305 lines
36 KiB
XML
305 lines
36 KiB
XML
<document id="AC612A815645F31F47F173368FB691C8" ID-DOI="10.11646/zootaxa.3641.4.2" ID-GBIF-Dataset="3b420769-6676-4089-be51-d5dd3c6e420e" ID-ISSN="1175-5326" ID-Zenodo-Dep="219655" ID-ZooBank="D9F4C67F-FE27-467B-B0DD-07A77E493538" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460595981066" checkinUser="plazi" docAuthor="Taphorn, Donald C., Armbruster, Jonathan W., Villa-Navarro, Francisco & Ray, C. Keith" docDate="2013" docId="0E26F513DE238466FF1EFA00FD499FAA" docLanguage="en" docName="zt03641p370.pdf" docOrigin="Zootaxa 3641 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ancistrus martini Schultz 1944" docType="treatment" docVersion="7" lastPageNumber="359" masterDocId="F21F8D6BDE2D8476FF89FFA7FFCE9842" masterDocTitle="Trans-Andean Ancistrus (Siluriformes: Loricariidae)" masterLastPageNumber="370" masterPageNumber="343" pageNumber="357" updateTime="1698326792420" updateUser="plazi">
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<mods:mods id="9CD8CB667314A423153628DD9878DB6D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3077BEF34699EBDE76F59C5635791A04">
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<mods:title id="09F3CC025BD1B1014CA35F72BC30E401">Trans-Andean Ancistrus (Siluriformes: Loricariidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B35E6D8CD7B881847ABEA7154C21C25C" type="personal">
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<mods:roleTerm id="C37618B5BC5FB9BB7DF30FD68D1F6DF8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0891B358728AFD3422DA79532114C20B">Taphorn, Donald C.</mods:namePart>
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</mods:name>
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<mods:name id="6FF0E9F31F16FF6ABF97FCF6704035E1" type="personal">
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<mods:roleTerm id="B010103B98B06C2CF9A245BF8156733A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9E49B639DBB57B812F956582F451E435">Armbruster, Jonathan W.</mods:namePart>
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</mods:name>
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<mods:name id="60C7F8091748AB37EF394B51B99C1527" type="personal">
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<mods:roleTerm id="1599F637BAB396A12F722C9931C97352">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ACC683BC8A70DF9762BC0E8DE1EBB05D">Villa-Navarro, Francisco</mods:namePart>
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</mods:name>
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<mods:name id="39DE9EA2EB60F00359B61E46C1D90E10" type="personal">
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<mods:roleTerm id="3F573F8630B0D58D70A8EFC76386ED37">Author</mods:roleTerm>
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<mods:namePart id="19F98F926A2AEFA636C58699551F12D8">Ray, C. Keith</mods:namePart>
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<mods:typeOfResource id="D7123AA98BC3D47408B2064C6C7D8E8A">text</mods:typeOfResource>
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<mods:title id="6672F2578D8EBA0212A85ACD6BEE2C36">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="83BE57AE105782A8A69C27E601BDA899">
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<mods:date id="9582BDF6C5CEC1F062B3FB031F5E0E2A">2013</mods:date>
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<mods:detail id="4C0715D8304B0CEA16D5A546409E3B83" type="volume">
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<mods:number id="78BADAE21A6D43C630DD87CE77A30804">3641</mods:number>
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</mods:detail>
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<mods:detail id="1758397977A969118AE98C5576441F54" type="issue">
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<mods:number id="790936719DFA59D6C7590051D644C402">4</mods:number>
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<mods:identifier id="9E90392AD8664B7CF37D3D4E945730DA" type="DOI">10.11646/zootaxa.3641.4.2</mods:identifier>
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<treatment id="0E26F513DE238466FF1EFA00FD499FAA" ID-DOI="http://doi.org/10.5281/zenodo.6157724" ID-GBIF-Taxon="119566644" ID-Zenodo-Dep="6157724" LSID="urn:lsid:plazi:treatment:0E26F513DE238466FF1EFA00FD499FAA" httpUri="http://treatment.plazi.org/id/0E26F513DE238466FF1EFA00FD499FAA" lastPageId="16" lastPageNumber="359" pageId="14" pageNumber="357">
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<subSubSection id="CE95178EDE238478FF1EFA00FD879E90" pageId="14" pageNumber="357" type="nomenclature">
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<paragraph id="86304405DE238478FF1EFA00FDDF9D83" blockId="14.[151,529,1447,1506]" box="[151,529,1447,1473]" pageId="14" pageNumber="357">
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<heading id="DD78F369DE238478FF1EFA00FDDF9D83" bold="true" box="[151,529,1447,1473]" fontSize="11" level="1" pageId="14" pageNumber="357" reason="1">
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<taxonomicName id="418F3F86DE238478FF1EFA00FDDF9D83" ID-CoL="DN43" authority="Schultz 1944" authorityName="Schultz" authorityYear="1944" box="[151,529,1447,1473]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="357" phylum="Chordata" rank="species" species="martini">
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||
