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<document id="C9C95F4996A791FC0A184CFEABE4836C" ID-DOI="10.5281/zenodo.209468" ID-GBIF-Dataset="2c269ccb-4369-4adc-86f4-e5a5225fcc15" ID-ISSN="1175-5326" ID-Zenodo-Dep="209468" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460474857246" checkinUser="plazi" docAuthor="Riehl, Torben, Wilson, George D. F. &amp; Hessler, Robert R." docDate="2012" docId="D602547EFFE25F1212C7FC75FA95FE81" docLanguage="en" docName="zt03277p026.pdf" docOrigin="Zootaxa 3277" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Macrostylis papillata Riehl, Wilson &amp; Hessler, 2012, n. sp." docType="treatment" docVersion="10" lastPageNumber="20" masterDocId="2A3B2C06FFEE5F011250FFBEFFECFFBE" masterDocTitle="New Macrostylidae Hansen, 1916 (Crustacea: Isopoda) from the Gay Head-Bermuda transect with special consideration of sexual dimorphism" masterLastPageNumber="26" masterPageNumber="1" pageNumber="13" updateTime="1698306533727" updateUser="plazi">
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<mods:title id="F29C68647404CA266EFC7EF58059F114">New Macrostylidae Hansen, 1916 (Crustacea: Isopoda) from the Gay Head-Bermuda transect with special consideration of sexual dimorphism</mods:title>
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<mods:namePart id="B8BB010DE3F9F33B248CC72E4D753DDD">Riehl, Torben</mods:namePart>
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<mods:namePart id="BFA5832B12ACBB7DBDD269A96F68B812">Wilson, George D. F.</mods:namePart>
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<mods:namePart id="5F281DE63675838185E4CBF3472BA77D">Hessler, Robert R.</mods:namePart>
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<mods:title id="3F0C8580A7A49E0FE37B058C52F646EC">Zootaxa</mods:title>
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<mods:date id="26F633EB8B751ADAF54FCABAFA10061F">2012</mods:date>
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<treatment id="D602547EFFE25F1212C7FC75FA95FE81" ID-DOI="http://doi.org/10.5281/zenodo.6176730" ID-GBIF-Taxon="119515900" ID-Zenodo-Dep="6176730" LSID="urn:lsid:zoobank.org:act:46083A44-BBF7-40F6-84E4-30F15EAA844D" httpUri="http://treatment.plazi.org/id/D602547EFFE25F1212C7FC75FA95FE81" lastPageId="19" lastPageNumber="20" pageId="12" pageNumber="13">
<subSubSection id="16B1B6E3FFE25F0D12C7FC75FE31FC5B" box="[151,477,971,997]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="5E14E568FFE25F0D12C7FC75FE31FC5B" blockId="12.[151,477,971,997]" box="[151,477,971,997]" pageId="12" pageNumber="13">
<heading id="055C5204FFE25F0D12C7FC75FE31FC5B" bold="true" box="[151,477,971,997]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="6CDF397AFFE25F0D12C7FC75FE31FC5B" bold="true" box="[151,477,971,997]" pageId="12" pageNumber="13">
<taxonomicName id="99AB9EEBFFE25F0D12C7FC75FE7FFC5B" ID-CoL="6R46L" box="[151,403,971,997]" class="Malacostraca" family="Macrostylidae" genus="Macrostylis" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="papillata" status="sp. nov.">
<emphasis id="6CDF397AFFE25F0D12C7FC75FE7FFC5B" bold="true" box="[151,403,971,997]" italics="true" pageId="12" pageNumber="13">Macrostylis papillata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77EC8401FFE25F0D13CBFC75FE31FC5B" box="[411,477,971,997]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="16B1B6E3FFE25F0D12C7FBADFAF3FBF0" pageId="12" pageNumber="13" type="etymology">
<paragraph id="5E14E568FFE25F0D12C7FBADFAF3FBF0" blockId="12.[151,1436,1042,1643]" pageId="12" pageNumber="13">
<emphasis id="6CDF397AFFE25F0D12C7FBADFEFAFB95" bold="true" box="[151,278,1043,1067]" pageId="12" pageNumber="13">Etymology</emphasis>
. The name 'papillata' is derived from the Latin word papilla, meaning 'wart' because this species is characterized by warty posterior margins of the cephalothorax' and the anterior four pereonites' tergites.
</paragraph>
</subSubSection>
<subSubSection id="16B1B6E3FFE25F131299FBE5FB39F856" lastPageId="18" lastPageNumber="19" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="5E14E568FFE25F0D1299FBE5FB9CFBCC" blockId="12.[151,1436,1042,1643]" box="[201,1136,1114,1139]" pageId="12" pageNumber="13">
<emphasis id="6CDF397AFFE25F0D1299FBE5FE8AFBCD" bold="true" box="[201,358,1114,1139]" pageId="12" pageNumber="13">
<typeStatus id="81105BCAFFE25F0D1299FBE5FEE8FBCD" box="[201,260,1115,1139]" pageId="12" pageNumber="13">Type</typeStatus>
fixation
</emphasis>
.
<typeStatus id="81105BCAFFE25F0D1324FBE4FE0DFBCC" box="[372,481,1114,1138]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
: ovigerous female,
<quantity id="9953488DFFE25F0D10EBFBE4FCFFFBCC" box="[699,787,1114,1138]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="mm" value="1.5">1.5 mm</quantity>
, AM P.86009, designated here.
</paragraph>
<paragraph id="5E14E568FFE25F0D1299FBC0FB23FAD0" blockId="12.[151,1436,1042,1643]" pageId="12" pageNumber="13">
<emphasis id="6CDF397AFFE25F0D1299FBC0FE00FB29" bold="true" box="[201,492,1150,1175]" pageId="12" pageNumber="13">
<typeStatus id="81105BCAFFE25F0D1299FBC0FEE8FB28" box="[201,260,1150,1174]" pageId="12" pageNumber="13">Type</typeStatus>
material examined
</emphasis>
.
<typeStatus id="81105BCAFFE25F0D1054FBC0FD9DFB28" box="[516,625,1150,1174]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
: ovigerous female,
<quantity id="9953488DFFE25F0D111EFBC0FC4BFB28" box="[846,935,1150,1174]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="mm" value="1.5">1.5 mm</quantity>
, AM P.86009, used for habitus illustrations,
<collectionCode id="38BA7DADFFE25F0D12C7FB1CFF0FFB04" box="[151,227,1186,1210]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
121.
<typeStatus id="81105BCAFFE25F0D1374FB1DFE79FB05" box="[292,405,1187,1211]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
: juvenile female,
<quantity id="9953488DFFE25F0D100DFB1CFD5AFB04" box="[605,694,1186,1210]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="13" unit="mm" value="1.3">1.3 mm</quantity>
, AM P86008, partly used for illustration of habitus and antennae,
<collectionCode id="38BA7DADFFE25F0D12C7FB78FF0FFB60" box="[151,227,1222,1246]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
121; non-ovigerous female,
<quantity id="9953488DFFE25F0D1073FB78FD97FB60" box="[547,635,1222,1246]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="mm" value="1.5">1.5 mm</quantity>
, AM P86010, used for habitus illustrations and dissected for illustration of appendages,
<collectionCode id="38BA7DADFFE25F0D131AFB54FE7AFABC" box="[330,406,1258,1282]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
121; terminal male,
<quantity id="9953488DFFE25F0D10D8FB54FD0EFABC" box="[648,738,1258,1282]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="13" unit="mm" value="1.3">1.3 mm</quantity>
, AM P86011, used for habitus illustrations and dissected for illustration of appendages,
<collectionCode id="38BA7DADFFE25F0D139EFAB0FDF6FA98" box="[462,538,1294,1318]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
121; ovigerous female,
<quantity id="9953488DFFE25F0D1166FAB0FC7DFA98" box="[822,913,1294,1318]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="mm" value="1.5">1.5 mm</quantity>
, AM P86013, MI 638, gold-coated for
<collectionCode id="38BA7DADFFE25F0D170BFAB0FA7BFA98" box="[1371,1431,1294,1318]" pageId="12" pageNumber="13">SEM</collectionCode>
,
<collectionCode id="38BA7DADFFE25F0D12C7FA8CFF0FFAF4" box="[151,227,1330,1354]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
121; terminal male, AM P86014, MI 635MI 637, dissected and gold-coated for
<collectionCode id="38BA7DADFFE25F0D162EFA8CFB56FAF4" box="[1150,1210,1330,1354]" pageId="12" pageNumber="13">SEM</collectionCode>
,
<collectionCode id="38BA7DADFFE25F0D1696FA8CFAFFFAF4" box="[1222,1299,1330,1354]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
121; immature male,
<quantity id="9953488DFFE25F0D135EFAE8FE8AFAD0" box="[270,358,1366,1390]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="13" unit="mm" value="1.3">1.3 mm</quantity>
, AM P86015, partly used for illustration of habitus and antennae,
<collectionCode id="38BA7DADFFE25F0D161AFAE8FB7AFAD0" box="[1098,1174,1366,1390]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
121.
</paragraph>
<paragraph id="5E14E568FFE25F0D1299FAC5FBE1FA08" blockId="12.[151,1436,1042,1643]" pageId="12" pageNumber="13">
<emphasis id="6CDF397AFFE25F0D1299FAC5FE85FA2D" bold="true" box="[201,361,1402,1427]" pageId="12" pageNumber="13">
<typeStatus id="81105BCAFFE25F0D1299FAC5FEEEFA2D" box="[201,258,1403,1427]" pageId="12" pageNumber="13">Type</typeStatus>
locality
</emphasis>
. Western North Atlantic abyssal plain between Long Island and
<collectingCountry id="26BCA5F8FFE25F0D16F3FAC4FAE2FA2C" box="[1187,1294,1402,1426]" name="Bermuda" pageId="12" pageNumber="13">Bermuda</collectingCountry>
:
<geoCoordinate id="3B9F83AFFFE25F0D1772FAC4FA7BFA2C" box="[1314,1431,1402,1426]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="92" value="35.833332">35°50.0'N</geoCoordinate>
;
<geoCoordinate id="3B9F83AFFFE25F0D12C7FA20FEE3FA08" box="[151,271,1438,1462]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="92" value="-65.183334">65°11.0'W</geoCoordinate>
;
<quantity id="9953488DFFE25F0D134BFA20FE9CFA08" box="[283,368,1438,1462]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="12" pageNumber="13" unit="m" value="4800.0">4800 m</quantity>
(
<collectionCode id="38BA7DADFFE25F0D132EFA20FE20FA08" box="[382,460,1438,1462]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
121),
<geoCoordinate id="3B9F83AFFFE25F0D1045FA20FD65FA08" box="[533,649,1438,1462]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="92" value="35.85">35°51.0'N</geoCoordinate>
;
<geoCoordinate id="3B9F83AFFFE25F0D10C5FA20FCE3FA08" box="[661,783,1438,1462]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="92" value="-64.97">64°58.2'W</geoCoordinate>
;
<quantity id="9953488DFFE25F0D114AFA20FC83FA08" box="[794,879,1438,1462]" metricMagnitude="3" metricUnit="m" metricValue="4.833" pageId="12" pageNumber="13" unit="m" value="4833.0">4833 m</quantity>
(
<collectionCode id="38BA7DADFFE25F0D112DFA20FC27FA08" box="[893,971,1438,1462]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
122).
</paragraph>
<paragraph id="5E14E568FFE25F0D1299FA7DFB58FA64" blockId="12.[151,1436,1042,1643]" box="[201,1204,1474,1499]" pageId="12" pageNumber="13">
<emphasis id="6CDF397AFFE25F0D1299FA7DFE15FA65" bold="true" box="[201,505,1474,1499]" pageId="12" pageNumber="13">
<typeStatus id="81105BCAFFE25F0D1299FA7DFEE8FA65" box="[201,260,1475,1499]" pageId="12" pageNumber="13">Type</typeStatus>
material Remarks
</emphasis>
. Collected during cruise R/V Atlantis
<date id="2A15C3A8FFE25F0D11F8FA7CFC0CFA64" box="[936,992,1474,1498]" pageId="12" pageNumber="13" value="1924-02">II-24</date>
(
<date id="2A15C3A8FFE25F0D11A0FA7CFB44FA64" box="[1008,1192,1474,1499]" pageId="12" pageNumber="13" value="1966-08-21">21 August 1966</date>
).
</paragraph>
<paragraph id="5E14E568FFE25F0D1299FA58FAFEF9D4" blockId="12.[151,1436,1042,1643]" pageId="12" pageNumber="13">
<emphasis id="6CDF397AFFE25F0D1299FA58FE60FA40" bold="true" box="[201,396,1510,1534]" pageId="12" pageNumber="13">Further records</emphasis>
. 1 terminal male, AM P.86016,
<collectionCode id="38BA7DADFFE25F0D1157FA58FCBFFA40" box="[775,851,1510,1534]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
<collectionCode id="38BA7DADFFE25F0D110DFA58FC6AFA40" box="[861,902,1510,1534]" pageId="12" pageNumber="13">LL</collectionCode>
1;
<specimenCount id="48AD2EE1FFE25F0D11F0FA58FBC9FA40" box="[928,1061,1510,1534]" pageId="12" pageNumber="13" type="generic">4 specimen</specimenCount>
, female and male, AM P.67254,
<collectionCode id="38BA7DADFFE25F0D12C7F9B4FF0FF99C" box="[151,227,1546,1570]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
58; 1 terminal male, AM P.83030,
<collectionCode id="38BA7DADFFE25F0D103BF9B4FD5BF99C" box="[619,695,1546,1570]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
83; 1 non-ovigerous female, AM P.86028,
<collectionCode id="38BA7DADFFE25F0D16C9F9B4FB09F99C" box="[1177,1253,1546,1570]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
85;
<specimenCount id="48AD2EE1FFE25F0D1747F9B4FA74F99C" box="[1303,1432,1546,1570]" pageId="12" pageNumber="13" type="generic">8 specimen</specimenCount>
, female and male, AM P.86029,
<collectionCode id="38BA7DADFFE25F0D1057F990FDBFF9F8" box="[519,595,1582,1606]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
95; 2 ovigerous female, AM P.86055,
<collectionCode id="38BA7DADFFE25F0D1648F990FB88F9F8" box="[1048,1124,1582,1606]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
120; 1 terminal male, AM P.86012,
<collectionCode id="38BA7DADFFE25F0D12AEF9ECFEA6F9D4" box="[254,330,1618,1642]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
125; terminal male,
<quantity id="9953488DFFE25F0D1063F9ECFD67F9D4" box="[563,651,1618,1642]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="13" unit="mm" value="1.3">1.3 mm</quantity>
, AM P.86012, MI 630, gold-coated for
<collectionCode id="38BA7DADFFE25F0D1616F9ECFB6EF9D4" box="[1094,1154,1618,1642]" pageId="12" pageNumber="13">SEM</collectionCode>
,
<collectionCode id="38BA7DADFFE25F0D16DDF9ECFB35F9D4" box="[1165,1241,1618,1642]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="12" pageNumber="13">WHOI</collectionCode>
125.
</paragraph>
<paragraph id="5E14E568FFE25F0D12C7F924FE9FF90C" blockId="12.[151,1437,1690,2002]" box="[151,371,1690,1714]" pageId="12" pageNumber="13">Description, female</paragraph>
<paragraph id="5E14E568FFE25F0D1299F900FE6EF86C" blockId="12.[151,1437,1690,2002]" pageId="12" pageNumber="13">
<emphasis id="6CDF397AFFE25F0D1299F900FE11F968" box="[201,509,1726,1750]" italics="true" pageId="12" pageNumber="13">
Body (
<figureCitation id="C690F9EDFFE25F0D1343F900FEB1F968" box="[275,349,1726,1750]" captionStart="FIGURE 8" captionStartId="13.[151,250,1725,1748]" captionTargetBox="[162,1436,195,1697]" captionTargetId="figure@13.[151,1436,195,1701]" captionTargetPageId="13" captionText="FIGURE 8. Macrostylis papillata n. sp. A, E, paratype juvenile Ψ (AM P 86008). B C, F G, paratype adult, non-ovigerous Ψ (AM P 860010). D, H, holotype ovigerous Ψ (AM P 86009). A D, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. D, uropod endopod, damaged. E F, left antennula and antenna, in situ, lateral. G, pleotelson, ventral. H, uropod, close-up, endopod damaged. Scales: A D = 0.5 mm, E F = 0.1 mm, G = 0.3 mm." httpUri="https://zenodo.org/record/209476/files/figure.png" pageId="12" pageNumber="13">Figs 8</figureCitation>
AD, 15AE).
</emphasis>
Length
<quantity id="9953488DFFE25F0D100DF900FD5BF968" box="[605,695,1726,1750]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="mm" value="1.5">1.5 mm</quantity>
, 4.5 width, subcylindrical, without setation.
<emphasis id="6CDF397AFFE25F0D16E2F900FAB5F968" box="[1202,1369,1726,1750]" italics="true" pageId="12" pageNumber="13">Ventral spines.</emphasis>
Pereonite 1 spine acute, prominent. Pereonite 3 spine blunt, small, closer to anterior segment border. Pereonite 4 spine directed posteriorly, acute, small, closer to posterior segment border. Pereonite 5 spine blunt, closer to posterior segment border. Pereonite 6 spine acute, prominent, closer to posterior segment border. Pereonite 7 spine small.
<emphasis id="6CDF397AFFE25F0D12C7F8F0FE02F8D8" box="[151,494,1870,1894]" italics="true" pageId="12" pageNumber="13">Imbricate ornamentation (IO).</emphasis>
Pereonite 1 IO along anterior tergite margin and medially on tergite from anterior to posterior, covering whole sternite; pereonites 2 and 3 IO in an hourglass-shaped band medially on tergite, wider in pereonite 3 than in pereonite 2, covering whole sternite; pereonite 47 and pleotelson IO covering all tergites, sternites and operculum.
</paragraph>
<caption id="0AD4B5E0FFE35F0C12C7F903FEF5F8F0" httpUri="https://zenodo.org/record/209476/files/figure.png" pageId="13" pageNumber="14" targetBox="[162,1436,195,1697]" targetPageId="13">
<paragraph id="5E14E568FFE35F0C12C7F903FEF5F8F0" blockId="13.[151,1436,1725,1870]" pageId="13" pageNumber="14">
<emphasis id="6CDF397AFFE35F0C12C7F903FCD7F96A" bold="true" box="[151,827,1725,1748]" pageId="13" pageNumber="14">
FIGURE 8.
<taxonomicName id="99AB9EEBFFE35F0C134BF903FE1AF96D" box="[283,502,1725,1747]" class="Malacostraca" family="Macrostylidae" genus="Macrostylis" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="papillata" status="sp. nov.">
<emphasis id="6CDF397AFFE35F0C134BF903FE1AF96D" bold="true" box="[283,502,1725,1747]" italics="true" pageId="13" pageNumber="14">Macrostylis papillata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77EC8401FFE35F0C13ADF900FDDBF96D" box="[509,567,1726,1747]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
A, E, paratype juvenile
</emphasis>
Ψ
<emphasis id="6CDF397AFFE35F0C110BF903FF50F94C" bold="true" pageId="13" pageNumber="14">(AM P86008). BC, FG, paratype adult, non-ovigerous</emphasis>
Ψ
<emphasis id="6CDF397AFFE35F0C128AF962FD63F94D" bold="true" box="[218,655,1756,1779]" pageId="13" pageNumber="14">(AM P860010). D, H, holotype ovigerous</emphasis>
Ψ
<emphasis id="6CDF397AFFE35F0C10FDF962FC6EF94F" bold="true" box="[685,898,1756,1779]" pageId="13" pageNumber="14">(AM P86009). AD,</emphasis>
dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated.
<emphasis id="6CDF397AFFE35F0C110EF945FC9AF8AE" bold="true" box="[862,886,1787,1808]" pageId="13" pageNumber="14">D,</emphasis>
uropod endopod, damaged.
<emphasis id="6CDF397AFFE35F0C16F5F945FB3AF8AE" bold="true" box="[1189,1238,1787,1808]" pageId="13" pageNumber="14">EF,</emphasis>
left antennula and antenna, in situ, lateral.
<emphasis id="6CDF397AFFE35F0C13DAF8A7FE72F88E" bold="true" box="[394,414,1817,1840]" pageId="13" pageNumber="14">G,</emphasis>
pleotelson, ventral.
<emphasis id="6CDF397AFFE35F0C1020F8A4FD65F891" bold="true" box="[624,649,1818,1839]" pageId="13" pageNumber="14">H,</emphasis>
uropod, close-up, endopod damaged. Scales: AD = 0.5 mm, EF = 0.1 mm, G = 0.3 mm.
</paragraph>
</caption>
<paragraph id="5E14E568FFE35F0F1299F8C5FBEAFE4A" blockId="13.[151,1436,1915,2011]" lastBlockId="14.[151,1437,151,2012]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="6CDF397AFFE35F0C1299F8C5FE94F82D" box="[201,376,1915,1939]" italics="true" pageId="13" pageNumber="14">Cephalothorax.</emphasis>
Length 0.82 width, 0.15 body length; frons in dorsal view convex, with wrinkles, frontal ridge absent; dorsal surface with array of setae, 1 pair on frons between anterior rims of antennulae articulations, 1 pair dorsally and 1 pair at back of cephalothorax. Posterolateral setae absent. Posterolateral corners acute. Posterior margin papillose.
<emphasis id="6CDF397AFFE05F0F1330FF27FE08FF0E" box="[352,484,153,176]" italics="true" pageId="14" pageNumber="15">Fossosome.</emphasis>
Length 0.85 width, 0.19 body length. Lateral tergite margins in dorsal view forming almost uninterrupted line, ventral surface without keel.
<emphasis id="6CDF397AFFE05F0F115AFF03FC7FFF6A" box="[778,915,188,212]" italics="true" pageId="14" pageNumber="15">Pereonite 1.</emphasis>
Anterior margin concave; posterolateral setae simple, posterior tergite margin papillose.
<emphasis id="6CDF397AFFE05F0F1029FF5FFCEEFF46" box="[633,770,224,248]" italics="true" pageId="14" pageNumber="15">Pereonite 2.</emphasis>
Posterolateral setae simple, posterior tergite margin papillose.
<emphasis id="6CDF397AFFE05F0F1284FEBBFEB1FEA2" box="[212,349,260,284]" italics="true" pageId="14" pageNumber="15">Pereonite 3.</emphasis>
Posterolateral setae simple, flexibly articulated, posterior tergite margin papillose.
<emphasis id="6CDF397AFFE05F0F1742FEBBFA70FEA2" box="[1298,1436,260,284]" italics="true" pageId="14" pageNumber="15">Pereonite 4.</emphasis>
Width 1.2 pereonite 5 width, length 0.66 width; pereonal collum present. Lateral margins in dorsal view simple convex. Posterior tergite margin papillose. Posterolateral margin rounded. Posterolateral setae simple, not robust, flexibly articulated.
<emphasis id="6CDF397AFFE05F0F13D0FECFFDE7FE39" box="[384,523,368,392]" italics="true" pageId="14" pageNumber="15">Pereonite 5.</emphasis>
Length 0.69 width. Posterolateral margin rounded. Tergite posterolateral setae bifid, robust.
<emphasis id="6CDF397AFFE05F0F137DFE2BFE59FE12" box="[301,437,404,428]" italics="true" pageId="14" pageNumber="15">Pereonite 6.</emphasis>
Length 0.72 width. Posterolateral margin produced posteriorly, rounded. Tergite posterolateral setae bifid, robust, flexibly articulated.
<emphasis id="6CDF397AFFE05F0F10CCFE07FCCFFE71" box="[668,803,440,464]" italics="true" pageId="14" pageNumber="15">Pereonite 7.</emphasis>
Length 0.63 width. Posterolateral margin produced posteriorly, rounded. Tergite posterolateral setae bifid, robust, flexibly articulated.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FDBEFCC4FDA9" blockId="14.[151,1437,151,2012]" box="[201,808,511,536]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FDBEFEAEFDA6" box="[201,322,512,536]" italics="true" pageId="14" pageNumber="15">Pleonite 1.</emphasis>
Tergal articulation with pleotelson present.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FD9AFB83FD19" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FD9AFDF2FD82" box="[201,542,548,572]" italics="true" pageId="14" pageNumber="15">
Pleotelson (
<figureCitation id="C690F9EDFFE05F0F1300FD9AFE76FD82" box="[336,410,548,572]" captionStart="FIGURE 8" captionStartId="13.[151,250,1725,1748]" captionTargetBox="[162,1436,195,1697]" captionTargetId="figure@13.[151,1436,195,1701]" captionTargetPageId="13" captionText="FIGURE 8. Macrostylis papillata n. sp. A, E, paratype juvenile Ψ (AM P 86008). B C, F G, paratype adult, non-ovigerous Ψ (AM P 860010). D, H, holotype ovigerous Ψ (AM P 86009). A D, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. D, uropod endopod, damaged. E F, left antennula and antenna, in situ, lateral. G, pleotelson, ventral. H, uropod, close-up, endopod damaged. Scales: A D = 0.5 mm, E F = 0.1 mm, G = 0.3 mm." httpUri="https://zenodo.org/record/209476/files/figure.png" pageId="14" pageNumber="15">Figs 8</figureCitation>
D, G, 15C).
</emphasis>
Ovoid, constricted anteriorly to uropod articulations, length 0.22 body length, 1.8 width, narrower than pereonite 7; statocysts present, dorsal slot-like apertures absent. Posterior apex convex, bluntly rounded. Posterior apex setae absent. Pleopodal cavity width 0.72 pleotelson width, preanal ridge width 0.44 pleotelson width. Anal opening terminal, tilted posteriorly relative to frontal plane.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FD0AFC9CFD72" blockId="14.[151,1437,151,2012]" box="[201,880,692,716]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FD0AFE49FD72" box="[201,421,692,716]" italics="true" pageId="14" pageNumber="15">
Labrum (
<figureCitation id="C690F9EDFFE05F0F1362FD0AFE68FD72" box="[306,388,692,716]" captionStart="FIGURE 14" captionStartId="21.[151,250,1059,1082]" captionTargetBox="[151,1436,193,1038]" captionTargetId="figure@21.[151,1436,193,1038]" captionTargetPageId="21" captionText="FIGURE 14. Macrostylis papillata n. sp. A D, paratype terminal ɗ (AM P 860012, MI 630). A, habitus dorsal. B, pleotelson, dorsal. C, cephalothorax, anterio-lateral view. D, anterior habitus. Scales: A = 0.5 mm, B D = 0.1 mm." httpUri="https://zenodo.org/record/209482/files/figure.png" pageId="14" pageNumber="15">Fig. 14</figureCitation>
F).
</emphasis>
Anterior margin in dorsal view concave.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FD66FEC2FC1A" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FD66FE30FD4E" box="[201,476,728,752]" italics="true" pageId="14" pageNumber="15">
Antennula (
<figureCitation id="C690F9EDFFE05F0F1301FD66FE76FD4E" box="[337,410,728,752]" captionStart="FIGURE 8" captionStartId="13.[151,250,1725,1748]" captionTargetBox="[162,1436,195,1697]" captionTargetId="figure@13.[151,1436,195,1701]" captionTargetPageId="13" captionText="FIGURE 8. Macrostylis papillata n. sp. A, E, paratype juvenile Ψ (AM P 86008). B C, F G, paratype adult, non-ovigerous Ψ (AM P 860010). D, H, holotype ovigerous Ψ (AM P 86009). A D, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. D, uropod endopod, damaged. E F, left antennula and antenna, in situ, lateral. G, pleotelson, ventral. H, uropod, close-up, endopod damaged. Scales: A D = 0.5 mm, E F = 0.1 mm, G = 0.3 mm." httpUri="https://zenodo.org/record/209476/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
EF).
</emphasis>
Length 0.25 head width, 0.25 antenna length, width 1.5 antenna width. Articles decreasing in size from proximal to distal. Article 1 squat, globular, widest but not longest, with 1 broom seta. Article 2 distinctly longer than wide, tubular, subequal article 1 length, with 1 broom seta. Article 3 distinctly longer than wide, tubular, length subequal article 1 length, with 1 simple seta. Article 4 squat, globular. Article 5 minute, squat, globular, with 1 simple seta. Terminal article with 1 aesthetasc, aesthetasc with intermediate belt of constrictions.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FC0EFB69FBB1" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FC0EFE57FC76" box="[201,443,944,968]" italics="true" pageId="14" pageNumber="15">
Antenna (
<figureCitation id="C690F9EDFFE05F0F1366FC0EFE95FC76" box="[310,377,944,968]" captionStart="FIGURE 8" captionStartId="13.[151,250,1725,1748]" captionTargetBox="[162,1436,195,1697]" captionTargetId="figure@13.[151,1436,195,1701]" captionTargetPageId="13" captionText="FIGURE 8. Macrostylis papillata n. sp. A, E, paratype juvenile Ψ (AM P 86008). B C, F G, paratype adult, non-ovigerous Ψ (AM P 860010). D, H, holotype ovigerous Ψ (AM P 86009). A D, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. D, uropod endopod, damaged. E F, left antennula and antenna, in situ, lateral. G, pleotelson, ventral. H, uropod, close-up, endopod damaged. Scales: A D = 0.5 mm, E F = 0.1 mm, G = 0.3 mm." httpUri="https://zenodo.org/record/209476/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
EF).
</emphasis>
Length 0.18 body length. Article 1 squat, globular. Article 2 squat, globular, longer than article 1. Article 3 elongate, longer than article 1. Article 4 shorter than articles 13 together, distally with 1 broom seta. Article 5 longer than article 4, distally with 4 broom setae. Flagellum with 4 articles.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FBA2FE43FB1E" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FBA2FE0FFB8A" box="[201,483,1052,1076]" italics="true" pageId="14" pageNumber="15">
Mandibles (
<figureCitation id="C690F9EDFFE05F0F131FFBA2FE72FB8A" box="[335,414,1052,1076]" captionStart="FIGURE 11" captionStartId="18.[151,250,1685,1708]" captionTargetBox="[168,1433,164,1674]" captionTargetId="figure@18.[152,1437,156,1677]" captionTargetPageId="18" captionText="FIGURE 11. Macrostylis papillata n. sp. A J, paratype non-ovigerous Ψ (AM P 860010). A, left mandible, medial view of incisor process and lacinia mobilis. B, left mandible, dorsal, setal row damaged. C, right mandible, dorsal, setal row and molar process damaged. D, right mandible, incisor process and lacinia mobilis, medial view. E, right maxillula, ventral, inner lobe broken off. F, right maxilliped, endite setation, ventral. G, right maxilliped, ventral. H, left maxilla, ventral. I, right pleopod III, ventral. J, operculum, ventral. Scales: A H = 0.05 mm, I J = 0.1 mm." httpUri="https://zenodo.org/record/209479/files/figure.png" pageId="14" pageNumber="15">Fig. 11</figureCitation>
AD).
</emphasis>
In medial view dorsoventrally flattened, with lateral setae; left mandible incisor process distal margin flattened and curved (shovel-like), with 4 cusps, lacinia mobilis grinding, with 4 cusps; right mandible incisior process with shovel-like appearance, with 3 cusps, lacinia mobilis grinding, clearly smaller than left lacinia, with 6 cusps.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FB12FCDFFB7A" blockId="14.[151,1437,151,2012]" box="[201,819,1196,1220]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FB12FE5BFB7A" box="[201,439,1196,1220]" italics="true" pageId="14" pageNumber="15">
Maxillula (
<figureCitation id="C690F9EDFFE05F0F1316FB12FE7AFB7A" box="[326,406,1196,1220]" captionStart="FIGURE 11" captionStartId="18.[151,250,1685,1708]" captionTargetBox="[168,1433,164,1674]" captionTargetId="figure@18.[152,1437,156,1677]" captionTargetPageId="18" captionText="FIGURE 11. Macrostylis papillata n. sp. A J, paratype non-ovigerous Ψ (AM P 860010). A, left mandible, medial view of incisor process and lacinia mobilis. B, left mandible, dorsal, setal row damaged. C, right mandible, dorsal, setal row and molar process damaged. D, right mandible, incisor process and lacinia mobilis, medial view. E, right maxillula, ventral, inner lobe broken off. F, right maxilliped, endite setation, ventral. G, right maxilliped, ventral. H, left maxilla, ventral. I, right pleopod III, ventral. J, operculum, ventral. Scales: A H = 0.05 mm, I J = 0.1 mm." httpUri="https://zenodo.org/record/209479/files/figure.png" pageId="14" pageNumber="15">Fig. 11</figureCitation>
E).
</emphasis>
Lateral lobe with 13 robust setae.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FB6EFC37FAB2" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FB6EFDEDFB56" box="[201,513,1232,1256]" italics="true" pageId="14" pageNumber="15">
Maxilla (
<figureCitation id="C690F9EDFFE05F0F1360FB6EFE92FB56" box="[304,382,1232,1256]" captionStart="FIGURE 11" captionStartId="18.[151,250,1685,1708]" captionTargetBox="[168,1433,164,1674]" captionTargetId="figure@18.[152,1437,156,1677]" captionTargetPageId="18" captionText="FIGURE 11. Macrostylis papillata n. sp. A J, paratype non-ovigerous Ψ (AM P 860010). A, left mandible, medial view of incisor process and lacinia mobilis. B, left mandible, dorsal, setal row damaged. C, right mandible, dorsal, setal row and molar process damaged. D, right mandible, incisor process and lacinia mobilis, medial view. E, right maxillula, ventral, inner lobe broken off. F, right maxilliped, endite setation, ventral. G, right maxilliped, ventral. H, left maxilla, ventral. I, right pleopod III, ventral. J, operculum, ventral. Scales: A H = 0.05 mm, I J = 0.1 mm." httpUri="https://zenodo.org/record/209479/files/figure.png" pageId="14" pageNumber="15">Fig. 11</figureCitation>
H, 15FG).
</emphasis>
Lateral lobe with 4 setae terminally, simple; middle endite with 5 setae terminally, simple; inner endite with 9 setae terminally, 4 monoserrate, 5 slim, simple.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FAA6FF15FAC6" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FAA6FDCDFA8E" box="[201,545,1304,1328]" italics="true" pageId="14" pageNumber="15">
Maxilliped (
<figureCitation id="C690F9EDFFE05F0F1302FAA6FE49FA8E" box="[338,421,1304,1328]" captionStart="FIGURE 11" captionStartId="18.[151,250,1685,1708]" captionTargetBox="[168,1433,164,1674]" captionTargetId="figure@18.[152,1437,156,1677]" captionTargetPageId="18" captionText="FIGURE 11. Macrostylis papillata n. sp. A J, paratype non-ovigerous Ψ (AM P 860010). A, left mandible, medial view of incisor process and lacinia mobilis. B, left mandible, dorsal, setal row damaged. C, right mandible, dorsal, setal row and molar process damaged. D, right mandible, incisor process and lacinia mobilis, medial view. E, right maxillula, ventral, inner lobe broken off. F, right maxilliped, endite setation, ventral. G, right maxilliped, ventral. H, left maxilla, ventral. I, right pleopod III, ventral. J, operculum, ventral. Scales: A H = 0.05 mm, I J = 0.1 mm." httpUri="https://zenodo.org/record/209479/files/figure.png" pageId="14" pageNumber="15">Figs 11</figureCitation>
FG, 15F).
</emphasis>
Basis endite length 3.5 width; epipod length 3.5 width, 1.2 basis-endite length; palp width subequal endite width, article 2 wider than article 1, article 2 wider than article 3, article 1 shorter than article 3.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FA3AFB94FA5A" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FA3AFE57FA22" box="[201,443,1412,1436]" italics="true" pageId="14" pageNumber="15">
Pereopod I (
<figureCitation id="C690F9EDFFE05F0F1306FA3AFE76FA22" box="[342,410,1412,1436]" captionStart="FIGURE 9" captionStartId="15.[151,250,1651,1674]" captionTargetBox="[151,1436,195,1625]" captionTargetId="figure@15.[151,1436,195,1628]" captionTargetPageId="15" captionText="FIGURE 9. Macrostylis papillata n. sp. A D, paratype non-ovigerous Ψ (AM P 860010). A, pereopod I, lateral, baso-ischial articulation damaged. B, pereopod II dactylus, lateral with close up of distally pappose fringe-like sensilla. C, pereopod II, lateral. D, pereopod III, lateral with close up of bisetulate split seta. Scale = 0.2 mm." httpUri="https://zenodo.org/record/209477/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
A).
</emphasis>
Length 0.33 body length. Ischium dorsal margin with 1 seta, split, bisetulate, latero-distally. Merus dorsal margin with 3 setae, bisetulate, distally, ventral margin with 2 setae, bisetulate, placed distally. Carpus dorsally with 1 seta, bisetulate, placed distally. Dactylus distally with 2 sensillae.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299FA4EFEF8F9CA" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299FA4EFE02F9B6" box="[201,494,1520,1544]" italics="true" pageId="14" pageNumber="15">
Pereopod II (
<figureCitation id="C690F9EDFFE05F0F1335FA4EFE40F9B6" box="[357,428,1520,1544]" captionStart="FIGURE 9" captionStartId="15.[151,250,1651,1674]" captionTargetBox="[151,1436,195,1625]" captionTargetId="figure@15.[151,1436,195,1628]" captionTargetPageId="15" captionText="FIGURE 9. Macrostylis papillata n. sp. A D, paratype non-ovigerous Ψ (AM P 860010). A, pereopod I, lateral, baso-ischial articulation damaged. B, pereopod II dactylus, lateral with close up of distally pappose fringe-like sensilla. C, pereopod II, lateral. D, pereopod III, lateral with close up of bisetulate split seta. Scale = 0.2 mm." httpUri="https://zenodo.org/record/209477/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
BC).
</emphasis>
Longer than pereopod I, length 0.39 body length. Ischium dorsally with 3 setae, bisetulate, placed distally. Merus dorsally with 3 setae, bisetulate, placed distally. Carpus dorsally with 3 setae, 1 bisetulate and 1 broom medially, 1 split distally, ventrally with 2 setae, 1 medially, 1 distally. Dactylus distally with 2 sensillae.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299F93EFD4FF896" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299F93EFE3CF926" box="[201,464,1664,1688]" italics="true" pageId="14" pageNumber="15">
Pereopod III (
<figureCitation id="C690F9EDFFE05F0F1338F93EFE40F926" box="[360,428,1664,1688]" captionStart="FIGURE 9" captionStartId="15.[151,250,1651,1674]" captionTargetBox="[151,1436,195,1625]" captionTargetId="figure@15.[151,1436,195,1628]" captionTargetPageId="15" captionText="FIGURE 9. Macrostylis papillata n. sp. A D, paratype non-ovigerous Ψ (AM P 860010). A, pereopod I, lateral, baso-ischial articulation damaged. B, pereopod II dactylus, lateral with close up of distally pappose fringe-like sensilla. C, pereopod II, lateral. D, pereopod III, lateral with close up of bisetulate split seta. Scale = 0.2 mm." httpUri="https://zenodo.org/record/209477/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
D).
