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<document id="712A4FB24D442080AB6C94EF3801500C" ID-CLB-Dataset="55748" ID-DOI="10.5281/zenodo.5301289" ID-GBIF-Dataset="55b2b0bc-d79f-48e3-8be1-81424185dea7" ID-ISSN="0253-116X" ID-Zenodo-Dep="5301289" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630101010072" checkinUser="marcus" docAuthor="Assing, V." docDate="2013" docId="1974879EFFEBFFF8FF04E9F7FCEAFC6D" docLanguage="en" docName="LinzerbiolBeitr.2013.45.2.1479-1520.pdf" docOrigin="Linzer biologische Beiträge 45 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Micrillus suturalis Assing 2013, nov.comb." docType="treatment" docVersion="9" lastPageNumber="1512" masterDocId="E54DFFE6FFF5FFD9FFB6EA15FFE8FFA6" masterDocTitle="On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="1520" masterPageNumber="1479" pageNumber="1509" updateTime="1699152599027" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CC89EFC9F6FADDFDEABA741815DD64FD">On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:date id="05ED162EA2EF898403645FFE6D653602">2013</mods:date>
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<treatment id="1974879EFFEBFFF8FF04E9F7FCEAFC6D" ID-DOI="http://doi.org/10.5281/zenodo.6512346" ID-GBIF-Taxon="194955322" ID-Zenodo-Dep="6512346" LSID="urn:lsid:plazi:treatment:1974879EFFEBFFF8FF04E9F7FCEAFC6D" httpUri="http://treatment.plazi.org/id/1974879EFFEBFFF8FF04E9F7FCEAFC6D" lastPageId="33" lastPageNumber="1512" pageId="30" pageNumber="1509">
<subSubSection id="D9C76503FFEBFFC7FF04E9F7FC96FC5E" box="[178,894,994,1016]" pageId="30" pageNumber="1509" type="nomenclature">
<paragraph id="91623688FFEBFFC7FF04E9F7FC96FC5E" blockId="30.[178,894,994,1016]" box="[178,894,994,1016]" pageId="30" pageNumber="1509">
<heading id="CA2A81E4FFEBFFC7FF04E9F7FC96FC5E" box="[178,894,994,1016]" fontSize="8" level="2" pageId="30" pageNumber="1509" reason="2">
<taxonomicName id="56DD4D0BFFEBFFC7FF04E9F7FDD8FC5E" ID-CoL="9PLZJ" authority="(CAMERON 1930)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1930" box="[178,560,994,1016]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="suturalis" status="comb. nov.">
<emphasis id="A3A9EA9AFFEBFFC7FF04E9F7FDD8FC5E" bold="true" box="[178,560,994,1016]" pageId="30" pageNumber="1509">
<emphasis id="A3A9EA9AFFEBFFC7FF04E9F7FE9BFC5E" bold="true" box="[178,371,994,1016]" italics="true" pageId="30" pageNumber="1509">Micrillus suturalis</emphasis>
(
<bibRefCitation id="F54C4B79FFEBFFC7FE37E9F7FE07FC51" author="CAMERON M" box="[385,495,994,1016]" pageId="30" pageNumber="1509" pagination="154 - 159" refId="ref22812" refString="CAMERON M. (1930): VIII. New Staphylinidae from the Malay Peninsula. - Journal of the Federated Malay States Museums 16: 154 - 159." type="journal article" year="1930">CAMERON</bibRefCitation>
1930)
</emphasis>
</taxonomicName>
,
<emphasis id="A3A9EA9AFFEBFFC7FD8BE9F7FD45FC5E" bold="true" box="[573,685,994,1016]" pageId="30" pageNumber="1509">
<taxonomicNameLabel id="B89A57E1FFEBFFC7FD8BE9F7FD45FC5E" box="[573,685,994,1016]" pageId="30" pageNumber="1509" rank="species">nov.comb.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="09E62A0DFFEBFFC7FD0CE9F7FCC3FC5E" box="[698,811,994,1016]" captionStart="Figs 48-55" captionStartId="29.[178,218,1520,1540]" captionTargetBox="[182,1085,213,1495]" captionTargetId="figure-61@29.[178,1085,212,1497]" captionTargetPageId="29" captionText="Figs 48-55: Micrillus suturalis (CAMERON) (holotypes of Scymbalium borneense CAMERON (48- 49, 51, 54-55) and S. javanicum CAMERON (50-53)): (48) habitus; (49-50) forebody; (51) male sternite VIII; (52-55) aedeagus in lateral and in ventral view. Scale bars: 48-50: 1.0 mm; 51-55: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5301309" httpUri="https://zenodo.org/record/5301309/files/figure.png" pageId="30" pageNumber="1509">Figs 48-55</figureCitation>
,
<figureCitation id="09E62A0DFFEBFFC7FC83E9F6FC9FFC5E" box="[821,887,994,1016]" captionStart="Map 3" captionStartId="17.[178,223,1423,1443]" captionTargetBox="[178,1085,552,1406]" captionTargetId="figure-217@17.[178,1085,552,1406]" captionTargetPageId="17" captionText="Map 3: Distributions of Micrillus indicus (EPPELSHEIM) (triangles), M. nigriceps (CAMERON) (diamonds), M. intermedius (CAMERON) (open and filled circles), and M. siamensis nov.sp. (filled circle) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.6512094" httpUri="https://zenodo.org/record/6512094/files/figure.png" pageId="30" pageNumber="1509">Map 3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFEBFFC7FF04EE1CFCDBFBF7" pageId="30" pageNumber="1509" type="reference_group">
<paragraph id="91623688FFEBFFC7FF04EE1CFD81FBBB" blockId="30.[178,1086,1033,1389]" box="[178,617,1033,1053]" pageId="30" pageNumber="1509">
<treatmentCitationGroup id="B1CD11A6FFEBFFC7FF04EE1CFD81FBBB" box="[178,617,1033,1053]" pageId="30" pageNumber="1509">
<treatmentCitation id="107C1099FFEBFFC7FF04EE1CFD8DFBBB" author="CAMERON M" box="[178,613,1033,1053]" page="155" pageId="30" pageNumber="1509" year="1930">
<taxonomicName id="56DD4D0BFFEBFFC7FF04EE1CFD8DFBBB" authority="CAMERON 1930: 155" authorityName="Cameron" authorityPageNumber="155" authorityYear="1930" box="[178,613,1033,1053]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="suturale">
<emphasis id="A3A9EA9AFFEBFFC7FF04EE1CFEF5FBBB" box="[178,285,1033,1053]" italics="true" pageId="30" pageNumber="1509">Scimbalium</emphasis>
[sic]
<emphasis id="A3A9EA9AFFEBFFC7FEE7EE1CFE73FBBB" box="[337,411,1033,1053]" italics="true" pageId="30" pageNumber="1509">suturale</emphasis>
<bibRefCitation id="F54C4B79FFEBFFC7FE17EE1CFD8DFBBB" author="CAMERON M" box="[417,613,1033,1053]" pageId="30" pageNumber="1509" pagination="154 - 159" refId="ref22812" refString="CAMERON M. (1930): VIII. New Staphylinidae from the Malay Peninsula. - Journal of the Federated Malay States Museums 16: 154 - 159." type="journal article" year="1930">CAMERON 1930: 155</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="91623688FFEBFFC7FF04EE36FD5FFB91" blockId="30.[178,1086,1033,1389]" box="[178,695,1059,1079]" pageId="30" pageNumber="1509">