<emphasis id="B4FB9817DE238478FF1EFA00FDDF9D83" bold="true" box="[151,529,1447,1473]" pageId="14" pageNumber="357">
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<emphasis id="B4FB9817DE238478FF1EFA00FEA29D83" bold="true" box="[151,364,1447,1473]" italics="true" pageId="14" pageNumber="357">Ancistrus martini</emphasis>
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Schultz 1944
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="86304405DE238478FF1EFA6DFEF59DA0" blockId="14.[151,529,1447,1506]" box="[151,315,1482,1506]" pageId="14" pageNumber="357">
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(
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<figureCitation id="1EB45880DE238478FF16FA6DFF239DA0" box="[159,237,1482,1506]" captionStart="FIGURE 8" captionStartId="16.[151,250,1506,1529]" captionTargetBox="[151,1435,251,1484]" captionTargetId="figure@16.[151,1436,251,1486]" captionTargetPageId="16" captionText="FIGURE 8. Ancistrus martini INHS 35367 specimen, 73.2 mm SL, Taguara Creek, Santa Ana River drainage, Lake Maracaibo Basin, 51 mm SL. Photograph by Nathan K. Lujan." httpUri="https://zenodo.org/record/219663/files/figure.png" pageId="14" pageNumber="357">Figs. 8</figureCitation>
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and
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<figureCitation id="1EB45880DE238478FEAAFA6DFEFC9DA0" box="[291,306,1482,1506]" captionStart="FIGURE 9" captionStartId="17.[151,250,869,892]" captionTargetBox="[188,1398,195,847]" captionTargetId="figure@17.[188,1398,195,847]" captionTargetPageId="17" captionText="FIGURE 9. Ancistrus martini, live, collected by and photographed by Gustavo Ballen from the upper Zulia River." httpUri="https://zenodo.org/record/219664/files/figure.png" pageId="14" pageNumber="357">9</figureCitation>
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)
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</paragraph>
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<paragraph id="86304405DE238478FF1EF9B4FC3F9E0C" blockId="14.[151,1437,1554,1614]" pageId="14" pageNumber="357">
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Images of the
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<typeStatus id="5934FAA7DE238478FEB1F9B5FE549E68" box="[312,410,1554,1578]" pageId="14" pageNumber="357" type="holotype">holotype</typeStatus>
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of this species, USNM
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<date id="F23162C5DE238478FD20F9B5FCCE9E68" box="[681,768,1554,1578]" pageId="14" pageNumber="357" value="64-12-10">121064</date>
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, are available at the All Catfish Species Inventory website: http://acsi.acnatsci.org/base/image_list.html?mode=genus&genus=
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<taxonomicName id="418F3F86DE238478FC0BF991FC3F9E0C" box="[898,1009,1590,1614]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="357" phylum="Chordata" rank="genus">Ancistrus</taxonomicName>
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</paragraph>
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<paragraph id="86304405DE238478FF1EF9D8FD879E90" blockId="14.[151,1436,1662,1992]" pageId="14" pageNumber="357">
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<taxonomicName id="418F3F86DE238478FF1EF9D8FD599ED6" authority="Schultz 1944: 302" authorityName="Schultz" authorityPageNumber="302" authorityYear="1944" box="[151,663,1662,1685]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="357" phylum="Chordata" rank="subSpecies" species="triradiatus" subSpecies="martini">
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<emphasis id="B4FB9817DE238478FF1EF9D8FE0D9ED7" box="[151,451,1663,1685]" italics="true" pageId="14" pageNumber="357">Ancistrus triradiatus martini</emphasis>
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, Schultz 1944: 302
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</taxonomicName>
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, pl. 9 (fig. C).
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<typeStatus id="5934FAA7DE238478FCC8F9D8FCBD9ED6" box="[833,883,1663,1684]" pageId="14" pageNumber="357">Type</typeStatus>
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locality: Río Táchira,
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<quantity id="4177E9E0DE238478FBE0F9D8FB6B9ED6" box="[1129,1189,1662,1684]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="357" unit="km" value="7.0">7 km</quantity>
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. north of San Antonio, Catatumbo system;
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<collectingCountry id="FE980495DE238478FE19F93AFE359EF1" box="[400,507,1693,1715]" name="Venezuela" pageId="14" pageNumber="357">Venezuela</collectingCountry>
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.
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<typeStatus id="5934FAA7DE238478FD8DF93AFDA89EF1" box="[516,614,1693,1715]" pageId="14" pageNumber="357" type="holotype">Holotype</typeStatus>
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: USNM
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<date id="F23162C5DE238478FD37F93AFCC39EF1" box="[702,781,1693,1715]" pageId="14" pageNumber="357" value="64-12-10">121064</date>
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||
.
