMax="3.0" valueMin="2.0">23</number>
short S-chaetae of
<typeStatus box="[1096,1144,1938,1962]" pageId="8" pageNumber="257">type</typeStatus>
k present externo-distally between sensorial field and
<collectionCode box="[462,525,1974,1998]" pageId="8" pageNumber="257">AIIO</collectionCode>
; longer S-chaetae of
<typeStatus box="[759,807,1974,1998]" pageId="8" pageNumber="257">type</typeStatus>
k present laterally to and mostly proximally to sensorial field. Basal mic present (
<number box="[433,445,2009,2033]" pageId="8" pageNumber="257" value="1.0">1</number>
dorsal,
<number box="[535,549,2009,2033]" pageId="8" pageNumber="257" value="2.0">2</number>
ventral,
<number box="[648,662,2009,2033]" pageId="8" pageNumber="257" value="1.0">1</number>
latero-external,
<number box="[846,860,2009,2033]" pageId="8" pageNumber="257" value="1.0">1</number>
latero-internal).
</paragraph>
<caption httpUri="https://zenodo.org/record/236230/files/figure.png" pageId="9" pageNumber="258" targetBox="[158,1428,202,1155]" targetPageId="9">
<paragraph blockId="9.[151,1436,1183,1236]" pageId="9" pageNumber="258">
<emphasis bold="true" box="[151,340,1183,1206]" pageId="9" pageNumber="258">FIGURES 1520.</emphasis>
<taxonomicName box="[346,724,1184,1206]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[346,479,1185,1206]" italics="true" pageId="9" pageNumber="258">Heteromurus</emphasis>
(
<emphasis box="[493,622,1184,1206]" italics="true" pageId="9" pageNumber="258">Verhoeffiella</emphasis>
)
<emphasis box="[637,724,1184,1206]" italics="true" pageId="9" pageNumber="258">absoloni</emphasis>
</taxonomicName>
, mouthparts: 15, labrum; 16, labial palp; 17, maxillary outer lobe; 18, maxilla; 19, mandible; 20, labial triangle. Scales: 40 µm (15), 20 µm (1620).
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/236231/files/figure.png" pageId="9" pageNumber="258" targetBox="[151,1436,1286,1831]" targetPageId="9">
<paragraph blockId="9.[151,1435,1852,1905]" pageId="9" pageNumber="258">
<emphasis bold="true" box="[151,340,1852,1875]" pageId="9" pageNumber="258">FIGURES 2122.</emphasis>
<taxonomicName box="[346,728,1852,1875]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[346,479,1854,1875]" italics="true" pageId="9" pageNumber="258">Heteromurus</emphasis>
(
<emphasis box="[494,623,1852,1874]" italics="true" pageId="9" pageNumber="258">Verhoeffiella</emphasis>
)
<emphasis box="[638,728,1852,1874]" italics="true" pageId="9" pageNumber="258">absoloni</emphasis>
</taxonomicName>
, optical microscope photos of Abd. IV and VI: 21, scales and long Slike chaetae on Abd. IV; 22, scales and serrated mes on Abd.VI. Scales 50 µm.
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="10" pageNumber="259" targetBox="[205,1361,209,1718]" targetPageId="10">
<paragraph blockId="10.[151,1436,1760,2027]" pageId="10" pageNumber="259">
<emphasis bold="true" box="[151,341,1760,1783]" pageId="10" pageNumber="259">FIGURES 2330.</emphasis>
<taxonomicName box="[347,726,1760,1783]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="259" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[347,480,1762,1783]" italics="true" pageId="10" pageNumber="259">Heteromurus</emphasis>
(
<emphasis box="[495,624,1760,1782]" italics="true" pageId="10" pageNumber="259">Verhoeffiella</emphasis>
)
<emphasis box="[639,726,1760,1782]" italics="true" pageId="10" pageNumber="259">absoloni</emphasis>
</taxonomicName>
: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a1, clubbed mac, a2, subcylindrical mac, b1, normal serrated mes, b2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a1, big serrated mac, a2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f1, short S-like chaetae on Th. III, f2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25).
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="11" pageNumber="260" targetBox="[152,1424,205,1914]" targetPageId="11">
<paragraph blockId="11.[151,1436,1915,2029]" pageId="11" pageNumber="260">
<emphasis bold="true" box="[151,341,1915,1938]" pageId="11" pageNumber="260">FIGURES 3136.</emphasis>
<taxonomicName box="[348,728,1915,1937]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="11" pageNumber="260" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[348,481,1916,1937]" italics="true" pageId="11" pageNumber="260">Heteromurus</emphasis>
(
<emphasis box="[496,625,1915,1937]" italics="true" pageId="11" pageNumber="260">Verhoeffiella</emphasis>
)
<emphasis box="[641,728,1915,1937]" italics="true" pageId="11" pageNumber="260">absoloni</emphasis>
</taxonomicName>
.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circlesanterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32).
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/236234/files/figure.png" pageId="12" pageNumber="261" targetBox="[151,1436,193,1322]" targetPageId="12">
<paragraph blockId="12.[151,1436,1344,1428]" pageId="12" pageNumber="261">
<emphasis bold="true" box="[151,339,1344,1367]" pageId="12" pageNumber="261">FIGURES 3739.</emphasis>
<taxonomicName box="[345,723,1345,1367]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="261" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[345,478,1346,1367]" italics="true" pageId="12" pageNumber="261">Heteromurus</emphasis>
(
<emphasis box="[492,621,1345,1367]" italics="true" pageId="12" pageNumber="261">Verhoeffiella</emphasis>
)
<emphasis box="[636,723,1345,1367]" italics="true" pageId="12" pageNumber="261">absoloni</emphasis>
</taxonomicName>
, optical microscope photos of legs: 37, internal structure of unguis; 38 trochanteral organ; 39, Tita I, posterior view; the oblique ridges visible on the the claw are an artefact. Scales: 50 µm (37, 38); 100 µm (39).
</paragraph>
</caption>
<paragraph blockId="12.[151,1436,1473,2038]" pageId="12" pageNumber="261">
Ant. IIIIV chaetotaxy with tendency to form whorls of chaetae, sometimes spiraling, without clear serial pattern across successive whorls (
<figureCitation box="[529,596,1509,1534]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="12" pageNumber="261">
Fig.
<number box="[582,596,1510,1534]" pageId="12" pageNumber="261" value="8.0">8</number>
</figureCitation>
) that are composed of
<number box="[853,867,1510,1534]" pageId="12" pageNumber="261" value="4.0">4</number>
<typeStatus box="[874,933,1510,1534]" pageId="12" pageNumber="261">types</typeStatus>
of chaetae variously interspersed (b, c, f and k,
<figureCitation box="[180,249,1545,1570]" captionStart="FIGURES 9 14" captionStartId="8.[151,264,1597,1620]" captionTargetBox="[169,1417,186,1574]" captionTargetId="figure@8.[151,1436,171,1589]" captionTargetPageId="8" captionText="FIGURES 9 14. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of antennal chaetae (chaetae types as in Fig. 2): 9, Ant. IV, with chaetae of types b, c and f; 10, Ant. Ia ventro-proximally, with chaetae of types g and l; 11, Ant. II, with its dorso-distal group of h-type chaetae (AIIO); 12, Ant. II, with its ventro-externo-distal sensorial field of i-type chaetae; 13, Ant. II ventro-externo-distally, with chaetae of type j; 14, Ant. IV laterally, with chaetae of type k. All scales 10 µm." httpUri="https://zenodo.org/record/236229/files/figure.png" pageId="12" pageNumber="261">
Fig.
<number box="[233,249,1545,1569]" pageId="12" pageNumber="261" value="9.0">9</number>
</figureCitation>
); dorsally with less c-type chaetae and all chaetae smaller in size compared to the ventral ones. Ant. IV with several thick subcylindrical S-chaetae (
<typeStatus box="[646,696,1582,1606]" pageId="12" pageNumber="261">type</typeStatus>
i) in distal part (
<figureCitation box="[882,951,1581,1606]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="12" pageNumber="261">
Fig.
<number box="[935,951,1582,1606]" pageId="12" pageNumber="261" value="7.0">7</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="12.[151,1436,1473,2038]" pageId="12" pageNumber="261">
<collectionCode box="[199,267,1617,1641]" pageId="12" pageNumber="261">AIIIO</collectionCode>
located dorso-externo-apically, with
<number box="[686,700,1617,1641]" pageId="12" pageNumber="261" value="2.0">2</number>
swollen S-chaetae side by side (
<typeStatus box="[1065,1115,1617,1641]" pageId="12" pageNumber="261">type</typeStatus>
h), surrounded internally by one thin short S-chaeta (
<typeStatus box="[423,473,1654,1678]" pageId="12" pageNumber="261">type</typeStatus>
k) and externally by three short external S-chaetae, of which
<number box="[1164,1178,1654,1678]" pageId="12" pageNumber="261" value="2.0">2</number>
pointed and bent (
<typeStatus box="[1386,1436,1654,1678]" pageId="12" pageNumber="261">type</typeStatus>
j), and
<number box="[233,247,1689,1713]" pageId="12" pageNumber="261" value="1.0">1</number>
thin (
<typeStatus box="[316,366,1689,1713]" pageId="12" pageNumber="261">type</typeStatus>
k); few additional S-chaetae of
<typeStatus box="[736,784,1690,1714]" pageId="12" pageNumber="261">type</typeStatus>
k present below (
<figureCitation box="[992,1062,1689,1714]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="12" pageNumber="261">
Fig.
