treatments-xml/data/0E/32/E7/0E32E7B8828E365249D5DF7F0D2B6CC0.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.81.13488" ID-PMC="PMC5523871" ID-Pensoft-Pub="1314-2003-81-47" ID-Pensoft-UUID="9264FFFAFFB05E16C86452418950FF97" ID-PubMed="28785165" ID-Zenodo-Dep="816510" ModsDocID="1314-2003-81-47" checkinTime="1502745051323" checkinUser="pensoft" docAuthor="Brambach, Fabian, Byng, James W. &amp; Culmsee, Heike" docDate="2017" docId="0E32E7B8828E365249D5DF7F0D2B6CC0" docLanguage="en" docName="PhytoKeys 81: 47-78" docOrigin="PhytoKeys 81" docPubDate="2017-06-15" docSource="http://dx.doi.org/10.3897/phytokeys.81.13488" docTitle="Syzygium devogelii Brambach, Byng &amp; Culmsee 2017, sp. nov." docType="treatment" docVersion="7" id="9264FFFAFFB05E16C86452418950FF97" lastPageNumber="64" masterDocId="9264FFFAFFB05E16C86452418950FF97" masterDocTitle="Five new species of Syzygium (Myrtaceae) from Sulawesi, Indonesia" masterLastPageNumber="78" masterPageNumber="47" pageNumber="62" updateTime="1668141982981" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five new species of Syzygium (Myrtaceae) from Sulawesi, Indonesia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brambach, Fabian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3203-2248</mods:nameIdentifier>
<mods:affiliation>Plant Ecology and Ecosystems Research, Albrecht-von-Haller Institute for Plant Sciences, University of Goettingen, Untere Karspuele 2, 37073 Goettingen, Germany</mods:affiliation>
<mods:nameIdentifier type="email">fbramba@gwdg.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Byng, James W.</mods:namePart>
<mods:affiliation>Plant Gateway, 5 Talbot Street, Hertford, Hertfordshire, SG 13 7 BX, UK &amp; Naturalis Biodiversity Center, Botany, P. O. Box 9517, 2300 RA, Leiden, The Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Culmsee, Heike</mods:namePart>
<mods:affiliation>Plant Ecology and Ecosystems Research, Albrecht-von-Haller Institute for Plant Sciences, University of Goettingen, Untere Karspuele 2, 37073 Goettingen, Germany &amp; DBU Natural Heritage, German Federal Foundation for the Environment, An der Bornau 2, 49090 Osnabrueck, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-06-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>81</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>47</mods:start>
<mods:end>78</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.81.13488</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.81.13488</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-81-47</mods:identifier>
<mods:identifier type="Pensoft-UUID">9264FFFAFFB05E16C86452418950FF97</mods:identifier>
<mods:identifier type="Zenodo-Dep">816510</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="132649809" LSID="urn:lsid:plazi:treatment:0E32E7B8828E365249D5DF7F0D2B6CC0" httpUri="http://treatment.plazi.org/id/0E32E7B8828E365249D5DF7F0D2B6CC0" lastPageId="17" lastPageNumber="64" pageId="15" pageNumber="62">
<subSubSection pageId="15" pageNumber="62" type="nomenclature">
<paragraph pageId="15" pageNumber="62">
<pageBreakToken pageId="15" pageNumber="62" start="start">3</pageBreakToken>
. 
<taxonomicName LSID="0E32E7B8-828E-3652-49D5-DF7F0D2B6CC0" authority="Brambach, Byng &amp; Culmsee" authorityName="Brambach, Byng &amp; Culmsee" authorityYear="2017" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium devogelii" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="species" species="devogelii" status="sp. nov.">Syzygium devogelii Brambach, Byng &amp; Culmsee</taxonomicName>
<taxonomicNameLabel pageId="15" pageNumber="62">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution map of four species of Syzygium in Central Sulawesi: Syzygium contiguum (orange diamonds), S. devogelii (yellow dots), S. eymae (light blue square), and S. galanthum (green triangles). Lore Lindu National Park is indicated by a black line. Map created with QGIS (QGIS Development Team 2016) using the digital elevation model of Jarvis et al. (2008)." figureDoi="10.3897/phytokeys.81.13488.figure4" httpUri="https://binary.pensoft.net/fig/140631" pageId="15" pageNumber="62">Figures 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Syzygium devogelii: a leafy twig with fruit b longitudinal section of ripe fruit with inner, flat side of cotyledon and echinate outer surface c flower bud, longitudinal section d flower bud, exterior view. a-b holotype de Vogel 5293 c-d Culmsee 1564." figureDoi="10.3897/phytokeys.81.13488.figure5" httpUri="https://binary.pensoft.net/fig/140632" pageId="15" pageNumber="62">, 5</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Leaves of all new species described. Variation of Syzygium balgooyi (a-e), S. contiguum (f), S. devogelii (g), S. eymae (h), and S. galanthum (i). a Brambach et al. 0283 b Brambach et al. 0681 c Brambach et al. 1333 d de Vogel 5413 [L. 2517563] e Culmsee r 2162 f Culmsee r 463 g Brambach et al. 0818 h Eyma 3624 [L. 2535689] i Brambach et al. 0533. Scale bar: 10 cm, valid for all leaves." figureDoi="10.3897/phytokeys.81.13488.figure8" httpUri="https://binary.pensoft.net/fig/140635" pageId="15" pageNumber="62">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="62" type="reference_group">
<paragraph pageId="15" pageNumber="62">
<taxonomicName authority="Brambach &amp; Byng &amp; Culmsee, 2017" authorityName="Brambach &amp; Byng &amp; Culmsee" authorityYear="2017" family="Myrtaceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="" order="Myrtales" pageId="15" pageNumber="62" rank="family" status="sp. nov.">Myrtaceae</taxonomicName>
&quot; sp. 10&quot; p.p. (
<bibRefCitation DOI="https://doi.org/10.3767/000651909X475473" author="Culmsee, H" journalOrPublisher="Blumea - Biodiversity, Evolution and Biogeography of Plants" pageId="25" pageNumber="72" pagination="119 - 123" refId="B18" refString="Culmsee, H, Pitopang, R, 2009. Tree diversity in sub-montane and lower montane primary rain forests in Central Sulawesi. Blumea - Biodiversity, Evolution and Biogeography of Plants 54: 119 - 123, DOI: https://doi.org/10.3767/000651909X475473" title="Tree diversity in sub-montane and lower montane primary rain forests in Central Sulawesi." url="https://doi.org/10.3767/000651909X475473" volume="54" year="2009">Culmsee and Pitopang 2009</bibRefCitation>
, see also 2017 (Erratum)).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="62" type="diagnosis">
<paragraph pageId="15" pageNumber="62">Diagnosis.</paragraph>
<paragraph pageId="15" pageNumber="62">
<taxonomicName authorityName="Brambach, Byng &amp; Culmsee" authorityYear="2017" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium devogelii" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="species" species="devogelii">
<emphasis italics="true" pageId="15" pageNumber="62">Syzygium devogelii</emphasis>
</taxonomicName>
is a species of treelets characterised by slender, narrowly winged young branchlets, medium-sized narrowly elliptic leaves, straight and distinct secondary veins connected by an intramarginal vein impressed above and prominent beneath, small flowers (5 
<normalizedToken originalValue="×">x</normalizedToken>
3 mm in bud) in terminal inflorescences that develop into rather large fruits (c. 20 
<normalizedToken originalValue="×">x</normalizedToken>
25 mm), mature seeds lacking a testa, and cotyledons with echinate outer surfaces. The species is morphologically similar to 
<taxonomicName authorityName="Masamune" authorityYear="1942" baseAuthorityName="Merr." class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium perspicuinervium" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="species" species="perspicuinervium">
<emphasis italics="true" pageId="15" pageNumber="62">Syzygium perspicuinervium</emphasis>
</taxonomicName>
(Merr.) 
<bibRefCitation author="Masamune, G" journalOrPublisher="Laurentius Salvius, Holmia [Stockholm]" pageId="27" pageNumber="74" refId="B39" refString="Masamune, G, 1942. Boruneo no kenka shokubutsu. Enumeratio phanerogamarum Bornearum. Taihoku Imperial University, Taihoku, Taiwan." title="Boruneo no kenka shokubutsu. Enumeratio phanerogamarum Bornearum. Taihoku Imperial University, Taihoku, Taiwan." year="1942">Masamune (1942</bibRefCitation>
, 537) but differs from that species in smaller leaves with fewer secondary veins and in flowers with distinct calyx lobes (vs calyx calyptrate). It is furthermore similar to 
<taxonomicName authorityName="Merr &amp; Perry" authorityYear="1939" baseAuthorityName="Duthie" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium valdevenosum" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="species" species="valdevenosum">
<emphasis italics="true" pageId="15" pageNumber="62">Syzygium valdevenosum</emphasis>
</taxonomicName>
(Duthie) 
<bibRefCitation DOI="https://doi.org/10.2307/25058505" author="Merrill, ED" journalOrPublisher="Memoirs of the American Academy of Arts and Sciences" pageId="27" pageNumber="74" pagination="135 - 202" refId="B47" refString="Merrill, ED, Perry, LM, 1939. The Myrtaceous genus Syzygium Gaertner in Borneo. Memoirs of the American Academy of Arts and Sciences 18: 135 - 202, DOI: https://doi.org/10.2307/25058505" title="The Myrtaceous genus Syzygium Gaertner in Borneo." url="https://doi.org/10.2307/25058505" volume="18" year="1939">Merrill and Perry (1939</bibRefCitation>
, 182) but differs in lateral veins which are impressed above (vs prominent), much smaller inflorescences, and smaller, obconical (vs infundibuliform) flowers. Floral formula B1 Bt2 K4* C4* A∞* 
<normalizedToken originalValue="Ĝ">Ĝ</normalizedToken>
(2)┼ Vx∞.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="62" type="materials_examined">
<paragraph pageId="15" pageNumber="62">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="15" pageNumber="62">
<materialsCitation collectingDate="1979-05-11" collectorName="de Vogel EF" country="INDONESIA" elevation="1350" latitude="-1.2916666" location="Mt Rorekautimbu" longLatPrecision="130" longitude="120.27167" specimenCount="1" stateProvince="Central Sulawesi" typeStatus="holotype">
<collectingCountry name="Indonesia">INDONESIA</collectingCountry>
. 
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Central Sulawesi</collectingRegion>
(
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Sulawesi Tengah</collectingRegion>
), LLNP, Kab. Poso, Kec. Lore Utara, west slope of 
<location LSID="urn:lsid:plazi:treatment:0E32E7B8828E365249D5DF7F0D2B6CC0:AB53FA149857E74CB1C4AEEC1884CC45" country="INDONESIA" latitude="-1.2916666" longLatPrecision="130" longitude="120.27167" name="Mt Rorekautimbu" stateProvince="Central Sulawesi">Mt Rorekautimbu</location>
, c. 
<geoCoordinate degrees="1" direction="south" minutes="17.5" orientation="latitude" precision="92" value="-1.2916666">1°17.5'S</geoCoordinate>
<geoCoordinate degrees="120" direction="east" minutes="16.3" orientation="longitude" precision="92" value="120.27167">120°16.3'E</geoCoordinate>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.35" unit="m" value="1350.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.35" unit="m" value="1350.0">1350 m</elevation>
</quantity>
, 
<collectingDate value="1979-05-11">11 May 1979</collectingDate>
: 
<emphasis italics="true" pageId="15" pageNumber="62">
<collectorName>de Vogel EF</collectorName>
5293
</emphasis>
(fruits; 
<typeStatus>holotype</typeStatus>
L [L.2535665]! [L.2535666]!; isotype K!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="62" type="description">
<paragraph pageId="15" pageNumber="62">Description.</paragraph>
<paragraph pageId="15" pageNumber="62">
<emphasis bold="true" pageId="15" pageNumber="62">Trees</emphasis>
, up to 13 m tall, diameter at breast height ≤ 13 cm, trunk ≤ 7 m tall. Outer 
<emphasis bold="true" pageId="15" pageNumber="62">bark</emphasis>
whitish to brown, mealy or peeling off in thin sheets, inner bark pale or dark red, wood cream-coloured. Young 
<emphasis bold="true" pageId="15" pageNumber="62">branchlets</emphasis>
1-2.5 
<normalizedToken originalValue="×">x</normalizedToken>
2-3 mm, 
<normalizedToken originalValue="±">+/-</normalizedToken>
flattened, angular or oblong in cross section with 4 narrow wings, epidermis dark red when young, drying reddish or yellowish brown, smooth; becoming rounded with 4 ridges, bark (yellowish) brown, peeling off in thin sheets.
</paragraph>
<paragraph pageId="15" pageNumber="62">
<emphasis bold="true" pageId="15" pageNumber="62">Leaves</emphasis>
(sub-)opposite. Petioles 7-16 
<normalizedToken originalValue="×">x</normalizedToken>
1-3 mm, channelled above, rounded beneath, epidermis drying smooth or with transverse cracks. Blades (12.5-) 14-19 (-22.5) 
<normalizedToken originalValue="×">x</normalizedToken>
(4-) 4.5-7 (-8.5) cm, ratio (2.1-) 2.6-3.3 (-4), narrowly elliptic (or lanceolate), base cuneate or obtuse, apex acuminate, margin revolute; chartaceous or coriaceous, red or pink when young, above, beneath, drying dull to satin, variable in colour from greyish brown and olive grey to very dusky red above, dull to satin and, dark reddish brown beneath; pellucid dots rather few, visible or not on both sides. Midrib channelled above, very prominent, rounded, smooth and drying darker than the lamina beneath. Secondary vein pairs (9-) 11-14 (-17), 5-22 mm apart, channelled or impressed above, prominent and drying darker than the lamina beneath, straight or slightly arching from the midrib; intersecondary veins sometimes present. Tertiary veins dense, 
<normalizedToken originalValue="±">+/-</normalizedToken>
ladder-like and perpendicular to the midrib, faint above, prominulous beneath. Inner intramarginal vein 2-9 mm from leaf margin, looping or not and prominent; outer intramarginal vein 0.5-1.5 mm from leaf margin, as prominent as tertiary venation.
</paragraph>
<paragraph pageId="15" pageNumber="62">
<emphasis bold="true" pageId="15" pageNumber="62">Inflorescences</emphasis>
terminal, dense metabotryoids, 2.5 cm long, peduncles 1 cm long, axes flattened, with 2 or 4 narrow wings, drying brown. Bracts c. 1.5 mm long, ovate, keeled, caducous; bracteoles 2 per flower, 1 mm long, similar to bracts.
</paragraph>
<paragraph pageId="15" pageNumber="62">
<emphasis bold="true" pageId="15" pageNumber="62">Flowers</emphasis>
c. 15 per inflorescence, within the inflorescence in triads, 4-merous, anthopodium absent, only known before anthesis, mature buds 5 
<normalizedToken originalValue="×">x</normalizedToken>
3 mm. Hypanthium c. 4 
<normalizedToken originalValue="×">x</normalizedToken>
3 mm, obconical, drying dark reddish brown, densely glandular-warty, hypanthium rim 2 mm long, glandular inside. Calyx lobes c. 1 
<normalizedToken originalValue="×">x</normalizedToken>
2 mm, broadly rounded. Petals c. 3 
<normalizedToken originalValue="×">x</normalizedToken>
3 mm, cucullate in bud. Stamens c. 100, filaments 2-3 mm long, anthers c. 0.4 mm long, ellipsoid. Ovary bilocular, surrounded by spongy tissue, ovules numerous per locule, ascending. Style 3-4 mm long, pointed.
</paragraph>
<paragraph pageId="15" pageNumber="62">
<emphasis bold="true" pageId="15" pageNumber="62">Fruits</emphasis>
1-seeded, c. 20 
<normalizedToken originalValue="×">x</normalizedToken>
25 mm, irregularly depressed globose, laterally compressed, green, drying black and, smooth, pericarp 
<normalizedToken originalValue="±">+/-</normalizedToken>
woody, 1 mm thick, hypanthium rim 1-2 mm long, 5-9 mm in diameter.
