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<mods:title id="6E01F02AC242FA0B43F5B232CABE1D21">Brachiopods from Late Jurassic-Early Cretaceous hydrocarbon seep deposits, central Spitsbergen, Svalbard</mods:title>
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<mods:namePart id="04C2FE7ADDB4F9D78B609EE7A339AFEC">Sandy, Michael R.</mods:namePart>
<mods:affiliation id="477B208ECD374670ADA11703C09086FD">Department of Geology, University of Dayton, Dayton, OH 45469 - 2364, USA. E-mail: msandy 1 @ udayton. edu</mods:affiliation>
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<mods:namePart id="49D9A928BDA1E984F8EE35060A21D6BD">Hryniewicz, Krzysztof</mods:namePart>
<mods:affiliation id="30DAAC4B252B0968C2548E70380D2FBD">Natural History Museum, University of Oslo, P. O. Box 1172, NO- 0318 Oslo, Norway. &amp; Current address: Institute of Paleobiology, Polish Academy of Sciences, ul. Twarda 51 / 55, 00 - 818 Warszawa, Poland.</mods:affiliation>
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<mods:namePart id="DB20731C5840EA24A0A3622CC4627125">Hammer, Øyvind</mods:namePart>
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<mods:namePart id="14BEDC4503E7C848F5616E54E7C4AAFC">Nakrem, Hans Arne</mods:namePart>
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<treatment id="466F6C33625EFFB6968547EAFBB033E6" ID-DOI="http://doi.org/10.5281/zenodo.4952536" ID-GBIF-Taxon="184851308" ID-Zenodo-Dep="4952536" LSID="urn:lsid:plazi:treatment:466F6C33625EFFB6968547EAFBB033E6" httpUri="http://treatment.plazi.org/id/466F6C33625EFFB6968547EAFBB033E6" lastPageId="9" lastPageNumber="510" pageId="6" pageNumber="507">
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<paragraph id="CE79DD25625EFFB9968547EAFD743650" blockId="6.[151,672,1529,1589]" box="[151,672,1529,1555]" pageId="6" pageNumber="507">
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<taxonomicName id="09C6A6A6625EFFB9968547EAFD743650" authority="Smirnova, 1987" authorityName="Smirnova. It" authorityYear="1987" box="[151,672,1529,1555]" class="Rhynchonellata" family="Norellidae" genus="Pseudomonticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="6" pageNumber="507" phylum="Brachiopoda" rank="species" species="varia">
<emphasis id="FCB20137625EFFB9968547EAFD743650" bold="true" box="[151,672,1529,1555]" pageId="6" pageNumber="507">
<emphasis id="FCB20137625EFFB9968547EAFE013650" bold="true" box="[151,469,1529,1555]" italics="true" pageId="6" pageNumber="507">Pseudomonticlarella varia</emphasis>
<bibRefCitation id="AA57A0D4625EFFB997CE47EAFD743650" author="Smirnova, T. N." box="[476,672,1529,1555]" pageId="6" pageNumber="507" pagination="39 - 49" refId="ref18461" refString="Smirnova, T. N. (1987) New taxa of late Volgian and early Hauterivian brachiopods of the Russian platform. Paleontological Journal, 21 (1), 39 - 49." type="journal article" year="1987">Smirnova, 1987</bibRefCitation>
</emphasis>
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<subSubSection id="86DC8EAE625EFFB99685440FFBFB36DB" pageId="6" pageNumber="507" type="description">
<paragraph id="CE79DD25625EFFB99685440FFEB13676" blockId="6.[151,672,1529,1589]" box="[151,357,1564,1589]" pageId="6" pageNumber="507">
(
<figureCitation id="56FDC1A0625EFFB9968D440FFE903676" box="[159,324,1564,1589]" captionStart="FIGURE 3" captionStartId="7.[151,250,1756,1778]" captionTargetBox="[261,1312,207,1726]" captionTargetId="figure-152@7.[260,1316,193,1726]" captionTargetPageId="7" captionText="FIGURE 3. 112. Pseudomonticlarella varia Smirnova. 14. PMO 227.416, seep 13, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 4). 58. PMO 227.419, seep 7, dorsal, ventral, lateral, and anterior views. 912. PMO 227.418, seep 13, dorsal, ventral, lateral, and anterior views. 1332. Ptilorhynchia mclachlani sp. nov. 1316. PMO 224.876, seep 9, dorsal, ventral, lateral, and anterior views. 1720, PMO 224.879, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5B). 2124. PMO 227.431, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5A). 2528. PMO 224.871, seep 9, dorsal, ventral, lateral, and anterior views. 2932. Holotype, PMO 224.882, seep 9, dorsal, ventral, lateral, and anterior views. Dotted line delimits photographs at the same magnification. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951754" httpUri="https://zenodo.org/record/4951754/files/figure.png" pageId="6" pageNumber="507">Figs. 3.13.12</figureCitation>