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<typeStatus id="5934FAA7DE238478FC9EF939FCB39EF1" box="[791,893,1694,1715]" pageId="14" pageNumber="357" type="paratype">Paratypes</typeStatus>
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: MCZ 37233 (1), USNM 120094 (1),
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<date id="F23162C5DE238478FB74F93AFA879EF1" box="[1277,1353,1693,1715]" pageId="14" pageNumber="357" value="65-12-10">121065</date>
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(3, now 2, 1 to MCZ); Pérez & Viloria (1994).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="CE95178EDE238478FF1EF97CFC619F8A" pageId="14" pageNumber="357" type="description">
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||
<paragraph id="86304405DE238478FF1EF97CFBB89F4D" blockId="14.[151,1436,1662,1992]" pageId="14" pageNumber="357">
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||
<taxonomicName id="418F3F86DE238478FF1EF97CFE2A9EB2" authority="Schultz 1944" authorityName="Schultz" authorityYear="1944" box="[151,484,1754,1776]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="357" phylum="Chordata" rank="species" species="martini">
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<emphasis id="B4FB9817DE238478FF1EF97CFE849EB2" box="[151,330,1755,1776]" italics="true" pageId="14" pageNumber="357">Ancistrus martini</emphasis>
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, Schultz 1944
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</taxonomicName>
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. Isbrücker (1980) raised subspecies to species level; Ferraris & Vari (1992) (
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<typeStatus id="5934FAA7DE238478FA84F97CFAF49EB2" box="[1293,1338,1755,1776]" pageId="14" pageNumber="357">type</typeStatus>
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catalog); Burgess 1989; Isbrücker (2001); Isbrücker (2002); Fisch-Muller (2003) Ferraris (2007) (list).
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</paragraph>
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<paragraph id="86304405DE238478FF1EF8BEFD719FC8" blockId="14.[151,1436,1662,1992]" pageId="14" pageNumber="357">
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||
<taxonomicName id="418F3F86DE238478FF1EF8BEFD689F6C" authority="Schultz 1944:305" authorityName="Schultz" authorityPageNumber="305" authorityYear="1944" box="[151,678,1816,1838]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="357" phylum="Chordata" rank="subSpecies" species="brevifilis" subSpecies="bodenhameri">
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<emphasis id="B4FB9817DE238478FF1EF8BEFE259F6C" box="[151,491,1816,1838]" italics="true" pageId="14" pageNumber="357">Ancistrus brevifilis bodenhameri,</emphasis>
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Schultz 1944:305
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</taxonomicName>
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, pl. 10 (fig. A).
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<typeStatus id="5934FAA7DE238478FCC6F8BEFC4F9F6C" box="[847,897,1817,1838]" pageId="14" pageNumber="357">Type</typeStatus>
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locality: Río
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<collectingCountry id="FE980495DE238478FB86F8BFFBB69F6C" box="[1039,1144,1816,1838]" name="Saint Pierre and Miquelon" pageId="14" pageNumber="357">San Pedro</collectingCountry>
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at the bridge south of Mene Grande, Motatán system;
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<collectingCountry id="FE980495DE238478FE46F891FDF49F0E" box="[463,570,1846,1868]" name="Venezuela" pageId="14" pageNumber="357">Venezuela</collectingCountry>
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.
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<typeStatus id="5934FAA7DE238478FDCDF891FD689F0E" box="[580,678,1846,1868]" pageId="14" pageNumber="357" type="holotype">Holotype</typeStatus>
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: USNM
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<date id="F23162C5DE238478FC89F891FC819F0E" box="[768,847,1846,1868]" pageId="14" pageNumber="357" value="66-12-10">121066</date>
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.
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<typeStatus id="5934FAA7DE238478FCD3F890FC0F9F0E" box="[858,961,1847,1868]" pageId="14" pageNumber="357" type="paratype">Paratypes</typeStatus>
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||
: MCZ 37241 [ex USNM
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<date id="F23162C5DE238478FB4FF891FADB9F0E" box="[1222,1301,1846,1868]" pageId="14" pageNumber="357" value="67-12-10">121067</date>
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] (3); UMMZ 141940 [ex USNM
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<date id="F23162C5DE238478FE16F8F2FE209F29" box="[415,494,1877,1899]" pageId="14" pageNumber="357" value="67-12-10">121067</date>
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||
] (6); USNM
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<date id="F23162C5DE238478FD0DF8F2FD1E9F29" box="[644,720,1877,1899]" pageId="14" pageNumber="357" value="67-12-10">121067</date>
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(132, now 123),
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<date id="F23162C5DE238478FC1DF8F2FC2E9F29" box="[916,992,1877,1899]" pageId="14" pageNumber="357" value="68-12-10">121068</date>
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||
–69 (1, 2),
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<date id="F23162C5DE238478FBE9F8F1FB629F29" box="[1120,1196,1878,1899]" pageId="14" pageNumber="357" value="99-12-12">121299</date>
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(1). Isbrücker (1980), relocated to species level, Pérez &
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<taxonomicName id="418F3F86DE238478FDA3F8D4FD759FC8" box="[554,699,1907,1930]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="370" phylum="Chordata" rank="species" species="galani" status="sp. nov.">Galani (1994)</taxonomicName>
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.
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</paragraph>
|
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<paragraph id="86304405DE238478FF1EF834FC619F8A" blockId="14.[151,1436,1662,1992]" pageId="14" pageNumber="357">
|
||
<taxonomicName id="418F3F86DE238478FF1EF834FDD59FEA" authority="Schultz 1944" authorityName="Schultz" authorityYear="1944" box="[151,539,1938,1960]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="357" phylum="Chordata" rank="species" species="bodenhameri">
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<emphasis id="B4FB9817DE238478FF1EF834FE459FEA" box="[151,395,1938,1960]" italics="true" pageId="14" pageNumber="357">Ancistrus bodenhameri,</emphasis>
|
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Schultz 1944
|
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</taxonomicName>
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||
. Ferraris & Vari (1992) list; Pérez & Viloria (1994); Isbrücker (1980); Burgess (1989); Isbrücker (2001); Isbrücker (2002); Fisch-Muller (2003); Ferraris (2007).
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CE95178EDE228479FF1EFF30FBF39AE5" pageId="15" pageNumber="358" type="diagnosis">
|
||
<paragraph id="86304405DE228479FF1EFF30FBF39AE5" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="358">
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||
<emphasis id="B4FB9817DE228479FF1EFF30FEDF98F2" bold="true" box="[151,273,151,176]" pageId="15" pageNumber="358">Diagnosis.</emphasis>
|
||
<taxonomicName id="418F3F86DE228479FE92FF3FFE2F98F2" box="[283,481,152,176]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="martini">
|
||
<emphasis id="B4FB9817DE228479FE92FF3FFE2F98F2" box="[283,481,152,176]" italics="true" pageId="15" pageNumber="358">Ancistrus martini</emphasis>
|
||
</taxonomicName>
|
||
can usually be distinguished from
|
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<taxonomicName id="418F3F86DE228479FCF1FF3FFB9B98F2" box="[888,1109,152,176]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="caucanus">
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||
<emphasis id="B4FB9817DE228479FCF1FF3FFB9B98F2" box="[888,1109,152,176]" italics="true" pageId="15" pageNumber="358">Ancistrus caucanus</emphasis>
|
||
</taxonomicName>
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||
by having a snout length to head length ratio of 55.5–65.3% (vs. 45.9–55.9%). Although pigmentation is notoriously variable among individuals of most
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<taxonomicName id="418F3F86DE228479FEFFFF47FE2E98BA" box="[374,480,224,248]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="genus">
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||
<emphasis id="B4FB9817DE228479FEFFFF47FE2E98BA" box="[374,480,224,248]" italics="true" pageId="15" pageNumber="358">Ancistrus</emphasis>
|
||
</taxonomicName>
|
||
species examined, most specimens of
|
||
<taxonomicName id="418F3F86DE228479FC18FF47FB9A98BA" box="[913,1108,224,248]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="martini">
|
||
<emphasis id="B4FB9817DE228479FC18FF47FB9A98BA" box="[913,1108,224,248]" italics="true" pageId="15" pageNumber="358">Ancistrus martini</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<emphasis id="B4FB9817DE228479FA0DFF47FEB2995E" italics="true" pageId="15" pageNumber="358">
|
||
<taxonomicName id="418F3F86DE228479FA0DFF47FECA995E" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="caucanus">A. caucanus</taxonomicName>
|
||
,
|
||
<taxonomicName id="418F3F86DE228479FE9BFEA3FEB2995E" box="[274,380,260,284]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="tolima">A. tolima</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="418F3F86DE228479FE4BFEA3FDB7995E" box="[450,633,260,284]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="vericaucanus">
|
||
<emphasis id="B4FB9817DE228479FE4BFEA3FDB7995E" box="[450,633,260,284]" italics="true" pageId="15" pageNumber="358">A. vericaucanus</emphasis>
|
||
</taxonomicName>
|
||
by having better defined light spots on body and fins in addition to relatively small dark spots on the fin rays and membranes that form six to nine irregular bands on the caudal fin (vs. three to five regular or irregular bands) and seven to nine spots along the pectoral-fin spine (vs. three to six spots). They also differ in the ratio of head-eye length to snout length: 53.9–72.1% in specimens greater than
|
||
<quantity id="4177E9E0DE228479FAAAFEC8FABA99C5" box="[1315,1396,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="358" unit="mm" value="60.0">60 mm</quantity>
|
||
SL (vs. 72.3–91.9%;
|
||
<figureCitation id="1EB45880DE228479FED4FE33FE5199EE" box="[349,415,404,428]" captionStart="FIGURE 3" captionStartId="6.[151,250,1323,1346]" captionTargetBox="[203,1405,226,1286]" captionTargetId="figure@6.[151,1436,193,1302]" captionTargetPageId="6" captionText="FIGURE 3. Diagnostic biplots for species of Ancistrus." httpUri="https://zenodo.org/record/219658/files/figure.png" pageId="15" pageNumber="358">Fig. 3</figureCitation>
|
||
C).
|
||
<taxonomicName id="418F3F86DE228479FE40FE33FDF499EE" box="[457,570,404,428]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="martini">
|
||
<emphasis id="B4FB9817DE228479FE40FE33FDF499EE" box="[457,570,404,428]" italics="true" pageId="15" pageNumber="358">A. martini</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="418F3F86DE228479FD46FE33FCA799EE" box="[719,873,404,428]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="centrolepis">
|
||
<emphasis id="B4FB9817DE228479FD46FE33FCA799EE" box="[719,873,404,428]" italics="true" pageId="15" pageNumber="358">A. centrolepis</emphasis>
|
||
</taxonomicName>
|
||
by having white spots on the body (vs. dark spots on body or body entirely gray), by lacking rows of enlarged odontodes on the lateral plates (vs. one to three rows of greatly enlarged odontodes) and by having the longest odontodes on the lateral plates less than half as long as the supporting plate is wide (vs. longest odontodes almost as long as the supporting plate is wide); from
|
||
<taxonomicName id="418F3F86DE228479FA99FDA7FAB69A5A" box="[1296,1400,512,536]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="galani">
|
||
<emphasis id="B4FB9817DE228479FA99FDA7FAB69A5A" box="[1296,1400,512,536]" italics="true" pageId="15" pageNumber="358">A. galani</emphasis>
|
||
</taxonomicName>
|
||
by having fully developed eyes and pigment (vs. eyes reduced and pigment absent or reduced); from
|
||
<taxonomicName id="418F3F86DE228479FB6FFD83FA839A7E" box="[1254,1357,548,572]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="tolima">
|
||
<emphasis id="B4FB9817DE228479FB6FFD83FA839A7E" box="[1254,1357,548,572]" italics="true" pageId="15" pageNumber="358">A. tolima</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B4FB9817DE228479FA0DFD83FEFB9A22" italics="true" pageId="15" pageNumber="358">
|
||
<taxonomicName id="418F3F86DE228479FA0DFD83FEE09A22" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="vericaucanus">A. vericaucanus</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
by usually having one preadipose plate (vs. two to three, rarely one in
|
||
<taxonomicName id="418F3F86DE228479FBDDFDEFFAC79A22" box="[1108,1289,584,608]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="vericaucanus">
|
||
<emphasis id="B4FB9817DE228479FBDDFDEFFAC79A22" box="[1108,1289,584,608]" italics="true" pageId="15" pageNumber="358">A. vericaucanus</emphasis>
|
||
</taxonomicName>
|
||
, and three to five in
|
||
<taxonomicName id="418F3F86DE228479FF63FDCBFE9C9AC6" box="[234,338,620,644]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="tolima">
|
||
<emphasis id="B4FB9817DE228479FF63FDCBFE9C9AC6" box="[234,338,620,644]" italics="true" pageId="15" pageNumber="358">A. tolima</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName id="418F3F86DE228479FE55FDCBFD8B9AC6" box="[476,581,620,644]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="358" phylum="Chordata" rank="species" species="tolima">
|
||
<emphasis id="B4FB9817DE228479FE55FDCBFD8B9AC6" box="[476,581,620,644]" italics="true" pageId="15" pageNumber="358">A. tolima</emphasis>
|
||
</taxonomicName>
|
||
by having the adipose fin spine at a 45° angle to the body and adipose-fin membrane present (vs. adipose-fin spine adnate and adipose-fin membrane absent).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CE95178EDE228466FF4EFD14FF2E9896" lastPageId="16" lastPageNumber="359" pageId="15" pageNumber="358" type="description">
|
||
<paragraph id="86304405DE228479FF4EFD14FB1A9A8E" blockId="15.[151,1437,151,2013]" box="[199,1236,691,717]" pageId="15" pageNumber="358">
|
||
<emphasis id="B4FB9817DE228479FF4EFD14FE979A8E" bold="true" box="[199,345,691,716]" pageId="15" pageNumber="358">Description.</emphasis>
|
||
N = 48. Morphometrics given in
|
||
<tableCitation id="CB0D71BEDE228479FD58FD13FCEB9A8E" box="[721,805,692,716]" captionStart="TABLE 3" captionStartId="11.[151,239,1747,1771]" captionTargetBox="[165,1428,1803,1991]" captionTargetPageId="11" captionText="TABLE 3. Selected morphometrics of Ancistrus chagresi and A. martini." httpUri="http://table.plazi.org/id/D2F0148DDE26847DFF1EF974FC6D9EA9" pageId="15" pageNumber="358" tableUuid="D2F0148DDE26847DFF1EF974FC6D9EA9">Table 3</tableCitation>
|
||
. Size range this study
|
||
<quantity id="4177E9E0DE228479FBA8FD13FB689A8E" box="[1057,1190,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="7.85" metricValueMax="11.1" metricValueMin="4.6" pageId="15" pageNumber="358" unit="mm" value="78.5" valueMax="111.0" valueMin="46.0">46–111 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
<paragraph id="86304405DE228479FF4EFD70FB759B1E" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="358">Body broadest anteriorly, greatest body width at base of pectoral fins then narrowing progressively to end of caudal peduncle. Head and body somewhat depressed, greatest body depth at or just posterior to pectoral-fin bases. Caudal peduncle deep, posteriorly compressed. Dorsal profile anteriorly convex from tip of snout to dorsal-fin origin; ventral surface of head straight and flat; chest and abdomen with gently convex profile.</paragraph>
|
||
<paragraph id="86304405DE228479FF4EFCC0FC4A9C86" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="358">Head broad, interorbital width about equal to depth of body. Snout obtuse, naked margin wider in males; snout length greater than one-half head length. Eye moderate, upper orbit margin not protruding from dorsal profile of head. Oral disk ovate, wider than long. Lips covered with minute papillae, larger near mouth. Lower lip moderate in size, not reaching gill aperture, its border covered with very small papillae. Maxillary barbel short, its length less than orbit diameter. Dentary tooth row straight, about same size as premaxillary tooth row. Teeth numerous (45– 100 per jaw ramus), asymmetrically bifid, medial cusp much larger and spatulate, lateral cusp minute and pointed, usually not reaching more than half length of medial cusp, but equal in worn teeth. Hypertrophied cheek odontodes strongly evertible, eight to 18 odontodes of various lengths with tips hooked anteriorly, bases encased in thick fleshy sheaths. Head smooth, bones on back of head not carinate; supraoccipital with margins between surrounding bones and plates usually clearly visible. Lateral plates not carinate.</paragraph>
|
||
<paragraph id="86304405DE228479FF4EFB68FC059D3B" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="358">Ventral surface of head and abdomen naked. Nuchal plate small and curved posterolaterally. No enlarged odontodes at edge of lateral plates. Five series of lateral plates anteriorly, three series on caudal peduncle, middorsal and mid-ventral plate series end on caudal peduncle beneath adipose fin. Last plate in median series slightly smaller than penultimate plate, base of caudal fin with about four small platelets after main series, then about eight roughly triangular platelets covering bases of caudal-fin rays.</paragraph>
|
||
<paragraph id="86304405DE228479FF4EFA23FBAD9E0D" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="358">Dorsal-fin origin situated anterior to vertical through pelvic-fin insertion. Dorsal fin moderately elevated, usually reaching preadipose plate when depressed; dorsal-fin base length noticeable greater than dorsal-adipose distance. Adipose fin well developed, spine stout, elevated from body at about a 45º angle. Pectoral fins reaching past base of pelvic fins, but failing to reach the middle of these fins, the spine shorter than head. Anal fin small, its origin at vertical through base of last ray of dorsal. Pelvic fins reaching well past anal-fin origin, inserted posterior to vertical through first branched dorsal-fin ray. Caudal fin with gently convex margin.</paragraph>
|
||
<paragraph id="86304405DE228479FF4EF9FBFB239EDA" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="358">Tiny odontodes present on body plates, largest on posterior margins of plates. All fin spines with small odontodes, more developed in pectoral-fin spine of males. All fin rays with tiny odontodes on rays.</paragraph>
|
||
<paragraph id="86304405DE228479FF4EF903FD209F0E" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="358">Mid-dorsal plates 5(1), 6(2), 7(3), 8(6), 9(9), 10(8), 11(3), 12(2), 16(2), 17(3), 18(5), 19(1); median plates 21(1), 22(7), 23(23), 24(14); mid-ventral plates 14(1), 16(18), 17(12), 18(8), 19(3) 20(2); plates bordering dorsalfin base 6(13), 7(28), 8(3), 9(1); plates between dorsal and adipose fins 6(3), 7(40), 8(2); preadipose plates: 1(45). Fin-ray formulae (N=44): dorsal i,7; pectoral i,6; pelvic i,5; anal i,3(1) i,4 (43); caudal i,14,i. Caudal procurrent spines: dorsal: 4(7), 5(37), ventral: 2(2), 3(28), 4(14).</paragraph>
|
||
<paragraph id="86304405DE228466FF4EF8F0FF2E9896" blockId="15.[151,1437,151,2013]" lastBlockId="16.[151,1436,151,212]" lastPageId="16" lastPageNumber="359" pageId="15" pageNumber="358">
|
||
<emphasis id="B4FB9817DE228479FF4EF8F0FE5F9F32" bold="true" box="[199,401,1879,1904]" pageId="15" pageNumber="358">Color in alcohol.</emphasis>
|
||
(
|
||
<figureCitation id="1EB45880DE228479FE2AF8F0FE279F2D" box="[419,489,1879,1903]" captionStart="FIGURE 8" captionStartId="16.[151,250,1506,1529]" captionTargetBox="[151,1435,251,1484]" captionTargetId="figure@16.[151,1436,251,1486]" captionTargetPageId="16" captionText="FIGURE 8. Ancistrus martini INHS 35367 specimen, 73.2 mm SL, Taguara Creek, Santa Ana River drainage, Lake Maracaibo Basin, 51 mm SL. Photograph by Nathan K. Lujan." httpUri="https://zenodo.org/record/219663/files/figure.png" pageId="15" pageNumber="358">Fig. 8</figureCitation>
|
||
). Base color brown, numerous light spots present on head, barbels and body, head spots smallest anteriorly, larger posteriorly. Flanks usually brown with large, light rounded spots. Abdomen usually tan or brown with light spots which vary in size, shape and number among individuals, sometimes running together to form vermiculations. Fins brownish with alternating dark and light spots on rays. In newer material, the dorsal, caudal, pelvic and pectoral fins have dark spots on spine and fin rays that may extend onto membranes to form bands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="D2F0148DDE3D8466FF1EFA45FD6E9E5A" httpUri="https://zenodo.org/record/219663/files/figure.png" pageId="16" pageNumber="359" targetBox="[151,1435,251,1484]" targetPageId="16">
|
||
<paragraph id="86304405DE3D8466FF1EFA45FD6E9E5A" blockId="16.[151,1436,1506,1560]" pageId="16" pageNumber="359">
|
||
<emphasis id="B4FB9817DE3D8466FF1EFA45FEDC9DBA" bold="true" box="[151,274,1506,1529]" pageId="16" pageNumber="359">FIGURE 8.</emphasis>
|
||
<taxonomicName id="418F3F86DE3D8466FE91FA43FE069DBB" box="[280,456,1508,1529]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="359" phylum="Chordata" rank="species" species="martini">
|
||
<emphasis id="B4FB9817DE3D8466FE91FA43FE069DBB" box="[280,456,1508,1529]" italics="true" pageId="16" pageNumber="359">Ancistrus martini</emphasis>
|
||
</taxonomicName>
|
||
INHS 35367 specimen, 73.2 mm SL, Taguara Creek, Santa Ana River drainage, Lake Maracaibo Basin, 51 mm SL. Photograph by Nathan K. Lujan.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="CE95178EDE3D8466FF4EF9E3FC399E1E" box="[199,1015,1604,1629]" pageId="16" pageNumber="359" type="distribution">
|
||
<paragraph id="86304405DE3D8466FF4EF9E3FC399E1E" blockId="16.[151,1436,1604,2025]" box="[199,1015,1604,1629]" pageId="16" pageNumber="359">
|
||
<emphasis id="B4FB9817DE3D8466FF4EF9E3FEAE9E1F" bold="true" box="[199,352,1604,1629]" pageId="16" pageNumber="359">Distribution.</emphasis>
|
||
<collectingCountry id="FE980495DE3D8466FEEEF9E3FE149E1E" box="[359,474,1604,1628]" name="Venezuela" pageId="16" pageNumber="359">Venezuela</collectingCountry>
|
||
and
|
||
<collectingCountry id="FE980495DE3D8466FD99F9E3FD4F9E1E" box="[528,641,1604,1628]" name="Colombia" pageId="16" pageNumber="359">Colombia</collectingCountry>
|
||
, rivers of Lake Maracaibo Basin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CE95178EDE3D8466FF4EF9CEFD499FAA" pageId="16" pageNumber="359" type="discussion">
|
||
<paragraph id="86304405DE3D8466FF4EF9CEFD499FAA" blockId="16.[151,1436,1604,2025]" pageId="16" pageNumber="359">
|
||
<emphasis id="B4FB9817DE3D8466FF4EF9CEFEFA9EC3" bold="true" box="[199,308,1641,1665]" pageId="16" pageNumber="359">Remarks</emphasis>
|
||
. Schultz described two new subspecies from the Lake Maracaibo Basin (1944):
|
||