<treatmentCitationGroup id="B1CD11A6FFEBFFC7FF04EE36FD5FFB91" box="[178,695,1059,1079]" pageId="30" pageNumber="1509">
<treatmentCitation id="107C1099FFEBFFC7FF04EE36FDA1FB91" author="CAMERON M" box="[178,585,1059,1079]" page="53" pageId="30" pageNumber="1509" year="1936">
<taxonomicName id="56DD4D0BFFEBFFC7FF04EE36FDA1FB91" authority="CAMERON 1936 b: 53" authorityName="Cameron" authorityPageNumber="53" authorityYear="1936" box="[178,585,1059,1079]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="javanicum">
<emphasis id="A3A9EA9AFFEBFFC7FF04EE36FE6AFB91" box="[178,386,1059,1079]" italics="true" pageId="30" pageNumber="1509">Scimbalium javanicum</emphasis>
<bibRefCitation id="F54C4B79FFEBFFC7FE31EE36FDA1FB91" author="CAMERON M" box="[391,585,1059,1079]" pageId="30" pageNumber="1509" pagination="25 - 54" refId="ref22944" refString="CAMERON M (1936 b) Fauna javanica. The Staphylinidae (Col.) collected by Mr. F. C. Drescher. - Tijdschrift voor Entomologie 79: 25 - 54." type="journal article" year="1936">
CAMERON 1936b:
<specimenCount id="87DBFD01FFEBFFC7FD84EE36FDA1FB91" box="[562,585,1059,1079]" pageId="30" pageNumber="1509" type="generic">53</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.;
<emphasis id="A3A9EA9AFFEBFFC7FDDEEE36FD5FFB91" bold="true" box="[616,695,1059,1079]" pageId="30" pageNumber="1509">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="91623688FFEBFFC7FF04EE28FCDBFBF7" blockId="30.[178,1086,1033,1389]" box="[178,819,1085,1105]" pageId="30" pageNumber="1509">
<treatmentCitationGroup id="B1CD11A6FFEBFFC7FF04EE28FCDBFBF7" box="[178,819,1085,1105]" pageId="30" pageNumber="1509">
<treatmentCitation id="107C1099FFEBFFC7FF04EE28FD33FBF7" author="CAMERON M" box="[178,731,1085,1105]" page="230" pageId="30" pageNumber="1509" year="1941">
<taxonomicName id="56DD4D0BFFEBFFC7FF04EE28FD33FBF7" authority="CAMERON 1941: 230" authorityName=", CAMERON" authorityPageNumber="230" authorityYear="1941" box="[178,731,1085,1105]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="borneense" subGenus="Schatzmayria">
<emphasis id="A3A9EA9AFFEBFFC7FF04EE28FEF5FBF7" box="[178,285,1085,1105]" italics="true" pageId="30" pageNumber="1509">Scimbalium</emphasis>
(
<emphasis id="A3A9EA9AFFEBFFC7FE9CEE28FE4EFBF7" box="[298,422,1085,1105]" italics="true" pageId="30" pageNumber="1509">Schatzmayria</emphasis>
)
<emphasis id="A3A9EA9AFFEBFFC7FE05EE28FDF8FBF7" box="[435,528,1085,1105]" italics="true" pageId="30" pageNumber="1509">borneense</emphasis>
<bibRefCitation id="F54C4B79FFEBFFC7FDA0EE28FD33FBF7" author="CAMERON M" box="[534,731,1085,1105]" pageId="30" pageNumber="1509" pagination="203 - 206" refId="ref23007" refString="CAMERON M. (1941): New species of Staphylinidae (Col.) from Borneo. - The Entomologist's Monthly Magazine 77: 203 - 206." type="journal article" year="1941">CAMERON 1941: 230</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<emphasis id="A3A9EA9AFFEBFFC7FD52EE28FCDBFBF7" bold="true" box="[740,819,1085,1105]" pageId="30" pageNumber="1509">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFEBFFC7FF04EE43FC3FFACB" pageId="30" pageNumber="1509" type="reference_group">
<paragraph id="91623688FFEBFFC7FF04EE43FDD7FB09" blockId="30.[178,1086,1033,1389]" pageId="30" pageNumber="1509">
Type material examined
<taxonomicName id="56DD4D0BFFEBFFC7FDF8EE43FD58FBCC" authorityName="Cameron" authorityYear="1930" box="[590,688,1110,1130]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="suturale">
<emphasis id="A3A9EA9AFFEBFFC7FDF8EE43FD58FBCC" box="[590,688,1110,1130]" italics="true" pageId="30" pageNumber="1509">S. suturale</emphasis>
</taxonomicName>
:
<materialsCitation id="21B53CD5FFEBFFC7FD08EE43FDD7FB09" ID-GBIF-Occurrence="3760604359" collectingDate="1928-03-30" collectionCode="BMNH" determinerName="V. Assing" location="Alor Star" municipality="Alor Star" pageId="30" pageNumber="1509" specimenCount="1" stateProvince="Malay Penin." typeStatus="holotype">
<typeStatus id="4E66882AFFEBFFC7FD08EE43FCFAFBCC" box="[702,786,1110,1130]" pageId="30" pageNumber="1509" type="holotype">Holotype</typeStatus>
: &quot;
<collectingRegion id="5319F86AFFEBFFC7FCF4EE43FC2AFBCC" box="[834,962,1110,1130]" pageId="30" pageNumber="1509">Malay Penin.</collectingRegion>
,
<collectingRegion id="5319F86AFFEBFFC7FC7DEE43FBE1FBCC" box="[971,1033,1110,1130]" country="Malaysia" name="Kedah" pageId="30" pageNumber="1509">Kedah</collectingRegion>
,
<collectingMunicipality id="7106ACF2FFEBFFC7FBA5EE43FF31FB27" name="Alor Star" pageId="30" pageNumber="1509" stateProvince="Kedah">Alor Star</collectingMunicipality>
, /
<date id="E5631048FFEBFFC7FF44EE78FE7DFB27" box="[242,405,1133,1153]" pageId="30" pageNumber="1509" value="1928-03-30">
<collectingDate id="F527E9A0FFEBFFC7FF44EE78FE7DFB27" box="[242,405,1133,1153]" pageId="30" pageNumber="1509" value="1928-03-30">March 30th 1928</collectingDate>
</date>
. /
<taxonomicName id="56DD4D0BFFEBFFC7FE18EE78FD4DFB27" authority="Cam." authorityName="Cameron" authorityYear="1930" box="[430,677,1133,1153]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="suturale">
<originalDetermination id="B2C38E74FFEBFFC7FE18EE78FD4DFB27" box="[430,677,1133,1153]" pageId="30" pageNumber="1509">Scimbalium suturale Cam.</originalDetermination>
</taxonomicName>
Type / Type / M. Cameron. Bequest. B.M. 1955-147. / Holotypus
<taxonomicName id="56DD4D0BFFEBFFC7FE38EE91FD58FB3E" authority="Cameron" authorityName="Cameron" authorityYear="1930" box="[398,688,1156,1176]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="suturale">Scymbalium suturale Cameron</taxonomicName>
, rev. V. Assing 2013 /
<taxonomicName id="56DD4D0BFFEBFFC7FC22EE91FEFFFB09" ID-CoL="9PLZJ" authority="(Cameron)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1930" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="suturalis">Micrillus suturalis (Cameron)</taxonomicName>
, det.