,
<figureCitation id="56FDC1A0625EFFB9975C440FFE893676" box="[334,349,1564,1589]" captionStart="FIGURE 4" captionStartId="8.[151,250,1704,1726]" captionTargetBox="[162,1434,188,1692]" captionTargetId="figure-173@8.[149,1434,184,1692]" captionTargetPageId="8" captionText="FIGURE 4. Transverse serial sections of Pseudomonticlarella varia Smirnova, seep 13, PMO 227.416. Sections 0.41.1 mm given at higher magnification below dotted black line. Serial sections prepared perpendicular to maximum length of the specimen, numbering of sections starts at 0.0 mm the initial section through the specimen. Dimensions of sectioned specimen: L 7.0+; W 8.1; T 4.1 mm (+ = dimension damaged/specimen broken in that direction). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951758" httpUri="https://zenodo.org/record/4951758/files/figure.png" pageId="6" pageNumber="507">4</figureCitation>
)
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<paragraph id="CE79DD25625EFFB996854477FB583639" blockId="6.[151,1164,1636,1688]" box="[151,1164,1636,1658]" pageId="6" pageNumber="507">
1987
<taxonomicName id="09C6A6A6625EFFB996E74477FDD6363A" authority="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little, 2014" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[245,514,1636,1658]" class="Rhynchonellata" family="Norellidae" genus="Pseudomonticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="6" pageNumber="507" phylum="Brachiopoda" rank="species" species="varia" status="sp. nov.">
<emphasis id="FCB20137625EFFB996E74477FDD6363A" box="[245,514,1636,1658]" italics="true" pageId="6" pageNumber="507">Pseudomonticlarella varia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E781BC4C625EFFB9941B4477FD553639" box="[521,641,1636,1658]" pageId="6" pageNumber="507" rank="species">new species</taxonomicNameLabel>
—Smirnova, p. 33, pl. 4, figs. 1, 2, 57, text-fig. 1.
</paragraph>
<paragraph id="CE79DD25625EFFB996854491FBFB36DB" blockId="6.[151,1164,1636,1688]" box="[151,1071,1666,1688]" pageId="6" pageNumber="507">
2002
<taxonomicName id="09C6A6A6625EFFB996E74491FDBF36DB" authority="Smirnova" authorityName="Smirnova. It" authorityYear="1987" box="[245,619,1666,1688]" class="Rhynchonellata" family="Norellidae" genus="Pseudomonticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="6" pageNumber="507" phylum="Brachiopoda" rank="species" species="varia">
<emphasis id="FCB20137625EFFB996E74491FDD636DB" box="[245,514,1666,1688]" italics="true" pageId="6" pageNumber="507">Pseudomonticlarella varia</emphasis>
Smirnova
</taxonomicName>
—Manceñido
<emphasis id="FCB20137625EFFB994EF4490FCE436DB" box="[765,816,1666,1688]" italics="true" pageId="6" pageNumber="507">et al.</emphasis>
, p. 1314, fig. 891.3a
<specimenCount id="D8C016AC625EFFB992014491FBFB36DB" box="[1043,1071,1666,1688]" pageId="6" pageNumber="507" type="generic">3p.</specimenCount>
</paragraph>
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<subSubSection id="86DC8EAE625EFFB9968544D6FD1D3741" pageId="6" pageNumber="507" type="materials_examined">
<paragraph id="CE79DD25625EFFB9968544D6FD1D3741" blockId="6.[151,1437,1733,2010]" pageId="6" pageNumber="507">
<emphasis id="FCB20137625EFFB9968544D6FE1F369E" box="[151,459,1733,1758]" inLineHeading="true" pageId="6" pageNumber="507" reason="1">
<emphasis id="FCB20137625EFFB9968544D6FE10369D" bold="true" box="[151,452,1733,1758]" pageId="6" pageNumber="507">Material and occurrence</emphasis>
.
