260 lines
25 KiB
XML
260 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1158.100518" ID-Pensoft-Pub="1313-2970-1158-1" ID-Pensoft-UUID="299C29A6693A5054BA17D1663FBCCEE0" ID-ZooBank="047C3E08B3F244F484050AD39F23E9E8" ModsDocID="1313-2970-1158-1" checkinTime="1681371110153" checkinUser="pensoft" docAuthor="dela Cruz, Ian Niel & Ohara, Masahiro" docDate="2023" docId="0EB5F543ECAF5E4D94ECB6C1959E4B83" docLanguage="en" docName="ZooKeys 1158: 1-26" docOrigin="ZooKeys 1158" docPubDate="2023-04-12" docSource="http://dx.doi.org/10.3897/zookeys.1158.100518" docTitle="Atholus philippinensis" docType="treatment" docVersion="1" id="299C29A6693A5054BA17D1663FBCCEE0" lastPageNumber="1" masterDocId="299C29A6693A5054BA17D1663FBCCEE0" masterDocTitle="Revision of the genus Atholus Thomson, 1859 (Coleoptera, Histeridae, Histerinae) from the Philippines with additional records" masterLastPageNumber="26" masterPageNumber="1" pageNumber="1" updateTime="1681371110153" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Atholus Thomson, 1859 (Coleoptera, Histeridae, Histerinae) from the Philippines with additional records</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>dela Cruz, Ian Niel</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2806-1337</mods:nameIdentifier>
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<mods:affiliation>Entomological Laboratory, Graduate School of Agriculture, Hokkaido University, N 9, W 9, Sapporo, 060 - 8589, Japan & Department of Biology, College of Mathematics and Natural Sciences, Caraga State University, Butuan City, 8600, Philippines</mods:affiliation>
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<mods:nameIdentifier type="email">histermushi@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ohara, Masahiro</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2806-1337</mods:nameIdentifier>
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<mods:affiliation>Entomological Laboratory, Graduate School of Agriculture, Hokkaido University, N 9, W 9, Sapporo, 060 - 8589, Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1158</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>26</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1158.100518</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1158.100518</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1158-1</mods:identifier>
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<mods:identifier type="ZooBank">047C3E08B3F244F484050AD39F23E9E8</mods:identifier>
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<mods:identifier type="Pensoft-UUID">299C29A6693A5054BA17D1663FBCCEE0</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:0EB5F543ECAF5E4D94ECB6C1959E4B83" httpUri="http://treatment.plazi.org/id/0EB5F543ECAF5E4D94ECB6C1959E4B83" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="5VWG2" LSID="0EB5F543-ECAF-5E4D-94EC-B6C1959E4B83" authority="(Marseul, 1854)" baseAuthorityName="Marseul" baseAuthorityYear="1854" class="Insecta" family="Histeridae" genus="Atholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atholus philippinensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="philippinensis">Atholus philippinensis (Marseul, 1854)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Philippine Atholus, dorsal habitus 1 A. philippinensis (Marseul, 1854) [IC- 21 - 18] 2 A. coelestis (Marseul, 1857) [IC- 21 - 20] 3 A. torquatus (Marseul, 1854) [IC- 21 - 49] 4 A. pirithous (Marseul, 1873) [IC- 21 - 47] 5 A. nitidissimus Desbordes, 1925 [syntype image] 6 A. bakeri (Bickhardt, 1914) [syntype image, No. 1639]. Scale bar: 1.00 mm." figureDoi="10.3897/zookeys.1158.100518.figures1-6" httpUri="https://binary.pensoft.net/fig/836675" pageId="0" pageNumber="1">Figs 1</figureCitation>
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<figureCitation captionStart="Figures 7, 8" captionStartId="F2" captionText="Figures 7, 8. Atholus philippinesis (Marseul, 1854) [IC- 21 - 18] 7 protibia, dorsal view 8 ditto, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures7-8" httpUri="https://binary.pensoft.net/fig/836676" pageId="0" pageNumber="1">, 7</figureCitation>
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<figureCitation captionStart="Figures 7, 8" captionStartId="F2" captionText="Figures 7, 8. Atholus philippinesis (Marseul, 1854) [IC- 21 - 18] 7 protibia, dorsal view 8 ditto, ventral view." figureDoi="10.3897/zookeys.1158.100518.figures7-8" httpUri="https://binary.pensoft.net/fig/836676" pageId="0" pageNumber="1">, 8</figureCitation>
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<figureCitation captionStart="Figures 35–37" captionStartId="F7" captionText="Figures 35 - 37. Atholus philippinensis (Marseul, 1854), female genitalia [IC- 21 - 18] 35 dorsal view 36 lateral view 37 dorsolateral view of gonocoxite. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1158.100518.figures35-37" httpUri="https://binary.pensoft.net/fig/836681" pageId="0" pageNumber="1">, 35-37</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Marseul" authorityYear="1854" class="Insecta" family="Histeridae" genus="Hister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hister philippinensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="philippinensis">Hister philippinensis</taxonomicName>
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Marseul, 1854: 547 [Malaisie (
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<normalizedToken originalValue="îles">iles</normalizedToken>
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Philippines)].