<determinerName id="64BEDA4CFFEBFFC7FEF1EE8EFE4AFB09" box="[327,418,1179,1199]" pageId="30" pageNumber="1509">V. Assing</determinerName>
2013&quot; (
<collectionCode id="F7CCAE4DFFEBFFC7FE5DEE8EFDDCFB09" box="[491,564,1179,1199]" pageId="30" pageNumber="1509">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="91623688FFEBFFC7FF04EEA4FE39FAAC" blockId="30.[178,1086,1033,1389]" pageId="30" pageNumber="1509">
<taxonomicName id="56DD4D0BFFEBFFC7FF04EEA4FED9FB63" authorityName="Cameron" authorityYear="1936" box="[178,305,1201,1221]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="javanicum">
<emphasis id="A3A9EA9AFFEBFFC7FF04EEA4FED9FB63" box="[178,305,1201,1221]" italics="true" pageId="30" pageNumber="1509">S. javanicum</emphasis>
</taxonomicName>
:
<materialsCitation id="21B53CD5FFEBFFC7FEF3EEA4FE39FAAC" ID-GBIF-Occurrence="3760604319" collectingDate="1929-03-22" collectionCode="BMNH" collectorName="F. C. Drescher &amp; V. Assing" county="Scimbalium" determinerName="V. Assing" location="Type" municipality="Type" pageId="30" pageNumber="1509" specimenCount="2" stateProvince="Jawa Timur" typeStatus="holotype">
<typeStatus id="4E66882AFFEBFFC7FEF3EEA4FE71FB63" box="[325,409,1201,1221]" pageId="30" pageNumber="1509" type="holotype">Holotype</typeStatus>
: &quot;F. C. Drescher, Preanger, Bandoeng [today Bandung] Dago,
<date id="E5631048FFEBFFC7FF04EEDDFEF1FB7A" box="[178,281,1224,1244]" pageId="30" pageNumber="1509" value="1929-03-22">
<collectingDate id="F527E9A0FFEBFFC7FF04EEDDFEF1FB7A" box="[178,281,1224,1244]" pageId="30" pageNumber="1509" value="1929-03-22">22.III.1929</collectingDate>
</date>
/
<collectingRegion id="5319F86AFFEBFFC7FE99EEDDFEB3FB7A" box="[303,347,1224,1244]" country="Indonesia" name="Jawa Timur" pageId="30" pageNumber="1509">Java</collectingRegion>
:
<collectorName id="3C28535EFFEBFFC7FED0EEDDFE01FB7A" box="[358,489,1224,1244]" pageId="30" pageNumber="1509">F.C. Drescher</collectorName>
, B.M. 1934.-264. /
<taxonomicName id="56DD4D0BFFEBFFC7FD13EEDDFC5BFB7A" authority="Cam." authorityName="Cameron" authorityYear="1936" box="[677,947,1224,1244]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="javanicum">Scimbalium javanicum Cam.</taxonomicName>
Type / Type / Holotypus
<taxonomicName id="56DD4D0BFFEBFFC7FEAEEECAFDA7FB55" authority="Cameron" authorityName="Cameron" authorityYear="1936" box="[280,591,1247,1267]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="javanicum">Scymbalium javanicum Cameron</taxonomicName>
, rev.
<collectorName id="3C28535EFFEBFFC7FD32EECAFD09FB55" box="[644,737,1247,1267]" pageId="30" pageNumber="1509">V. Assing</collectorName>
2013/
<taxonomicName id="56DD4D0BFFEBFFC7FC93EECAFBD1FB55" ID-CoL="9PLZJ" authority="(Cameron)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1930" box="[805,1081,1247,1267]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="suturalis">Micrillus suturalis (Cameron)</taxonomicName>
, det.
<determinerName id="64BEDA4CFFEBFFC7FF6FEEE3FEDCFAAC" box="[217,308,1270,1290]" pageId="30" pageNumber="1509">V. Assing</determinerName>
2013&quot; (
<collectionCode id="F7CCAE4DFFEBFFC7FECBEEE3FE2EFAAC" box="[381,454,1270,1290]" pageId="30" pageNumber="1509">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="91623688FFEBFFC7FF04EF19FD02FAE8" blockId="30.[178,1086,1033,1389]" pageId="30" pageNumber="1509">
<taxonomicName id="56DD4D0BFFEBFFC7FF04EF19FEC0FA86" authorityName=", CAMERON" authorityYear="1941" box="[178,296,1292,1312]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="borneense">
<emphasis id="A3A9EA9AFFEBFFC7FF04EF19FEC0FA86" box="[178,296,1292,1312]" italics="true" pageId="30" pageNumber="1509">S. borneense</emphasis>
</taxonomicName>
:
<materialsCitation id="21B53CD5FFEBFFC7FE81EF19FD02FAE8" ID-GBIF-Occurrence="3760604424" collectionCode="BMNH" collectorName="M. Cameron. Bequest. &amp; V. Assing" determinerName="V. Assing" pageId="30" pageNumber="1509" specimenCount="2" typeStatus="holotype">
<typeStatus id="4E66882AFFEBFFC7FE81EF19FE63FA86" box="[311,395,1292,1312]" pageId="30" pageNumber="1509" type="holotype">Holotype</typeStatus>
: &quot;Pontianak, Borneo /
<taxonomicName id="56DD4D0BFFEBFFC7FD36EF19FCD8FA86" authority="Cam." authorityName=", CAMERON" authorityYear="1941" box="[640,816,1292,1312]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="borneense">S. borneense Cam.</taxonomicName>
Type / Type /
<collectorName id="3C28535EFFEBFFC7FC74EF19FEEAFA91" pageId="30" pageNumber="1509">M. Cameron. Bequest.</collectorName>
B.M. 1955-147. / Holotypus
<taxonomicName id="56DD4D0BFFEBFFC7FD95EF36FCB4FA91" authority="Cameron" authorityName=", CAMERON" authorityYear="1941" box="[547,860,1315,1335]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="borneense">Scymbalium borneense Cameron</taxonomicName>
, rev.
<collectorName id="3C28535EFFEBFFC7FC20EF36FC1DFA91" box="[918,1013,1315,1335]" pageId="30" pageNumber="1509">V. Assing</collectorName>
2013 /
<taxonomicName id="56DD4D0BFFEBFFC7FF04EF2FFE2AFAE8" ID-CoL="9PLZJ" authority="(Cameron)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1930" box="[178,450,1338,1358]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="suturalis">Micrillus suturalis (Cameron)</taxonomicName>
, det.
<determinerName id="64BEDA4CFFEBFFC7FE45EF2FFDA6FAE8" box="[499,590,1338,1358]" pageId="30" pageNumber="1509">V. Assing</determinerName>
2013&quot; (
<collectionCode id="F7CCAE4DFFEBFFC7FD20EF2FFD37FAE8" box="[662,735,1338,1358]" pageId="30" pageNumber="1509">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="91623688FFEBFFC7FF04EF4CFC3FFACB" blockId="30.[178,1086,1033,1389]" box="[178,983,1369,1389]" pageId="30" pageNumber="1509">
Additional material examined:
<materialsCitation id="21B53CD5FFEBFFC7FD12EF4CFC3FFACB" ID-GBIF-Occurrence="3760604407" box="[676,983,1369,1389]" pageId="30" pageNumber="1509">
<collectingCountry id="E9CA7618FFEBFFC7FD12EF4CFD14FACB" box="[676,764,1369,1389]" name="Thailand" pageId="30" pageNumber="1509">Thailand</collectingCountry>
: 1,
<collectingRegion id="5319F86AFFEBFFC7FC9DEF4CFC92FACB" box="[811,890,1369,1389]" country="Thailand" name="Krung Thep Mahanakhon" pageId="30" pageNumber="1509">Bangkok</collectingRegion>
(BMNH).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFEBFFC6FF04EF6EFCDEFD9C" lastPageId="31" lastPageNumber="1510" pageId="30" pageNumber="1509" type="description">
<paragraph id="91623688FFEBFFC7FF04EF6EFEC2FA4F" blockId="30.[178,1085,1403,1513]" pageId="30" pageNumber="1509">
Comment: The original description of
<taxonomicName id="56DD4D0BFFEBFFC7FD32EF6EFCB4FA37" authorityName="Cameron" authorityYear="1930" box="[644,860,1403,1425]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="suturale">
<emphasis id="A3A9EA9AFFEBFFC7FD32EF6EFCB4FA37" box="[644,860,1403,1425]" italics="true" pageId="30" pageNumber="1509">Scymbalium suturale</emphasis>
</taxonomicName>
is based on a unique
<typeStatus id="4E66882AFFEBFFC7FF04EF8DFEE3FA08" box="[178,267,1432,1454]" pageId="30" pageNumber="1509" type="holotype">holotype</typeStatus>
from &quot;Malay Peninsula.
<collectingRegion id="5319F86AFFEBFFC7FDA1EF8DFDB4FA08" box="[535,604,1432,1454]" country="Malaysia" name="Kedah" pageId="30" pageNumber="1509">Kedah</collectingRegion>
; Alor Star&quot; (
<bibRefCitation id="F54C4B79FFEBFFC7FD5EEF8DFC79FA08" author="CAMERON M" box="[744,913,1432,1454]" pageId="30" pageNumber="1509" pagination="154 - 159" refId="ref22812" refString="CAMERON M. (1930): VIII. New Staphylinidae from the Malay Peninsula. - Journal of the Federated Malay States Museums 16: 154 - 159." type="journal article" year="1930">CAMERON 1930</bibRefCitation>
).
<collectingMunicipality id="7106ACF2FFEBFFC7FC10EF8DFC25FA08" box="[934,973,1432,1454]" pageId="30" pageNumber="1509">The</collectingMunicipality>
specimen, unfortunately a female in rather poor condition, was located in the
<location id="94026053FFEBFFC7FCEBEFA3FC51FA6A" LSID="urn:lsid:plazi:treatment:1974879EFFEBFFF8FF04E9F7FCEAFC6D:94026053FFEBFFC7FCEBEFA3FC51FA6A" box="[861,953,1462,1484]" municipality="The" name="Cameron" pageId="30" pageNumber="1509" stateProvince="Kedah">Cameron</location>
collection at the BMNH.
</paragraph>
<paragraph id="91623688FFEBFFC6FF04EFEDFF1BFE25" blockId="30.[178,1085,1528,1609]" lastBlockId="31.[178,1086,218,387]" lastPageId="31" lastPageNumber="1510" pageId="30" pageNumber="1509">
The original description of
<taxonomicName id="56DD4D0BFFEBFFC7FE7AEFECFDB8F9A8" authorityName="Cameron" authorityYear="1936" box="[460,592,1529,1550]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="javanicum">
<emphasis id="A3A9EA9AFFEBFFC7FE7AEFECFDB8F9A8" box="[460,592,1529,1550]" italics="true" pageId="30" pageNumber="1509">S. javanicum</emphasis>
</taxonomicName>
is based on a unique
<typeStatus id="4E66882AFFEBFFC7FC85EFEDFC64F9A8" box="[819,908,1528,1550]" pageId="30" pageNumber="1509" type="holotype">holotype</typeStatus>
from &quot;Bandoeng Dago, Preanger&quot; (Java) (
<bibRefCitation id="F54C4B79FFEBFFC7FE00EC03FD82F98A" author="CAMERON M" box="[438,618,1558,1580]" pageId="30" pageNumber="1509" pagination="25 - 54" refId="ref22944" refString="CAMERON M (1936 b) Fauna javanica. The Staphylinidae (Col.) collected by Mr. F. C. Drescher. - Tijdschrift voor Entomologie 79: 25 - 54." type="journal article" year="1936">CAMERON 1936b</bibRefCitation>
), that of
<taxonomicName id="56DD4D0BFFEBFFC7FD79EC03FCBBF98A" authorityName=", CAMERON" authorityYear="1941" box="[719,851,1558,1580]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1509" phylum="Arthropoda" rank="species" species="borneense">
<emphasis id="A3A9EA9AFFEBFFC7FD79EC03FCBBF98A" box="[719,851,1558,1580]" italics="true" pageId="30" pageNumber="1509">S. borneense</emphasis>
</taxonomicName>
on a unique
<typeStatus id="4E66882AFFEBFFC7FC52EC03FBD5F98A" box="[996,1085,1558,1580]" pageId="30" pageNumber="1509" type="holotype">holotype</typeStatus>
from &quot;W. Borneo: Pontianak&quot; (
<bibRefCitation id="F54C4B79FFEBFFC7FDB4EC26FD44F9EF" author="CAMERON M" box="[514,684,1587,1609]" pageId="30" pageNumber="1509" pagination="203 - 206" refId="ref23007" refString="CAMERON M. (1941): New species of Staphylinidae (Col.) from Borneo. - The Entomologist's Monthly Magazine 77: 203 - 206." type="journal article" year="1941">CAMERON 1941</bibRefCitation>
). Both
<typeStatus id="4E66882AFFEBFFC7FD49EC26FC8AF9EF" box="[767,866,1587,1609]" pageId="30" pageNumber="1509" type="holotype">holotypes</typeStatus>
were located in the Cameron collection at the BMNH. In the description of
<taxonomicName id="56DD4D0BFFEAFFC6FD4EEACEFBD5FF56" authority=", CAMERON (1941)" authorityName=", CAMERON" authorityYear="1941" box="[760,1085,218,240]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="borneense">
<emphasis id="A3A9EA9AFFEAFFC6FD4EEACEFC93FF56" box="[760,891,218,240]" italics="true" pageId="31" pageNumber="1510">S. borneense</emphasis>
,
<bibRefCitation id="F54C4B79FFEAFFC6FC3CEACFFBD5FF56" author="CAMERON M" box="[906,1085,218,240]" pageId="31" pageNumber="1510" pagination="203 - 206" refId="ref23007" refString="CAMERON M. (1941): New species of Staphylinidae (Col.) from Borneo. - The Entomologist's Monthly Magazine 77: 203 - 206." type="journal article" year="1941">CAMERON (1941)</bibRefCitation>
</taxonomicName>
compares the species with
<taxonomicName id="56DD4D0BFFEAFFC6FE77EAEDFCD4FEA8" authority="CAMERON 1930" authorityName="Cameron" authorityYear="1930" box="[449,828,248,270]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="suturale">
<emphasis id="A3A9EA9AFFEAFFC6FE77EAEDFD7EFEA8" box="[449,662,248,270]" italics="true" pageId="31" pageNumber="1510">Scymbalium suturale</emphasis>
<bibRefCitation id="F54C4B79FFEAFFC6FD2AEAEDFCD4FEA8" author="CAMERON M" box="[668,828,248,270]" pageId="31" pageNumber="1510" pagination="154 - 159" refId="ref22812" refString="CAMERON M. (1930): VIII. New Staphylinidae from the Malay Peninsula. - Journal of the Federated Malay States Museums 16: 154 - 159." type="journal article" year="1930">CAMERON 1930</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="E9CA7618FFEAFFC6FCCCEAEDFC32FEA8" box="[890,986,248,270]" name="Malaysia" pageId="31" pageNumber="1510">Malaysia</collectingCountry>
, but there is no reference to
<taxonomicName id="56DD4D0BFFEAFFC6FEDBEB03FE18FE8D" authorityName="Cameron" authorityYear="1936" box="[365,496,278,299]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="javanicum">
<emphasis id="A3A9EA9AFFEAFFC6FEDBEB03FE18FE8D" box="[365,496,278,299]" italics="true" pageId="31" pageNumber="1510">S. javanicum</emphasis>
</taxonomicName>
. An examination of the type material, however, revealed that both
<typeStatus id="4E66882AFFEAFFC6FEA2EB27FE9EFEEE" box="[276,374,306,328]" pageId="31" pageNumber="1510" type="holotype">holotypes</typeStatus>
are undoubtedly conspecific; the male primary and secondary sexual characters are identical (
<figureCitation id="09E62A0DFFEAFFC6FE18EB45FDC9FEC0" box="[430,545,336,358]" captionStart="Figs 48-55" captionStartId="29.[178,218,1520,1540]" captionTargetBox="[182,1085,213,1495]" captionTargetId="figure-61@29.[178,1085,212,1497]" captionTargetPageId="29" captionText="Figs 48-55: Micrillus suturalis (CAMERON) (holotypes of Scymbalium borneense CAMERON (48- 49, 51, 54-55) and S. javanicum CAMERON (50-53)): (48) habitus; (49-50) forebody; (51) male sternite VIII; (52-55) aedeagus in lateral and in ventral view. Scale bars: 48-50: 1.0 mm; 51-55: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5301309" httpUri="https://zenodo.org/record/5301309/files/figure.png" pageId="31" pageNumber="1510">Figs 52-55</figureCitation>
), and no significant external differences were found either.
</paragraph>
<paragraph id="91623688FFEAFFC6FF04EB87FCDEFD9C" blockId="31.[178,1085,402,570]" pageId="31" pageNumber="1510">
The types of
<taxonomicName id="56DD4D0BFFEAFFC6FEF7EB86FE2EFE0E" authorityName="Cameron" authorityYear="1936" box="[321,454,403,424]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="javanicum">
<emphasis id="A3A9EA9AFFEAFFC6FEF7EB86FE2EFE0E" box="[321,454,403,424]" italics="true" pageId="31" pageNumber="1510">S. javanicum</emphasis>
</taxonomicName>
and
<taxonomicName id="56DD4D0BFFEAFFC6FE48EB86FD6BFE0E" authorityName=", CAMERON" authorityYear="1941" box="[510,643,402,424]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="borneense">
<emphasis id="A3A9EA9AFFEAFFC6FE48EB86FD6BFE0E" box="[510,643,402,424]" italics="true" pageId="31" pageNumber="1510">S. borneense</emphasis>
</taxonomicName>
are distinguished from the
<typeStatus id="4E66882AFFEAFFC6FC1FEB87FBEAFE0E" box="[937,1026,402,424]" pageId="31" pageNumber="1510" type="holotype">holotype</typeStatus>
of
<taxonomicName id="56DD4D0BFFEAFFC6FB9CEB86FEEDFE63" authorityName="Cameron" authorityYear="1930" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="suturale">
<emphasis id="A3A9EA9AFFEAFFC6FB9CEB86FEEDFE63" italics="true" pageId="31" pageNumber="1510">S. suturale</emphasis>
</taxonomicName>
only by the more pronounced microsculpture on the pronotum. However, the additional specimen seen from
<collectingCountry id="E9CA7618FFEAFFC6FE4DEBD8FDBDFE45" box="[507,597,461,483]" name="Thailand" pageId="31" pageNumber="1510">Thailand</collectingCountry>
is intermediate in this respect. Since no additional evidence was found suggesting that
<taxonomicName id="56DD4D0BFFEAFFC6FDD7EBFEFD0BFDA6" authorityName="Cameron" authorityYear="1936" box="[609,739,491,512]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="javanicum">
<emphasis id="A3A9EA9AFFEAFFC6FDD7EBFEFD0BFDA6" box="[609,739,491,512]" italics="true" pageId="31" pageNumber="1510">S. javanicum</emphasis>
</taxonomicName>
represents a distinct species, both
<taxonomicName id="56DD4D0BFFEAFFC6FF04E81DFEDFFDBB" authorityName="Cameron" authorityYear="1936" box="[178,311,520,541]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="javanicum">
<emphasis id="A3A9EA9AFFEAFFC6FF04E81DFEDFFDBB" box="[178,311,520,541]" italics="true" pageId="31" pageNumber="1510">S. javanicum</emphasis>
</taxonomicName>
and
<taxonomicName id="56DD4D0BFFEAFFC6FED9E81DFE1BFDBB" authorityName=", CAMERON" authorityYear="1941" box="[367,499,519,541]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="borneense">
<emphasis id="A3A9EA9AFFEAFFC6FED9E81DFE1BFDBB" box="[367,499,519,541]" italics="true" pageId="31" pageNumber="1510">S. borneense</emphasis>
</taxonomicName>
are placed in synonymy with
<taxonomicName id="56DD4D0BFFEAFFC6FC83E81DFC4DFDBB" authorityName="Cameron" authorityYear="1930" box="[821,933,519,541]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="suturale">
<emphasis id="A3A9EA9AFFEAFFC6FC83E81DFC4DFDBB" box="[821,933,519,541]" italics="true" pageId="31" pageNumber="1510">S. suturale</emphasis>
</taxonomicName>
. Based on the undilated protarsomeres I-IV,
<taxonomicName id="56DD4D0BFFEAFFC6FE52E830FDB9FD9C" authorityName="Cameron" authorityYear="1930" box="[484,593,548,570]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="suturale">
<emphasis id="A3A9EA9AFFEAFFC6FE52E830FDB9FD9C" box="[484,593,548,570]" italics="true" pageId="31" pageNumber="1510">S. suturale</emphasis>
</taxonomicName>
is moved to
<taxonomicName id="56DD4D0BFFEAFFC6FD65E831FCD8FD9C" box="[723,816,548,570]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="genus">
<emphasis id="A3A9EA9AFFEAFFC6FD65E831FCD8FD9C" box="[723,816,548,570]" italics="true" pageId="31" pageNumber="1510">Micrillus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFEAFFC6FF04E85FFD4CFA38" pageId="31" pageNumber="1510" type="description">
<paragraph id="91623688FFEAFFC6FF04E85FFDD0FD54" blockId="31.[178,1086,586,754]" pageId="31" pageNumber="1510">
Redescription: Body length 6.5-7.0 mm, length of forebody
<quantity id="56259B6DFFEAFFC6FC0DE85FFBD2FDC6" box="[955,1082,586,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="31" pageNumber="1510" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.5-3.6 mm</quantity>
. Habitus as in
<figureCitation id="09E62A0DFFEAFFC6FEFCE872FE71FDDB" box="[330,409,615,637]" captionStart="Figs 48-55" captionStartId="29.[178,218,1520,1540]" captionTargetBox="[182,1085,213,1495]" captionTargetId="figure-61@29.[178,1085,212,1497]" captionTargetPageId="29" captionText="Figs 48-55: Micrillus suturalis (CAMERON) (holotypes of Scymbalium borneense CAMERON (48- 49, 51, 54-55) and S. javanicum CAMERON (50-53)): (48) habitus; (49-50) forebody; (51) male sternite VIII; (52-55) aedeagus in lateral and in ventral view. Scale bars: 48-50: 1.0 mm; 51-55: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5301309" httpUri="https://zenodo.org/record/5301309/files/figure.png" pageId="31" pageNumber="1510">Fig. 48</figureCitation>
. Coloration: head blackish; pronotum and elytra dark-brown to blackish-brown, except for the narrowly and indistinctly dark-reddish posterior margins and suture of the elytra; abdomen blackish, with the paratergites, the posterior margins of segments III-VI, the posterior portion of segment VII, and segments VIII-X dark-reddish; legs and antennae pale reddish-brown.
</paragraph>
<paragraph id="91623688FFEAFFC6FF04E914FEE6FC61" blockId="31.[178,1086,769,967]" pageId="31" pageNumber="1510">
Head (
<figureCitation id="09E62A0DFFEAFFC6FF40E914FE80FCB1" box="[246,360,769,791]" captionStart="Figs 48-55" captionStartId="29.[178,218,1520,1540]" captionTargetBox="[182,1085,213,1495]" captionTargetId="figure-61@29.[178,1085,212,1497]" captionTargetPageId="29" captionText="Figs 48-55: Micrillus suturalis (CAMERON) (holotypes of Scymbalium borneense CAMERON (48- 49, 51, 54-55) and S. javanicum CAMERON (50-53)): (48) habitus; (49-50) forebody; (51) male sternite VIII; (52-55) aedeagus in lateral and in ventral view. Scale bars: 48-50: 1.0 mm; 51-55: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5301309" httpUri="https://zenodo.org/record/5301309/files/figure.png" pageId="31" pageNumber="1510">Figs 49-50</figureCitation>
) approximately 1.1 times as long as broad and with subparallel lateral margins; posterior angles moderately marked; punctation extremely fine, sparse, and barely noticeable in median dorsal portion, dense and more distinct in posterior and lateral portions; interstices with shallow microsculpture. Eyes slightly more than half the length of postocular region in dorsal view. Antennae slender, approximately
<quantity id="56259B6DFFEAFFC6FC71E963FBEAFC2A" box="[967,1026,886,908]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="1510" unit="mm" value="3.0">3 mm</quantity>
long; antennomeres IV-XI distinctly oblong; antennomere X approximately 1.5 times as long as broad.
</paragraph>
<paragraph id="91623688FFEAFFC6FF04E9C2FE6CFBC7" blockId="31.[178,1086,982,1121]" pageId="31" pageNumber="1510">
Pronotum (
<figureCitation id="09E62A0DFFEAFFC6FE92E9C3FE7DFC4A" box="[292,405,982,1004]" captionStart="Figs 48-55" captionStartId="29.[178,218,1520,1540]" captionTargetBox="[182,1085,213,1495]" captionTargetId="figure-61@29.[178,1085,212,1497]" captionTargetPageId="29" captionText="Figs 48-55: Micrillus suturalis (CAMERON) (holotypes of Scymbalium borneense CAMERON (48- 49, 51, 54-55) and S. javanicum CAMERON (50-53)): (48) habitus; (49-50) forebody; (51) male sternite VIII; (52-55) aedeagus in lateral and in ventral view. Scale bars: 48-50: 1.0 mm; 51-55: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5301309" httpUri="https://zenodo.org/record/5301309/files/figure.png" pageId="31" pageNumber="1510">Figs 49-50</figureCitation>
) 1.11-1.14 times as long as broad and 1.05-1.09 times as broad as head; general shape rather variable: widest near anterior angles and converging posteriad from anterior fourth or with subparallel lateral margins in anterior half (dorsal view); posterior margin weakly to distinctly concave; punctation fine and very dense; interstices with microsculpture.
</paragraph>
<paragraph id="91623688FFEAFFC6FF04EE64FC5EFB79" blockId="31.[178,1086,1137,1247]" pageId="31" pageNumber="1510">
Elytra (
<figureCitation id="09E62A0DFFEAFFC6FEB0EE64FE97FB21" box="[262,383,1137,1159]" captionStart="Figs 48-55" captionStartId="29.[178,218,1520,1540]" captionTargetBox="[182,1085,213,1495]" captionTargetId="figure-61@29.[178,1085,212,1497]" captionTargetPageId="29" captionText="Figs 48-55: Micrillus suturalis (CAMERON) (holotypes of Scymbalium borneense CAMERON (48- 49, 51, 54-55) and S. javanicum CAMERON (50-53)): (48) habitus; (49-50) forebody; (51) male sternite VIII; (52-55) aedeagus in lateral and in ventral view. Scale bars: 48-50: 1.0 mm; 51-55: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5301309" httpUri="https://zenodo.org/record/5301309/files/figure.png" pageId="31" pageNumber="1510">Figs 49-50</figureCitation>
) 0.91-0.94 times as long as pronotum; surface matt and with microsculpture; punctation extremely fine and dense. Hind wings fully developed. Metatarsomere I longer than the combined length of II and III, but at least slightly shorter than the combined length of II-IV. Protarsomeres I-IV not distinctly dilated.
</paragraph>
<paragraph id="91623688FFEAFFC6FF04EEFBFE1BFA98" blockId="31.[178,1085,1262,1342]" pageId="31" pageNumber="1510">Abdomen narrower than elytra; punctation dense and rather fine; anterior tergites with distinct, posterior tergites with shallow to indistinct microsculpture; posterior margin of tergite VII with palisade fringe.</paragraph>
<paragraph id="91623688FFEAFFC6FF71EF5BFD4CFA38" blockId="31.[178,1086,1358,1438]" pageId="31" pageNumber="1510">
: sternite VIII (
<figureCitation id="09E62A0DFFEAFFC6FED6EF5BFE42FAC2" box="[352,426,1358,1380]" captionStart="Figs 48-55" captionStartId="29.[178,218,1520,1540]" captionTargetBox="[182,1085,213,1495]" captionTargetId="figure-61@29.[178,1085,212,1497]" captionTargetPageId="29" captionText="Figs 48-55: Micrillus suturalis (CAMERON) (holotypes of Scymbalium borneense CAMERON (48- 49, 51, 54-55) and S. javanicum CAMERON (50-53)): (48) habitus; (49-50) forebody; (51) male sternite VIII; (52-55) aedeagus in lateral and in ventral view. Scale bars: 48-50: 1.0 mm; 51-55: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5301309" httpUri="https://zenodo.org/record/5301309/files/figure.png" pageId="31" pageNumber="1510">Fig. 51</figureCitation>
) strongly oblong, posterior excision broad and moderately deep; aedeagus (
<figureCitation id="09E62A0DFFEAFFC6FEA9EF7EFE79FA27" box="[287,401,1387,1409]" captionStart="Figs 48-55" captionStartId="29.[178,218,1520,1540]" captionTargetBox="[182,1085,213,1495]" captionTargetId="figure-61@29.[178,1085,212,1497]" captionTargetPageId="29" captionText="Figs 48-55: Micrillus suturalis (CAMERON) (holotypes of Scymbalium borneense CAMERON (48- 49, 51, 54-55) and S. javanicum CAMERON (50-53)): (48) habitus; (49-50) forebody; (51) male sternite VIII; (52-55) aedeagus in lateral and in ventral view. Scale bars: 48-50: 1.0 mm; 51-55: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5301309" httpUri="https://zenodo.org/record/5301309/files/figure.png" pageId="31" pageNumber="1510">Figs 52-55</figureCitation>
) approximately
<quantity id="56259B6DFFEAFFC6FD8BEF7EFD65FA27" box="[573,653,1387,1409]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="31" pageNumber="1510" unit="mm" value="1.3">1.3 mm</quantity>
long, slender, distinctly asymmetric, and with conspicuously shaped spines in internal sac.
</paragraph>
</subSubSection>
<subSubSection id="D9C76503FFEAFFC6FF04EFB8FC80F9BD" pageId="31" pageNumber="1510" type="description">
<paragraph id="91623688FFEAFFC6FF04EFB8FC80F9BD" blockId="31.[178,1085,1453,1563]" pageId="31" pageNumber="1510">
Comparative notes: This species is readily distinguished from other
<taxonomicName id="56DD4D0BFFEAFFC6FF04EFDEFEE7FA47" box="[178,271,1483,1505]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1510" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A3A9EA9AFFEAFFC6FF04EFDEFEE7FA47" box="[178,271,1483,1505]" italics="true" pageId="31" pageNumber="1510">Micrillus</emphasis>
</taxonomicName>
species known from the
<collectingRegion id="5319F86AFFEAFFC6FDAEEFDEFD5DFA47" box="[536,693,1483,1505]" country="Egypt" name="Ash Sharqiyah" pageId="31" pageNumber="1510">Oriental region</collectingRegion>
by distinctly larger body size, darker coloration, and by the completely different morphology of the aedeagus (much larger, more slender, internal sac with series of strongly sclerotized spines).
</paragraph>
</subSubSection>
<caption id="C5A26600FFD5FFF9FF04EFE5FE50F9EE" ID-DOI="http://doi.org/10.5281/zenodo.5301311" ID-Zenodo-Dep="5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="32" pageNumber="1511" startId="32.[178,218,1520,1540]" targetBox="[190,1085,222,1489]" targetPageId="32">
<paragraph id="91623688FFD5FFF9FF04EFE5FE50F9EE" blockId="32.[178,1085,1520,1608]" pageId="32" pageNumber="1511">
<emphasis id="A3A9EA9AFFD5FFF9FF04EFE5FEF1F9A2" bold="true" box="[178,281,1520,1540]" pageId="32" pageNumber="1511">Figs 56-62</emphasis>
:
<taxonomicName id="56DD4D0BFFD5FFF9FE9CEFE5FD79F9A2" authority="(MOTSCHULSKY)" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[298,657,1520,1540]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1511" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="A3A9EA9AFFD5FFF9FE9CEFE5FE0FF9A2" box="[298,487,1520,1540]" italics="true" pageId="32" pageNumber="1511">Scymbalium badium</emphasis>
(MOTSCHULSKY)
</taxonomicName>
, lectotype (
<emphasis id="A3A9EA9AFFD5FFF9FCB2EFE5FCD0F9A2" bold="true" box="[772,824,1520,1540]" pageId="32" pageNumber="1511">56-60</emphasis>
) and
<taxonomicName id="56DD4D0BFFD5FFF9FCC3EFE5FC03F9A2" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[885,1003,1520,1540]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1511" phylum="Arthropoda" rank="species" species="nepalense" status="sp. nov.">
<emphasis id="A3A9EA9AFFD5FFF9FCC3EFE5FC03F9A2" box="[885,1003,1520,1540]" italics="true" pageId="32" pageNumber="1511">S. nepalense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFD5FFF9FC43EFE5FBD2F9A2" box="[1013,1082,1520,1540]" pageId="32" pageNumber="1511" rank="species">nov.sp.</taxonomicNameLabel>
, holotype (
<emphasis id="A3A9EA9AFFD5FFF9FEA4EC12FEAFF9BD" bold="true" box="[274,327,1543,1563]" pageId="32" pageNumber="1511">61-62</emphasis>
): (
<emphasis id="A3A9EA9AFFD5FFF9FED3EC12FE94F9BD" bold="true" box="[357,380,1543,1563]" pageId="32" pageNumber="1511">56</emphasis>
) forebody; (
<emphasis id="A3A9EA9AFFD5FFF9FE40EC12FDD2F9BD" bold="true" box="[502,570,1543,1563]" pageId="32" pageNumber="1511">57, 61)</emphasis>
median dorsal portion of head; (
<emphasis id="A3A9EA9AFFD5FFF9FCCEEC12FC5CF9BD" bold="true" box="[888,948,1543,1563]" pageId="32" pageNumber="1511">58, 62</emphasis>
) male sternite VIII; (
<emphasis id="A3A9EA9AFFD5FFF9FF58EC08FECAF997" bold="true" box="[238,290,1565,1585]" pageId="32" pageNumber="1511">59-60</emphasis>
) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm.
</paragraph>
</caption>
<caption id="C5A26600FFD4FFF8FF04E957FC12FCCA" ID-DOI="http://doi.org/10.5281/zenodo.5301317" ID-Zenodo-Dep="5301317" httpUri="https://zenodo.org/record/5301317/files/figure.png" pageId="33" pageNumber="1512" startId="33.[178,218,834,854]" targetBox="[191,1085,214,809]" targetPageId="33">
<paragraph id="91623688FFD4FFF8FF04E957FC12FCCA" blockId="33.[178,1085,834,876]" pageId="33" pageNumber="1512">
<emphasis id="A3A9EA9AFFD4FFF8FF04E957FEFEFCF0" bold="true" box="[178,278,834,854]" pageId="33" pageNumber="1512">Figs 63-66</emphasis>
:
<taxonomicName id="56DD4D0BFFD4FFF8FE94E957FE1BFCF0" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[290,499,834,854]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="1512" phylum="Arthropoda" rank="species" species="nepalense" status="sp. nov.">
<emphasis id="A3A9EA9AFFD4FFF8FE94E957FE1BFCF0" box="[290,499,834,854]" italics="true" pageId="33" pageNumber="1512">Scymbalium nepalense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFD4FFF8FE4FE957FDD5FCF0" box="[505,573,834,854]" pageId="33" pageNumber="1512" rank="species">nov.sp.</taxonomicNameLabel>
, holotype: (
<emphasis id="A3A9EA9AFFD4FFF8FD1CE957FD29FCF0" bold="true" box="[682,705,834,854]" pageId="33" pageNumber="1512">63</emphasis>
) habitus; (
<emphasis id="A3A9EA9AFFD4FFF8FC93E957FCD4FCF0" bold="true" box="[805,828,834,854]" pageId="33" pageNumber="1512">64</emphasis>
) male sternite VIII; (
<emphasis id="A3A9EA9AFFD4FFF8FBB6E957FBDEFCF0" bold="true" box="[1024,1078,834,854]" pageId="33" pageNumber="1512">65-66</emphasis>
) aedeagus in lateral and in ventral view. Scale bars: 63: 1.0 mm; 64: 0.5 mm; 65-66: 0.2 mm.
</paragraph>
</caption>
<subSubSection id="D9C76503FFD4FFF8FF04E96EFCEAFC6D" pageId="33" pageNumber="1512" type="description">
<paragraph id="91623688FFD4FFF8FF04E96EFCEAFC6D" blockId="33.[178,1085,891,971]" pageId="33" pageNumber="1512">
Distribution and natural history:
<taxonomicName id="56DD4D0BFFD4FFF8FCA1E96EFC3EFC37" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1930" box="[791,982,891,913]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="1512" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="A3A9EA9AFFD4FFF8FCA1E96EFC3EFC37" box="[791,982,891,913]" italics="true" pageId="33" pageNumber="1512">Micrillus suturalis</emphasis>
</taxonomicName>
is apparently widespread, but not common in the Oriental region. It is currently known from Java, Borneo, Peninsular
<collectingCountry id="E9CA7618FFD4FFF8FE02E9A0FDFBFC6D" box="[436,531,949,971]" name="Malaysia" pageId="33" pageNumber="1512">Malaysia</collectingCountry>
, and
<collectingCountry id="E9CA7618FFD4FFF8FDFFE9A0FD4BFC6D" box="[585,675,949,971]" name="Thailand" pageId="33" pageNumber="1512">Thailand</collectingCountry>
(
<figureCitation id="09E62A0DFFD4FFF8FD06E9A0FD1DFC6D" box="[688,757,949,971]" captionStart="Map 4" captionStartId="27.[178,223,1091,1111]" captionTargetBox="[178,1085,212,1066]" captionTargetId="figure-174@27.[178,1085,212,1066]" captionTargetPageId="27" captionText="Map 4: Distributions of Micrillus distortus nov.sp. (triangles) and M. suturalis (CAMERON) (circles) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.5301305" httpUri="https://zenodo.org/record/5301305/files/figure.png" pageId="33" pageNumber="1512">Map 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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