</paragraph>
<paragraph pageId="15" pageNumber="62">
<emphasis bold="true" pageId="15" pageNumber="62">Seeds</emphasis>
c. 15 
<normalizedToken originalValue="×">x</normalizedToken>
20 mm, transverse ellipsoid, testa adhering to the pericarp, spongy inside and adhering to the outer surface of the cotyledons, cotyledons 
<normalizedToken originalValue="±">+/-</normalizedToken>
half-globose, facing surfaces undulate, outer surfaces densely echinate, protuberances obscured by spongy testa tissue.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="62" type="etymology">
<paragraph pageId="15" pageNumber="62">Etymology.</paragraph>
<paragraph pageId="15" pageNumber="62">The species is named after Eduard Ferdinand de Vogel (*1942). Ed de Vogel is a renowned authority on Malesian orchids, especially those from New Guinea. His contributions to the flora of Sulawesi are perhaps less well known: with almost 2000 specimens of excellent quality collected there in 1973-74 and 1979 - among them the type specimen of this species - he was one of the most prolific plant collectors on the island during the 20th century.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="62" type="phenology">
<paragraph pageId="15" pageNumber="62">Phenology.</paragraph>
<paragraph pageId="15" pageNumber="62">Flowering was recorded in August, fruiting in May.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="62" type="distribution">
<paragraph pageId="15" pageNumber="62">Distribution and habitat.</paragraph>
<paragraph pageId="15" pageNumber="62">
<taxonomicName authorityName="Brambach, Byng &amp; Culmsee" authorityYear="2017" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium devogelii" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="species" species="devogelii">
<emphasis italics="true" pageId="15" pageNumber="62">Syzygium devogelii</emphasis>
</taxonomicName>
is endemic to the province of Central Sulawesi, currently known to occur in lower montane forest at two localities in LLNP from 1350-1400 m elevation (Figure 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution map of four species of Syzygium in Central Sulawesi: Syzygium contiguum (orange diamonds), S. devogelii (yellow dots), S. eymae (light blue square), and S. galanthum (green triangles). Lore Lindu National Park is indicated by a black line. Map created with QGIS (QGIS Development Team 2016) using the digital elevation model of Jarvis et al. (2008)." figureDoi="10.3897/phytokeys.81.13488.figure4" httpUri="https://binary.pensoft.net/fig/140631" pageId="15" pageNumber="62">4</figureCitation>
). In the Bariri NE inventory plot, it was fairly common, growing on mid-slope terraces with Rhodic Ferralsols (
<bibRefCitation author="IUSS Working Group WRB" journalOrPublisher="FAO, Rome" pageId="26" pageNumber="73" publicationUrl="http://www.fao.org/3/a-i3794e.pdf" refId="B33" refString="IUSS Working Group WRB, 2014. World Reference Base for Soil Resources 2014. International soil classification system for naming soils and creating legends for soil maps. FAO, Rome, http://www.fao.org/3/a-i3794e.pdf" title="World Reference Base for Soil Resources 2014. International soil classification system for naming soils and creating legends for soil maps." url="http://www.fao.org/3/a-i3794e.pdf" year="2014">IUSS Working Group WRB 2014</bibRefCitation>
) derived from acid plutonic rocks. The forest there was dominated by 
<taxonomicName authorityName="Brambach &amp; Byng &amp; Culmsee" authorityYear="2017" family="Fagaceae" kingdom="Plantae" lsidName="" order="Myrtales" pageId="15" pageNumber="62" rank="family">Fagaceae</taxonomicName>
, 
<taxonomicName authorityName="Brambach &amp; Byng &amp; Culmsee" authorityYear="2017" family="Myrtaceae" kingdom="Plantae" lsidName="" order="Myrtales" pageId="15" pageNumber="62" rank="family">Myrtaceae</taxonomicName>
, 
<taxonomicName authorityName="Brambach &amp; Byng &amp; Culmsee" authorityYear="2017" family="Burseraceae" kingdom="Plantae" lsidName="" order="Myrtales" pageId="15" pageNumber="62" rank="family">Burseraceae</taxonomicName>
, 
<taxonomicName authorityName="Brambach &amp; Byng &amp; Culmsee" authorityYear="2017" family="Lauraceae" kingdom="Plantae" lsidName="" order="Myrtales" pageId="15" pageNumber="62" rank="family">Lauraceae</taxonomicName>
, and 
<taxonomicName authorityName="Brambach &amp; Byng &amp; Culmsee" authorityYear="2017" family="Elaeocarpaceae" kingdom="Plantae" lsidName="" order="Myrtales" pageId="15" pageNumber="62" rank="family">Elaeocarpaceae</taxonomicName>
(families with top five FIV) and contained six other species of 
<taxonomicName authorityName="Brambach &amp; Byng &amp; Culmsee" authorityYear="2017" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="62">Syzygium</emphasis>
</taxonomicName>
: 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. acuminatissimum" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="species" species="acuminatissimum">
<emphasis italics="true" pageId="15" pageNumber="62">S. acuminatissimum</emphasis>
</taxonomicName>
, 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. baeuerlenii" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="species" species="baeuerlenii">S. aff. baeuerlenii</taxonomicName>
(F.Muell.) Craven and Biffin (in 
<bibRefCitation DOI="https://doi.org/10.3767/000651906X622382" author="Craven, LA" journalOrPublisher="Blumea - Biodiversity, Evolution and Biogeography of Plants" pageId="25" pageNumber="72" pagination="131 - 142" refId="B16" refString="Craven, LA, Biffin, E, Ashton, PS, 2006. Acmena, Acmenosperma, Cleistocalyx, Piliocalyx and Waterhousea Formally Transferred to Syzygium (Myrtaceae). Blumea - Biodiversity, Evolution and Biogeography of Plants 51: 131 - 142, DOI: https://doi.org/10.3767/000651906X622382" title="Acmena, Acmenosperma, Cleistocalyx, Piliocalyx and Waterhousea Formally Transferred to Syzygium (Myrtaceae)." url="https://doi.org/10.3767/000651906X622382" volume="51" year="2006">Craven et al. 2006</bibRefCitation>
, 135), 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. lineatum" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="species" species="lineatum">
<emphasis italics="true" pageId="15" pageNumber="62">S. lineatum</emphasis>
</taxonomicName>
, 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. zeylanicum" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="species" species="zeylanicum">
<emphasis italics="true" pageId="15" pageNumber="62">S. zeylanicum</emphasis>
</taxonomicName>
(L.) de 
<bibRefCitation author="Candolle, AP de" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="25" pageNumber="72" publicationUrl="http://biodiversitylibrary.org/page/152793" refId="B12" refString="Candolle, AP de, 1828. Prodromus systematis naturalis regni vegetabilis 3. Treuttel &amp; Wuertz, Paris. http://biodiversitylibrary.org/page/152793" title="Prodromus systematis naturalis regni vegetabilis 3. Treuttel &amp; Wuertz, Paris." url="http://biodiversitylibrary.org/page/152793" year="1828">Candolle (1828</bibRefCitation>
, 260), and two undetermined species (
<bibRefCitation author="Brambach, F" journalOrPublisher="Tectona" pageId="25" pageNumber="72" refId="B9" refString="Brambach, F, Byng, JW, Culmsee, H, in press. Diversity, endemism, and composition of tropical mountain forest communities in Sulawesi, Indonesia, in relation to elevation and soil properties. Perspectives in Plant Ecology, Evolution and Systematics." title="Diversity, endemism, and composition of tropical mountain forest communities in Sulawesi, Indonesia, in relation to elevation and soil properties. Perspectives in Plant Ecology, Evolution and Systematics." year="in press">Brambach et al. in press</bibRefCitation>
). See 
<bibRefCitation DOI="https://doi.org/10.3767/000651909X475473" author="Culmsee, H" journalOrPublisher="Blumea - Biodiversity, Evolution and Biogeography of Plants" pageId="25" pageNumber="72" pagination="119 - 123" refId="B18" refString="Culmsee, H, Pitopang, R, 2009. Tree diversity in sub-montane and lower montane primary rain forests in Central Sulawesi. Blumea - Biodiversity, Evolution and Biogeography of Plants 54: 119 - 123, DOI: https://doi.org/10.3767/000651909X475473" title="Tree diversity in sub-montane and lower montane primary rain forests in Central Sulawesi." url="https://doi.org/10.3767/000651909X475473" volume="54" year="2009">Culmsee and Pitopang (2009)</bibRefCitation>
for more information on the floristics of the Bariri forest.
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="63" pageId="15" pageNumber="62" type="conservation status">
<paragraph pageId="15" pageNumber="62">Conservation status.</paragraph>
<paragraph lastPageId="16" lastPageNumber="63" pageId="15" pageNumber="62">
<taxonomicName authorityName="Brambach, Byng &amp; Culmsee" authorityYear="2017" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium devogelii" order="Myrtales" pageId="15" pageNumber="62" phylum="Tracheophyta" rank="species" species="devogelii">
<emphasis italics="true" pageId="15" pageNumber="62">Syzygium devogelii</emphasis>
</taxonomicName>
has a limited geographical distribution and seems to be restricted to lower montane forest within a narrow elevational belt. Known from only two localities, the EOO and AOO cannot be estimated reliably for the species. Because of the low collection density in Central Sulawesi, we believe that the species is more widespread and common than it currently appears. Deforestation has been recorded close to the type locality (using the Global Forest Change website, 
<bibRefCitation DOI="https://doi.org/10.1126/science.1244693" author="Hansen, MC" journalOrPublisher="Science" pageId="26" pageNumber="73" pagination="850 - 853" refId="B27" refString="Hansen, MC, Potapov, PV, Moore, R, Hancher, M, Turubanova, SA, Tyukavina, A, Thau, D, Stehman, SV, Goetz, SJ, Loveland, TR, Kommareddy, A, Egorov, A, Chini, L, Justice, CO, Townshend, JRG, 2013. High-Resolution Global Maps of 21st-Century Forest Cover Change. Science 342: 850 - 853, DOI: https://doi.org/10.1126/science.1244693" title="High-Resolution Global Maps of 21 st-Century Forest Cover Change." url="https://doi.org/10.1126/science.1244693" volume="342" year="2013">Hansen et al. 2013</bibRefCitation>
). Given the apparent narrow geographical and elevational distribution, ongoing deforestation and the recommendation to use a precautionary attitude 
<pageBreakToken pageId="16" pageNumber="63" start="start">in</pageBreakToken>
conservation assessments (
<bibRefCitation author="IUCN, Standards and Petitions Subcommittee" journalOrPublisher="Australian Journal of Botany Supplementary Series" pageId="26" pageNumber="73" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B32" refString="IUCN, Standards and Petitions Subcommittee, 2017. Guidelines for Using the IUCN Red List Categories and Criteria. Version 13. Prepared by the Standards and Petitions Subcommittee. IUCN, 87 pp. http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 13. Prepared by the Standards and Petitions Subcommittee. IUCN, 87 pp." url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2017">IUCN Standards and Petitions Subcommittee 2014</bibRefCitation>
) we propose a preliminary extinction risk assessment of 
<normalizedToken originalValue="“Endangered”">&quot;Endangered&quot;</normalizedToken>
(EN B1ab(i,ii,iii)).