<taxonomicName id="418F3F86DE3D8466FB4EF9CFFEE59EE7" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="359" phylum="Chordata" rank="subSpecies" species="brevifilis" subSpecies="bodenhameri">
|
||
<emphasis id="B4FB9817DE3D8466FB4EF9CFFEE59EE7" italics="true" pageId="16" pageNumber="359">Ancistrus brevifilis bodenhameri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="418F3F86DE3D8466FEEDF92AFD649EE7" box="[356,682,1677,1701]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="359" phylum="Chordata" rank="subSpecies" species="triradiatus" subSpecies="martini">
|
||
<emphasis id="B4FB9817DE3D8466FEEDF92AFD649EE7" box="[356,682,1677,1701]" italics="true" pageId="16" pageNumber="359">Ancistrus triradiatus martini</emphasis>
|
||
</taxonomicName>
|
||
, each of which has been subsequently considered as valid species (Fisch-Muller, 2003). However, our morphometric analysis failed to detect significant differences between them. Schultz diagnosed his subspecies based on the color pattern of the ventrum: with separate spots in
|
||
<taxonomicName id="418F3F86DE3D8466FA9EF972FEE59F52" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="359" phylum="Chordata" rank="subSpecies" species="brevifilis" subSpecies="bodenhameri">
|
||
<emphasis id="B4FB9817DE3D8466FA9EF972FEE59F52" italics="true" pageId="16" pageNumber="359">A. brevifilis bodenhameri</emphasis>
|
||
</taxonomicName>
|
||
and spots sometimes running together to form vermiculations in
|
||
<taxonomicName id="418F3F86DE3D8466FBA4F95FFAE89F52" box="[1069,1318,1784,1808]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="359" phylum="Chordata" rank="subSpecies" species="triradiatus" subSpecies="martini">
|
||
<emphasis id="B4FB9817DE3D8466FBA4F95FFAE89F52" box="[1069,1318,1784,1808]" italics="true" pageId="16" pageNumber="359">A. triradiatus martini</emphasis>
|
||
</taxonomicName>
|
||
. We have found great individual variation in the pigmentation pattern of the ventrum of specimens from the Lake Maracaibo Basin and so that character is not useful for distinguishing them. Based on both morphometric and pigmentation pattern differences we consider the Maracaibo species as distinct from both
|
||
<taxonomicName id="418F3F86DE3D8466FB8EF8C2FB6E9F3F" box="[1031,1184,1893,1917]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="359" phylum="Chordata" rank="species" species="triradiatus">
|
||
<emphasis id="B4FB9817DE3D8466FB8EF8C2FB6E9F3F" box="[1031,1184,1893,1917]" italics="true" pageId="16" pageNumber="359">A. triradiatus</emphasis>
|
||
</taxonomicName>
|
||
(which occurs on the southern flank of the Andes in the Orinoco River Basin) and
|
||
<taxonomicName id="418F3F86DE3D8466FCC2F82FFC009FE2" box="[843,974,1928,1952]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="359" phylum="Chordata" rank="species" species="brevifilis">
|
||
<emphasis id="B4FB9817DE3D8466FCC2F82FFC009FE2" box="[843,974,1928,1952]" italics="true" pageId="16" pageNumber="359">A. brevifilis</emphasis>
|
||
</taxonomicName>
|
||
(which occurs in the Tuy River Basin of coastal Caribbean central
|
||
<collectingCountry id="FE980495DE3D8466FE48F80BFDF79F86" box="[449,569,1964,1988]" name="Venezuela" pageId="16" pageNumber="359">Venezuela</collectingCountry>
|
||
). We therefore consider
|
||
<taxonomicName id="418F3F86DE3D8466FCDAF80AFAAB9F86" authority="Schultz 1944" authorityName="Schultz" authorityYear="1944" box="[851,1381,1964,1989]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="359" phylum="Chordata" rank="subSpecies" species="brevifilis" subSpecies="bodenhameri">
|
||
<emphasis id="B4FB9817DE3D8466FCDAF80AFB089F87" box="[851,1222,1965,1989]" italics="true" pageId="16" pageNumber="359">Ancistrus brevifilis bodenhameri</emphasis>
|
||
Schultz 1944
|
||
</taxonomicName>
|
||
as a synonym of
|
||
<taxonomicName id="418F3F86DE3D8466FEAAF877FD4D9FAA" authority="Schultz 1944" authorityName="Schultz" authorityYear="1944" box="[291,643,2000,2024]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="359" phylum="Chordata" rank="species" species="martini">
|
||
<emphasis id="B4FB9817DE3D8466FEAAF877FE2B9FAA" box="[291,485,2000,2024]" italics="true" pageId="16" pageNumber="359">Ancistrus martini</emphasis>
|
||
Schultz 1944
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |