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<document id="A525EDE48AB5F31B4AE258108C36B26D" ID-DOI="10.11646/zootaxa.1647.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5103860" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626285615291" checkinUser="felipe" docAuthor="Desjardins, Christopher A." docDate="2007" docId="D40DA74BDE76545EAE8F6030FE01BC68" docLanguage="en" docName="zt01647p088.pdf" docOrigin="Zootaxa 1647 (1)" docSource="https://www.biotaxa.org/Zootaxa/article/view/zootaxa.1647.1.1" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Chimaerolelaps Desjardins 2007, New Genus" docType="treatment" docVersion="3" lastPageNumber="34" masterDocId="2834DF33DE56547FAE186577FFCCB81A" masterDocTitle="Phylogenetics and classification of the world genera of Diparinae (Hymenoptera: Pteromalidae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="33" updateTime="1699008634511" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="55CD6EFEF2A877EFF9E73CBE993B7596">Phylogenetics and classification of the world genera of Diparinae (Hymenoptera: Pteromalidae)</mods:title>
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<mods:namePart id="CF66C5E32A00BC277974C66B5B6B04A4">Desjardins, Christopher A.</mods:namePart>
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<subSubSection id="14BE45D6DE76545FAE8F6030FD4DBD7B" box="[151,641,1351,1377]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="5C1B165DDE76545FAE8F6030FD4DBD7B" blockId="32.[151,641,1351,1412]" box="[151,641,1351,1377]" pageId="32" pageNumber="33">
<heading id="0753A131DE76545FAE8F6030FD4DBD7B" bold="true" box="[151,641,1351,1377]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="6ED0CA4FDE76545FAE8F6030FD4DBD7B" bold="true" box="[151,641,1351,1377]" pageId="32" pageNumber="33">
<taxonomicName id="9BA46DDEDE76545FAE8F6030FE29BD7B" authority="Desjardins" authorityName="Desjardins" authorityYear="2007" box="[151,485,1351,1377]" class="Insecta" family="Pteromalidae" genus="Chimaerolelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6ED0CA4FDE76545FAE8F6030FE9ABD7B" bold="true" box="[151,342,1351,1377]" italics="true" pageId="32" pageNumber="33">Chimaerolelaps</emphasis>
Desjardins
</taxonomicName>
,
<taxonomicNameLabel id="75E37734DE76545FAFEA6030FD4DBD7B" box="[498,641,1351,1377]" pageId="32" pageNumber="33" rank="genus">New Genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="14BE45D6DE76545FAE8F601DFF31BD9E" box="[151,253,1386,1412]" pageId="32" pageNumber="33" type="description">
<paragraph id="5C1B165DDE76545FAE8F601DFF31BD9E" blockId="32.[151,641,1351,1412]" box="[151,253,1386,1412]" pageId="32" pageNumber="33">
(
<figureCitation id="C49F0AD8DE76545FAE87601DFF39BD9E" box="[159,245,1386,1412]" captionStart="FIGURES 5964" captionStartId="48.[151,269,1769,1793]" captionTargetBox="[259,1346,273,1723]" captionTargetId="figure-136@48.[217,1369,242,1749]" captionTargetPageId="48" captionText="FIGURES 5964. Lateral habitus views: 59, Neapterolelaps viridescens; 60, N. mitteri; 61, Cerodipara sabensis; 62, Chimaerolelaps villosa; 63, Conophorisca littoriticus; 64, C. grisselli." figureDoi="http://doi.org/10.5281/zenodo.5103899" httpUri="https://zenodo.org/record/5103899/files/figure.png" pageId="32" pageNumber="33">Fig. 62</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="14BE45D6DE76545FAE8F60CDFC4EBDCE" box="[151,898,1466,1492]" pageId="32" pageNumber="33" type="type_taxon">
<paragraph id="5C1B165DDE76545FAE8F60CDFC4EBDCE" blockId="32.[151,1436,1466,2012]" box="[151,898,1466,1492]" pageId="32" pageNumber="33">
<emphasis id="6ED0CA4FDE76545FAE8F60CDFE8CBDCE" bold="true" box="[151,320,1466,1492]" pageId="32" pageNumber="33">
<typeStatus id="831FA8FFDE76545FAE8F60CDFF18BDCE" box="[151,212,1466,1492]" pageId="32" pageNumber="33">Type</typeStatus>
Species:
</emphasis>
<taxonomicName id="9BA46DDEDE76545FAF5060CCFD12BDCE" authority="Desjardins" authorityName="Desjardins" authorityYear="2007" box="[328,734,1466,1492]" class="Insecta" family="Pteromalidae" genus="Chimaerolelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="villosa" status="sp. nov.">
<emphasis id="6ED0CA4FDE76545FAF5060CCFD9BBDCE" box="[328,599,1467,1492]" italics="true" pageId="32" pageNumber="33">Chimaerolelaps villosa</emphasis>
Desjardins
</taxonomicName>
,
<taxonomicNameLabel id="75E37734DE76545FACF260CDFC4EBDCE" box="[746,898,1466,1492]" pageId="32" pageNumber="33" rank="species">New Species</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="14BE45D6DE76545FAEDE6095FCBABEDE" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="5C1B165DDE76545FAEDE6095FCBABEDE" blockId="32.[151,1436,1466,2012]" pageId="32" pageNumber="33">
<emphasis id="6ED0CA4FDE76545FAEDE6095FE84BDE6" bold="true" box="[198,328,1506,1532]" pageId="32" pageNumber="33">Diagnosis:</emphasis>
<taxonomicName id="9BA46DDEDE76545FAF486094FDC6BDE6" authorityName="Desjardins" authorityYear="2007" box="[336,522,1507,1532]" class="Insecta" family="Pteromalidae" genus="Chimaerolelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6ED0CA4FDE76545FAF486094FDC6BDE6" box="[336,522,1507,1532]" italics="true" pageId="32" pageNumber="33">Chimaerolelaps</emphasis>
</taxonomicName>
is unique among diparines in 2 ways. First, it has 4 pairs of scutellar bristles, while all other diparines have at most 2 pairs. Second, it has an elongate anellus which is longer than broad (
<figureCitation id="C49F0AD8DE76545FAE876345FF36BE56" box="[159,250,1586,1612]" captionStart="FIGURES 5964" captionStartId="48.[151,269,1769,1793]" captionTargetBox="[259,1346,273,1723]" captionTargetId="figure-136@48.[217,1369,242,1749]" captionTargetPageId="48" captionText="FIGURES 5964. Lateral habitus views: 59, Neapterolelaps viridescens; 60, N. mitteri; 61, Cerodipara sabensis; 62, Chimaerolelaps villosa; 63, Conophorisca littoriticus; 64, C. grisselli." figureDoi="http://doi.org/10.5281/zenodo.5103899" httpUri="https://zenodo.org/record/5103899/files/figure.png" pageId="32" pageNumber="33">Fig. 62</figureCitation>
), while all other diparines have an anellus that is broader than long (
<figureCitation id="C49F0AD8DE76545FAA2F6345FB07BE56" box="[1079,1227,1586,1612]" captionStart="FIGURES 5964" captionStartId="48.[151,269,1769,1793]" captionTargetBox="[259,1346,273,1723]" captionTargetId="figure-136@48.[217,1369,242,1749]" captionTargetPageId="48" captionText="FIGURES 5964. Lateral habitus views: 59, Neapterolelaps viridescens; 60, N. mitteri; 61, Cerodipara sabensis; 62, Chimaerolelaps villosa; 63, Conophorisca littoriticus; 64, C. grisselli." figureDoi="http://doi.org/10.5281/zenodo.5103899" httpUri="https://zenodo.org/record/5103899/files/figure.png" pageId="32" pageNumber="33">Figs. 59, 60</figureCitation>
).
<taxonomicName id="9BA46DDEDE76545FAAFA6344FA50BE56" authorityName="Desjardins" authorityYear="2007" box="[1250,1436,1587,1612]" class="Insecta" family="Pteromalidae" genus="Chimaerolelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6ED0CA4FDE76545FAAFA6344FA50BE56" box="[1250,1436,1587,1612]" italics="true" pageId="32" pageNumber="33">Chimaerolelaps</emphasis>
</taxonomicName>
may superficially resemble
<taxonomicName id="9BA46DDEDE76545FAFC6632CFDE0BE6E" box="[478,556,1627,1652]" class="Insecta" family="Pteromalidae" genus="Lelaps" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6ED0CA4FDE76545FAFC6632CFDE0BE6E" box="[478,556,1627,1652]" italics="true" pageId="32" pageNumber="33">Lelaps</emphasis>
</taxonomicName>
or
<taxonomicName id="9BA46DDEDE76545FAC4E632CFD29BE6E" authorityName="Boucek. Although" authorityYear="1954" box="[598,741,1627,1652]" class="Insecta" family="Pteromalidae" genus="Netomocera" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6ED0CA4FDE76545FAC4E632CFD29BE6E" box="[598,741,1627,1652]" italics="true" pageId="32" pageNumber="33">Netomocera</emphasis>
</taxonomicName>
.
<taxonomicName id="9BA46DDEDE76545FACED632CFC63BE6E" authorityName="Desjardins" authorityYear="2007" box="[757,943,1627,1652]" class="Insecta" family="Pteromalidae" genus="Chimaerolelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6ED0CA4FDE76545FACED632CFC63BE6E" box="[757,943,1627,1652]" italics="true" pageId="32" pageNumber="33">Chimaerolelaps</emphasis>
</taxonomicName>
has a clypeal margin which is protruding and straight, while
<taxonomicName id="9BA46DDEDE76545FAF6363F4FE05BE86" box="[379,457,1667,1692]" class="Insecta" family="Pteromalidae" genus="Lelaps" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6ED0CA4FDE76545FAF6363F4FE05BE86" box="[379,457,1667,1692]" italics="true" pageId="32" pageNumber="33">Lelaps</emphasis>
</taxonomicName>
has a median clypeal tooth (
<figureCitation id="C49F0AD8DE76545FAD3863F5FCBBBE86" box="[800,887,1666,1692]" captionStart="FIGURES 1116" captionStartId="36.[151,269,1802,1826]" captionTargetBox="[217,1369,434,1778]" captionTargetId="figure-212@36.[217,1369,434,1778]" captionTargetPageId="36" captionText="FIGURES 1116. Conophorisca grisselli: 11, mesosoma (lateral view); 12, metasoma (lateral view); 13, cercus. Lelaps sp. A: 14, antenna; 15, clava; 16, clypeus, mct = median clypeal tooth." figureDoi="http://doi.org/10.5281/zenodo.5103877" httpUri="https://zenodo.org/record/5103877/files/figure.png" pageId="32" pageNumber="33">Fig. 16</figureCitation>
, mct).
<taxonomicName id="9BA46DDEDE76545FADD363F4FB49BE86" authorityName="Desjardins" authorityYear="2007" box="[971,1157,1667,1692]" class="Insecta" family="Pteromalidae" genus="Chimaerolelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6ED0CA4FDE76545FADD363F4FB49BE86" box="[971,1157,1667,1692]" italics="true" pageId="32" pageNumber="33">Chimaerolelaps</emphasis>
</taxonomicName>
also has a symmetrical flagellum, while the flagellum in
<taxonomicName id="9BA46DDEDE76545FAC0763DCFD62BEDE" authorityName="Boucek. Although" authorityYear="1954" box="[543,686,1707,1732]" class="Insecta" family="Pteromalidae" genus="Netomocera" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6ED0CA4FDE76545FAC0763DCFD62BEDE" box="[543,686,1707,1732]" italics="true" pageId="32" pageNumber="33">Netomocera</emphasis>
</taxonomicName>
is asymmetrical.
</paragraph>
</subSubSection>
<subSubSection id="14BE45D6DE76545EAEDE63A5FA58BAA0" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="5C1B165DDE76545EAEDE63A5FA58BAA0" blockId="32.[151,1436,1466,2012]" lastBlockId="33.[151,1436,152,1138]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="6ED0CA4FDE76545FAEDE63A5FE1CBEF6" bold="true" box="[198,464,1746,1772]" pageId="32" pageNumber="33">Description: Female.</emphasis>
<emphasis id="6ED0CA4FDE76545FAFC663A4FDEBBEF6" box="[478,551,1747,1772]" italics="true" pageId="32" pageNumber="33">Head:</emphasis>
Occipital margin rounded; occipital carina present; upper face without strong, transversely carinate sculpture; eyes not posteriorly extended beyond occipital margin; inner eye margins uniformly convex; eyes bare; scrobe present and scrobal channel parallel-sided dorsal to toruli; dorsal margin of scrobe rounded; toruli not on shelf, junction of upper and lower face rounded; antennae symmetrically clavate; antennal formula 11173; anellus longer than broad; pedicel, first funicular segment, second funicular segment subequal in length; claval apex without thick tuft of micropilosity; apical clypeal margin protruding and straight; malar groove present; single pair of strong, dark bristles on vertex present.
<emphasis id="6ED0CA4FDE76545EAB5F62B4FF1FB8AB" italics="true" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">Mesos- oma:</emphasis>
dorsum of mesosoma with strong, dark bristles (1 pair median scutal, 1 pair lateral scutal, 3 pairs anterior scutellar, 1 pair posterior scutellar); pronotum short, collar-like; notauli not arched and meeting posterior scutal margin at lateral edge of scutoscutellar suture; lateral lobes of scutum similar in color to remainder of scutum; posterior scutal margin without setose groove; scutellum large, convex, and sharply sloped posteriorly; axillae convex and not reduced; posterior notal wing process present, pointed; frenal sulcus present; metanotum present as narrow, sculptured band; propodeum at least 1.5X longer than high; propodeum with dorsoventrally flattened spine near anterior margin; plicae absent; suture between postspiracular area and metapleuron diagonal; propodeal foramen circular, open in one plane; longitudinal invagination of pronotum, mesepimeron and metapleuron absent; prepectus elongate, reaching tegula; tegula normal, flap-like; axillary wing sclerite not visible; acropleuron normal, not expanded; mesopleuron with smooth and sculptured regions posteriorly; metacoxa posteriorly convex, with transverse striations; metacoxa posteriorly without thick vertical brush of setae; 2 metatibial spurs, &lt;1.5X width of tibia at point of insertion.
<emphasis id="6ED0CA4FDE77545EAA2C6727FB74BA73" box="[1076,1208,592,617]" italics="true" pageId="33" pageNumber="34">Metasoma:</emphasis>
Petiole cylindrical, with single pair of setae, at least 2X as long as broad; GT1 expanded, covering at least half of metasoma length; GT1 rounded lateral to petiole insertion; cercal setae elongate; cercal brush present.
<emphasis id="6ED0CA4FDE77545EAAD067E8FADCBAA3" bold="true" box="[1224,1296,671,697]" pageId="33" pageNumber="34">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="14BE45D6DE77545EAEDE67B0FF22BB28" pageId="33" pageNumber="34" type="discussion">
<paragraph id="5C1B165DDE77545EAEDE67B0FF22BB28" blockId="33.[151,1436,152,1138]" pageId="33" pageNumber="34">
<emphasis id="6ED0CA4FDE77545EAEDE67B0FE9FBAFB" bold="true" box="[198,339,711,737]" pageId="33" pageNumber="34">Discussion:</emphasis>
<taxonomicName id="9BA46DDEDE77545EAF4567BFFDDBBAFB" authorityName="Desjardins" authorityYear="2007" box="[349,535,712,737]" class="Insecta" family="Pteromalidae" genus="Chimaerolelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="6ED0CA4FDE77545EAF4567BFFDDBBAFB" box="[349,535,712,737]" italics="true" pageId="33" pageNumber="34">Chimaerolelaps</emphasis>
</taxonomicName>
is positioned phylogenetically as sister-group to (or near in some analyses) the
<taxonomicName id="9BA46DDEDE77545EAEDE6787FED6BB13" authorityName="Walker" authorityYear="1833" box="[198,282,752,777]" class="Insecta" family="Pteromalidae" genus="Dipara" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="6ED0CA4FDE77545EAEDE6787FED6BB13" box="[198,282,752,777]" italics="true" pageId="33" pageNumber="34">Dipara</emphasis>
</taxonomicName>
clade, although the only synapomorphy for this clade is the presence of petiolar setae in the female.
</paragraph>
</subSubSection>
<subSubSection id="14BE45D6DE77545EAEDE6648FD60BBE0" pageId="33" pageNumber="34" type="etymology">
<paragraph id="5C1B165DDE77545EAEDE6648FE27BBC8" blockId="33.[151,1436,152,1138]" pageId="33" pageNumber="34">
<emphasis id="6ED0CA4FDE77545EAEDE6648FE9BBB43" bold="true" box="[198,343,831,857]" pageId="33" pageNumber="34">Etymology:</emphasis>
<emphasis id="6ED0CA4FDE77545EAF786637FE1BBB43" box="[352,471,832,857]" italics="true" pageId="33" pageNumber="34">chimaero-</emphasis>
, to represent the odd assemblage of ancestral and derived features present in this taxon, and
<emphasis id="6ED0CA4FDE77545EAF01661FFEA5BB9B" box="[281,361,872,897]" italics="true" pageId="33" pageNumber="34">-lelaps</emphasis>
, to ally the genus with
<taxonomicName id="9BA46DDEDE77545EAC65661FFD39BB98" authorityName="Thomson" authorityYear="1876" box="[637,757,872,898]" class="Insecta" family="Pteromalidae" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="subFamily" subFamily="Diparinae">Diparinae</taxonomicName>
. The genus is identified in some collections with a manuscript name. As the manuscript name was somewhat misleading with regards to the phylogenetic position of the group, it is not used here.
</paragraph>
<paragraph id="5C1B165DDE77545EAEDF66A8FD60BBE0" blockId="33.[151,1436,152,1138]" box="[199,684,991,1018]" pageId="33" pageNumber="34">
<emphasis id="6ED0CA4FDE77545EAEDF66A8FE76BBE3" bold="true" box="[199,442,991,1017]" pageId="33" pageNumber="34">Number of Species:</emphasis>
1 described species.
</paragraph>
</subSubSection>
<subSubSection id="14BE45D6DE77545EAEDF6170FE34BC38" box="[199,504,1031,1058]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="5C1B165DDE77545EAEDF6170FE34BC38" blockId="33.[151,1436,152,1138]" box="[199,504,1031,1058]" pageId="33" pageNumber="34">
<emphasis id="6ED0CA4FDE77545EAEDF6170FEA5BC3B" bold="true" box="[199,361,1031,1057]" pageId="33" pageNumber="34">Distribution:</emphasis>
<collectingCountry id="24B356CDDE77545EAF69617FFE39BC38" box="[369,501,1032,1058]" name="Costa Rica" pageId="33" pageNumber="34">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="14BE45D6DE77545EAEDF6158FE55BC50" box="[199,409,1071,1098]" pageId="33" pageNumber="34" type="biology_ecology">
<paragraph id="5C1B165DDE77545EAEDF6158FE55BC50" blockId="33.[151,1436,152,1138]" box="[199,409,1071,1098]" pageId="33" pageNumber="34">
<emphasis id="6ED0CA4FDE77545EAEDF6158FEDABC53" bold="true" box="[199,278,1071,1097]" pageId="33" pageNumber="34">Hosts:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="14BE45D6DE77545EAEDF6120FE01BC68" box="[199,461,1111,1138]" pageId="33" pageNumber="34" type="key">
<paragraph id="5C1B165DDE77545EAEDF6120FE01BC68" blockId="33.[151,1436,152,1138]" box="[199,461,1111,1138]" pageId="33" pageNumber="34">
<emphasis id="6ED0CA4FDE77545EAEDF6120FE49BC6B" bold="true" box="[199,389,1111,1137]" pageId="33" pageNumber="34">Key to Species:</emphasis>
none.
</paragraph>
</subSubSection>
</treatment>
</document>