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Gemminger & Harold" authorityYear="1868" class="Insecta" family="Histeridae" genus="Hister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hister philippensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="philippensis">Hister philippensis</taxonomicName>
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(sic):
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.9089" author="Gemminger, M" journalOrPublisher="Carl Gerold's Sohn, Wien" pageId="0" pageNumber="1" refId="B17" refString="Gemminger, M, Harold, E, 1868. Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus. Monachii, sumptu E.H. Gummi, 1868-1876. [753-978 + [5] pp.] https://doi.org/10.5962/bhl.title.9089" title="Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus. Monachii, sumptu E. H. Gummi, 1868 - 1876. [753 - 978 + [5] pp.]" url="https://doi.org/10.5962/bhl.title.9089" year="1868">Gemminger and Harold 1868</bibRefCitation>
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: 771.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Marseul" authorityYear="1854" class="Insecta" family="Histeridae" genus="Hister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hister (Atholus) philippinensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="philippinensis" subGenus="Atholus">Hister (Atholus) philippinensis</taxonomicName>
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:
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<bibRefCitation author="Bickhardt, H" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="1" refId="B3" refString="Bickhardt, H, 1910. Histeridae. In: Junk W, Schenkling S (Eds) Coleopterorum Catalogus, pars 24. W. Junk, Berlin" title="Histeridae. In: Junk W, Schenkling S (Eds) Coleopterorum Catalogus, pars 24. W." year="1910">Bickhardt 1910</bibRefCitation>
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: 54 [catalogued]; 1913: 173 [Hoozan, Taihorin]; 1917: 194 [catalogued];
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<bibRefCitation author="Miwa, Y" journalOrPublisher="Department of Agriculture Government Research Institute, Formosa, Japan, Report" pageId="0" pageNumber="1" pagination="49 - 58" refId="B44" refString="Miwa, Y, 1931. Histeridae and Niponiidae. In: A systematic catalogue of Formosan Coleoptera. Department of Agriculture Government Research Institute, Formosa, Japan, Report 55: 49 - 58" title="Histeridae and Niponiidae. In: A systematic catalogue of Formosan Coleoptera." volume="55" year="1931">Miwa 1931</bibRefCitation>
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: 57 [Hoozan, Taihorin];
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<bibRefCitation author="Kamiya, K" journalOrPublisher="Scientia Agricola" pageId="0" pageNumber="1" pagination="21 - 32" refId="B22" refString="Kamiya, K, Takagi, S, 1938. A list of Japanese Histeridae. Scientia Agricola 19 (1): 21 - 32" title="A list of Japanese Histeridae." volume="19" year="1938">Kamiya and Takagi 1938</bibRefCitation>
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: 31.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Lewis" authorityYear="1901" class="Insecta" family="Histeridae" genus="Hister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hister sectator" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sectator">Hister sectator</taxonomicName>
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Lewis, 1901: 375, synonymized by Bickhardt, 1917: 194.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Insecta" family="Histeridae" genus="Atholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atholus sectator" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sectator">Atholus sectator</taxonomicName>
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:
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<bibRefCitation DOI="https://doi.org/10.1080/00222930608562636" author="Lewis, G" journalOrPublisher="Annals & Magazine of Natural History" pageId="0" pageNumber="1" pagination="397 - 403" refId="B30" refString="Lewis, G, 1906. On new species of Histeridae and notices of others. Annals & Magazine of Natural History 18 (7): 397 - 403, DOI: https://doi.org/10.1080/00222930608562636" title="On new species of Histeridae and notices of others." url="https://doi.org/10.1080/00222930608562636" volume="18" year="1906">Lewis 1906</bibRefCitation>
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: 402.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Marseul" baseAuthorityYear="1854" class="Insecta" family="Histeridae" genus="Atholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atholus philippinensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="philippinensis">Atholus philippinensis</taxonomicName>
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:
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<bibRefCitation DOI="https://doi.org/10.1080/00222930608562636" author="Lewis, G" journalOrPublisher="Annals & Magazine of Natural History" pageId="0" pageNumber="1" pagination="397 - 403" refId="B30" refString="Lewis, G, 1906. On new species of Histeridae and notices of others. Annals & Magazine of Natural History 18 (7): 397 - 403, DOI: https://doi.org/10.1080/00222930608562636" title="On new species of Histeridae and notices of others." url="https://doi.org/10.1080/00222930608562636" volume="18" year="1906">Lewis 1906</bibRefCitation>
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: 402;
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<bibRefCitation DOI="https://doi.org/10.1080/00222931508693684" author="Lewis, G" journalOrPublisher="Annals & Magazine of Natural History" pageId="0" pageNumber="1" pagination="54 - 56" refId="B31" refString="Lewis, G, 1915. On new species of Histeridae and notices of others. Annals & Magazine of Natural History 16 (8): 54 - 56, DOI: https://doi.org/10.1080/00222931508693684" title="On new species of Histeridae and notices of others." url="https://doi.org/10.1080/00222931508693684" volume="16" year="1915">1915</bibRefCitation>
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: 55;
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<bibRefCitation author="Mazur, S" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="0" pageNumber="1" pagination="1 - 376" refId="B36" refString="Mazur, S, 1984. A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54 (3-4): 1 - 376" title="A world catalogue of Histeridae." volume="54" year="1984">Mazur 1984</bibRefCitation>
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: 215;
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<bibRefCitation author="Mazur, S" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="0" pageNumber="1" refId="B37" refString="Mazur, S, 1997. A world catalogue of Histeridae (Coleoptera: Histeroidea). Genus International Journal of Invertebrate Taxonomy (Supplement), Polskie Towarzystwo Taksonomiczne 1-373." title="A world catalogue of Histeridae (Coleoptera: Histeroidea). Genus International Journal of Invertebrate Taxonomy (Supplement), Polskie Towarzystwo Taksonomiczne 1 - 373." year="1997">1997</bibRefCitation>
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||
: 132;
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<bibRefCitation author="Mazur, S" journalOrPublisher="Warsaw University of Life Sciences SGGW Press, Warsaw" pageId="0" pageNumber="1" refId="B40" refString="Mazur, S, 2011. A Concise Catalogue of the Histeridae (Insecta: Coleoptera). Warsaw University of Life Sciences SGGW Press, Warsaw" title="A Concise Catalogue of the Histeridae (Insecta: Coleoptera)." year="2011">2011</bibRefCitation>
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: 106 [catalogued];
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<bibRefCitation author="Ohara, M" journalOrPublisher="Insecta Matsumurana, New series" pageId="0" pageNumber="1" pagination="75 - 132" refId="B51" refString="Ohara, M, 1999. A revision of the superfamily Histeroidea of Japan (Coleoptera), Supplementum 1. Insecta Matsumurana, New series 55: 75 - 132" title="A revision of the superfamily Histeroidea of Japan (Coleoptera), Supplementum 1." volume="55" year="1999">
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<normalizedToken originalValue="Ôhara">Ohara</normalizedToken>
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1999
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</bibRefCitation>
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b: 32-36 [Taiwan].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" location="Mindanao Island" municipality="Agusan del Norte" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
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<specimenCount type="male">3 ♂♂</specimenCount>
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,
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<specimenCount type="female">3 ♀♀</specimenCount>
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.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<location LSID="urn:lsid:plazi:treatment:0EB5F543ECAF5E4D94ECB6C1959E4B83:CF5D9E3E3C9F3E78FDF7B09C43A27A03" municipality="Agusan del Norte" name="Mindanao Island">Mindanao Island</location>
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</emphasis>
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,
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<collectingMunicipality>Agusan del Norte</collectingMunicipality>
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, Butuan, Taligaman
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</materialsCitation>
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,
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<specimenCount type="male">3 ♂♂</specimenCount>
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,
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<specimenCount type="female">3 ♀♀</specimenCount>
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[IC-21-18],
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<geoCoordinate degrees="08.56894" direction="north" orientation="latitude" precision="1" value="8.56894">08.56894°N</geoCoordinate>
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,
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<geoCoordinate degrees="125.38534" direction="east" orientation="longitude" precision="1" value="125.38534">125.38534°E</geoCoordinate>
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<quantity metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">60 m</quantity>
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a.s.l.,
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<date value="2021-06-02">2021-VI-02</date>
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[AN-21-IDC-002], I.N. DELA CRUZ leg.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Marseul" baseAuthorityYear="1854" class="Insecta" family="Histeridae" genus="Atholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atholus philippinensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="philippinensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Atholus philippinensis</emphasis>
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</taxonomicName>
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(Marseul, 1854) (Fig.