</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="64" pageId="16" pageNumber="63" type="notes">
<paragraph pageId="16" pageNumber="63">Notes.</paragraph>
<paragraph pageId="16" pageNumber="63">
Most species of 
<taxonomicName authorityName="Brambach &amp; Byng &amp; Culmsee" authorityYear="2017" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium" order="Myrtales" pageId="16" pageNumber="63" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="16" pageNumber="63">Syzygium</emphasis>
</taxonomicName>
are reported to have cotyledons with rather smooth outer surfaces, unlike the peculiar echinate cotyledons of 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. devogelii" order="Myrtales" pageId="16" pageNumber="63" phylum="Tracheophyta" rank="species" species="devogelii">
<emphasis italics="true" pageId="16" pageNumber="63">S. devogelii</emphasis>
</taxonomicName>
. We here interpret the tissue covering the outer surface of the cotyledons (Figure 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Syzygium devogelii: a leafy twig with fruit b longitudinal section of ripe fruit with inner, flat side of cotyledon and echinate outer surface c flower bud, longitudinal section d flower bud, exterior view. a-b holotype de Vogel 5293 c-d Culmsee 1564." figureDoi="10.3897/phytokeys.81.13488.figure5" httpUri="https://binary.pensoft.net/fig/140632" pageId="16" pageNumber="63">5</figureCitation>
) and obscuring its protuberances as derived from the testa, as reported for the Australian species 
<taxonomicName authorityName="B. P. M. Hyland" authorityYear="1983" baseAuthorityName="F. M. Bail." class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium bungadinnia" order="Myrtales" pageId="16" pageNumber="63" phylum="Tracheophyta" rank="species" species="bungadinnia">
<emphasis italics="true" pageId="16" pageNumber="63">Syzygium bungadinnia</emphasis>
</taxonomicName>
(F.M.Bailey) 
<bibRefCitation DOI="https://doi.org/10.1071/BT8309001" author="Hyland, BP" journalOrPublisher="Australian Journal of Botany Supplementary Series" pageId="26" pageNumber="73" pagination="1 - 164" refId="B30" refString="Hyland, BP, 1983. A Revision of Syzygium and Allied Genera (Myrtaceae) in Australia. Australian Journal of Botany Supplementary Series 13: 1 - 164, DOI: https://doi.org/10.1071/BT8309001" title="A Revision of Syzygium and Allied Genera (Myrtaceae) in Australia." url="https://doi.org/10.1071/BT8309001" volume="13" year="1983">Hyland (1983</bibRefCitation>
, 64), but closer examinations of fruit and seed structures are necessary to corroborate this interpretation.
</paragraph>
<paragraph pageId="17" pageNumber="64">
<pageBreakToken pageId="17" pageNumber="64" start="start">Juvenile</pageBreakToken>
specimens of 
<taxonomicName authorityName="Brambach, Byng &amp; Culmsee" authorityYear="2017" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium balgooyi" order="Myrtales" pageId="17" pageNumber="64" phylum="Tracheophyta" rank="species" species="balgooyi">
<emphasis italics="true" pageId="17" pageNumber="64">Syzygium balgooyi</emphasis>
</taxonomicName>
are similar to 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. devogelii" order="Myrtales" pageId="17" pageNumber="64" phylum="Tracheophyta" rank="species" species="devogelii">
<emphasis italics="true" pageId="17" pageNumber="64">S. devogelii</emphasis>
</taxonomicName>
in their leaf shape, colour, and venation. In fact, both species were treated as one morphotype in 
<bibRefCitation DOI="https://doi.org/10.3767/000651909X475473" author="Culmsee, H" journalOrPublisher="Blumea - Biodiversity, Evolution and Biogeography of Plants" pageId="25" pageNumber="72" pagination="119 - 123" refId="B18" refString="Culmsee, H, Pitopang, R, 2009. Tree diversity in sub-montane and lower montane primary rain forests in Central Sulawesi. Blumea - Biodiversity, Evolution and Biogeography of Plants 54: 119 - 123, DOI: https://doi.org/10.3767/000651909X475473" title="Tree diversity in sub-montane and lower montane primary rain forests in Central Sulawesi." url="https://doi.org/10.3767/000651909X475473" volume="54" year="2009">Culmsee and Pitopang (2009</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.3767/000651917X694813" author="Culmsee, H" journalOrPublisher="Blumea - Biodiversity, Evolution and Biogeography of Plants" pageId="25" pageNumber="72" refId="B19" refString="Culmsee, H, Pitopang, R, 2017. Erratum: Tree diversity in sub-montane and lower montane primary rain forests in Central Sulawesi. Blumea - Biodiversity, Evolution and Biogeography of Plants 62: 6. https://doi.org/10.3767/000651917X694813" title="Erratum: Tree diversity in sub-montane and lower montane primary rain forests in Central Sulawesi. Blumea - Biodiversity, Evolution and Biogeography of Plants 62: 6." url="https://doi.org/10.3767/000651917X694813" year="2017">2017</bibRefCitation>
). Besides the very different flowers, they can, however, be distinguished by the shape of the young branchlets: strongly flattened and with rounded ridges in 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. balgooyi" order="Myrtales" pageId="17" pageNumber="64" phylum="Tracheophyta" rank="species" species="balgooyi">
<emphasis italics="true" pageId="17" pageNumber="64">S. balgooyi</emphasis>
</taxonomicName>
(Figure 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Morphological characters of Syzygium balgooyi and S. schumannianum. Syzygium balgooyi (a-k): a c. 20 m tall trunk b upper leaf surface; c lower leaf surface d bark with slash e ridged shoot apex with subopposite leaves f flowers at different stages during anthesis and fruit g trunk base with steep narrow buttresses h inflorescence with flower buds i shoot with young leaves j dried flowers before and during anthesis and longitudinal section of flower k dried fruit and longitudinal section of fruit showing two cotyledons. Syzygium schumannianum (l-m): l dried flowers before and during anthesis m dried fruit. a-b and h-i Brambach et al. 1564 c Brambach et al. 0861 d-e Brambach et al. 0628 f sighting on Mt Katopas by FB g Brambach et al. 0889; j holotype van Balgooy 3956 [L. 2517558] k de Vogel 5413 [L. 2517563] l Wiakabu et al. LAE 50571 [L. 2535534] m Brass 13610 [L. 2524420]. All scale bars: 1 cm." figureDoi="10.3897/phytokeys.81.13488.figure1" httpUri="https://binary.pensoft.net/fig/140628" pageId="17" pageNumber="64">1e</figureCitation>
) vs 
<normalizedToken originalValue="±">+/-</normalizedToken>
flattened with 4 narrow wings in 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. devogelii" order="Myrtales" pageId="17" pageNumber="64" phylum="Tracheophyta" rank="species" species="devogelii">
<emphasis italics="true" pageId="17" pageNumber="64">S. devogelii</emphasis>
</taxonomicName>
(Figure 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Syzygium devogelii: a leafy twig with fruit b longitudinal section of ripe fruit with inner, flat side of cotyledon and echinate outer surface c flower bud, longitudinal section d flower bud, exterior view. a-b holotype de Vogel 5293 c-d Culmsee 1564." figureDoi="10.3897/phytokeys.81.13488.figure5" httpUri="https://binary.pensoft.net/fig/140632" pageId="17" pageNumber="64">5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="64" type="materials_examined">
<paragraph pageId="17" pageNumber="64">Additional specimens examined</paragraph>
<paragraph pageId="17" pageNumber="64">
<emphasis bold="true" pageId="17" pageNumber="64">
(
<typeStatus>Paratypes</typeStatus>
). 
<materialsCitation collectingDate="2017-01-01" collectingDateMax="2017-12-31" collectingDateMin="2017-01-01" country="INDONESIA" location="Central Sulawesi" specimenCount="1" stateProvince="Central Sulawesi">
<collectingCountry name="Indonesia">INDONESIA</collectingCountry>
. 
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Central Sulawesi</collectingRegion>
(
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Sulawesi Tengah</collectingRegion>
)
</materialsCitation>
, 
<materialsCitation collectingDate="2006-09-09" collectingDateMax="2011-08-18" collectingDateMin="2006-09-09" collectorName="Bariri NE, Culmsee H, Brambach F, Mangopo H, Firdaus, Faber M, Tiranda R" country="Indonesia" elevation="100" latitude="-1.6566668" location="Bariri" longLatPrecision="130" longitude="120.175" specimenCount="1" stateProvince="Kec. Lore Tengah">
LLNP, Kab. Poso, 
<collectingRegion inferredBySuffix="true">Kec. Lore Tengah</collectingRegion>
: 
<locationDeviation location="Bariri">
<quantity metricMagnitude="3" metricUnit="m" metricValue="9.0" unit="km" value="9.0">9 km</quantity>
NW of Bariri
</locationDeviation>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
</quantity>
east of climate tower, tree-inventory plot 
<collectorName>
<location LSID="urn:lsid:plazi:treatment:0E32E7B8828E365249D5DF7F0D2B6CC0:B3F572D7E0CDECB8DA1CEECEC7EA5AA4" country="Indonesia" latitude="-1.6566668" longLatPrecision="130" longitude="120.175" name="Bariri" stateProvince="Kec. Lore Tengah">Bariri</location>
NE
</collectorName>
, 
<geoCoordinate degrees="1" direction="south" minutes="39.4" orientation="latitude" precision="92" value="-1.6566668">1°39.4'S</geoCoordinate>
, 
<geoCoordinate degrees="120" direction="east" minutes="10.5" orientation="longitude" precision="92" value="120.175">120°10.5'E</geoCoordinate>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</elevation>
</quantity>
, 
<collectingDate value="2006-09-09">9 Sep 2006</collectingDate>
: 
<emphasis italics="true" pageId="17" pageNumber="64">
<collectorName>Culmsee H</collectorName>
1333
</emphasis>
(sterile; BO [BO-1938455]!, CEB) &amp; 
<emphasis italics="true" pageId="17" pageNumber="64">1378</emphasis>
(sterile; CEB, K [K000993491]!); ibid. loco, 
<collectingDate value="2011-08-18">18 Aug 2011</collectingDate>
: 
<emphasis italics="true" pageId="17" pageNumber="64">
<collectorName>Brambach F</collectorName>
, 
<collectorName>Mangopo H</collectorName>
, 
<collectorName>Firdaus</collectorName>
, 
<collectorName>Faber M</collectorName>
, 
<collectorName>Tiranda R</collectorName>
0818
</emphasis>
(sterile; BO [BO-1938442]!, CEB, GOET [GOET020015]!) &amp; 
<emphasis italics="true" pageId="17" pageNumber="64">0845</emphasis>
(sterile; BO [BO-1938443]!, CEB, L!)
</materialsCitation>
</emphasis>
.
</paragraph>
<paragraph pageId="17" pageNumber="64">
<materialsCitation collectingDate="2006-07" collectorName="Bariri S, Culmsee H" elevation="80" latitude="-1.6583333" location="Bariri" longLatPrecision="130" longitude="120.17333" specimenCount="1">
<locationDeviation location="Bariri">
<quantity metricMagnitude="3" metricUnit="m" metricValue="9.0" unit="km" value="9.0">9 km</quantity>
NW of Bariri
</locationDeviation>
, 
<quantity metricMagnitude="1" metricUnit="m" metricValue="8.0" unit="m" value="80.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="8.0" unit="m" value="80.0">80 m</elevation>
</quantity>
south of climate tower, tree-inventory plot 
<collectorName>
<location LSID="urn:lsid:plazi:treatment:0E32E7B8828E365249D5DF7F0D2B6CC0:FA2B3F5398D52D6254B2CFA138779C56" latitude="-1.6583333" longLatPrecision="130" longitude="120.17333" name="Bariri">Bariri</location>
S
</collectorName>
, 
<geoCoordinate degrees="1" direction="south" minutes="39.5" orientation="latitude" precision="92" value="-1.6583333">1°39.5'S</geoCoordinate>
, 
<geoCoordinate degrees="120" direction="east" minutes="10.4" orientation="longitude" precision="92" value="120.17333">120°10.4'E</geoCoordinate>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</elevation>
</quantity>
, 
<collectingDate value="2006-07">Jul 2006</collectingDate>
: 
<emphasis italics="true" pageId="17" pageNumber="64">
<collectorName>Culmsee H</collectorName>
1252
</emphasis>
(sterile; CEB, GOET [GOET020016]!) &amp; 
<emphasis italics="true" pageId="17" pageNumber="64">1564</emphasis>
(flower buds; CEB, L!)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/phytokeys.81.13488.figure4" httpUri="https://binary.pensoft.net/fig/140631" pageId="17" pageNumber="64" start="Figure 4" startId="F4">
<paragraph pageId="17" pageNumber="64">
<emphasis bold="true" pageId="17" pageNumber="64">Figure 4.</emphasis>
Distribution map of four species of 
<taxonomicName authorityName="Brambach &amp; Byng &amp; Culmsee" authorityYear="2017" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium" order="Myrtales" pageId="17" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="17" pageNumber="64">Syzygium</emphasis>
</taxonomicName>
in 
<collectingRegion country="Indonesia" name="Sulawesi Tengah">Central Sulawesi</collectingRegion>
: 
<taxonomicName authorityName="Brambach, Byng &amp; Culmsee" authorityYear="2017" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syzygium contiguum" order="Myrtales" pageId="17" pageNumber="64" phylum="Tracheophyta" rank="species" species="contiguum">
<emphasis italics="true" pageId="17" pageNumber="64">Syzygium contiguum</emphasis>
</taxonomicName>
(orange diamonds), 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. devogelii" order="Myrtales" pageId="17" pageNumber="64" phylum="Tracheophyta" rank="species" species="devogelii">
<emphasis italics="true" pageId="17" pageNumber="64">S. devogelii</emphasis>
</taxonomicName>
(yellow dots), 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. eymae" order="Myrtales" pageId="17" pageNumber="64" phylum="Tracheophyta" rank="species" species="eymae">
<emphasis italics="true" pageId="17" pageNumber="64">S. eymae</emphasis>
</taxonomicName>
(light blue square), and 
<taxonomicName class="Magnoliopsida" family="Myrtaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. galanthum" order="Myrtales" pageId="17" pageNumber="64" phylum="Tracheophyta" rank="species" species="galanthum">
<emphasis italics="true" pageId="17" pageNumber="64">S. galanthum</emphasis>
</taxonomicName>
(green triangles). Lore Lindu National Park is indicated by a black line. Map created with QGIS (
<bibRefCitation author="QGIS Development Team" journalOrPublisher="Taxon" pageId="28" pageNumber="75" publicationUrl="http://www.qgis.org" refId="B54" refString="QGIS Development Team, 2016. QGIS Geographic Information System. Version 2.14.2-Essen. Open Source Geospatial Foundation Project. http://www.qgis.org" title="QGIS Geographic Information System. Version 2.14.2 - Essen. Open Source Geospatial Foundation Project." url="http://www.qgis.org" year="2016">QGIS Development Team 2016</bibRefCitation>
) using the digital elevation model of 
<bibRefCitation author="Jarvis, A" journalOrPublisher="FAO, Rome" pageId="26" pageNumber="73" publicationUrl="http://srtm.csi.cgiar.org" refId="B34" refString="Jarvis, A, Reuter, HI, Nelson, A, Guevara, E, 2008. Hole-filled SRTM for the globe, Version 4. http://srtm.csi.cgiar.org" title="Hole-filled SRTM for the globe, Version 4." url="http://srtm.csi.cgiar.org" year="2008">Jarvis et al. (2008)</bibRefCitation>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>