</emphasis>
Length 0.41 body length. Ischium with no seta proximo-dorsally, dorsal lobe tapering; proximally with 1 bisetulate seta; apex with 1 prominent seta; apical seta robust, bifid, bent towards proximal, spine-like; distally with 1 bisetulate seta. Merus dorsally with 4 setae, 2 bisetulate, 2 split, bisetulate, ventrally with 2 setae, simple. Carpus dorsally with 5 setae, 1 bisetulate, 2 split, bisetulate, 1 broom, 1 split, bisetulate, ventrally with 3 setae, simple. Dactylus with 2 sensillae.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299F88AFD17F8F2" blockId="14.[151,1437,151,2012]" box="[201,763,1844,1868]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299F88AFE35F8F2" box="[201,473,1844,1868]" italics="true" pageId="14" pageNumber="15">
Pereopod IV (
<figureCitation id="C690F9EDFFE05F0F1336F88AFE55F8F2" box="[358,441,1844,1868]" captionStart="FIGURE 10" captionStartId="16.[151,250,1751,1774]" captionTargetBox="[151,1436,195,1727]" captionTargetId="figure@16.[151,1436,195,1727]" captionTargetPageId="16" captionText="FIGURE 10. Macrostylis papillata n. sp. A D, paratype non-ovigerous Ψ (AM P 860010). A, pereopod IV, posterior. B, pereopod V, lateral. C, pereopod VI, lateral. D, pereopod VII, lateral. Scale = 0.2 mm." httpUri="https://zenodo.org/record/209478/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
A).
</emphasis>
Length 0.24 body length.
</paragraph>
<paragraph id="5E14E568FFE05F0F1299F8E6FEBCF862" blockId="14.[151,1437,151,2012]" pageId="14" pageNumber="15">
<emphasis id="6CDF397AFFE05F0F1299F8E6FE3CF8CE" box="[201,464,1880,1904]" italics="true" pageId="14" pageNumber="15">
Pereopod V (
<figureCitation id="C690F9EDFFE05F0F130DF8E6FE5CF8CE" box="[349,432,1880,1904]" captionStart="FIGURE 10" captionStartId="16.[151,250,1751,1774]" captionTargetBox="[151,1436,195,1727]" captionTargetId="figure@16.[151,1436,195,1727]" captionTargetPageId="16" captionText="FIGURE 10. Macrostylis papillata n. sp. A D, paratype non-ovigerous Ψ (AM P 860010). A, pereopod IV, posterior. B, pereopod V, lateral. C, pereopod VI, lateral. D, pereopod VII, lateral. Scale = 0.2 mm." httpUri="https://zenodo.org/record/209478/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
B).
</emphasis>
Length 0.33 body length. Ischium mid-ventrally with 1 seta, bisetulate, disto-ventrally with 2 setae, bisetulate. Merus disto-dorsally with 2 setae, 1 short, bisetulate, 1 long, bisetulate, disto-ventrally with 2 setae, 1 long, bisetulate, 1 short, bisetulate. Carpus disto-dorsally with 1 seta, bisetulate, disto-ventrally with 2 setae, bisetulate.
</paragraph>
<caption id="0AD4B5E0FFE15F0E12C7F9CDFC39F979" httpUri="https://zenodo.org/record/209477/files/figure.png" pageId="15" pageNumber="16" targetBox="[151,1436,195,1625]" targetPageId="15">
<paragraph id="5E14E568FFE15F0E12C7F9CDFC39F979" blockId="15.[151,1435,1651,1735]" pageId="15" pageNumber="16">
<emphasis id="6CDF397AFFE15F0E12C7F9CDFC94F934" bold="true" box="[151,888,1651,1674]" pageId="15" pageNumber="16">
FIGURE 9.
<taxonomicName id="99AB9EEBFFE15F0E1349F9CAFE1FF934" box="[281,499,1652,1674]" class="Malacostraca" family="Macrostylidae" genus="Macrostylis" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="papillata" status="sp. nov.">
<emphasis id="6CDF397AFFE15F0E1349F9CAFE1FF934" bold="true" box="[281,499,1652,1674]" italics="true" pageId="15" pageNumber="16">Macrostylis papillata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77EC8401FFE15F0E13AAF9CAFDDFF937" box="[506,563,1652,1673]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
AD, paratype non-ovigerous
</emphasis>
Ψ
<emphasis id="6CDF397AFFE15F0E11C5F9CDFBBBF937" bold="true" box="[917,1111,1651,1674]" pageId="15" pageNumber="16">(AM P860010). A,</emphasis>
pereopod I, lateral, baso-ischial articulation damaged.
<emphasis id="6CDF397AFFE15F0E1329F92DFE7CF916" bold="true" box="[377,400,1683,1704]" pageId="15" pageNumber="16">B,</emphasis>
pereopod II dactylus, lateral with close up of distally pappose fringe-like sensilla.
<emphasis id="6CDF397AFFE15F0E1688F92CFB1EF917" bold="true" box="[1240,1266,1682,1705]" pageId="15" pageNumber="16">C,</emphasis>
pereopod II, lateral.
<emphasis id="6CDF397AFFE15F0E1299F90CFF0EF979" bold="true" box="[201,226,1714,1735]" pageId="15" pageNumber="16">D,</emphasis>
pereopod III, lateral with close up of bisetulate split seta. Scale
<emphasis id="6CDF397AFFE15F0E113CF90CFC96F979" bold="true" box="[876,890,1714,1735]" pageId="15" pageNumber="16">=</emphasis>
0.2 mm.
</paragraph>
</caption>
<paragraph id="5E14E568FFE15F0E1297F951FC06F8CD" blockId="15.[151,1436,1775,2015]" pageId="15" pageNumber="16">
<emphasis id="6CDF397AFFE15F0E1297F951FE33F8B9" box="[199,479,1775,1799]" italics="true" pageId="15" pageNumber="16">
Pereopod VI (
<figureCitation id="C690F9EDFFE15F0E1338F951FE52F8B9" box="[360,446,1775,1799]" captionStart="FIGURE 10" captionStartId="16.[151,250,1751,1774]" captionTargetBox="[151,1436,195,1727]" captionTargetId="figure@16.[151,1436,195,1727]" captionTargetPageId="16" captionText="FIGURE 10. Macrostylis papillata n. sp. A D, paratype non-ovigerous Ψ (AM P 860010). A, pereopod IV, posterior. B, pereopod V, lateral. C, pereopod VI, lateral. D, pereopod VII, lateral. Scale = 0.2 mm." httpUri="https://zenodo.org/record/209478/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
C).
</emphasis>
Length 0.41 body length; ischium mid-ventrally with 2 setae, bisetulate, disto-ventrally with 2 setae, bisetulate; merus disto-dorsally with 2 setae, bisetulate, disto-ventrally with 2 setae, bisetulate; disto-dorsally with 3 setae, 1 broom, 1 prominent, split, bisetulate, 1 small, bisetulate, mid-ventrally with 1 seta, bisetulate, disto-ventrally with 2 setae, 1 short, bisetulate, 1 long, bisetulate.
</paragraph>
<paragraph id="5E14E568FFE15F111299F8C1FD69F827" blockId="15.[151,1436,1775,2015]" lastBlockId="16.[151,1436,1885,2017]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="6CDF397AFFE15F0E1299F8C1FE0AF829" box="[201,486,1919,1943]" italics="true" pageId="15" pageNumber="16">
Pereopod VII (
<figureCitation id="C690F9EDFFE15F0E1320F8C1FE2FF829" box="[368,451,1919,1943]" captionStart="FIGURE 10" captionStartId="16.[151,250,1751,1774]" captionTargetBox="[151,1436,195,1727]" captionTargetId="figure@16.[151,1436,195,1727]" captionTargetPageId="16" captionText="FIGURE 10. Macrostylis papillata n. sp. A D, paratype non-ovigerous Ψ (AM P 860010). A, pereopod IV, posterior. B, pereopod V, lateral. C, pereopod VI, lateral. D, pereopod VII, lateral. Scale = 0.2 mm." httpUri="https://zenodo.org/record/209478/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
D).
</emphasis>
Length less than pereopod VI length, 0.33 body length. Basis length 4.3 width; with no elongate setae. Ischium length 3.5 width, disto-ventrally with 1 seta, bisetulate. Merus length 3.0 width, distodorsally with 1 seta, bisetulate, disto-ventrally with 1 seta, short, bisetulate. Carpus length 6.0 width, disto-dorsally with 2 setae, 1 broom, 1 split, bisetulate, disto-ventrally with 2 setae, 1 short, bisetulate, 1 long bisetulate. Propodus length 4.0 width. Dactylus length 4.0 width.
</paragraph>
<caption id="0AD4B5E0FFFE5F1112C7F969FC13F8B2" httpUri="https://zenodo.org/record/209478/files/figure.png" pageId="16" pageNumber="17" targetBox="[151,1436,195,1727]" targetPageId="16">
<paragraph id="5E14E568FFFE5F1112C7F969FC13F8B2" blockId="16.[151,1435,1751,1805]" pageId="16" pageNumber="17">
<emphasis id="6CDF397AFFFE5F1112C7F969FC76F950" bold="true" box="[151,922,1751,1774]" pageId="16" pageNumber="17">
FIGURE 10.
<taxonomicName id="99AB9EEBFFFE5F11137BF969FDE4F953" box="[299,520,1751,1773]" class="Malacostraca" family="Macrostylidae" genus="Macrostylis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="papillata" status="sp. nov.">
<emphasis id="6CDF397AFFFE5F11137BF969FDE4F953" bold="true" box="[299,520,1751,1773]" italics="true" pageId="16" pageNumber="17">Macrostylis papillata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77EC8401FFFE5F111042F966FDA1F953" box="[530,589,1752,1773]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
AD, paratype non-ovigerous
</emphasis>
Ψ
<emphasis id="6CDF397AFFFE5F1111EDF969FB69F953" bold="true" box="[957,1157,1751,1774]" pageId="16" pageNumber="17">(AM P860010). A,</emphasis>
pereopod IV, posterior.
<emphasis id="6CDF397AFFFE5F1117D4F966FA77F953" bold="true" box="[1412,1435,1752,1773]" pageId="16" pageNumber="17">B,</emphasis>
pereopod V, lateral.
<emphasis id="6CDF397AFFFE5F111333F948FE97F8B3" bold="true" box="[355,379,1782,1805]" pageId="16" pageNumber="17">C,</emphasis>
pereopod VI, lateral.
<emphasis id="6CDF397AFFFE5F11100AF949FD9EF8B2" bold="true" box="[602,626,1783,1804]" pageId="16" pageNumber="17">D,</emphasis>
pereopod VII, lateral. Scale = 0.2 mm.
</paragraph>
</caption>
<paragraph id="5E14E568FFFE5F111299F818FC8CF85F" blockId="16.[151,1436,1885,2017]" pageId="16" pageNumber="17">
<emphasis id="6CDF397AFFFE5F111299F818FD3DF800" box="[201,721,1958,1982]" italics="true" pageId="16" pageNumber="17">
Operculum (female pleopod II;
<figureCitation id="C690F9EDFFFE5F111062F819FD96F800" box="[562,634,1958,1982]" captionStart="FIGURE 8" captionStartId="13.[151,250,1725,1748]" captionTargetBox="[162,1436,195,1697]" captionTargetId="figure@13.[151,1436,195,1701]" captionTargetPageId="13" captionText="FIGURE 8. Macrostylis papillata n. sp. A, E, paratype juvenile Ψ (AM P 86008). B C, F G, paratype adult, non-ovigerous Ψ (AM P 860010). D, H, holotype ovigerous Ψ (AM P 86009). A D, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. D, uropod endopod, damaged. E F, left antennula and antenna, in situ, lateral. G, pleotelson, ventral. H, uropod, close-up, endopod damaged. Scales: A D = 0.5 mm, E F = 0.1 mm, G = 0.3 mm." httpUri="https://zenodo.org/record/209476/files/figure.png" pageId="16" pageNumber="17">Figs 8</figureCitation>
G, 11J).
</emphasis>
Stout, length 1.5 width, 0.48 pleotelson length, ovoid, without keel, with 10 pappose setae on apex, extending to anal opening.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FF26FE85FF49" blockId="17.[151,1436,151,319]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FF26FE21FF0E" box="[201,461,152,176]" italics="true" pageId="17" pageNumber="18">
Pleopod III (
<figureCitation id="C690F9EDFFFF5F101330FF26FE58FF0E" box="[352,436,152,176]" captionStart="FIGURE 11" captionStartId="18.[151,250,1685,1708]" captionTargetBox="[168,1433,164,1674]" captionTargetId="figure@18.[152,1437,156,1677]" captionTargetPageId="18" captionText="FIGURE 11. Macrostylis papillata n. sp. A J, paratype non-ovigerous Ψ (AM P 860010). A, left mandible, medial view of incisor process and lacinia mobilis. B, left mandible, dorsal, setal row damaged. C, right mandible, dorsal, setal row and molar process damaged. D, right mandible, incisor process and lacinia mobilis, medial view. E, right maxillula, ventral, inner lobe broken off. F, right maxilliped, endite setation, ventral. G, right maxilliped, ventral. H, left maxilla, ventral. I, right pleopod III, ventral. J, operculum, ventral. Scales: A H = 0.05 mm, I J = 0.1 mm." httpUri="https://zenodo.org/record/209479/files/figure.png" pageId="17" pageNumber="18">
Fig.
<date id="2A15C3A8FFFF5F1013C9FF26FE58FF0E" box="[409,436,152,176]" pageId="17" pageNumber="18" value="2011-01">11</date>
</figureCitation>
I).
</emphasis>
Length 2.2 width, protopod length 2.0 width, 0.55 pleopod III length; exopod with fringe of fine setae, about as long as pleopod III exopod width, with simple seta subterminally, exopod length 0.73 pleopod III length.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FEBAFC2BFE81" blockId="17.[151,1436,151,319]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FEBAFE57FEA2" box="[201,443,260,284]" italics="true" pageId="17" pageNumber="18">
Uropod (
<figureCitation id="C690F9EDFFFF5F10137FFEBAFE98FEA2" box="[303,372,260,284]" captionStart="FIGURE 8" captionStartId="13.[151,250,1725,1748]" captionTargetBox="[162,1436,195,1697]" captionTargetId="figure@13.[151,1436,195,1701]" captionTargetPageId="13" captionText="FIGURE 8. Macrostylis papillata n. sp. A, E, paratype juvenile Ψ (AM P 86008). B C, F G, paratype adult, non-ovigerous Ψ (AM P 860010). D, H, holotype ovigerous Ψ (AM P 86009). A D, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. D, uropod endopod, damaged. E F, left antennula and antenna, in situ, lateral. G, pleotelson, ventral. H, uropod, close-up, endopod damaged. Scales: A D = 0.5 mm, E F = 0.1 mm, G = 0.3 mm." httpUri="https://zenodo.org/record/209476/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
D, H).
</emphasis>
Protopod length 1.3 pleotelson length; inserting on pleotelson on posterior margin. Protopod distal margin blunt, endopod insertion terminal, length 24.0 width.
</paragraph>
<paragraph id="5E14E568FFFF5F1012C7FED1FE2EFE39" blockId="17.[151,1436,367,1940]" box="[151,450,367,391]" pageId="17" pageNumber="18">Description, terminal male</paragraph>
<paragraph id="5E14E568FFFF5F101299FE2AFD44FE4A" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FE2AFDA7FE12" box="[201,587,404,428]" italics="true" pageId="17" pageNumber="18">
Body (
<figureCitation id="C690F9EDFFFF5F101343FE2AFE85FE12" box="[275,361,404,428]" captionStart="FIGURE 12" captionStartId="19.[151,250,1919,1942]" captionTargetBox="[164,1435,385,1893]" captionTargetId="figure@19.[151,1436,375,1893]" captionTargetPageId="19" captionText="FIGURE 12. Macrostylis papillata n. sp. A D, paratype terminal ɗ (AM P 860011). E G, paratype subadult ɗ (AM P 860015). A B, E, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. C, G, antennula and antenna, in situ, lateral. D, F, pleotelson, ventral. Scales: A B, E = 0.5 mm, C, G = 0.1 mm, D, F = 0.3 mm." httpUri="https://zenodo.org/record/209480/files/figure.png" pageId="17" pageNumber="18">Figs 12</figureCitation>
AB, E, 14AB, D).
</emphasis>
Length
<quantity id="9953488DFFFF5F1010FAFE2AFCEFFE12" box="[682,771,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="17" pageNumber="18" unit="mm" value="1.3">1.3 mm</quantity>
, 5.0 width.
<emphasis id="6CDF397AFFFF5F1011DBFE2AFBDDFE12" box="[907,1073,404,428]" italics="true" pageId="17" pageNumber="18">Ventral spines.</emphasis>
Pereonite 3 spine acute, prominent, located closer to anterior segment border. Pereonite 4 spine directed ventrally and posteriorly, blunt, prominent, located closer to posterior segment boder.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FDBEFC3EFD72" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FDBEFE94FDA6" box="[201,376,512,536]" italics="true" pageId="17" pageNumber="18">Cephalothorax.</emphasis>
Frons with wrinkles, frontal ridge present, as cluster of slight transversal scratches between antennulae articulations; length/width ratio subequal to female, 0.16 body length; with conspicious array of setae, posterolateral corners acute, posterolateral setae absent, posterior margins papillose.
<emphasis id="6CDF397AFFFF5F101626FDF7FB16FDDE" box="[1142,1274,585,608]" italics="true" pageId="17" pageNumber="18">Fossosome.</emphasis>
Length/width ratio greater than in female, length 0.94 width, length/body-length ratio subequal to female; keeled.
<emphasis id="6CDF397AFFFF5F101742FDD3FA70FD3A" box="[1298,1436,620,644]" italics="true" pageId="17" pageNumber="18">Pereonite 4.</emphasis>
Narrower than pereonite 5, length/width ratio subequal to female; pereonal collum present. Lateral margins in dorsal view medially convex. Posterolateral margin not produced posteriorly.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FD66FBDCFD51" blockId="17.[151,1436,367,1940]" box="[201,1072,727,752]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FD66FE52FD4E" box="[201,446,728,752]" italics="true" pageId="17" pageNumber="18">
Pleonite 1 (
<figureCitation id="C690F9EDFFFF5F10131BFD66FE72FD4E" box="[331,414,728,752]" captionStart="FIGURE 14" captionStartId="21.[151,250,1059,1082]" captionTargetBox="[151,1436,193,1038]" captionTargetId="figure@21.[151,1436,193,1038]" captionTargetPageId="21" captionText="FIGURE 14. Macrostylis papillata n. sp. A D, paratype terminal ɗ (AM P 860012, MI 630). A, habitus dorsal. B, pleotelson, dorsal. C, cephalothorax, anterio-lateral view. D, anterior habitus. Scales: A = 0.5 mm, B D = 0.1 mm." httpUri="https://zenodo.org/record/209482/files/figure.png" pageId="17" pageNumber="18">Fig. 14</figureCitation>
B).
</emphasis>
Tergal and sternal articulations with pleotelson present.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FD42FE7CFCC1" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FD42FDB2FCAA" box="[201,606,764,788]" italics="true" pageId="17" pageNumber="18">
Pleotelson (
<figureCitation id="C690F9EDFFFF5F10131EFD42FE49FCAA" box="[334,421,764,788]" captionStart="FIGURE 12" captionStartId="19.[151,250,1919,1942]" captionTargetBox="[164,1435,385,1893]" captionTargetId="figure@19.[151,1436,375,1893]" captionTargetPageId="19" captionText="FIGURE 12. Macrostylis papillata n. sp. A D, paratype terminal ɗ (AM P 860011). E G, paratype subadult ɗ (AM P 860015). A B, E, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. C, G, antennula and antenna, in situ, lateral. D, F, pleotelson, ventral. Scales: A B, E = 0.5 mm, C, G = 0.1 mm, D, F = 0.3 mm." httpUri="https://zenodo.org/record/209480/files/figure.png" pageId="17" pageNumber="18">Figs 12</figureCitation>
B, DF, 14AB).
</emphasis>
In dorsal view, approximately rectangular, length/width ratio in male subequal to female, 0.23 body length, width subequal pereonite 7 width. Posterior apex convex, more obtusely-angled compared to female, without setae on margin, pleopodal cavity width 0.75 pleotelson width, preanal ridge width 0.43 pleotelson width.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FC32FDC3FB8A" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FC32FDF2FC1A" box="[201,542,908,932]" italics="true" pageId="17" pageNumber="18">
Antennula (
<figureCitation id="C690F9EDFFFF5F10131BFC32FE4DFC1A" box="[331,417,908,932]" captionStart="FIGURE 12" captionStartId="19.[151,250,1919,1942]" captionTargetBox="[164,1435,385,1893]" captionTargetId="figure@19.[151,1436,375,1893]" captionTargetPageId="19" captionText="FIGURE 12. Macrostylis papillata n. sp. A D, paratype terminal ɗ (AM P 860011). E G, paratype subadult ɗ (AM P 860015). A B, E, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. C, G, antennula and antenna, in situ, lateral. D, F, pleotelson, ventral. Scales: A B, E = 0.5 mm, C, G = 0.1 mm, D, F = 0.3 mm." httpUri="https://zenodo.org/record/209480/files/figure.png" pageId="17" pageNumber="18">Figs 12</figureCitation>
C, G, 14C).
</emphasis>
Length 0.35 head width, 0.27 antenna length, width 1.2 antenna width; terminal article with 2 aesthetascs, penultimate article with 2 aesthetascs, aesthetascs with intermediate belt of constrictions; article 1 squat, globular, longest and widest, 1 broom seta, article 2 squat, globular, shorter than article 1, 2 broom setae, article 3 squat, globular, shorter than article 1, 1 broom seta, article 4 squat, globular, minute, article 5 squat, globular, minute, with 1 simple seta.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FBFEFBA4FB1E" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FBFEFDE5FBE6" box="[201,521,1088,1112]" italics="true" pageId="17" pageNumber="18">
Antenna (
<figureCitation id="C690F9EDFFFF5F101365FBFEFE60FBE6" box="[309,396,1088,1112]" captionStart="FIGURE 12" captionStartId="19.[151,250,1919,1942]" captionTargetBox="[164,1435,385,1893]" captionTargetId="figure@19.[151,1436,375,1893]" captionTargetPageId="19" captionText="FIGURE 12. Macrostylis papillata n. sp. A D, paratype terminal ɗ (AM P 860011). E G, paratype subadult ɗ (AM P 860015). A B, E, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. C, G, antennula and antenna, in situ, lateral. D, F, pleotelson, ventral. Scales: A B, E = 0.5 mm, C, G = 0.1 mm, D, F = 0.3 mm." httpUri="https://zenodo.org/record/209480/files/figure.png" pageId="17" pageNumber="18">Figs 12</figureCitation>
C, G, 14C).
</emphasis>
Length 0.22 body length, flagellum of 4 articles, article 1 elongate, article 2 elongate, longer than article 1, article 3 elongate, longer than article 1, article 4 shorter than articles 13 together, 1 broom seta, article 5 longer than article 4, distally with 1 simple seta, 4 broom setae.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FB12FCF5FB56" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FB12FEA2FB7A" box="[201,334,1196,1220]" italics="true" pageId="17" pageNumber="18">Pereopod I.</emphasis>
Length 0.37 body length. Merus setation as in female. Carpus dorsally with 2 setae, 1 broom, 1 bisetulate, carpus ventrally with 3 setae, 1 simple, 2 split.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FB4AFBE1FAEA" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FB4AFEB5FAB2" box="[201,345,1268,1292]" italics="true" pageId="17" pageNumber="18">Pereopod II.</emphasis>
Ischium setation as in female. Merus dorsally with 4 setae, dorso-distally, bisetulate, ventrally with 2 setae, along margin, distally, bisetulate. Carpus dorsally with 4 setae, 2 bisetulate along margin, 2 split distally, bisetulate, ventrally with 3 setae, 1 bisetulate, 1 broom, 1 split, bisetulate.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FADEFD6CFAC6" blockId="17.[151,1436,367,1940]" box="[201,640,1376,1400]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FADEFEB3FAC6" box="[201,351,1376,1400]" italics="true" pageId="17" pageNumber="18">Pereopod III.</emphasis>
Length 0.44 body length.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FA3AFC9DFA5A" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FA3AFEBDFA22" box="[201,337,1412,1436]" italics="true" pageId="17" pageNumber="18">Pereopod V.</emphasis>
Length 0.35 body length. Merus disto-dorsally with 2 setae; 1 short, bisetulate, 1 long, bisetulate; mid-ventrally with 1 seta; bisetulate; disto-ventrally with 2 setae; 1 short, bisetulate, 1 long, bisetulate. Carpus disto-ventrally with 2 setae, 1 short, bisetulate, 1 long, bisetulate.
</paragraph>
<paragraph id="5E14E568FFFF5F101299FA4EFB49F9B6" blockId="17.[151,1436,367,1940]" box="[201,1189,1520,1544]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299FA4EFEB2F9B6" box="[201,350,1520,1544]" italics="true" pageId="17" pageNumber="18">Pereopod VI.</emphasis>
Length 0.51 body length; ischium, merus and carpus setation as in female.
</paragraph>
<paragraph id="5E14E568FFFF5F101299F9AAFC0FF926" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299F9AAFE8BF992" box="[201,359,1556,1580]" italics="true" pageId="17" pageNumber="18">Pereopod VII.</emphasis>
Length 0.35 body length, less than pereopod VI length, segment L/W ratios sexually dimorphic; basis length 4.3 width; ischium length 2.8 width, setation as in female; merus length 1.7 width, disto-dorsally with 1 seta, bisetulate, disto-ventrally with 2 setae, 1 short, bisetulate, 1 long, bisetulate; carpus length 5 width, carpus setation as in female; propodus length 4.0 width; dactylus length 3.0 width.
</paragraph>
<paragraph id="5E14E568FFFF5F101299F91AFC7EF896" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299F91AFE57F902" box="[201,443,1700,1724]" italics="true" pageId="17" pageNumber="18">
Pleopod I (
<figureCitation id="C690F9EDFFFF5F101318F91AFE76F902" box="[328,410,1700,1724]" captionStart="FIGURE 13" captionStartId="20.[151,250,1098,1121]" captionTargetBox="[172,1436,199,1077]" captionTargetId="figure@20.[151,1436,193,1077]" captionTargetPageId="20" captionText="FIGURE 13. Macrostylis papillata n. sp. A C, paratype terminal ɗ (AM P 860011). A, pleopods I, ventral. B, right pleopod II, dorsal, with indicated endopod musculature and sperm duct. C, left pleopod III, dorsal. Scale = 0.05 mm." httpUri="https://zenodo.org/record/209481/files/figure.png" pageId="17" pageNumber="18">Fig. 13</figureCitation>
A).
</emphasis>
Length 0.58 pleotelson length, distally with fringe-like sensillae.
<emphasis id="6CDF397AFFFF5F1016F1F91AFA70F902" box="[1185,1436,1700,1724]" italics="true" pageId="17" pageNumber="18">
Pleopod II (
<figureCitation id="C690F9EDFFFF5F101779F91AFA90F902" box="[1321,1404,1700,1724]" captionStart="FIGURE 13" captionStartId="20.[151,250,1098,1121]" captionTargetBox="[172,1436,199,1077]" captionTargetId="figure@20.[151,1436,193,1077]" captionTargetPageId="20" captionText="FIGURE 13. Macrostylis papillata n. sp. A C, paratype terminal ɗ (AM P 860011). A, pleopods I, ventral. B, right pleopod II, dorsal, with indicated endopod musculature and sperm duct. C, left pleopod III, dorsal. Scale = 0.05 mm." httpUri="https://zenodo.org/record/209481/files/figure.png" pageId="17" pageNumber="18">Fig. 13</figureCitation>
B).
</emphasis>
Protopod apex tapering, with setae on proximal lateral margin, 3 pappose setae altogether, with 5 pappose setae distally. Endopod distance of insertion from protopod distal margin 0.38 protopod length. Stylet weakly curved, not extending to distal margin of protopod, length 0.52 protopod length.
</paragraph>
<paragraph id="5E14E568FFFF5F101299F88AFEE8F82A" blockId="17.[151,1436,367,1940]" pageId="17" pageNumber="18">
<emphasis id="6CDF397AFFFF5F101299F88AFE4EF8F2" box="[201,418,1844,1868]" italics="true" pageId="17" pageNumber="18">
Uropod (
<figureCitation id="C690F9EDFFFF5F10137FF88AFE6DF8F2" box="[303,385,1844,1868]" captionStart="FIGURE 12" captionStartId="19.[151,250,1919,1942]" captionTargetBox="[164,1435,385,1893]" captionTargetId="figure@19.[151,1436,375,1893]" captionTargetPageId="19" captionText="FIGURE 12. Macrostylis papillata n. sp. A D, paratype terminal ɗ (AM P 860011). E G, paratype subadult ɗ (AM P 860015). A B, E, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. C, G, antennula and antenna, in situ, lateral. D, F, pleotelson, ventral. Scales: A B, E = 0.5 mm, C, G = 0.1 mm, D, F = 0.3 mm." httpUri="https://zenodo.org/record/209480/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
F).
</emphasis>
Length 2.0 pleotelson length; protopod length/width ratio greater than in female, length1.5 width, with endopod inserting terminally; endopod length 0.31 protopod length, 11.5 width, width less than protopod.
</paragraph>
<caption id="0AD4B5E0FFFC5F1312C7F92BFCB2F898" httpUri="https://zenodo.org/record/209479/files/figure.png" pageId="18" pageNumber="19" targetBox="[168,1433,164,1674]" targetPageId="18">
<paragraph id="5E14E568FFFC5F1312C7F92BFCB2F898" blockId="18.[151,1436,1685,1830]" pageId="18" pageNumber="19">
<emphasis id="6CDF397AFFFC5F1312C7F92BFC6FF912" bold="true" box="[151,899,1685,1708]" pageId="18" pageNumber="19">
FIGURE 11.
<taxonomicName id="99AB9EEBFFFC5F131376F92BFDEDF915" box="[294,513,1685,1707]" class="Malacostraca" family="Macrostylidae" genus="Macrostylis" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="papillata" status="sp. nov.">
<emphasis id="6CDF397AFFFC5F131376F92BFDEDF915" bold="true" box="[294,513,1685,1707]" italics="true" pageId="18" pageNumber="19">Macrostylis papillata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77EC8401FFFC5F131058F928FDAEF915" box="[520,578,1686,1707]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
AJ, paratype non-ovigerous
</emphasis>
Ψ
<emphasis id="6CDF397AFFFC5F1311F3F92BFB8AF915" bold="true" box="[931,1126,1685,1708]" pageId="18" pageNumber="19">(AM P860010). A,</emphasis>
left mandible, medial view of incisor process and lacinia mobilis.
<emphasis id="6CDF397AFFFC5F131052F90BFDF4F974" bold="true" box="[514,536,1717,1738]" pageId="18" pageNumber="19">B,</emphasis>
left mandible, dorsal, setal row damaged.
<emphasis id="6CDF397AFFFC5F131198F90AFC0CF975" bold="true" box="[968,992,1716,1739]" pageId="18" pageNumber="19">C,</emphasis>
right mandible, dorsal, setal row and molar process damaged.
<emphasis id="6CDF397AFFFC5F131305F96DFE81F956" bold="true" box="[341,365,1747,1768]" pageId="18" pageNumber="19">D,</emphasis>
right mandible, incisor process and lacinia mobilis, medial view.
<emphasis id="6CDF397AFFFC5F13164AF96DFBDDF956" bold="true" box="[1050,1073,1747,1768]" pageId="18" pageNumber="19">E,</emphasis>
right maxillula, ventral, inner lobe broken off.
<emphasis id="6CDF397AFFFC5F13135CF94CFEF1F8B9" bold="true" box="[268,285,1778,1799]" pageId="18" pageNumber="19">F,</emphasis>
right maxilliped, endite setation, ventral.
<emphasis id="6CDF397AFFFC5F131097F94FFD37F8B6" bold="true" box="[711,731,1777,1800]" pageId="18" pageNumber="19">G,</emphasis>
right maxilliped, ventral.
<emphasis id="6CDF397AFFFC5F1311B5F94CFC13F8B9" bold="true" box="[997,1023,1778,1799]" pageId="18" pageNumber="19">H,</emphasis>
left maxilla, ventral.
<emphasis id="6CDF397AFFFC5F131688F94CFB05F8B9" bold="true" box="[1240,1257,1778,1799]" pageId="18" pageNumber="19">I,</emphasis>
right pleopod III, ventral.
<emphasis id="6CDF397AFFFC5F1312BAF8AFFF11F898" bold="true" box="[234,253,1809,1830]" pageId="18" pageNumber="19">J,</emphasis>
operculum, ventral. Scales: AH = 0.05 mm, IJ = 0.1 mm.
</paragraph>
</caption>
<paragraph id="5E14E568FFFC5F1312C7F8F5FB39F856" blockId="18.[151,1436,1867,2025]" pageId="18" pageNumber="19">
other authors,
<taxonomicName id="99AB9EEBFFFC5F13136AF8F2FD99F8DD" authority="Birstein, 1963" authorityName="Birstein" authorityYear="1963" box="[314,629,1867,1891]" class="Malacostraca" family="Macrostylidae" genus="Macrostylis" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="6CDF397AFFFC5F13136AF8F2FE21F8DD" box="[314,461,1867,1891]" italics="true" pageId="18" pageNumber="19">M. reticulata</emphasis>
<bibRefCitation id="3A3A9899FFFC5F131385F8F5FD99F8DD" author="Birstein" box="[469,629,1867,1891]" pageId="18" pageNumber="19" refString="Birstein, Y. A. (1963) Deep Sea Isopod Crustaceans of the Northwestern Part of the Pacific Ocean. Institute of Oceanology, Academy of Sciences, USSR: Moscow. (In Russian with English summary)." type="book" year="1963">Birstein, 1963</bibRefCitation>
</taxonomicName>
has strongly developed imbricate ornamentation and could thus potentially show marginal wartyness as well. This latter species is substantially different from
<taxonomicName id="99AB9EEBFFFC5F131622F8D0FB17F83B" box="[1138,1275,1901,1925]" class="Malacostraca" family="Macrostylidae" genus="Macrostylis" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="papillata" status="sp. nov.">
<emphasis id="6CDF397AFFFC5F131622F8D0FB17F83B" box="[1138,1275,1901,1925]" italics="true" pageId="18" pageNumber="19">M. papillata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77EC8401FFFC5F131752F8D3FAD1F83A" box="[1282,1341,1901,1924]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
because it has the ornamentation on all somites. Both species differ in the shape of their pereopod III ischium, in that
<taxonomicName id="99AB9EEBFFFC5F13172DF831FF10F879" class="Malacostraca" family="Macrostylidae" genus="Macrostylis" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="papillata">
<emphasis id="6CDF397AFFFC5F13172DF831FF10F879" italics="true" pageId="18" pageNumber="19">M. papillata</emphasis>
</taxonomicName>
has an narrow dorsal projection bearing a robust proximally curved seta with two flanking bisetulate setae, whereas
<taxonomicName id="99AB9EEBFFFC5F13136FF86CFE3DF857" box="[319,465,2001,2025]" class="Malacostraca" family="Macrostylidae" genus="Macrostylis" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="6CDF397AFFFC5F13136FF86CFE3DF857" box="[319,465,2001,2025]" italics="true" pageId="18" pageNumber="19">M. reticulata</emphasis>
</taxonomicName>
has a more rounded projection with only 3 straight non-robust setae.
</paragraph>
</subSubSection>
<subSubSection id="16B1B6E3FFFD5F121299FF26FA95FE81" pageId="19" pageNumber="20" type="discussion">
<paragraph id="5E14E568FFFD5F121299FF26FA95FE81" blockId="19.[151,1436,151,320]" pageId="19" pageNumber="20">
<emphasis id="6CDF397AFFFD5F121299FF26FED1FF0E" bold="true" box="[201,317,152,176]" pageId="19" pageNumber="20">Remarks.</emphasis>
<taxonomicName id="99AB9EEBFFFD5F121316FF27FE3EFF0E" box="[326,466,152,176]" class="Malacostraca" family="Macrostylidae" genus="Macrostylis" kingdom="Animalia" order="Isopoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="papillata">
<emphasis id="6CDF397AFFFD5F121316FF27FE3EFF0E" box="[326,466,152,176]" italics="true" pageId="19" pageNumber="20">M. papillata</emphasis>
</taxonomicName>
differs from any previously described macrostylid owing to the presence of a tergal pleonite 1 articulation with the pleotelson. Furthermore, the ridges that create the imbricate ornamentation in this species overlap posteriorly with the margin of the pereonites 14 and head. As a result, the margins of these somites have a warty appearance that is most evident in the SEM images (
<figureCitation id="C690F9EDFFFD5F121188FEBAFBB6FEA2" box="[984,1114,260,284]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStartId-0="21.[151,250,1059,1082]" captionStartId-1="22.[151,250,1592,1615]" captionTargetBox-0="[151,1436,193,1038]" captionTargetBox-1="[151,1436,195,1567]" captionTargetId-0="figure@21.[151,1436,193,1038]" captionTargetId-1="figure@22.[151,1436,195,1568]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURE 14. Macrostylis papillata n. sp. A D, paratype terminal ɗ (AM P 860012, MI 630). A, habitus dorsal. B, pleotelson, dorsal. C, cephalothorax, anterio-lateral view. D, anterior habitus. Scales: A = 0.5 mm, B D = 0.1 mm." captionText-1="FIGURE 15. Macrostylis papillata n. sp. A D, paratype ovigerous Ψ (AM P 860013, MI 638). E, paratype terminal ɗ (AM P 860012, MI 630). F G, paratype subadult Ψ (AM P 860014, MI 635). A, habitus, lateral. B D habitus, dorsal. E, habitus, lateral. F, mouthparts, ventral. G, maxilla, ventral, close up. Scales: A E = 0.5 mm, F G = 0.1 mm." httpUri-0="https://zenodo.org/record/209482/files/figure.png" httpUri-1="https://zenodo.org/record/209483/files/figure.png" pageId="19" pageNumber="20">Figs 1415</figureCitation>
), but can be seen in the light microscope (
<figureCitation id="C690F9EDFFFD5F121377FE99FE96FE81" box="[295,378,295,319]" captionStart="FIGURE 12" captionStartId="19.[151,250,1919,1942]" captionTargetBox="[164,1435,385,1893]" captionTargetId="figure@19.[151,1436,375,1893]" captionTargetPageId="19" captionText="FIGURE 12. Macrostylis papillata n. sp. A D, paratype terminal ɗ (AM P 860011). E G, paratype subadult ɗ (AM P 860015). A B, E, dorsal and lateral habitus, cuticle ornamentation and appendages mostly omitted, uropods missing where not illustrated. C, G, antennula and antenna, in situ, lateral. D, F, pleotelson, ventral. Scales: A B, E = 0.5 mm, C, G = 0.1 mm, D, F = 0.3 mm." httpUri="https://zenodo.org/record/209480/files/figure.png" pageId="19" pageNumber="20">Fig. 12</figureCitation>
B). Although this subtlety of the imbricate ornamentation may not have been fully noted by
</paragraph>
</subSubSection>
</treatment>
</document>