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Philippine Atholus, dorsal habitus 1 A. philippinensis (Marseul, 1854) [IC- 21 - 18] 2 A. coelestis (Marseul, 1857) [IC- 21 - 20] 3 A. torquatus (Marseul, 1854) [IC- 21 - 49] 4 A. pirithous (Marseul, 1873) [IC- 21 - 47] 5 A. nitidissimus Desbordes, 1925 [syntype image] 6 A. bakeri (Bickhardt, 1914) [syntype image, No. 1639]. Scale bar: 1.00 mm." figureDoi="10.3897/zookeys.1158.100518.figures1-6" httpUri="https://binary.pensoft.net/fig/836675" pageId="0" pageNumber="1">1</figureCitation>
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) is easily distinguished from other Philippine congeners by entire dorsal elytral striae 1-3 (fourth stria is incomplete), and dense punctation of propygidium and pygidium. Among Philippine species, it is the largest one in size, with its markedly wider elytra and posterior angles of pronotum. The number of denticles of the outer sublateral margin of protibia is four.
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</paragraph>
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<caption doi="10.3897/zookeys.1158.100518.figures1-6" httpUri="https://binary.pensoft.net/fig/836675" pageId="0" pageNumber="1" start="Figures 1–6" startId="F1">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figures 1-6.</emphasis>
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Philippine
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<taxonomicName authorityName="Thomson" authorityYear="1859" class="Insecta" family="Histeridae" genus="Atholus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atholus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Atholus</emphasis>
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</taxonomicName>
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, dorsal habitus
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<emphasis bold="true" pageId="0" pageNumber="1">1</emphasis>
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<taxonomicName lsidName="A. philippinensis" pageId="0" pageNumber="1" rank="species" species="philippinensis">
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<emphasis italics="true" pageId="0" pageNumber="1">A. philippinensis</emphasis>
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</taxonomicName>
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(Marseul, 1854) [IC-21-18]
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<emphasis bold="true" pageId="0" pageNumber="1">2</emphasis>
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<taxonomicName lsidName="A. coelestis" pageId="0" pageNumber="1" rank="species" species="coelestis">
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<emphasis italics="true" pageId="0" pageNumber="1">A. coelestis</emphasis>
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</taxonomicName>
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(Marseul, 1857) [IC-21-20]
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<emphasis bold="true" pageId="0" pageNumber="1">3</emphasis>
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<taxonomicName lsidName="A. torquatus" pageId="0" pageNumber="1" rank="species" species="torquatus">
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<emphasis italics="true" pageId="0" pageNumber="1">A. torquatus</emphasis>
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</taxonomicName>
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(Marseul, 1854) [IC-21-49]
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<emphasis bold="true" pageId="0" pageNumber="1">4</emphasis>
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<taxonomicName lsidName="A. pirithous" pageId="0" pageNumber="1" rank="species" species="pirithous">
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<emphasis italics="true" pageId="0" pageNumber="1">A. pirithous</emphasis>
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</taxonomicName>
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(Marseul, 1873) [IC-21-47]
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<emphasis bold="true" pageId="0" pageNumber="1">5</emphasis>
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<taxonomicName lsidName="A. nitidissimus" pageId="0" pageNumber="1" rank="species" species="nitidissimus">
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<emphasis italics="true" pageId="0" pageNumber="1">A. nitidissimus</emphasis>
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</taxonomicName>
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Desbordes, 1925 [syntype image]
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<emphasis bold="true" pageId="0" pageNumber="1">6</emphasis>
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<taxonomicName lsidName="A. bakeri" pageId="0" pageNumber="1" rank="species" species="bakeri">
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<emphasis italics="true" pageId="0" pageNumber="1">A. bakeri</emphasis>
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</taxonomicName>
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(Bickhardt, 1914) [syntype image, No. 1639]. Scale bar: 1.00 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">Additional description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
|
||
: anterior portion of valvifers (Figs
|
||
<figureCitation captionStart="Figures 35–37" captionStartId="F7" captionText="Figures 35 - 37. Atholus philippinensis (Marseul, 1854), female genitalia [IC- 21 - 18] 35 dorsal view 36 lateral view 37 dorsolateral view of gonocoxite. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1158.100518.figures35-37" httpUri="https://binary.pensoft.net/fig/836681" pageId="0" pageNumber="1">35</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 35–37" captionStartId="F7" captionText="Figures 35 - 37. Atholus philippinensis (Marseul, 1854), female genitalia [IC- 21 - 18] 35 dorsal view 36 lateral view 37 dorsolateral view of gonocoxite. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1158.100518.figures35-37" httpUri="https://binary.pensoft.net/fig/836681" pageId="0" pageNumber="1">36</figureCitation>
|
||
) paddle-shaped; gonocoxite (Fig.
|
||
<figureCitation captionStart="Figures 35–37" captionStartId="F7" captionText="Figures 35 - 37. Atholus philippinensis (Marseul, 1854), female genitalia [IC- 21 - 18] 35 dorsal view 36 lateral view 37 dorsolateral view of gonocoxite. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1158.100518.figures35-37" httpUri="https://binary.pensoft.net/fig/836681" pageId="0" pageNumber="1">37</figureCitation>
|
||
) slightly elongate, almost twice as long as broad, shovel-like; inner and outer surfaces differentiated; inner face moderately separated from outer face by elevated lateral ridge; sclerotized setae on apical half of outer face short and somewhat dense; inner face with short and sparse setae; apex of gonocoxite with two teeth; gonostyli present, freely articulated; spermathecae multiple, consisting of four sacs; sacs gradually enlarged and elongate, not sclerotized.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Philippines; Malaysia; Indonesia (Sumatra, Borneo, Java); Myanmar, Vietnam; India (Meghalaya); China (Hainan); Taiwan (
|
||
<bibRefCitation author="Mazur, S" journalOrPublisher="Warsaw University of Life Sciences SGGW Press, Warsaw" pageId="0" pageNumber="1" refId="B40" refString="Mazur, S, 2011. A Concise Catalogue of the Histeridae (Insecta: Coleoptera). Warsaw University of Life Sciences SGGW Press, Warsaw" title="A Concise Catalogue of the Histeridae (Insecta: Coleoptera)." year="2011">Mazur 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
This species occurs in decaying banana stumps and are often found along with some species of
|
||
<taxonomicName authorityName="Lewis" authorityYear="1892" class="Insecta" family="Histeridae" genus="Platylister" kingdom="Animalia" lsidName="Platylister" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Platylister</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Bickhardt" authorityYear="1914" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Platysomatini">Platysomatini</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Gyllenhal" authorityYear="1808" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Histerinae">Histerinae</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Gyllenhal" authorityYear="1808" class="Insecta" family="Histeridae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The protibial teeth of
|
||
<taxonomicName lsidName="A. philippinensis" pageId="0" pageNumber="1" rank="species" species="philippinensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">A. philippinensis</emphasis>
|
||
</taxonomicName>
|
||
(Marseul, 1854) are not as prominent as they are in other species. Moreover, in comparison to the description of
|
||
<bibRefCitation author="Ohara, M" journalOrPublisher="Insecta Matsumurana, New series" pageId="0" pageNumber="1" pagination="75 - 132" refId="B51" refString="Ohara, M, 1999. A revision of the superfamily Histeroidea of Japan (Coleoptera), Supplementum 1. Insecta Matsumurana, New series 55: 75 - 132" title="A revision of the superfamily Histeroidea of Japan (Coleoptera), Supplementum 1." volume="55" year="1999">
|
||
<normalizedToken originalValue="Ôhara">Ohara</normalizedToken>
|
||
(1999
|
||
</bibRefCitation>
|
||
b), the number of denticles may vary, ranging from 9-11 on the outer margin, one on the inner apical angle, and four or five on the outer sublateral margin. This species was already re-described based on specimens of Taiwan and western Kalimantan, Indonesia (
|
||
<normalizedToken originalValue="Ôhara">Ohara</normalizedToken>
|
||
, 1999b), including the illustrations of male genitalia and spermatheca of female.
|
||
<bibRefCitation author="Ohara, M" journalOrPublisher="Insecta Matsumurana, New series" pageId="0" pageNumber="1" pagination="75 - 132" refId="B51" refString="Ohara, M, 1999. A revision of the superfamily Histeroidea of Japan (Coleoptera), Supplementum 1. Insecta Matsumurana, New series 55: 75 - 132" title="A revision of the superfamily Histeroidea of Japan (Coleoptera), Supplementum 1." volume="55" year="1999">
|
||
<normalizedToken originalValue="Ôhara">Ohara</normalizedToken>
|
||
(1999
|
||
</bibRefCitation>
|
||
b) also provided a figure of the spermatheca; we add illustrations of the female gonocoxite and valvifers here (Figs
|
||
<figureCitation captionStart="Figures 35–37" captionStartId="F7" captionText="Figures 35 - 37. Atholus philippinensis (Marseul, 1854), female genitalia [IC- 21 - 18] 35 dorsal view 36 lateral view 37 dorsolateral view of gonocoxite. Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1158.100518.figures35-37" httpUri="https://binary.pensoft.net/fig/836681" pageId="0" pageNumber="1">35-37</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |