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178 lines
25 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.362.6300" ID-PMC="PMC3867105" ID-Pensoft-Pub="1313-2970-362-29" ID-Pensoft-UUID="EA63FFD35254FFFC603C7A5E8D08FFA0" ID-PubMed="24363599" ID-Zenodo-Dep="578109" ID-ZooBank="569812E982B54BCF84E0D4C43AEE8273" ModsDocID="1313-2970-362-29" checkinTime="1451246566598" checkinUser="pensoft" docAuthor="Erwin, Terry L. & Amundson, Lauren M." docDate="2013" docId="D487F44CE4C35D43BA9C4E0B5E931F2C" docLanguage="en" docName="ZooKeys 362: 29-54" docOrigin="ZooKeys 362" docPubDate="2013-12-13" docSource="http://dx.doi.org/10.3897/zookeys.362.6300" docTitle="Pseudomorpha (Pseudomorpha) pima Amundson & Erwin 2013, sp. n." docType="treatment" docUuid="0040741D-5DEE-4BE0-8C86-7C3242251E8D" docUuidSource="ZooBank" docVersion="3" id="EA63FFD35254FFFC603C7A5E8D08FFA0" lastPageNumber="40" masterDocId="EA63FFD35254FFFC603C7A5E8D08FFA0" masterDocTitle="Beetles that live with ants (Carabidae, Pseudomorphini, Pseudomorpha Kirby, 1825): A revision of the santarita species group" masterLastPageNumber="54" masterPageNumber="29" pageNumber="39" updateTime="1668157171424" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Beetles that live with ants (Carabidae, Pseudomorphini, Pseudomorpha Kirby, 1825): A revision of the santarita species group</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Erwin, Terry L.</mods:namePart>
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<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
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<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Amundson, Lauren M.</mods:namePart>
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<mods:affiliation>Hyper-diversity Group Summer Intern, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-12-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>362</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.362.6300</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.362.6300</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-362-29</mods:identifier>
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<mods:identifier type="ZooBank">569812E982B54BCF84E0D4C43AEE8273</mods:identifier>
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<mods:identifier type="Pensoft-UUID">EA63FFD35254FFFC603C7A5E8D08FFA0</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578109</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182235298" LSID="urn:lsid:zoobank.org:act:0040741D-5DEE-4BE0-8C86-7C3242251E8D" httpUri="http://treatment.plazi.org/id/D487F44CE4C35D43BA9C4E0B5E931F2C" lastPageId="11" lastPageNumber="40" pageId="10" pageNumber="39">
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<subSubSection pageId="10" pageNumber="39" type="nomenclature">
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<paragraph pageId="10" pageNumber="39">
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<taxonomicName LSID="http://zoobank.org/0040741D-5DEE-4BE0-8C86-7C3242251E8D" authority="Amundson & Erwin" authorityName="Amundson & Erwin" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pseudomorpha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Pseudomorpha (Pseudomorpha) pima" order="Coleoptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="pima" status="sp. n." subGenus="Pseudomorpha">Pseudomorpha (Pseudomorpha) pima Amundson & Erwin</taxonomicName>
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<taxonomicNameLabel pageId="10" pageNumber="39">sp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="10" pageNumber="39">Figures 4</figureCitation>
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="10" pageNumber="39">, 10</figureCitation>
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<figureCitation captionStart="Figure 13–17" captionStartId="F3" captionText="Figure 13 - 17. 13 Pseudomorpha huachinera sp. n., male paratype, ADP 110961; Pajarito Mountains, AZ. Male genitalia, median lobe and parameres labeled as in repose in male, A right lateral aspect, B left lateral aspect, and C dorsal aspects. 14 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Harshaw Creek, Patagonia Mountains, AZ 15 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, AZ 16 Pseudomorpha santarita sp. n., male paratype, ADP 110388; Santa Rita Ranch, Santa Rita Mountains, AZ 17 Pseudomorpha pima sp. n., female paratype, ADP 109219; Bog Springs, Santa Rita Mountains, AZ. Ovipositor sclerites, Legend: gc 1 gonocoxite 1; gc 2 gonocoxite 2; lt laterotergite; b base of gonocoxite 2; bl blade of gonocoxite 2. Scale lines = 0.5 mm." httpUri="https://binary.pensoft.net/fig/27291" pageId="10" pageNumber="39">, 17</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F4" captionText="Figure 18. Distribution map for species of the santarita group of Pseudomorpha." httpUri="https://binary.pensoft.net/fig/27292" pageId="10" pageNumber="39">, 18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="39" type="holotype">
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<paragraph pageId="10" pageNumber="39">Holotype.</paragraph>
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<paragraph pageId="10" pageNumber="39">
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<emphasis bold="true" pageId="10" pageNumber="39">USA</emphasis>
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: Arizona,
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<emphasis italics="true" pageId="10" pageNumber="39">Pima County</emphasis>
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, Santa Rita Mountains, Lower Madera Canyon,
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<geoCoordinate degrees="31.745" direction="north" orientation="latitude" precision="55" value="31.745">31.745°N</geoCoordinate>
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,
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<geoCoordinate degrees="110.919" direction="west" orientation="longitude" precision="55" value="-110.919">110.919°W</geoCoordinate>
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, 1174m, 14-16 July 1978 (W.H. Tyson) (CAS: 8111006, female). A paratype is listed below; see other specimens examined.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="39" type="derivation of specific epithet">
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<paragraph pageId="10" pageNumber="39">Derivation of specific epithet.</paragraph>
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<paragraph pageId="10" pageNumber="39">
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The epithet "
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<emphasis italics="true" pageId="10" pageNumber="39">pima</emphasis>
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" is a singular feminine noun used in apposition and is the name of Aztecan descended peoples that live along the Gila and Salt rivers in southern Arizona.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="39" type="proposed english vernacular name">
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<paragraph pageId="10" pageNumber="39">Proposed english vernacular name.</paragraph>
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<paragraph pageId="10" pageNumber="39">Pima False-form beetle.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="39" type="diagnosis">
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<paragraph pageId="10" pageNumber="39">Diagnosis.</paragraph>
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<paragraph pageId="10" pageNumber="39">Color tone of dorsum piceous with head slightly darker over eyes; body robust and rectangulate, lateral margins of elytra tapering to an apically truncated and laterally slightly rounded apex; dorsum mostly glabrous with irregularly and wide-spaced short erect setae; pronotum with lateral margins moderately explanate, about coequal in width to that of elytra across humeri, disc markedly convex and planar; elytral interneurs moderately impressed and visible under low magnification, interneurs slightly convex; 10 umbilicate setae present near lateral margin, dorsal edge of epipleuron lined with long laterally erect setae.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="40" pageId="10" pageNumber="39" type="description">
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<paragraph pageId="10" pageNumber="39">Description.</paragraph>
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<paragraph lastPageId="11" lastPageNumber="40" pageId="10" pageNumber="39">
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(
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="10" pageNumber="39">Figs 4</figureCitation>
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,
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="10" pageNumber="39">10</figureCitation>
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,
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<figureCitation captionStart="Figure 13–17" captionStartId="F3" captionText="Figure 13 - 17. 13 Pseudomorpha huachinera sp. n., male paratype, ADP 110961; Pajarito Mountains, AZ. Male genitalia, median lobe and parameres labeled as in repose in male, A right lateral aspect, B left lateral aspect, and C dorsal aspects. 14 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Harshaw Creek, Patagonia Mountains, AZ 15 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, AZ 16 Pseudomorpha santarita sp. n., male paratype, ADP 110388; Santa Rita Ranch, Santa Rita Mountains, AZ 17 Pseudomorpha pima sp. n., female paratype, ADP 109219; Bog Springs, Santa Rita Mountains, AZ. Ovipositor sclerites, Legend: gc 1 gonocoxite 1; gc 2 gonocoxite 2; lt laterotergite; b base of gonocoxite 2; bl blade of gonocoxite 2. Scale lines = 0.5 mm." httpUri="https://binary.pensoft.net/fig/27291" pageId="10" pageNumber="39">17</figureCitation>
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).
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<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Tables with measurements and ratios for Pseudomorpha pima sp. n.; all measurements are in millimeters." httpUri="http://table.plazi.org/id/911E406A35C6B2A49F11FBC681934C3E" pageId="10" pageNumber="39" tableUuid="911E406A35C6B2A49F11FBC681934C3E">Table 4</tableCitation>
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.
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<emphasis italics="true" pageId="11" pageNumber="40">
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<pageBreakToken pageId="11" pageNumber="40" start="start">Size</pageBreakToken>
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</emphasis>
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: Large for genus, ABL = 6.3 to 6.8 mm, SBL = 6.3 to 6.7 mm, TW = 5.1 to 5.7 mm. Preocular lobe-eye ratio: 0.56 to 0.57. Pronotum ratio (L/W): 0.31. Elytron ratio (L/W): 1.5 to 1.6.
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<emphasis italics="true" pageId="11" pageNumber="40">Color</emphasis>
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: Dorsum piceous, slightly lighter brown along explanate edges of dorsum.
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<emphasis italics="true" pageId="11" pageNumber="40">Luster</emphasis>
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:Dorsum dull, slightly matte.
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<emphasis italics="true" pageId="11" pageNumber="40">Microsculpture</emphasis>
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: Small isodiametric sculpticells throughout dorsal surface.
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<emphasis bold="true" pageId="11" pageNumber="40">Head:</emphasis>
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Genal lobe obsolete, rim posteriad and below eye bearing at least five robust setae directed perpendicular to head; preocular lobe distinct, moderately prominent, and more or less straight (
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="11" pageNumber="40">Fig. 10</figureCitation>
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); eye not exceeding preocular lobe/gena boundary, shallowly arcuate; clypeus fused to frons with pigmented furrow entire and visible, bisetose, setae laterad on margin; labrum with four setae projecting anteriorly (
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<figureCitation captionStart="Figure 7–12" captionStartId="F2" captionText="Figure 7 - 12. 7 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Head, dorsal planar aspect 8 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Head, dorsal planar aspect 9 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Head, dorsal planar aspect 10 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Head, dorsal planar aspect 11 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Head, dorsal planar aspect 12 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. A Head, dorsal planar aspect." httpUri="https://binary.pensoft.net/fig/27290" pageId="11" pageNumber="40">Fig. 10</figureCitation>
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); antennal flagellum markedly setose, antennomeres 1-3 bisetose.
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<emphasis bold="true" pageId="11" pageNumber="40">Prothorax:</emphasis>
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Pronotum (
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="11" pageNumber="40">Fig. 4</figureCitation>
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) mostly glabrous with irregularly and wide-spaced short erect setae, apex slightly arcuate medially and narrower than ocular boundary, disk markedly convex and planar, width slightly wider than elytra across humeri, base and apex fringed with more or less evenly spaced setae, median line ending about
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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before basal margin, lateral margins of pronotum with wide explanate sides, anterior angle 89.22°; prosternal apex fringed with short, evenly spaced setae.
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<emphasis bold="true" pageId="11" pageNumber="40">Pterothorax:</emphasis>
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Scutellum (normally) visible, moderate sized, narrowly rounded apically; elytra smooth (
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<figureCitation captionStart="Figure 1–6" captionStartId="F1" captionText="Figure 1 - 6. 1 Pseudomorpha huachinera sp. n., male holotype, ADP 110112; Arroyo El Cocono, Sierra Huachinera, Sonora, Mexico. Habitus, dorsal aspect, ABL = 5.88 mm 2 Pseudomorpha patagonia sp. n., male holotype, ADP 110694; Madera Canyon, Santa Rita Mountains, AZ. Habitus, dorsal aspect, ABL = 4.39 mm 3 Pseudomorpha penablanca sp. n., female holotype, CAS 8111005; Pena Blanca Lake, Arizona. Habitus, dorsal aspect, ABL = 6.37 mm 4 Pseudomorpha pima sp. n., female holotype, CAS 8111006; Madera Canyon (lower), Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 6.76 mm 5 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, Arizona. Habitus, dorsal aspect, ABL = 5.78 mm 6 Pseudomorpha santarita sp. n., male holotype, ADP 110817; Pajarito Mountains, AZ. Habitus, dorsal aspect, ABL = 5.33 mm." httpUri="https://binary.pensoft.net/fig/27289" pageId="11" pageNumber="40">Fig. 4</figureCitation>
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), interneurs very shallow, clearly visible under low magnification, markedly zig-zagged, intervals slightly convex on disc, lateral margin very slightly sinuate at basal third, 10 umbilicate erect setae on the ventrally directed curvature of the elytral lateral portion.
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<emphasis bold="true" pageId="11" pageNumber="40">Abdomen:</emphasis>
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All sterna sparsely setiferous, sternum III densely so; male unknown; female with 2 pairs of 4 setae on sternum, and numerous longer setae on sterna IV, V, and VI.
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<emphasis bold="true" pageId="11" pageNumber="40">Legs:</emphasis>
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Legs flattened, setiferous, tibia bearing fringed ring of setae on distal end, femur with distinct lateral sulcus, femora and tibiae sparsely setose.
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<emphasis bold="true" pageId="11" pageNumber="40">Female Genitalia:</emphasis>
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(see
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<figureCitation captionStart="Figure 13–17" captionStartId="F3" captionText="Figure 13 - 17. 13 Pseudomorpha huachinera sp. n., male paratype, ADP 110961; Pajarito Mountains, AZ. Male genitalia, median lobe and parameres labeled as in repose in male, A right lateral aspect, B left lateral aspect, and C dorsal aspects. 14 Pseudomorpha patagonia sp. n., male paratype, ADP 110694; Harshaw Creek, Patagonia Mountains, AZ 15 Pseudomorpha santacruz sp. n., male holotype, ADP 111870; Madera Canyon, Santa Rita Mountains, AZ 16 Pseudomorpha santarita sp. n., male paratype, ADP 110388; Santa Rita Ranch, Santa Rita Mountains, AZ 17 Pseudomorpha pima sp. n., female paratype, ADP 109219; Bog Springs, Santa Rita Mountains, AZ. Ovipositor sclerites, Legend: gc 1 gonocoxite 1; gc 2 gonocoxite 2; lt laterotergite; b base of gonocoxite 2; bl blade of gonocoxite 2. Scale lines = 0.5 mm." httpUri="https://binary.pensoft.net/fig/27291" pageId="11" pageNumber="40">Fig. 17</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="40" type="dispersal potential">
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<paragraph pageId="11" pageNumber="40">Dispersal potential.</paragraph>
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<paragraph pageId="11" pageNumber="40">These beetles are macropterous and have been recorded at lights, hence capable of flight; they are swift and agile runners. Accordingly, the species may be expected to be more broadly distributed across a wider geographical range than current records indicate. Female adults are attracted to lights.</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="40" type="way of life">
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<paragraph pageId="11" pageNumber="40">Way of life.</paragraph>
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<paragraph pageId="11" pageNumber="40">
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Adults are likely found in ant nests and the surrounding vicinity; females are ovoviviparous (
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<bibRefCitation author="Liebherr, JK" journalOrPublisher="Journal of Natural History" pageId="16" pageNumber="45" pagination="1079 - 1086" publicationUrl="10.1080/00222938500770681" refId="B18" refString="Liebherr, JK, Kavanaugh, DH, 1985. Ovoviviparity in carabid beetles of the genus Pseudomorpha (Insecta: Coleoptera). Journal of Natural History 19: 1079 - 1086, 10.1080/00222938500770681" title="Ovoviviparity in carabid beetles of the genus Pseudomorpha (Insecta: Coleoptera)." url="10.1080/00222938500770681" volume="19" year="1985">Liebherr and Kavanaugh 1985</bibRefCitation>
|
||
); larvae are ant nest inquilines (
|
||
<bibRefCitation author="Erwin, TL" journalOrPublisher="The Coleopterists Bulletin" pageId="16" pageNumber="45" pagination="53 - 68" refId="B8" refString="Erwin, TL, 1981. A synopsis of the immature stages of Pseudomorphini (Coleoptera: Carabidae) with notes on tribal affinities and behavior in relation to life with ants. The Coleopterists Bulletin 35 (1): 53 - 68" title="A synopsis of the immature stages of Pseudomorphini (Coleoptera: Carabidae) with notes on tribal affinities and behavior in relation to life with ants." volume="35" year="1981">Erwin 1981</bibRefCitation>
|
||
). Members of
|
||
<taxonomicName authorityName="Amundson & Erwin" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pseudomorpha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="pima">
|
||
<emphasis italics="true" pageId="11" pageNumber="40">Pseudomorpha pima</emphasis>
|
||
</taxonomicName>
|
||
occur at midland and upland altitudes near the Sonoran Desert. Adults are active in early to mid-July, a very hot month in this area. For images of habitats at the paratype locality (Bog Springs) see: http://www.meetup.com/phoenix-atheists/events/109850282/.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="40" type="other specimens examined">
|
||
<paragraph pageId="11" pageNumber="40">Other specimens examined.</paragraph>
|
||
<paragraph pageId="11" pageNumber="40">
|
||
<emphasis bold="true" pageId="11" pageNumber="40">USA</emphasis>
|
||
: Arizona,
|
||
<emphasis italics="true" pageId="11" pageNumber="40">Pima County</emphasis>
|
||
, Santa Rita Mountains, Bog Springs,
|
||
<geoCoordinate degrees="31.726" direction="north" orientation="latitude" precision="55" value="31.726">31.726°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="110.874" direction="west" orientation="longitude" precision="55" value="-110.874">110.874°W</geoCoordinate>
|
||
, 1524m, 3 July 1958 (J. von Bloeker Jr.) (SBNHM: ADP109219, female paratype).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="40" type="geographic distribution">
|
||
<paragraph pageId="11" pageNumber="40">Geographic distribution.</paragraph>
|
||
<paragraph pageId="11" pageNumber="40">
|
||
(
|
||
<figureCitation captionStart="Figure 18" captionStartId="F4" captionText="Figure 18. Distribution map for species of the santarita group of Pseudomorpha." httpUri="https://binary.pensoft.net/fig/27292" pageId="11" pageNumber="40">Fig. 18</figureCitation>
|
||
). This species is currently known from Arizona.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |