treatments-xml/data/4B/77/99/4B77999825FC505B97288CF823CA656D.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.157.54697" ID-GBIF-Dataset="64a99b7b-daa1-40d2-8076-8b69bedd0e1b" ID-PMC="PMC7467942" ID-Pensoft-Pub="1314-2003-157-217" ID-Pensoft-UUID="0CE239D67002528CBE89A83B04E4C21A" ID-PubMed="32934460" ModsDocID="1314-2003-157-217" checkinTime="1598489615053" checkinUser="pensoft" docAuthor="Chen, Wen-Hong, Guo, Shi-Wei, Nguyen, Hieu Quang, Chen, Li &amp; Shui, Yu-Min" docDate="2020" docId="4B77999825FC505B97288CF823CA656D" docLanguage="en" docName="PhytoKeys 157: 217-226" docOrigin="PhytoKeys 157" docSource="http://dx.doi.org/10.3897/phytokeys.157.54697" docTitle="Paraboea sinensis subsp. var. var. glabrissima W. H. Chen &amp; Y. M. Shui 2020, var. nov." docType="treatment" docVersion="4" id="0CE239D67002528CBE89A83B04E4C21A" lastPageNumber="217" masterDocId="0CE239D67002528CBE89A83B04E4C21A" masterDocTitle="Two new taxa of Gesneriaceae in the karst regions in North Vietnam" masterLastPageNumber="226" masterPageNumber="217" pageNumber="217" updateTime="1668139984387" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Two new taxa of Gesneriaceae in the karst regions in North Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Wen-Hong</mods:namePart>
<mods:affiliation>CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, 132 Lanhei Road, Kunming 650201, Yunnan, China &amp; Karst Conservation Initiative of Yunnan, Kunming 650201, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guo, Shi-Wei</mods:namePart>
<mods:affiliation>CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, 132 Lanhei Road, Kunming 650201, Yunnan, China &amp; University of the Chinese Academy of Sciences, Beijing 100049, China &amp; Karst Conservation Initiative of Yunnan, Kunming 650201, Yunnan, China</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6987-1240</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Hieu Quang</mods:namePart>
<mods:affiliation>Center for Plant Conservation of Vietnam (CPC), Vietnam Union of Science and Technology Associations, 25 / 32 Lane 191, Lac Long Quan Rd., Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Li</mods:namePart>
<mods:affiliation>CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, 132 Lanhei Road, Kunming 650201, Yunnan, China &amp; School of Life Sciences, Yunnan University, Kunming 650091, Yunnan, China</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6030-2439</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shui, Yu-Min</mods:namePart>
<mods:affiliation>CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, 132 Lanhei Road, Kunming 650201, Yunnan, China &amp; Karst Conservation Initiative of Yunnan, Kunming 650201, Yunnan, China</mods:affiliation>
<mods:nameIdentifier type="email">ymshui@mail.kib.ac.cn</mods:nameIdentifier>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>157</mods:number>
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<mods:extent unit="page">
<mods:start>217</mods:start>
<mods:end>226</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.157.54697</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.157.54697</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-157-217</mods:identifier>
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<treatment ID-GBIF-Taxon="167369783" LSID="urn:lsid:plazi:treatment:4B77999825FC505B97288CF823CA656D" httpUri="http://treatment.plazi.org/id/4B77999825FC505B97288CF823CA656D" lastPageNumber="217" pageId="0" pageNumber="217">
<subSubSection pageId="0" pageNumber="217" type="nomenclature">
<paragraph pageId="0" pageNumber="217">
<taxonomicName LSID="4B779998-25FC-505B-9728-8CF823CA656D" authority="W. H. Chen &amp; Y. M. Shui" authorityName="W. H. Chen &amp; Y. M. Shui" authorityYear="2020" class="Magnoliopsida" family="Gesneriaceae" genus="Paraboea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paraboea sinensis subsp. var. var. glabrissima" order="Lamiales" pageId="0" pageNumber="217" phylum="Tracheophyta" rank="variety" species="sinensis" status="var. nov." subSpecies="var." variety="glabrissima">Paraboea sinensis var. glabrissima W.H.Chen &amp; Y.M.Shui</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="217">var. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paraboea sinensis var. glabrissima W. H. Chen &amp; Y. M. Shui, var. nov. (A-F) and P. sinensis (Oliv.) Burtt var. sinensis (G &amp; H) A habit, arrows indicate the young leaves with thin brown pannose B inflorescence with twisted fruits, arrows showing the indumentum of abaxial surface of leaf (pe = peduncle, abl = abaxial leaf) C face view of flower D dorsal view of the calyx, the arrow showing the coherent position E birds-eye view of opened corolla F stamens and staminodes, arrows showing the staminodes G habitat H inflorescence." figureDoi="10.3897/phytokeys..54697.figure2" httpUri="https://binary.pensoft.net/fig/446116" pageId="0" pageNumber="217">Figure 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="materials_examined">
<paragraph pageId="0" pageNumber="217">Type.</paragraph>
<paragraph pageId="0" pageNumber="217">
Vientam, Pho Tho Province: Xuan Son County, Xuan Son National Park, 
<geoCoordinate degrees="21" direction="north" minutes="07" orientation="latitude" precision="1" seconds="49.3" value="21.130362">21°07'49.3&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="57" orientation="longitude" precision="15" seconds="09" value="104.9525">104°57'09&quot;E</geoCoordinate>
, 463 m a.s.l., 8 April 2016, 
<emphasis italics="true" pageId="0" pageNumber="217">Y. M. Shui</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="217">W.H. Chen</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="217">C. Liu</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="217">H.Q. Nguyen</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="217">H.T. Nguyen</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="217">N.Q. Chuong CK909</emphasis>
(holotype, KUN!; isotype, CPC!=Herbarium of Center for Plant Conservation of Vietnam).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="diagnosis">
<paragraph pageId="0" pageNumber="217">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="217">
The new variety is similar to 
<taxonomicName authority="(Oliv.) Burtt var. sinensis" class="Magnoliopsida" family="Gesneriaceae" genus="Paraboea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paraboea sinensis subsp. var. var. sinensis" order="Lamiales" pageId="0" pageNumber="217" phylum="Tracheophyta" rank="variety" species="sinensis" subSpecies="var." variety="sinensis">Paraboea sinensis (Oliv.) Burtt var. sinensis</taxonomicName>
in its morphology of habit, calyx, corolla and fruits, but differs in its young leaf abaxially, stem and peduncle sparsely and thin pannose (vs. dense and thick pannose), acute top of leaves (vs. acuminate), pistil glandular-pubescent or pubescent (vs. glabrous).
</paragraph>
<paragraph pageId="0" pageNumber="217">
Subshrubs. Stem erect, 50-80 cm tall, ca. 0.3 cm in diam., with many branches, pannose when young. Leaves opposite, equal to subequal in pairs on the stem; petiole 1-10 cm long, pannose when young; blade herbaceous, slightly asymmetric, oblong to obovate, 9-19 
<normalizedToken originalValue="×">x</normalizedToken>
3.5-8 cm, base cuneate, apex acute, margin denticulate from the base, adaxially glabrous, abaxially sparsely pannose when young; venation penninerved, lateral veins 7-12 on each side of the midrib. Cymes axillary near branch apices; peduncle 1.5-4.5 cm long, sparsely pannose; bracts caducous; pedicel 1.4-2 cm long, sparsely pannose; bracteoles caducous. Calyx zygomorphic, 2-lipped, adaxial calyx ca. 1 cm long, 3-lobed to the middle, lobes rounded ca. 0.5 
<normalizedToken originalValue="×">x</normalizedToken>
0.5 cm, abaxial calyx 2-lobed to the base, lobes obovate, ca. 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
0.6 cm, apex round, margin entire, outside glabrous, inside glabrous. Corolla campanulate, zygomorphic, 2-2.3 cm long, ca. 1.3 cm wide at the throat, both sides glabrous, tube 1-1.4 cm long; limb 2-lipped; adaxial lip 2-lobed, lobes broadly ovate, ca. 0.9 
<normalizedToken originalValue="×">x</normalizedToken>
1.5 cm; abaxial lip 3-lobed, lobes broadly ovate, middle lobe ca. 0.7 
<normalizedToken originalValue="×">x</normalizedToken>
1.2 cm. Stamens 2, adnate to the corolla base; anthers glabrous; filaments ca. 1 cm long, pubescent, staminodes 3, glandular-pubescent, lateral ones 2.5-3.0 mm long, adnate to the corolla tube base, the middle one ca. 1 mm long, adnate to the corolla tube ca. 2 mm above the base. Disc absent. Pistil ca. 1.2 cm long; ovary linear, ca. 0.8 cm long, sparsely glandular-pubescent or pubescent; style linear, ca. 0.4 cm long, glandular-pubescent; stigma 1, capitate. Capsule linear, spirally twisted, 2.5-3.7 cm long, glabrous.
</paragraph>
<caption doi="10.3897/phytokeys..54697.figure2" httpUri="https://binary.pensoft.net/fig/446116" pageId="0" pageNumber="217" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="217">
<emphasis bold="true" pageId="0" pageNumber="217">Figure 2.</emphasis>
<taxonomicName authorityName="W. H. Chen &amp; Y. M. Shui" authorityYear="2020" class="Magnoliopsida" family="Gesneriaceae" genus="Paraboea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paraboea sinensis subsp. var. var. glabrissima" order="Lamiales" pageId="0" pageNumber="217" phylum="Tracheophyta" rank="variety" species="sinensis" subSpecies="var." variety="glabrissima">Paraboea sinensis var. glabrissima</taxonomicName>
W.H.Chen &amp; Y.M.Shui, var. nov. (
<emphasis bold="true" pageId="0" pageNumber="217">A-F</emphasis>
) and 
<taxonomicName authority="(Oliv.) Burtt var. sinensis" lsidName="P. sinensis subsp. var. var. sinensis" pageId="0" pageNumber="217" rank="variety" species="sinensis" subSpecies="var." variety="sinensis">P. sinensis (Oliv.) Burtt var. sinensis</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="217">G &amp; H</emphasis>
) 
<emphasis bold="true" pageId="0" pageNumber="217">A</emphasis>
habit, arrows indicate the young leaves with thin brown pannose 
<emphasis bold="true" pageId="0" pageNumber="217">B</emphasis>
inflorescence with twisted fruits, arrows showing the indumentum of abaxial surface of leaf (pe = peduncle, abl = abaxial leaf) 
<emphasis bold="true" pageId="0" pageNumber="217">C</emphasis>
face view of flower 
<emphasis bold="true" pageId="0" pageNumber="217">D</emphasis>
dorsal view of the calyx, the arrow showing the coherent position 
<emphasis bold="true" pageId="0" pageNumber="217">E</emphasis>
birds-eye view of opened corolla 
<emphasis bold="true" pageId="0" pageNumber="217">F</emphasis>
stamens and staminodes, arrows showing the staminodes 
<emphasis bold="true" pageId="0" pageNumber="217">G</emphasis>
habitat 
<emphasis bold="true" pageId="0" pageNumber="217">H</emphasis>
inflorescence.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="phenology">
<paragraph pageId="0" pageNumber="217">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="217">Flowering from March to April, fruiting from April to May.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="etymology">
<paragraph pageId="0" pageNumber="217">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="217">
The word &quot; 
<taxonomicName lsidName="glabrissima" pageId="0" pageNumber="217" rank="species" species="glabrissima">
<emphasis italics="true" pageId="0" pageNumber="217">glabrissima</emphasis>
</taxonomicName>
&quot; indicates the glabrous surface across the whole plant except the young leaves (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paraboea sinensis var. glabrissima W. H. Chen &amp; Y. M. Shui, var. nov. (A-F) and P. sinensis (Oliv.) Burtt var. sinensis (G &amp; H) A habit, arrows indicate the young leaves with thin brown pannose B inflorescence with twisted fruits, arrows showing the indumentum of abaxial surface of leaf (pe = peduncle, abl = abaxial leaf) C face view of flower D dorsal view of the calyx, the arrow showing the coherent position E birds-eye view of opened corolla F stamens and staminodes, arrows showing the staminodes G habitat H inflorescence." figureDoi="10.3897/phytokeys..54697.figure2" httpUri="https://binary.pensoft.net/fig/446116" pageId="0" pageNumber="217">2A</figureCitation>
), which become glabrous as soon as the leaves become mature. In this manner, it is different from the original variety of 
<taxonomicName authority="var. sinensis" class="Magnoliopsida" family="Gesneriaceae" genus="Paraboea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paraboea sinensis subsp. var. var. sinensis" order="Lamiales" pageId="0" pageNumber="217" phylum="Tracheophyta" rank="variety" species="sinensis" subSpecies="var." variety="sinensis">Paraboea sinensis var. sinensis</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="distribution">
<paragraph pageId="0" pageNumber="217">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="217">The new variety only occurs in Xuan Son County, Pho Tho Province and grows in soil, on rocks in limestone forests.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="provisional conservation status">
<paragraph pageId="0" pageNumber="217">Provisional conservation status.</paragraph>
<paragraph pageId="0" pageNumber="217">
Based on field surveys and detailed discussions with Vietnamese colleagues, including Hieu Quang Nguyen, this new variety has only been observed in the Xuan Son National Park. There were only a total of 30 mature individuals, so we provisionally considered it as Critically Endangered (CR): B1b (v) + 2b (v). (
<bibRefCitation author="IUCN" journalOrPublisher="IUCN, Gland, Switzerland and Cambridge" pageId="0" pageNumber="217" publicationUrl="https://portals.iucn.org/library/sites/library/files/documents/RL-2001-001-2nd.pdf" refId="B8" refString="IUCN, 2012. IUCN Red List Categories and Criteria, Version 3.1. 2nd edition. IUCN, Gland, Switzerland and Cambridge, https://portals.iucn.org/library/sites/library/files/documents/RL-2001-001-2nd.pdf" title="IUCN Red List Categories and Criteria, Version 3.1. 2 nd edition." url="https://portals.iucn.org/library/sites/library/files/documents/RL-2001-001-2nd.pdf" year="2012">IUCN 2012</bibRefCitation>
; IUCN Standards and Petitions Subcommittee 2017).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="notes">
<paragraph pageId="0" pageNumber="217">Note.</paragraph>
<paragraph pageId="0" pageNumber="217">
This new variety is very similar to the original variety of 
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Paraboea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paraboea sinensis" order="Lamiales" pageId="0" pageNumber="217" phylum="Tracheophyta" rank="species" species="sinensis">
<emphasis italics="true" pageId="0" pageNumber="217">Paraboea sinensis</emphasis>
</taxonomicName>
in subshrub habit, obovate calyces and twisted fruits, but different in the almost glabrous habit (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison of Paraboea sinensis var. glabrissima W. H. Chen &amp; Y. M. Shui and P. sinensis var. sinensis." httpUri="http://table.plazi.org/id/9A491EF809496D80E389160F5A81F5F2" pageId="0" pageNumber="217" tableUuid="9A491EF809496D80E389160F5A81F5F2">1</tableCitation>
; 
<bibRefCitation author="Wang, WT" editor="Wu, ZY" journalOrPublisher="18. Science Press (Beijing) and Missouri Botanical Garden Press (St. Louis)" pageId="0" pageNumber="217" pagination="268 - 272" refId="B28" refString="Wang, WT, Pan, KY, Li, ZY, Weitman, AL, Skog, LE, 1998. Gesneriaceae. In: Wu, ZY, Raven, PH, Eds., Flora of China Vol. 18. Science Press (Beijing) and Missouri Botanical Garden Press (St. Louis): 268 - 272" title="Gesneriaceae." volumeTitle="Flora of China Vol." year="1998">Wang et al. 1998</bibRefCitation>
; 
<bibRefCitation author="Xu, ZR" journalOrPublisher="Edinburgh Journal of Botany" pageId="0" pageNumber="217" pagination="161 - 347" refId="B31" refString="Xu, ZR, Burtt, BL, Skog, LE, Middleton, DJ, 2008. A revision of Paraboea (Gesneriaceae). Edinburgh Journal of Botany 65 (2): 161 - 347" title="A revision of Paraboea (Gesneriaceae)." volume="65" year="2008">Xu et al. 2008</bibRefCitation>
). Furthermore, the new variety is covered by pannose indumenti on the young leaves (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The distribution of Paraboea sinensis var. glabrissima W. H. Chen &amp; Y. M. Shui var. nov. (triangle) and Primulina xuansonensis W. H. Chen &amp; Y. M. Shui sp. nov. (square)." figureDoi="10.3897/phytokeys..54697.figure1" httpUri="https://binary.pensoft.net/fig/446115" pageId="0" pageNumber="217">1A</figureCitation>
), but quickly becomes glabrous on the mature ones, while pannose indumenti consistently covers the original variety on both young and mature leaves (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The distribution of Paraboea sinensis var. glabrissima W. H. Chen &amp; Y. M. Shui var. nov. (triangle) and Primulina xuansonensis W. H. Chen &amp; Y. M. Shui sp. nov. (square)." figureDoi="10.3897/phytokeys..54697.figure1" httpUri="https://binary.pensoft.net/fig/446115" pageId="0" pageNumber="217">1G, H</figureCitation>
). Additionally, the top of the leaf is acute rather than acuminate in the original variety. It is necessary to explore the morphology diversity and genetic differentiation of 
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Paraboea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paraboea sinensis" order="Lamiales" pageId="0" pageNumber="217" phylum="Tracheophyta" rank="species" species="sinensis">
<emphasis italics="true" pageId="0" pageNumber="217">Paraboea sinensis</emphasis>
</taxonomicName>
in the future, a wide-distributed species in the genus.
</paragraph>
<caption ID-Table-UUID="9A491EF809496D80E389160F5A81F5F2" httpUri="http://table.plazi.org/id/9A491EF809496D80E389160F5A81F5F2" pageId="0" pageNumber="217" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="217">
<emphasis bold="true" pageId="0" pageNumber="217">Table 1.</emphasis>
Morphological comparison of 
<taxonomicName authorityName="W. H. Chen &amp; Y. M. Shui" authorityYear="2020" class="Magnoliopsida" family="Gesneriaceae" genus="Paraboea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paraboea sinensis subsp. var. var. glabrissima" order="Lamiales" pageId="0" pageNumber="217" phylum="Tracheophyta" rank="variety" species="sinensis" subSpecies="var." variety="glabrissima">Paraboea sinensis var. glabrissima</taxonomicName>
W.H. Chen &amp; Y.M. Shui and 
<taxonomicName authority="var. sinensis" lsidName="P. sinensis subsp. var. var. sinensis" pageId="0" pageNumber="217" rank="variety" species="sinensis" subSpecies="var." variety="sinensis">P. sinensis var. sinensis</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="217">
<table pageId="0" pageNumber="217">
<tr pageId="0" pageNumber="217">
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">Characters</th>
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
<taxonomicName lsidName="P. sinensis subsp. var. var. glabrissima" pageId="0" pageNumber="217" rank="variety" species="sinensis" subSpecies="var." variety="glabrissima">P. sinensis var. glabrissima</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
<taxonomicName authority="var. sinensis" lsidName="P. sinensis subsp. var. var. sinensis" pageId="0" pageNumber="217" rank="variety" species="sinensis" subSpecies="var." variety="sinensis">P. sinensis var. sinensis</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Indumenti</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Young leaf abaxially, stem and peduncle sparsely and thin pannose, mature glabrous</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Young and mature leaf abaxially, stem and peduncle dense and thick pannose</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Base of leaves</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Cuneate</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Broadly cuneate to round</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Top of leaves</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Acute</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Acuminate</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Pistil</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Glandular pubescent or pubescent</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Glabrous</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>