<number box="[1046,1062,1689,1714]" pageId="12" pageNumber="261" value="6.0">6</number>
</figureCitation>
). Ant. III ventro-externo-distally with sensorial field constituted of about
<number box="[621,690,1726,1750]" pageId="12" pageNumber="261" value="25.0" valueMax="30.0" valueMin="20.0">2030</number>
i-type chaetae, and
<number box="[928,970,1726,1750]" pageId="12" pageNumber="261" value="2.0" valueMax="3.0" valueMin="1.0">13</number>
small pointed j-type chaetae. Basal mic absent or undifferentiated on Ant. III and absent on Ant. IV. Scales absent on Ant. IIIIV. Ant. IV with large slightly bifurcate subapical organite and a subapical papilla (
<figureCitation box="[831,899,1797,1822]" captionStart="FIGURES 2 8" captionStartId="7.[151,264,1704,1727]" captionTargetBox="[235,1327,204,1639]" captionTargetId="figure@7.[223,1332,193,1651]" captionTargetPageId="7" captionText="FIGURES 2 8. Heteromurus (Verhoeffiella) absoloni, antennal chaetae: 2, types of antennal chaetae: a—serrated mac, bthick serrated mes, c—thin serrated mes, d—pointed smooth mac, e—pointed smooth mes, f— acuminate smooth mes, gpointed smooth mic, h—swollen S-chaeta, i—thick subcylindrical S-chaeta, j—short pointed bent S-chaeta, k—thin short Schaeta, l—thin, long S-chaetae; 3, Ant. Ia, ventral side with g-type of chaetae, and sensorial field (circled by dotted line) composed of i- and l-types of chaetae (i in black); 4, Ant. Ib, ventro-externo-distal sensorial field (i-type chaetae in black), pseudopores (cross-circles) and the 2 h-type chaetae of the dorsal side in dotted line; 5, Ant. II, dorso-externo-apical organite (AIIO) with acuminate thin mes (f) and swollen S-chaetae (h); 6, Ant. III, dorso-externo-apical organite (AIIIO), with sockets of f-type and b-type chaetae; 7, Ant. IV, distal part: hollow circles—thick serrated mes (b), triangles—thin serrated mes (c), black circles—acuminate smooth mes (f), x—fallen chaetae, hollow lines—thin short S-chaetae (k), filled lines—thick subcylindrical S-chaetae (i); 8, whorls of Ant. IV, middle part; symbols same as in Fig. 7, in addition asterisk—thin short Schaetae (k). All scales 20 µm." httpUri="https://zenodo.org/record/236228/files/figure.png" pageId="12" pageNumber="261">
Fig.
<number box="[883,899,1798,1822]" pageId="12" pageNumber="261" value="7.0">7</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="12.[151,1436,1473,2038]" box="[199,1317,1833,1858]" pageId="12" pageNumber="261">When Ant. III and IV are fused, the described pattern of Ant. III distal chaetae is strongly modified.</paragraph>
<paragraph blockId="12.[151,1436,1473,2038]" lastBlockId="13.[151,1437,151,501]" lastPageId="13" lastPageNumber="262" pageId="12" pageNumber="261">
<emphasis bold="true" box="[199,342,1869,1894]" pageId="12" pageNumber="261">Mouthparts</emphasis>
(
<figureCitation box="[360,494,1869,1894]" captionStart="FIGURES 15 20" captionStartId="9.[151,264,1183,1206]" captionTargetBox="[158,1428,202,1155]" captionTargetId="figure@9.[151,1436,193,1162]" captionTargetPageId="9" captionText="FIGURES 15 20. Heteromurus (Verhoeffiella) absoloni, mouthparts: 15, labrum; 16, labial palp; 17, maxillary outer lobe; 18, maxilla; 19, mandible; 20, labial triangle. Scales: 40 µm (15), 20 µm (16 20)." httpUri="https://zenodo.org/record/236230/files/figure.png" pageId="12" pageNumber="261">
Figs
<number box="[420,494,1869,1894]" pageId="12" pageNumber="261" value="17.5" valueMax="20.0" valueMin="15.0">1520</number>
</figureCitation>
). Labral formula
<number box="[698,712,1869,1894]" pageId="12" pageNumber="261" value="4.0">4</number>
/5,5,4. No labral papillae. Prelabral and labral chaetae smooth, acuminate, of similar size. Labrum margin with inverted medial U-form intrusion devoid of primary granules, distal smooth area large (
<figureCitation box="[438,523,1941,1966]" captionStart="FIGURES 15 20" captionStartId="9.[151,264,1183,1206]" captionTargetBox="[158,1428,202,1155]" captionTargetId="figure@9.[151,1436,193,1162]" captionTargetPageId="9" captionText="FIGURES 15 20. Heteromurus (Verhoeffiella) absoloni, mouthparts: 15, labrum; 16, labial palp; 17, maxillary outer lobe; 18, maxilla; 19, mandible; 20, labial triangle. Scales: 40 µm (15), 20 µm (16 20)." httpUri="https://zenodo.org/record/236230/files/figure.png" pageId="12" pageNumber="261">
Fig.
<number box="[492,523,1941,1966]" pageId="12" pageNumber="261" value="15.0">15</number>
</figureCitation>
). Ventro-distally on labrum two unequal combs with one apical row of about
<number box="[1408,1436,1942,1966]" pageId="12" pageNumber="261" value="10.0">10</number>
and
<number box="[198,226,1977,2002]" pageId="12" pageNumber="261" value="15.0">15</number>
teeth respectively, and a pair of sinuous rods (
<figureCitation box="[748,832,1977,2002]" captionStart="FIGURES 15 20" captionStartId="9.[151,264,1183,1206]" captionTargetBox="[158,1428,202,1155]" captionTargetId="figure@9.[151,1436,193,1162]" captionTargetPageId="9" captionText="FIGURES 15 20. Heteromurus (Verhoeffiella) absoloni, mouthparts: 15, labrum; 16, labial palp; 17, maxillary outer lobe; 18, maxilla; 19, mandible; 20, labial triangle. Scales: 40 µm (15), 20 µm (16 20)." httpUri="https://zenodo.org/record/236230/files/figure.png" pageId="12" pageNumber="261">
Fig.
<number box="[801,832,1977,2002]" pageId="12" pageNumber="261" value="15.0">15</number>
</figureCitation>
). Labial triangle with
<number box="[1081,1137,1977,2001]" pageId="12" pageNumber="261" value="10.5" valueMax="12.0" valueMin="9.0">912</number>
chaetae, all smooth (N=
<number box="[1414,1428,1977,2001]" pageId="12" pageNumber="261" value="9.0">9</number>
) and no scales (
<figureCitation box="[314,398,2013,2038]" captionStart="FIGURES 15 20" captionStartId="9.[151,264,1183,1206]" captionTargetBox="[158,1428,202,1155]" captionTargetId="figure@9.[151,1436,193,1162]" captionTargetPageId="9" captionText="FIGURES 15 20. Heteromurus (Verhoeffiella) absoloni, mouthparts: 15, labrum; 16, labial palp; 17, maxillary outer lobe; 18, maxilla; 19, mandible; 20, labial triangle. Scales: 40 µm (15), 20 µm (16 20)." httpUri="https://zenodo.org/record/236230/files/figure.png" pageId="12" pageNumber="261">
Fig.
<number box="[366,398,2014,2038]" pageId="12" pageNumber="261" value="20.0">20</number>
</figureCitation>
); lateral area (submentum) with
<number box="[759,773,2013,2038]" pageId="12" pageNumber="261" value="5.0">5</number>
smooth chaetae. Labial palp with
<number box="[1156,1170,2013,2038]" pageId="12" pageNumber="261" value="5.0">5</number>
proximal chaetae and
<number box="[1422,1436,2013,2038]" pageId="12" pageNumber="261" value="5.0">5</number>
papillae (A,B,C,D,E); guard chaeta of papilla A integrated in papilla B complex which thus appears to have
<number box="[1352,1366,151,176]" pageId="13" pageNumber="262" value="5.0">5</number>
guard chaetae (
<bibRefCitation author="Fjellberg" box="[252,483,188,213]" pageId="13" pageNumber="262" refString="Fjellberg, A. (1999) The Labial Palp in Collembola. Zoologischer Anzeiger, 237, 309 - 330." type="journal article" year="1999">
Fjellberg
<number box="[364,424,188,212]" pageId="13" pageNumber="262" value="1999.0">1999</number>
:
<number box="[437,483,188,213]" pageId="13" pageNumber="262" value="325.0">325</number>
</bibRefCitation>
); papilla C without guard chaeta; papilla D with
<number box="[1052,1066,188,212]" pageId="13" pageNumber="262" value="4.0">4</number>
guard chaetae; papilla E with
<number box="[1422,1436,188,212]" pageId="13" pageNumber="262" value="4.0">4</number>
guard chaetae and well developed labial process (
<figureCitation box="[732,819,223,248]" captionStart="FIGURES 15 20" captionStartId="9.[151,264,1183,1206]" captionTargetBox="[158,1428,202,1155]" captionTargetId="figure@9.[151,1436,193,1162]" captionTargetPageId="9" captionText="FIGURES 15 20. Heteromurus (Verhoeffiella) absoloni, mouthparts: 15, labrum; 16, labial palp; 17, maxillary outer lobe; 18, maxilla; 19, mandible; 20, labial triangle. Scales: 40 µm (15), 20 µm (16 20)." httpUri="https://zenodo.org/record/236230/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[788,819,223,248]" pageId="13" pageNumber="262" value="16.0">16</number>
</figureCitation>
). Maxillary outer lobe with one basal chaeta, simple maxillary palp, three sublobal hairs and one smaller chaeta (
<figureCitation box="[855,942,260,285]" captionStart="FIGURES 15 20" captionStartId="9.[151,264,1183,1206]" captionTargetBox="[158,1428,202,1155]" captionTargetId="figure@9.[151,1436,193,1162]" captionTargetPageId="9" captionText="FIGURES 15 20. Heteromurus (Verhoeffiella) absoloni, mouthparts: 15, labrum; 16, labial palp; 17, maxillary outer lobe; 18, maxilla; 19, mandible; 20, labial triangle. Scales: 40 µm (15), 20 µm (16 20)." httpUri="https://zenodo.org/record/236230/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[911,942,260,284]" pageId="13" pageNumber="262" value="17.0">17</number>
</figureCitation>
); basal fold with one chaeta. Maxilla with strong tridentate claw,
<number box="[417,431,295,320]" pageId="13" pageNumber="262" value="5.0">5</number>
lamellae and a thin rod, long and bent (
<figureCitation box="[910,996,295,320]" captionStart="FIGURES 15 20" captionStartId="9.[151,264,1183,1206]" captionTargetBox="[158,1428,202,1155]" captionTargetId="figure@9.[151,1436,193,1162]" captionTargetPageId="9" captionText="FIGURES 15 20. Heteromurus (Verhoeffiella) absoloni, mouthparts: 15, labrum; 16, labial palp; 17, maxillary outer lobe; 18, maxilla; 19, mandible; 20, labial triangle. Scales: 40 µm (15), 20 µm (16 20)." httpUri="https://zenodo.org/record/236230/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[966,996,296,320]" pageId="13" pageNumber="262" value="18.0">18</number>
</figureCitation>
, after Fjellberg
<number box="[1185,1245,296,320]" pageId="13" pageNumber="262" value="2007.0">2007</number>
:
<number box="[1260,1306,295,320]" pageId="13" pageNumber="262" value="160.0">160</number>
). Mandible asymmetrical with
<number box="[367,381,332,357]" pageId="13" pageNumber="262" value="5.0">5</number>
teeth on right and
<number box="[596,610,332,356]" pageId="13" pageNumber="262" value="4.0">4</number>
on left on all examined specimens (
<figureCitation box="[1020,1104,332,357]" captionStart="FIGURES 15 20" captionStartId="9.[151,264,1183,1206]" captionTargetBox="[158,1428,202,1155]" captionTargetId="figure@9.[151,1436,193,1162]" captionTargetPageId="9" captionText="FIGURES 15 20. Heteromurus (Verhoeffiella) absoloni, mouthparts: 15, labrum; 16, labial palp; 17, maxillary outer lobe; 18, maxilla; 19, mandible; 20, labial triangle. Scales: 40 µm (15), 20 µm (16 20)." httpUri="https://zenodo.org/record/236230/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[1073,1104,332,356]" pageId="13" pageNumber="262" value="19.0">19</number>
</figureCitation>
). Clypeus with many pointed smooth and serrated mesochaetae. About
<number box="[629,657,368,392]" pageId="13" pageNumber="262" value="10.0">10</number>
+
<number box="[673,701,368,392]" pageId="13" pageNumber="262" value="10.0">10</number>
smooth post-labial chaetae (PLQ of
<bibRefCitation author="Mari-Mutt" box="[1128,1311,367,392]" pageId="13" pageNumber="262" refString="Mari-Mutt, J. A. (1985) Three new species of Heteromurus (Alloscopus) and descriptive notes for species of the subgenus (Collembola: Entomobryidae). The Florida Entomologist, 68 (2), 335 - 346. http: // dx. doi. org / 10.2307 / 3494370" type="journal article" year="1985">
Mari-Mutt
<number box="[1255,1311,367,392]" pageId="13" pageNumber="262" value="1985.0">1985</number>
</bibRefCitation>
<emphasis box="[1320,1426,369,392]" italics="true" pageId="13" pageNumber="262">pro parte</emphasis>
) interspersed with scales.
</paragraph>
<paragraph blockId="13.[151,1437,151,501]" pageId="13" pageNumber="262">
<emphasis bold="true" box="[199,390,439,464]" pageId="13" pageNumber="262">Head and body</emphasis>
(
<figureCitation box="[409,544,439,464]" captionStart-0="FIGURES 21 22" captionStart-1="FIGURES 23 30" captionStartId-0="9.[151,264,1852,1875]" captionStartId-1="10.[151,264,1760,1783]" captionTargetBox-0="[151,1436,1286,1831]" captionTargetBox-1="[205,1361,209,1718]" captionTargetId-0="figure@9.[151,1436,1286,1831]" captionTargetId-1="figure@10.[194,1380,193,1738]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 21 22. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of Abd. IV and VI: 21, scales and long Slike chaetae on Abd. IV; 22, scales and serrated mes on Abd. VI. Scales 50 µm." captionText-1="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri-0="https://zenodo.org/record/236231/files/figure.png" httpUri-1="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Figs
<number box="[470,544,440,464]" pageId="13" pageNumber="262" value="25.5" valueMax="30.0" valueMin="21.0">2130</number>
</figureCitation>
). Chaetae are rather diverse on head (
<figureCitation box="[995,1081,439,464]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[1052,1081,440,464]" pageId="13" pageNumber="262" value="27.0">27</number>
</figureCitation>
) and body (
<figureCitation box="[1228,1316,439,464]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[1285,1316,440,464]" pageId="13" pageNumber="262" value="28.0">28</number>
</figureCitation>
), with the following categories usually relatively easy to recognize:
</paragraph>
<paragraph blockId="13.[151,1436,548,933]" box="[151,1436,548,573]" pageId="13" pageNumber="262">
serrated mac, either clubbed (
<emphasis box="[529,591,549,572]" italics="true" pageId="13" pageNumber="262">sensu</emphasis>
<bibRefCitation author="Stach" box="[598,726,548,573]" pageId="13" pageNumber="262" refString="Stach, J. (1960) The Apterygotan fauna of Poland in relation to the World-fauna of this group of Insects. Tribe: Orchesellini. Polska Akademia Nauk, Instytut Zoologiczny, Oddzial w Krakowie, Panstwowe Wydawnictwo Naukowe, 151 pp." type="book" year="1960">
Stach
<number box="[666,726,548,573]" pageId="13" pageNumber="262" value="1960.0">1960</number>
</bibRefCitation>
: plate I) or subcylindrical-pointed (
<figureCitation box="[1119,1205,548,573]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Figs
<number box="[1175,1205,548,572]" pageId="13" pageNumber="262" value="27.0">27</number>
</figureCitation>
a
<number box="[1218,1227,548,572]" pageId="13" pageNumber="262" value="1.0">1</number>
a
<number box="[1256,1270,548,572]" pageId="13" pageNumber="262" value="2.0">2</number>
and
<number box="[1324,1355,548,572]" pageId="13" pageNumber="262" value="28.0">28</number>
a
<number box="[1368,1377,548,572]" pageId="13" pageNumber="262" value="1.0">1</number>
a
<number box="[1408,1423,548,572]" pageId="13" pageNumber="262" value="2.0">2</number>
);
</paragraph>
<paragraph blockId="13.[151,1436,548,933]" box="[151,828,583,608]" pageId="13" pageNumber="262">
serrated mes on head and body (
<figureCitation box="[560,645,583,608]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Figs
<number box="[616,645,583,608]" pageId="13" pageNumber="262" value="27.0">27</number>
</figureCitation>
b
<number box="[660,669,583,608]" pageId="13" pageNumber="262" value="1.0">1</number>
b
<number box="[700,714,583,608]" pageId="13" pageNumber="262" value="2.0">2</number>
and
<number box="[769,800,583,608]" pageId="13" pageNumber="262" value="28.0">28</number>
b);
</paragraph>
<paragraph blockId="13.[151,1436,548,933]" box="[151,1314,620,645]" pageId="13" pageNumber="262">
thin mes slightly serrated on its distal half and associated with thin S-chaetae (
<figureCitation box="[1076,1288,620,645]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Figs 25, 26 and
<number box="[1257,1288,620,644]" pageId="13" pageNumber="262" value="28.0">28</number>
</figureCitation>
c);
</paragraph>
<paragraph blockId="13.[151,1436,548,933]" box="[151,617,655,680]" pageId="13" pageNumber="262">
smooth mes, only on head (
<figureCitation box="[507,591,655,680]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[560,591,656,680]" pageId="13" pageNumber="262" value="27.0">27</number>
</figureCitation>
c);
</paragraph>
<paragraph blockId="13.[151,1436,548,933]" pageId="13" pageNumber="262">
trichobothria on head (
<figureCitation box="[469,555,692,717]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[526,555,692,717]" pageId="13" pageNumber="262" value="27.0">27</number>
</figureCitation>
d) and tergites, often fallen in our specimens; those of head (
<number box="[1291,1302,692,716]" pageId="13" pageNumber="262" value="1.0">1</number>
+
<number box="[1322,1333,692,716]" pageId="13" pageNumber="262" value="1.0">1</number>
) located laterally and rather short (
<number box="[488,517,727,752]" pageId="13" pageNumber="262" value="50.0">50</number>
µm), those of tergites longer;
</paragraph>
<paragraph blockId="13.[151,1436,548,933]" box="[151,1136,764,789]" pageId="13" pageNumber="262">
S-chaetae, limited to tergites, smooth, very short, thick, pointed (ms,
<figureCitation box="[973,1098,764,789]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">Figs 24, 28</figureCitation>
d);
</paragraph>
<paragraph blockId="13.[151,1436,548,933]" box="[151,1393,799,824]" pageId="13" pageNumber="262">
thin S-chaetae, limited to tergites, smooth, short and hyaline, subcylindrical, blunt (sens,
<figureCitation box="[1196,1366,799,824]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Figs
<number box="[1251,1280,799,824]" pageId="13" pageNumber="262" value="24.0">24</number>
26, 28
</figureCitation>
e);
</paragraph>
<paragraph blockId="13.[151,1436,548,933]" pageId="13" pageNumber="262">
smooth, thin, blunt, short to long S-like chaetae, longer, less transparent and slightly thicker than thin Schaetae, only on Abd. IV (
<figureCitation box="[496,576,871,896]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[548,576,872,896]" pageId="13" pageNumber="262" value="28.0">28</number>
</figureCitation>
f
<number box="[596,606,871,896]" pageId="13" pageNumber="262" value="1.0">1</number>
and f
<number box="[673,688,871,896]" pageId="13" pageNumber="262" value="2.0">2</number>
);
</paragraph>
<paragraph blockId="13.[151,1436,548,933]" box="[151,1070,908,933]" pageId="13" pageNumber="262">
mic,
<number box="[255,269,908,932]" pageId="13" pageNumber="262" value="2.0">2</number>
minute on each anal valve, and
<number box="[630,644,908,932]" pageId="13" pageNumber="262" value="2.0">2</number>
+
<number box="[659,673,908,932]" pageId="13" pageNumber="262" value="2.0">2</number>
on female genital plate (
<figureCitation box="[952,1032,908,933]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Fig.
<quantity box="[1004,1032,908,932]" metricMagnitude="-2" metricUnit="kg" metricValue="2.8" pageId="13" pageNumber="262" unit="g" value="28.0">28</quantity>
</figureCitation>
g);
</paragraph>
<paragraph blockId="13.[151,1437,980,1941]" pageId="13" pageNumber="262">
Scales on dorsal and ventral side of head and body, rounded to oval-elongated, sometimes truncated, hyaline, overlapping, of various size (from
<number box="[537,641,1015,1040]" pageId="13" pageNumber="262" value="57.5" valueMax="100.0" valueMin="15.0">15 to 100</number>
µm long), with a dense cover of short spicules arranged in more or less regular lines (
<figureCitation box="[305,436,1052,1077]" captionStart="FIGURES 21 22" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[151,1436,1286,1831]" captionTargetId="figure@9.[151,1436,1286,1831]" captionTargetPageId="9" captionText="FIGURES 21 22. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of Abd. IV and VI: 21, scales and long Slike chaetae on Abd. IV; 22, scales and serrated mes on Abd. VI. Scales 50 µm." httpUri="https://zenodo.org/record/236231/files/figure.png" pageId="13" pageNumber="262">Figs 21, 22</figureCitation>
).
</paragraph>
<paragraph blockId="13.[151,1437,980,1941]" pageId="13" pageNumber="262">
Chaetae, scales and trichobothria of dorsal side of head and body usually detached in microscopic preparations or on skins retrieved after
<collectionCode LSID="urn:lsid:biocol.org:col:15536" box="[442,503,1124,1148]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="13" pageNumber="262">DNA</collectionCode>
extraction. Chaetae have circular or oval sockets, those of mac and large mes easily recognizable by their size, but those of tergite S-chaetae not distinguishable from those of small mes. Scale sockets are characterized by cupuliform morphology of their socket. Trichobothria have small round socket often with minute point in the middle, but often indistinguishable from mes sockets.
</paragraph>
<paragraph blockId="13.[151,1437,980,1941]" pageId="13" pageNumber="262">
Head dorsally with macrochaetae arranged as in
<figureCitation box="[761,845,1268,1293]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[815,845,1268,1292]" pageId="13" pageNumber="262" value="23.0">23</number>
</figureCitation>
, and dense cover of small to large scales and short serrated mes; laterally with scales mixed with medium size serrated mes; between antennae with few smooth mes; posteriorly with row of serrated mes regularly interspersed with large scales; ventrally with smooth and serrated mes and small to medium-size scales;
<number box="[580,591,1376,1400]" pageId="13" pageNumber="262" value="1.0">1</number>
+
<number box="[612,623,1376,1400]" pageId="13" pageNumber="262" value="1.0">1</number>
trichobothria posterior to ocular area.
</paragraph>
<paragraph blockId="13.[151,1437,980,1941]" pageId="13" pageNumber="262">
Dorsal chaetotaxy illustrated in
<figureCitation box="[559,642,1412,1437]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[611,642,1412,1436]" pageId="13" pageNumber="262" value="23.0">23</number>
</figureCitation>
. Pseudopore formula 1,1/1,1,1,1. Trichobothrial formula
<number box="[1288,1297,1412,1436]" pageId="13" pageNumber="262" value="1.0">1</number>
/0,0/0,2,3,2. Macrochaeta formula (
<number box="[412,425,1448,1472]" pageId="13" pageNumber="262" value="3.0">3</number>
+
<number box="[442,457,1448,1472]" pageId="13" pageNumber="262" value="4.0">4</number>
),(
<number box="[478,491,1448,1472]" pageId="13" pageNumber="262" value="3.0">3</number>
+
<number box="[508,521,1448,1472]" pageId="13" pageNumber="262" value="3.0">3</number>
)/0,2,3,(
<number box="[615,628,1448,1472]" pageId="13" pageNumber="262" value="3.0">3</number>
+0+
<number box="[676,691,1448,1472]" pageId="13" pageNumber="262" value="4.0">4</number>
+
<number box="[708,723,1448,1472]" pageId="13" pageNumber="262" value="4.0">4</number>
). Macrochaeta arrangement stable; multiplets
<emphasis box="[1253,1315,1449,1472]" italics="true" pageId="13" pageNumber="262">sensu</emphasis>
<bibRefCitation author="Szeptycki" pageId="13" pageNumber="262" refString="Szeptycki, A. (1979) Chaetotaxy of the Entomobryidae and its phylogenetical significance. Morpho-systematic studies on Collembola. IV. Polska Akademia Nauk, Zaklad Zoologii Systematycznej i Doswiadczalnej, Panstwowe Wydawnictwo Naukowe, Warszawa-Krakow, 219 pp." type="book" year="1979">
Szeptycki
<number box="[151,207,1484,1508]" pageId="13" pageNumber="262" value="1979.0">1979</number>
</bibRefCitation>
only present anteriorly on Th. II; Th. II with
<number box="[709,723,1484,1508]" pageId="13" pageNumber="262" value="3.0">3</number>
anterior and line of
<number box="[947,961,1484,1508]" pageId="13" pageNumber="262" value="4.0">4</number>
posterior mac; Th. III with
<number box="[1269,1283,1484,1508]" pageId="13" pageNumber="262" value="3.0">3</number>
posterior and
<number box="[151,165,1520,1544]" pageId="13" pageNumber="262" value="3.0">3</number>
lateral mac; Abd. IV with
<number box="[462,476,1520,1544]" pageId="13" pageNumber="262" value="3.0">3</number>
medial,
<number box="[572,586,1520,1544]" pageId="13" pageNumber="262" value="4.0">4</number>
antero-lateral and
<number box="[795,809,1520,1544]" pageId="13" pageNumber="262" value="4.0">4</number>
postero-lateral mac; Abd. V with
<number box="[1188,1278,1519,1544]" pageId="13" pageNumber="262" value="14.0" valueMax="16.0" valueMin="12.0">12 to 16</number>
mac; Abd. VI with
<number box="[209,236,1556,1580]" pageId="13" pageNumber="262" value="11.0">11</number>
mac in two rows, including an uneven one in the anterior row (
<figureCitation box="[963,1047,1556,1581]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[1016,1047,1556,1580]" pageId="13" pageNumber="262" value="23.0">23</number>
</figureCitation>
). Serrated mes present on anterior part on Th. II, dorsally on Abd. V and VI (
<figureCitation box="[639,722,1591,1616]" captionStart="FIGURES 21 22" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[151,1436,1286,1831]" captionTargetId="figure@9.[151,1436,1286,1831]" captionTargetPageId="9" captionText="FIGURES 21 22. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of Abd. IV and VI: 21, scales and long Slike chaetae on Abd. IV; 22, scales and serrated mes on Abd. VI. Scales 50 µm." httpUri="https://zenodo.org/record/236231/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[693,722,1592,1616]" pageId="13" pageNumber="262" value="22.0">22</number>
</figureCitation>
) and very few laterally on all tergites. Many rather long S-like chaetae on Abd. IV,
<number box="[393,422,1628,1653]" pageId="13" pageNumber="262" value="50.0">50</number>
% longer than mes of other tergites, arranged symmetrically (
<figureCitation box="[1140,1227,1628,1653]" captionStart="FIGURES 21 22" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[151,1436,1286,1831]" captionTargetId="figure@9.[151,1436,1286,1831]" captionTargetPageId="9" captionText="FIGURES 21 22. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of Abd. IV and VI: 21, scales and long Slike chaetae on Abd. IV; 22, scales and serrated mes on Abd. VI. Scales 50 µm." httpUri="https://zenodo.org/record/236231/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[1196,1227,1628,1652]" pageId="13" pageNumber="262" value="21.0">21</number>
</figureCitation>
). Sens difficult to observe, often detached or hidden by scales,
<number box="[658,700,1664,1688]" pageId="13" pageNumber="262" value="3.5" valueMax="4.0" valueMin="3.0">34</number>
times longer than mac socket; ms very short, about the length of mac socket. S-chaetotaxy given in
<tableCitation box="[555,627,1700,1725]" captionStart="TABLE 2" captionStartId="15.[151,239,1265,1288]" captionText="TABLE 2. Tergite S-chaetotaxy of H. (V.) absoloni from the type locality. Mesochaetae are mentioned when they are in tandem with sens, with (sens + mesochaeta) given between brackets." pageId="13" pageNumber="262">
Tab.
<number box="[613,627,1700,1724]" pageId="13" pageNumber="262" value="2.0">2</number>
</tableCitation>
and
<figureCitation box="[687,818,1700,1725]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="13" pageNumber="262">
Figs
<number box="[745,818,1700,1725]" pageId="13" pageNumber="262" value="24.5" valueMax="26.0" valueMin="23.0">2326</number>
</figureCitation>
. Sens associated with thin serrated mes in tandem on Abd. IIIII (also possibly in other locations). S-chaetotaxy: ms formula
<number box="[972,981,1736,1760]" pageId="13" pageNumber="262" value="1.0">1</number>
,?
<number box="[997,1006,1736,1760]" pageId="13" pageNumber="262" value="1.0">1</number>
/1,0,1,0,0 (ms on Th. III observed in two specimens, possibly an aberration as it would be unique in Entomobryoidea); sens formula, 2,2/1,3,3,3,3, with S-like chaetae at least
<number box="[401,429,1808,1832]" pageId="13" pageNumber="262" value="10.0">10</number>
+
<number box="[445,473,1808,1832]" pageId="13" pageNumber="262" value="10.0">10</number>
on Abd. IV.
</paragraph>
<paragraph blockId="13.[151,1437,980,1941]" pageId="13" pageNumber="262">
Scales rounded, of various sizes: medium-size on all tergites except Abd. VI where they are smaller (
<figureCitation box="[1338,1423,1844,1869]" captionStart="FIGURES 21 22" captionStartId="9.[151,264,1852,1875]" captionTargetBox="[151,1436,1286,1831]" captionTargetId="figure@9.[151,1436,1286,1831]" captionTargetPageId="9" captionText="FIGURES 21 22. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of Abd. IV and VI: 21, scales and long Slike chaetae on Abd. IV; 22, scales and serrated mes on Abd. VI. Scales 50 µm." httpUri="https://zenodo.org/record/236231/files/figure.png" pageId="13" pageNumber="262">
Fig.
<number box="[1391,1423,1844,1868]" pageId="13" pageNumber="262" value="22.0">22</number>
</figureCitation>
); larger posteriorly (at least twice as large as the anterior ones) on all tergites except Abd. VI. No special arrangement or morphological modification of scales and chaetae around mac and trichobothria.
</paragraph>
<caption httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="14" pageNumber="263" targetBox="[223,1359,202,1857]" targetPageId="14">
<paragraph blockId="14.[151,1436,1886,2031]" pageId="14" pageNumber="263">
<emphasis bold="true" box="[151,341,1886,1909]" pageId="14" pageNumber="263">FIGURES 4045.</emphasis>
<taxonomicName box="[347,727,1887,1909]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="14" pageNumber="263" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[347,480,1888,1909]" italics="true" pageId="14" pageNumber="263">Heteromurus</emphasis>
(
<emphasis box="[496,625,1887,1909]" italics="true" pageId="14" pageNumber="263">Verhoeffiella</emphasis>
)
<emphasis box="[640,727,1887,1909]" italics="true" pageId="14" pageNumber="263">absoloni</emphasis>
</taxonomicName>
: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42).
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/236236/files/figure.png" pageId="15" pageNumber="264" targetBox="[151,1436,193,1146]" targetPageId="15">
<paragraph blockId="15.[151,1436,1167,1220]" pageId="15" pageNumber="264">
<emphasis bold="true" box="[151,341,1167,1190]" pageId="15" pageNumber="264">FIGURES 4649.</emphasis>
<taxonomicName box="[349,730,1168,1190]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[349,482,1169,1190]" italics="true" pageId="15" pageNumber="264">Heteromurus</emphasis>
(
<emphasis box="[498,627,1168,1190]" italics="true" pageId="15" pageNumber="264">Verhoeffiella</emphasis>
)
<emphasis box="[643,730,1168,1190]" italics="true" pageId="15" pageNumber="264">absoloni</emphasis>
</taxonomicName>
, optical microscope photos of the dens: 46, dens, ventro-distal area; 47, dens; 48, dens thickening; 49, dens internal spines. Scales: 100 µm (4647); 50 µm (49); 10 µm (48).
</paragraph>
</caption>
<caption pageId="15" pageNumber="264">
<paragraph blockId="15.[151,1436,1264,1320]" pageId="15" pageNumber="264">
<emphasis bold="true" box="[151,266,1265,1288]" pageId="15" pageNumber="264">TABLE 2.</emphasis>
Tergite S-chaetotaxy of
<taxonomicName box="[531,700,1264,1288]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="15" pageNumber="264" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[531,558,1265,1288]" italics="true" pageId="15" pageNumber="264">H.</emphasis>
(
<emphasis box="[573,591,1265,1288]" italics="true" pageId="15" pageNumber="264">V.</emphasis>
)
<emphasis box="[608,700,1264,1288]" italics="true" pageId="15" pageNumber="264">absoloni</emphasis>
</taxonomicName>
from the type locality. Mesochaetae are mentioned when they are in tandem with sens, with (sens+mesochaeta) given between brackets.
</paragraph>
</caption>
<paragraph blockId="15.[159,974,1348,1990]" pageId="15" pageNumber="264">
Area on tergite S-chaetae and mesochetae ThII antero-lateral (
<number box="[669,680,1389,1411]" pageId="15" pageNumber="264" value="1.0">1</number>
ms+
<number box="[734,742,1389,1411]" pageId="15" pageNumber="264" value="1.0">1</number>
sens) ThII postero-lateral
<number box="[659,671,1430,1452]" pageId="15" pageNumber="264" value="1.0">1</number>
sens
</paragraph>
<paragraph blockId="15.[159,974,1348,1990]" pageId="15" pageNumber="264">
ThIII antero-lateral (
<number box="[669,680,1472,1494]" pageId="15" pageNumber="264" value="1.0">1</number>
ms+
<number box="[734,742,1472,1494]" pageId="15" pageNumber="264" value="1.0">1</number>
sens) ThIII postero-lateral
<number box="[659,671,1513,1535]" pageId="15" pageNumber="264" value="1.0">1</number>
sens
</paragraph>
<paragraph blockId="15.[159,974,1348,1990]" pageId="15" pageNumber="264">
Abd. I lateral
<number box="[659,671,1554,1576]" pageId="15" pageNumber="264" value="1.0">1</number>
ms,
<number box="[720,732,1554,1576]" pageId="15" pageNumber="264" value="1.0">1</number>
sens Abd. II median
<number box="[659,671,1596,1618]" pageId="15" pageNumber="264" value="1.0">1</number>
sens
</paragraph>
<paragraph blockId="15.[159,974,1348,1990]" pageId="15" pageNumber="264">
Abd. II medio-lateral (
<number box="[669,680,1637,1659]" pageId="15" pageNumber="264" value="1.0">1</number>
sens+
<number box="[748,756,1637,1659]" pageId="15" pageNumber="264" value="1.0">1</number>
mesochaeta) Abd. II lateral
<number box="[659,671,1678,1700]" pageId="15" pageNumber="264" value="1.0">1</number>
sens
</paragraph>
<paragraph blockId="15.[159,974,1348,1990]" box="[159,722,1720,1742]" pageId="15" pageNumber="264">
Abd. III median
<number box="[659,671,1720,1742]" pageId="15" pageNumber="264" value="1.0">1</number>
sens
</paragraph>
<paragraph blockId="15.[159,974,1348,1990]" pageId="15" pageNumber="264">
Abd.III medio-lateral (
<number box="[669,680,1761,1783]" pageId="15" pageNumber="264" value="1.0">1</number>
sens+
<number box="[748,756,1761,1783]" pageId="15" pageNumber="264" value="1.0">1</number>
mesochaeta) Abd. III lateral (
<number box="[669,680,1802,1824]" pageId="15" pageNumber="264" value="1.0">1</number>
ms+
<number box="[734,742,1802,1824]" pageId="15" pageNumber="264" value="1.0">1</number>
sens) Abd. IV median
<number box="[659,671,1844,1866]" pageId="15" pageNumber="264" value="2.0">2</number>
sens,&gt;
<number box="[749,774,1844,1866]" pageId="15" pageNumber="264" value="10.0">10</number>
long S-like chaetae Abd. IV postero-lateral
<number box="[659,671,1885,1907]" pageId="15" pageNumber="264" value="1.0">1</number>
sens
</paragraph>
<paragraph blockId="15.[159,974,1348,1990]" box="[159,722,1926,1948]" pageId="15" pageNumber="264">
Abd. V median
<number box="[659,671,1926,1948]" pageId="15" pageNumber="264" value="2.0">2</number>
sens
</paragraph>
<paragraph blockId="15.[159,974,1348,1990]" box="[159,722,1968,1990]" pageId="15" pageNumber="264">
Abd. V lateral
<number box="[659,671,1968,1990]" pageId="15" pageNumber="264" value="1.0">1</number>
sens
</paragraph>
<paragraph blockId="16.[151,1436,151,573]" pageId="16" pageNumber="265">
Ventral tube with smooth pointed and serrated mes and mac. Posteriorly with about
<number box="[1150,1192,151,176]" pageId="16" pageNumber="265" value="125.0">125</number>
sockets (serrated and smooth mes in unknown proportion, impossible to recognize from their socket once they have fallen) and
<number box="[1394,1436,188,213]" pageId="16" pageNumber="265" value="6.5" valueMax="7.0" valueMin="6.0">67</number>
larger sockets (possibly mac); no scale observed except a single scale in place in one specimen. Anteriorly with
<number pageId="16" pageNumber="265" value="47.5" valueMax="49.0" valueMin="46.0">46 49</number>
+
<number box="[212,282,260,285]" pageId="16" pageNumber="265" value="47.5" valueMax="49.0" valueMin="46.0">4649</number>
sockets (serrated or smooth mes). Each latero-distal flap with
<number box="[998,1026,260,285]" pageId="16" pageNumber="265" value="45.0">45</number>
mes, about
<number box="[1166,1194,260,284]" pageId="16" pageNumber="265" value="12.0">12</number>
of them serrated and others smooth (
<figureCitation box="[324,408,295,320]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[376,408,296,320]" pageId="16" pageNumber="265" value="29.0">29</number>
</figureCitation>
). Genital plate of female with
<number box="[749,763,296,320]" pageId="16" pageNumber="265" value="2.0">2</number>
+
<number box="[779,793,296,320]" pageId="16" pageNumber="265" value="2.0">2</number>
mic (
<figureCitation box="[857,940,295,320]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[909,940,296,320]" pageId="16" pageNumber="265" value="30.0">30</number>
</figureCitation>
), that of adult male not observed.
</paragraph>
<paragraph blockId="16.[151,1436,151,573]" pageId="16" pageNumber="265">
<emphasis bold="true" box="[199,255,332,357]" pageId="16" pageNumber="265">Legs</emphasis>
(
<figureCitation box="[270,402,332,357]" captionStart-0="FIGURES 31 36" captionStart-1="FIGURES 37 39" captionStartId-0="11.[151,264,1915,1938]" captionStartId-1="12.[151,264,1344,1367]" captionTargetBox-0="[152,1424,205,1914]" captionTargetBox-1="[151,1436,193,1322]" captionTargetId-0="figure@11.[151,1436,193,1914]" captionTargetId-1="figure@12.[151,1436,193,1324]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." captionText-1="FIGURES 37 39. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of legs: 37, internal structure of unguis; 38 trochanteral organ; 39, Tita I, posterior view; the oblique ridges visible on the the claw are an artefact. Scales: 50 µm (37, 38); 100 µm (39)." httpUri-0="https://zenodo.org/record/236233/files/figure.png" httpUri-1="https://zenodo.org/record/236234/files/figure.png" pageId="16" pageNumber="265">
Figs
<number box="[328,402,332,356]" pageId="16" pageNumber="265" value="35.0" valueMax="39.0" valueMin="31.0">3139</number>
</figureCitation>
). Unguis relatively thin and elongated, with one inner pair of equal basal teeth at
<number box="[1323,1352,332,357]" pageId="16" pageNumber="265" value="25.0">25</number>
% from its base and one inner unpaired tooth at
<number box="[611,640,367,392]" pageId="16" pageNumber="265" value="50.0">50</number>
% from its base (
<figureCitation box="[836,970,367,392]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">Figs 31, 32</figureCitation>
). Large hyaline lamella covers claw, and bears
<number box="[217,229,404,428]" pageId="16" pageNumber="265" value="1.0">1</number>
+
<number box="[250,262,404,428]" pageId="16" pageNumber="265" value="1.0">1</number>
small latero-basal teeth (
<figureCitation box="[545,677,404,429]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">Figs 31, 32</figureCitation>
). Internal structure of unguis illustrated in
<figureCitation box="[1158,1241,404,429]" captionStart="FIGURES 37 39" captionStartId="12.[151,264,1344,1367]" captionTargetBox="[151,1436,193,1322]" captionTargetId="figure@12.[151,1436,193,1324]" captionTargetPageId="12" captionText="FIGURES 37 39. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of legs: 37, internal structure of unguis; 38 trochanteral organ; 39, Tita I, posterior view; the oblique ridges visible on the the claw are an artefact. Scales: 50 µm (37, 38); 100 µm (39)." httpUri="https://zenodo.org/record/236234/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[1210,1241,404,428]" pageId="16" pageNumber="265" value="37.0">37</number>
</figureCitation>
. Unguis (internal edge): unguiculus =
<number box="[395,439,439,464]" pageId="16" pageNumber="265" value="1.6">1.6</number>
:
<number box="[443,454,439,464]" pageId="16" pageNumber="265" value="1.0">1</number>
. Unguiculus robust, pointed, about
<number box="[876,905,439,464]" pageId="16" pageNumber="265" value="75.0">75</number>
% as long as inner edge of unguis, with two external ridges, without basal lamella (
<figureCitation box="[592,725,476,501]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">Figs 31, 32</figureCitation>
). Small outer tooth, sometimes difficult to see at
<number box="[1287,1316,476,501]" pageId="16" pageNumber="265" value="50.0">50</number>
% from its base on anterior ridge of unguiculus I, II and posterior ridge of unguiculus III (
<figureCitation box="[1032,1163,511,536]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">Figs 31, 32</figureCitation>
).
</paragraph>
<paragraph blockId="16.[151,1436,151,573]" box="[199,1208,548,573]" pageId="16" pageNumber="265">Legs with ordinary chaetae and scales, often detached, including the following categories:</paragraph>
<paragraph blockId="16.[151,1436,620,933]" box="[151,1355,620,645]" pageId="16" pageNumber="265">
serrated mac, cylindrical-pointed, on subcoxae
<number box="[728,744,620,644]" pageId="16" pageNumber="265" value="2.0">2</number>
, coxae and tibiotarsi (similar to
<figureCitation box="[1106,1189,620,645]" captionStart="FIGURES 23 30" captionStartId="10.[151,264,1760,1783]" captionTargetBox="[205,1361,209,1718]" captionTargetId="figure@10.[194,1380,193,1738]" captionTargetPageId="10" captionText="FIGURES 23 30. Heteromurus (Verhoeffiella) absoloni: 23, body chaetotaxy; chaeta notation after Szeptycki (1979); chaetal groups circled with dotted lines; circles—mac, x—trichobothria, crossed circles—pseudopores, lines—S-chaetae, a—anterior mac, an—antennal mac, m—medial mac, sut—sutural mac, p—posterior mac, al—antero-lateral mac, pl—postero-lateral mac; S-like chaetae not given on Abd. IV; 24, sensillar complex of Th. II; 25, sensillar complex of Abd. II; 26, part of sensillar complex of Abd. III; 27, head chaetae: a 1, clubbed mac, a 2, subcylindrical mac, b 1, normal serrated mes, b 2, small serrated mes, c, smooth mes, d, trichobothria; 28, tergite chaetae: a 1, big serrated mac, a 2, lateral pointed mac, b, serrated mes, c, thin serrated mes, d, short pointed S-chaetae, e, thin subcylindrical S-chaetae, f 1, short S-like chaetae on Th. III, f 2, long S-like chaetae on Abd. IV, g, mic; 29, latero-distal flap of ventral tube, anterior side; sockets—smooth mes; 30, female genital plate. Scales: 500 µm (23); 40 µm (29); 20 µm (27, 28, 30); 10 µm (24, 25)." httpUri="https://zenodo.org/record/236232/files/figure.png" pageId="16" pageNumber="265">
Figs
<number box="[1161,1189,620,644]" pageId="16" pageNumber="265" value="27.0">27</number>
</figureCitation>
a
<number box="[1208,1222,620,644]" pageId="16" pageNumber="265" value="2.0">2</number>
and
<number box="[1277,1305,620,644]" pageId="16" pageNumber="265" value="28.0">28</number>
a
<number box="[1326,1342,620,644]" pageId="16" pageNumber="265" value="2.0">2</number>
);
</paragraph>
<paragraph blockId="16.[151,1436,620,933]" box="[151,744,655,680]" pageId="16" pageNumber="265">
serrated mes, thick, on all leg segments (
<figureCitation box="[654,736,655,680]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[707,736,656,680]" pageId="16" pageNumber="265" value="31.0">31</number>
</figureCitation>
)
</paragraph>
<paragraph blockId="16.[151,1436,620,933]" box="[151,741,692,717]" pageId="16" pageNumber="265"> smooth mac, pointed, on femur, trochanter, coxa</paragraph>
<paragraph blockId="16.[151,1436,620,933]" box="[151,903,727,752]" pageId="16" pageNumber="265">
smooth mes, pointed, on Tita, femur, trochanter, coxa (
<figureCitation box="[813,894,727,752]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[865,894,728,752]" pageId="16" pageNumber="265" value="31.0">31</number>
</figureCitation>
)
</paragraph>
<paragraph blockId="16.[151,1436,620,933]" pageId="16" pageNumber="265">
tenent hairs, smooth, acuminate, of similar length but thinner than smooth mes, located dorso-apically on Tita (
<figureCitation box="[209,290,799,824]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[261,290,800,824]" pageId="16" pageNumber="265" value="31.0">31</number>
</figureCitation>
)
</paragraph>
<paragraph blockId="16.[151,1436,620,933]" box="[151,1318,836,861]" pageId="16" pageNumber="265">
smooth, pointed mes, inserted perpendicular to integument, forming the trochanteral organ (
<figureCitation box="[1228,1310,836,861]" captionStart="FIGURES 37 39" captionStartId="12.[151,264,1344,1367]" captionTargetBox="[151,1436,193,1322]" captionTargetId="figure@12.[151,1436,193,1324]" captionTargetPageId="12" captionText="FIGURES 37 39. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of legs: 37, internal structure of unguis; 38 trochanteral organ; 39, Tita I, posterior view; the oblique ridges visible on the the claw are an artefact. Scales: 50 µm (37, 38); 100 µm (39)." httpUri="https://zenodo.org/record/236234/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[1281,1310,836,860]" pageId="16" pageNumber="265" value="38.0">38</number>
</figureCitation>
)
</paragraph>
<paragraph blockId="16.[151,1436,620,933]" box="[151,484,871,896]" pageId="16" pageNumber="265">
mic of pretarsus (
<figureCitation box="[395,476,871,896]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[447,476,872,896]" pageId="16" pageNumber="265" value="31.0">31</number>
</figureCitation>
)
</paragraph>
<paragraph blockId="16.[151,1436,620,933]" box="[151,895,908,933]" pageId="16" pageNumber="265"> scales small, elongated, subequal, present on all leg segments.</paragraph>
<paragraph blockId="16.[151,1437,980,2013]" pageId="16" pageNumber="265">
Detailed distribution of the different
<typeStatus box="[631,690,981,1005]" pageId="16" pageNumber="265">types</typeStatus>
of chaetae on each leg segment as follows: pretarsus with
<number box="[1392,1404,980,1004]" pageId="16" pageNumber="265" value="1.0">1</number>
+
<number box="[1424,1436,980,1004]" pageId="16" pageNumber="265" value="1.0">1</number>
minute mic (
<figureCitation box="[295,379,1015,1040]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[348,379,1016,1040]" pageId="16" pageNumber="265" value="31.0">31</number>
</figureCitation>
). Tibiotarsus with a large number of serrated mes, few serrated mac, smooth mes, thin tenent hair and scales (
<figureCitation box="[330,414,1052,1077]" captionStart="FIGURES 37 39" captionStartId="12.[151,264,1344,1367]" captionTargetBox="[151,1436,193,1322]" captionTargetId="figure@12.[151,1436,193,1324]" captionTargetPageId="12" captionText="FIGURES 37 39. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of legs: 37, internal structure of unguis; 38 trochanteral organ; 39, Tita I, posterior view; the oblique ridges visible on the the claw are an artefact. Scales: 50 µm (37, 38); 100 µm (39)." httpUri="https://zenodo.org/record/236234/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[382,414,1052,1076]" pageId="16" pageNumber="265" value="39.0">39</number>
</figureCitation>
). Distal whorl of Tita with
<number box="[717,786,1052,1076]" pageId="16" pageNumber="265" value="12.5" valueMax="14.0" valueMin="11.0">1114</number>
(usually
<number box="[890,919,1052,1076]" pageId="16" pageNumber="265" value="13.0">13</number>
) chaetae including 5, 5,
<number box="[1193,1207,1052,1076]" pageId="16" pageNumber="265" value="4.0">4</number>
smooth mes and the tenent hair on leg I, II, III. Serrated mac as 1, 3,
<number box="[689,703,1088,1112]" pageId="16" pageNumber="265" value="3.0">3</number>
on Tita I, II, III (
<figureCitation box="[894,1020,1087,1112]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[946,1020,1087,1112]" pageId="16" pageNumber="265" value="34.5" valueMax="36.0" valueMin="33.0">3336</number>
</figureCitation>
). Beside the apical ones, smooth mes present ventrally in two longitudinal rows on Tita I, II, III, anterior one with 3, 36, 5
<number box="[1115,1128,1124,1149]" pageId="16" pageNumber="265" value="7.0">7</number>
chaetae, posterior one with
<number box="[151,166,1159,1184]" pageId="16" pageNumber="265" value="4.0">4</number>
5, 34, 6
<number box="[292,305,1159,1184]" pageId="16" pageNumber="265" value="7.0">7</number>
chaetae (
<figureCitation box="[413,497,1159,1184]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[466,497,1160,1184]" pageId="16" pageNumber="265" value="33.0">33</number>
</figureCitation>
). Few scales present in the proximal third of Tita I, more numerous in the proximal half of Tita II, and many distributed on the whole length of Tita III. Tita of leg III longer than those of legs I and II; length
<number box="[228,280,1232,1256]" pageId="16" pageNumber="265" value="0.4">0.40</number>
,
<number box="[291,343,1232,1256]" pageId="16" pageNumber="265" value="0.49">0.49</number>
,
<quantity box="[354,454,1231,1256]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="16" pageNumber="265" unit="mm" value="0.58">0.58 mm</quantity>
for Tita I, II, III.
</paragraph>
<paragraph blockId="16.[151,1437,980,2013]" pageId="16" pageNumber="265">
Femur with many serrated mes (some almost as large as serrated mac), smooth mac, smooth mes, and scales. Ventrally about
<number box="[332,342,1304,1328]" pageId="16" pageNumber="265" value="1.0">1</number>
2, 12, 1 smooth macrochaetae on femora I, II, III and few additional smooth mes on femora I (
<figureCitation box="[159,292,1340,1365]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Figs
<number box="[217,292,1340,1365]" pageId="16" pageNumber="265" value="35.0" valueMax="36.0" valueMin="34.0">3436</number>
</figureCitation>
). Biggest smooth mac as long as
<number box="[677,789,1340,1365]" pageId="16" pageNumber="265" value="0.4" valueMax="0.45" valueMin="0.35">0.350.45</number>
length of femur. Group of ventro-proximal serrated mes without interspersed scales (
<figureCitation box="[468,552,1375,1400]" captionStart="FIGURES 37 39" captionStartId="12.[151,264,1344,1367]" captionTargetBox="[151,1436,193,1322]" captionTargetId="figure@12.[151,1436,193,1324]" captionTargetPageId="12" captionText="FIGURES 37 39. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of legs: 37, internal structure of unguis; 38 trochanteral organ; 39, Tita I, posterior view; the oblique ridges visible on the the claw are an artefact. Scales: 50 µm (37, 38); 100 µm (39)." httpUri="https://zenodo.org/record/236234/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[520,552,1376,1400]" pageId="16" pageNumber="265" value="38.0">38</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="16.[151,1437,980,2013]" pageId="16" pageNumber="265">
Trochanter with many chaetae ranging in size from mes to mac including many serrated mes,
<number box="[1246,1285,1412,1436]" pageId="16" pageNumber="265" value="1.5" valueMax="2.0" valueMin="1.0">12</number>
smooth mac, and smooth mes. Smooth chaetae about
<number box="[603,659,1448,1472]" pageId="16" pageNumber="265" value="9.5" valueMax="10.0" valueMin="9.0">910</number>
on trochanter I,
<number box="[845,888,1447,1472]" pageId="16" pageNumber="265" value="4.5" valueMax="6.0" valueMin="3.0">36</number>
on trochanter II, many on trochanter III in adult, of which
<number box="[255,360,1484,1509]" pageId="16" pageNumber="265" value="77.5" valueMax="100.0" valueMin="55.0">55 to 100</number>
forming the trochanteral organ of leg III (
<figureCitation box="[826,909,1484,1509]" captionStart="FIGURES 37 39" captionStartId="12.[151,264,1344,1367]" captionTargetBox="[151,1436,193,1322]" captionTargetId="figure@12.[151,1436,193,1324]" captionTargetPageId="12" captionText="FIGURES 37 39. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of legs: 37, internal structure of unguis; 38 trochanteral organ; 39, Tita I, posterior view; the oblique ridges visible on the the claw are an artefact. Scales: 50 µm (37, 38); 100 µm (39)." httpUri="https://zenodo.org/record/236234/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[878,909,1484,1508]" pageId="16" pageNumber="265" value="38.0">38</number>
</figureCitation>
). Scales in small number on trochanter I, II and in large number on trochanter III.
</paragraph>
<paragraph blockId="16.[151,1437,980,2013]" pageId="16" pageNumber="265">
Coxa with pseudopores, thick serrated mes, smooth mes, smooth mac, multiplets of serrated chaetae and scales. Pseudopores as
<number box="[410,420,1592,1616]" pageId="16" pageNumber="265" value="1.0">1</number>
2, 36, 6
<number box="[551,578,1591,1616]" pageId="16" pageNumber="265" value="10.0">10</number>
on coxae I, II, III. Smooth mac as
<number box="[968,984,1592,1616]" pageId="16" pageNumber="265" value="2.0">2</number>
, 0, 0 on dorso-anterior side of coxa I, II, III; multiplet of
<number box="[345,386,1628,1653]" pageId="16" pageNumber="265" value="6.0" valueMax="7.0" valueMin="5.0">57</number>
dorso-posterior serrated chaetae on coxa I, two multiplets (dorso-anterior one and dorsoposterior one) of
<number box="[350,392,1663,1688]" pageId="16" pageNumber="265" value="6.5" valueMax="8.0" valueMin="5.0">58</number>
serrated chaetae on coxa II; chaetae in multiplets of coxae I and II decreasing in size from mac (distally) to mes (proximally) (
<figureCitation box="[552,684,1700,1725]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Figs
<number box="[609,684,1700,1725]" pageId="16" pageNumber="265" value="34.5" valueMax="35.0" valueMin="34.0">3435</number>
</figureCitation>
). Dorsally on coxa III, a transversal line of
<number box="[1174,1188,1700,1724]" pageId="16" pageNumber="265" value="3.0">3</number>
serrated mac and
<number box="[1394,1436,1700,1725]" pageId="16" pageNumber="265" value="6.5" valueMax="8.0" valueMin="5.0">58</number>
serrated mes (
<figureCitation box="[307,391,1735,1760]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[359,391,1735,1760]" pageId="16" pageNumber="265" value="36.0">36</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="16.[151,1437,980,2013]" pageId="16" pageNumber="265">
Separation between subcoxae
<number box="[552,566,1772,1796]" pageId="16" pageNumber="265" value="1.0">1</number>
and
<number box="[630,644,1772,1796]" pageId="16" pageNumber="265" value="2.0">2</number>
indistinct. Subcoxae with scales. Subcoxa I with
<number box="[1239,1253,1772,1796]" pageId="16" pageNumber="265" value="3.0">3</number>
dorso-posterior serrated mac (
<figureCitation box="[308,392,1807,1832]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[360,392,1808,1832]" pageId="16" pageNumber="265" value="34.0">34</number>
</figureCitation>
). Subcoxa II with
<number box="[596,637,1807,1832]" pageId="16" pageNumber="265" value="5.0" valueMax="6.0" valueMin="4.0">46</number>
dorso-anterior serrated chaetae ranging in size from mac to mes and
<number pageId="16" pageNumber="265" value="4.0" valueMax="5.0" valueMin="3.0">3 5</number>
dorso-posterior serrated mac (
<figureCitation box="[517,603,1844,1869]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[572,603,1844,1869]" pageId="16" pageNumber="265" value="35.0">35</number>
</figureCitation>
). Subcoxa III, with group of
<number box="[942,983,1844,1869]" pageId="16" pageNumber="265" value="4.5" valueMax="5.0" valueMin="4.0">45</number>
dorso-posterior serrated mac and some serrated mes (
<figureCitation box="[307,391,1879,1904]" captionStart="FIGURES 31 36" captionStartId="11.[151,264,1915,1938]" captionTargetBox="[152,1424,205,1914]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 31 36. Heteromurus (Verhoeffiella) absoloni.: 31, unguis and unguiculus of leg III, posterior view (scale 20 µm); 32, unguis and unguiculus of leg II (scale 20 µm); 33, Tita II, posterior view chaetae, hollow circles anterior chaetae; 34, leg I, posterior view; 35, Leg II, posterior view, hollow circles—anterior chaetae; 36, leg III, anterior view. Scales: 200 µm (34 36); 100 µm (33); 20 µm (31, 32)." httpUri="https://zenodo.org/record/236233/files/figure.png" pageId="16" pageNumber="265">
Fig.
<number box="[359,391,1879,1904]" pageId="16" pageNumber="265" value="36.0">36</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="16.[151,1437,980,2013]" lastBlockId="17.[151,1437,151,1149]" lastPageId="17" lastPageNumber="266" pageId="16" pageNumber="265">
<emphasis bold="true" box="[199,270,1916,1941]" pageId="16" pageNumber="265">Furca</emphasis>
(
<figureCitation box="[290,426,1916,1941]" captionStart-0="FIGURES 40 45" captionStart-1="FIGURES 46 49" captionStartId-0="14.[151,264,1886,1909]" captionStartId-1="15.[151,264,1167,1190]" captionTargetBox-0="[223,1359,202,1857]" captionTargetBox-1="[151,1436,193,1146]" captionTargetId-0="figure@14.[200,1367,193,1868]" captionTargetId-1="figure@15.[151,1436,193,1146]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." captionText-1="FIGURES 46 49. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of the dens: 46, dens, ventro-distal area; 47, dens; 48, dens thickening; 49, dens internal spines. Scales: 100 µm (46 47); 50 µm (49); 10 µm (48)." httpUri-0="https://zenodo.org/record/236235/files/figure.png" httpUri-1="https://zenodo.org/record/236236/files/figure.png" pageId="16" pageNumber="265">
Figs
<number box="[352,426,1916,1940]" pageId="16" pageNumber="265" value="44.5" valueMax="49.0" valueMin="40.0">4049</number>
</figureCitation>
). Furca with pseudopores, smooth mac, different kinds of serrated mes and scales of various morphologies. Pseudopores as
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dorsally on each side of manubrium, among distal group of chaetae. Manubrium dorsally with two longitudinal stripes of serrated mes, each one interspersed with
<number box="[1262,1290,1988,2012]" pageId="16" pageNumber="265" value="14.0">14</number>
±
<number box="[1328,1342,1988,2012]" pageId="16" pageNumber="265" value="2.0">2</number>
smooth pointed mac (
<figureCitation box="[305,390,151,176]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[358,390,152,176]" pageId="17" pageNumber="266" value="40.0">40</number>
</figureCitation>
); between these stripes, loose cover of scales except in basal
<number box="[1083,1093,152,176]" pageId="17" pageNumber="266" value="1.0">1</number>
/
<number box="[1105,1119,152,176]" pageId="17" pageNumber="266" value="3.0">3</number>
where few serrated mes are present; externally, each longitudinal stripe of mes is bordered by a very narrow stripe devoid of chaetae and scales. Cross-section of manubrium as shown in
<figureCitation box="[630,714,223,248]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[683,714,224,248]" pageId="17" pageNumber="266" value="41.0">41</number>
</figureCitation>
. Manubrium covered ventrally by medium size scales, without chaetae, except an unpaired ventro-basal serrated and thick mac and
<number box="[914,928,260,284]" pageId="17" pageNumber="266" value="2.0">2</number>
+
<number box="[944,958,260,284]" pageId="17" pageNumber="266" value="2.0">2</number>
apical serrated mes near the axis (
<figureCitation box="[1339,1423,260,285]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[1391,1423,260,284]" pageId="17" pageNumber="266" value="42.0">42</number>
</figureCitation>
); scales evenly distributed, except on two ventro-distal areas where scale cover is more dense with larger and more elongated scales (
<figureCitation box="[349,433,332,357]" captionStart="FIGURES 46 49" captionStartId="15.[151,264,1167,1190]" captionTargetBox="[151,1436,193,1146]" captionTargetId="figure@15.[151,1436,193,1146]" captionTargetPageId="15" captionText="FIGURES 46 49. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of the dens: 46, dens, ventro-distal area; 47, dens; 48, dens thickening; 49, dens internal spines. Scales: 100 µm (46 47); 50 µm (49); 10 µm (48)." httpUri="https://zenodo.org/record/236236/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[401,433,332,357]" pageId="17" pageNumber="266" value="46.0">46</number>
</figureCitation>
). Distal manubrial thickening with
<number box="[829,893,332,357]" pageId="17" pageNumber="266" value="3.5" valueMax="5.0" valueMin="2.0">2 to 5</number>
small teeth on each side (
<figureCitation box="[1183,1267,332,357]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[1235,1267,332,356]" pageId="17" pageNumber="266" value="42.0">42</number>
</figureCitation>
).
</paragraph>
<paragraph blockId="17.[151,1437,151,1149]" pageId="17" pageNumber="266">
Dens not articulated, but constituted of a short basal part hardly annulated, a long medial part annulated dorsally and a short, thinner, non annulated distal part about
<number box="[849,863,404,428]" pageId="17" pageNumber="266" value="4.0">4</number>
times as long as mucro (
<figureCitation box="[1154,1288,404,429]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">Figs 43, 45</figureCitation>
); covered by various chaetae and scales arranged in a complex pattern as shown in schematic cross-sections (
<figureCitation box="[1222,1306,439,464]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[1274,1306,440,464]" pageId="17" pageNumber="266" value="44.0">44</number>
</figureCitation>
). All dental chaetae serrated except one smooth mac basally on dorsal part of each dens (
<figureCitation box="[1003,1087,476,501]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[1055,1087,476,500]" pageId="17" pageNumber="266" value="40.0">40</number>
</figureCitation>
). Two
<typeStatus box="[1162,1221,477,501]" pageId="17" pageNumber="266">types</typeStatus>
of serrated chaetae can be recognized (
<figureCitation box="[367,450,511,536]" captionStart="FIGURES 46 49" captionStartId="15.[151,264,1167,1190]" captionTargetBox="[151,1436,193,1146]" captionTargetId="figure@15.[151,1436,193,1146]" captionTargetPageId="15" captionText="FIGURES 46 49. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of the dens: 46, dens, ventro-distal area; 47, dens; 48, dens thickening; 49, dens internal spines. Scales: 100 µm (46 47); 50 µm (49); 10 µm (48)." httpUri="https://zenodo.org/record/236236/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[418,450,512,536]" pageId="17" pageNumber="266" value="47.0">47</number>
</figureCitation>
); the thickest ones arranged dorso-laterally in two longitudinal rows on basal and medial part of dens; the thinner ones, ventro-laterally, very long on basal part and shorter on medial part, arranged in two longitudinal rows, with two irregular lines of serrated chaetae internally on basal part (
<figureCitation box="[1120,1204,583,608]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[1172,1204,584,608]" pageId="17" pageNumber="266" value="44.0">44</number>
</figureCitation>
). Small serrated mes present between rows and ventral scales. Ventrally between two rows of thin serrated chaeta, dense cover of medium size elongated scales, small and rounded on basal part of dens, increasing in size laterally to large, fusiform, pointed apically (
<figureCitation box="[458,543,692,717]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[511,543,692,716]" pageId="17" pageNumber="266" value="44.0">44</number>
</figureCitation>
); scales absent in distal non-annulated part (
<figureCitation box="[1041,1126,692,717]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[1094,1126,692,717]" pageId="17" pageNumber="266" value="45.0">45</number>
</figureCitation>
). Dens thickening with two faint transversal structures (
<figureCitation box="[470,559,727,752]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">Figs 42</figureCitation>
,
<figureCitation box="[574,606,728,752]" captionStart="FIGURES 46 49" captionStartId="15.[151,264,1167,1190]" captionTargetBox="[151,1436,193,1146]" captionTargetId="figure@15.[151,1436,193,1146]" captionTargetPageId="15" captionText="FIGURES 46 49. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of the dens: 46, dens, ventro-distal area; 47, dens; 48, dens thickening; 49, dens internal spines. Scales: 100 µm (46 47); 50 µm (49); 10 µm (48)." httpUri="https://zenodo.org/record/236236/files/figure.png" pageId="17" pageNumber="266">48</figureCitation>
). One short and thick internal spine basally on each dens (
<figureCitation box="[1287,1377,727,752]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">Figs 40</figureCitation>
,
<figureCitation box="[1392,1424,728,752]" captionStart="FIGURES 46 49" captionStartId="15.[151,264,1167,1190]" captionTargetBox="[151,1436,193,1146]" captionTargetId="figure@15.[151,1436,193,1146]" captionTargetPageId="15" captionText="FIGURES 46 49. Heteromurus (Verhoeffiella) absoloni, optical microscope photos of the dens: 46, dens, ventro-distal area; 47, dens; 48, dens thickening; 49, dens internal spines. Scales: 100 µm (46 47); 50 µm (49); 10 µm (48)." httpUri="https://zenodo.org/record/236236/files/figure.png" pageId="17" pageNumber="266">49</figureCitation>
). Ratio manubrium: dens: mucro =
<number box="[551,583,764,788]" pageId="17" pageNumber="266" value="10.0">10</number>
:
<number box="[598,629,764,788]" pageId="17" pageNumber="266" value="14.0">14</number>
:
<number box="[645,683,764,789]" pageId="17" pageNumber="266" value="0.5">0.5</number>
. Mucro bidentate with one basal spine, sometimes not observed (probably detached) (
<figureCitation box="[391,475,799,824]" captionStart="FIGURES 40 45" captionStartId="14.[151,264,1886,1909]" captionTargetBox="[223,1359,202,1857]" captionTargetId="figure@14.[200,1367,193,1868]" captionTargetPageId="14" captionText="FIGURES 40 45. Heteromurus (Verhoeffiella) absoloni: 40, distribution of smooth chaetae on manubrium and proximal part of dens, dorsal view; 41, schematic cross section of manubrium; 42, manubrium and basal part of the dens, ventro-distal sclerification, chaetae and scale insertions; 43, dens, dorsal side; 44, schematic cross sections of dens: a, proximal part, b, medial part, c, distal part; 45, distal part of the dens and mucro. Scales: 200 µm (40), 100 µm (41, 43); 50 µm (44); 40 µm (45); 20 µm (42)." httpUri="https://zenodo.org/record/236235/files/figure.png" pageId="17" pageNumber="266">
Fig.
<number box="[443,475,799,824]" pageId="17" pageNumber="266" value="45.0">45</number>
</figureCitation>
). Tenaculum with
<number box="[682,696,800,824]" pageId="17" pageNumber="266" value="1.0">1</number>
serrated chaeta and
<number box="[925,939,800,824]" pageId="17" pageNumber="266" value="4.0">4</number>
teeth on each ramus.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="266" type="discussion">
<paragraph blockId="17.[151,1437,151,1149]" pageId="17" pageNumber="266">
<emphasis bold="true" box="[199,332,836,861]" pageId="17" pageNumber="266">Discussion.</emphasis>
The extensive re-description of
<taxonomicName box="[708,891,836,861]" class="Entognatha" family="Entomobryidae" genus="Heteromurus" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="266" phylum="Arthropoda" rank="species" species="Absoloni" subGenus="Verhoeffiella">
<emphasis box="[708,728,837,860]" italics="true" pageId="17" pageNumber="266">H</emphasis>
. (
<emphasis box="[755,768,837,860]" italics="true" pageId="17" pageNumber="266">V</emphasis>
.)
<emphasis box="[795,891,836,861]" italics="true" pageId="17" pageNumber="266">absoloni</emphasis>
</taxonomicName>
given here is the first detailed description of a species of the subgenus, all existing ones being very succinct and lacking most characters used in modern taxonomy of
<taxonomicName box="[314,490,908,933]" class="Entognatha" family="Entomobryidae" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="266" phylum="Arthropoda" rank="family">Entomobryidae</taxonomicName>
. Several morphological features, described for the first time for the subgenus, revealed interesting characters, e.g. the presence of a chaeta ms on Th. III. It was observed in two specimens only, but S-chaetae are often very difficult to detect; if it is not an aberration, it would be unique among Entomobryoidea. The detailed description of antennal chaetotaxy shows a huge diversity in chaetae morphological
<typeStatus box="[1298,1360,1016,1040]" pageId="17" pageNumber="266">types</typeStatus>
. Their distribution on antennae follows recognizable but loose patterns, as chaetae-to-chaetae homologies are not possible, with a few exceptions, between adults of a same species or between whorls of Ant. IIIIV. The descriptive framework set up in this paper will be used as a reference in the ongoing subgenus revision.
</paragraph>
</subSubSection>
</treatment>
</document>