</emphasis>
Seep 7 (PMO 227.419), seep 13 (PMO 227.416418), seep 10 [damaged specimen, anterior commissure not present, PMO 224.923?].
</paragraph>
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<subSubSection id="86DC8EAE625EFFB796D5451EFC5937AC" lastPageId="8" lastPageNumber="509" pageId="6" pageNumber="507" type="description">
<paragraph id="CE79DD25625EFFB996D5451EFDF437D1" blockId="6.[151,1437,1733,2010]" pageId="6" pageNumber="507">
<emphasis id="FCB20137625EFFB996D5451EFE8D3765" bold="true" box="[199,345,1805,1830]" inLineHeading="true" pageId="6" pageNumber="507" reason="1">Description.</emphasis>
Small (less than
<quantity id="093E70C0625EFFB9940F451EFD813766" box="[541,597,1805,1830]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="507" unit="cm" value="1.0">1 cm</quantity>
in length, width, and thickness), smooth rhynchonellide that shows traces of capillate ornament in places. However, the decortication of specimens makes it difficult to verify the ornament of the shell exterior. The anterior commissure is bisulcate with weakly developed sulcii at the commissure; central sulcus shallower than lateral sulcii.
</paragraph>
<paragraph id="CE79DD25625EFFB796D5458EFC5937AC" blockId="6.[151,1437,1733,2010]" lastBlockId="8.[151,1436,1862,2031]" lastPageId="8" lastPageNumber="509" pageId="6" pageNumber="507">
The following internal characters may be visible from external examination, either through translucent shell or because of decortication: dental lamellae in the ventral umbo; a median septum extending at least half the length of the dorsal valve. Internal characters were also determined from serial sections (
<figureCitation id="56FDC1A06250FFB792284555FB57371C" box="[1082,1155,1862,1887]" captionStart="FIGURE 4" captionStartId="8.[151,250,1704,1726]" captionTargetBox="[162,1434,188,1692]" captionTargetId="figure-173@8.[149,1434,184,1692]" captionTargetPageId="8" captionText="FIGURE 4. Transverse serial sections of Pseudomonticlarella varia Smirnova, seep 13, PMO 227.416. Sections 0.41.1 mm given at higher magnification below dotted black line. Serial sections prepared perpendicular to maximum length of the specimen, numbering of sections starts at 0.0 mm the initial section through the specimen. Dimensions of sectioned specimen: L 7.0+; W 8.1; T 4.1 mm (+ = dimension damaged/specimen broken in that direction). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951758" httpUri="https://zenodo.org/record/4951758/files/figure.png" pageId="8" pageNumber="509">Fig. 4</figureCitation>
): a small septalium-like structure is present due to the concave ventral surface of the hinge plates (
<figureCitation id="56FDC1A06250FFB792134579FB9D37C0" box="[1025,1097,1898,1923]" captionStart="FIGURE 4" captionStartId="8.[151,250,1704,1726]" captionTargetBox="[162,1434,188,1692]" captionTargetId="figure-173@8.[149,1434,184,1692]" captionTargetPageId="8" captionText="FIGURE 4. Transverse serial sections of Pseudomonticlarella varia Smirnova, seep 13, PMO 227.416. Sections 0.41.1 mm given at higher magnification below dotted black line. Serial sections prepared perpendicular to maximum length of the specimen, numbering of sections starts at 0.0 mm the initial section through the specimen. Dimensions of sectioned specimen: L 7.0+; W 8.1; T 4.1 mm (+ = dimension damaged/specimen broken in that direction). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951758" httpUri="https://zenodo.org/record/4951758/files/figure.png" pageId="8" pageNumber="509">Fig. 4</figureCitation>
, sections
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); crura develop on the inner margins of the hinge plates (sections 0.50.7); crura arch ventrally (sections 0.71.1). Dental lamellae were not seen in the serial sections (ventral valve umbo of the sectioned specimen was damaged,
<figureCitation id="56FDC1A06250FFB7936245A1FF3A37AC" captionStart="FIGURE 3" captionStartId="7.[151,250,1756,1778]" captionTargetBox="[261,1312,207,1726]" captionTargetId="figure-152@7.[260,1316,193,1726]" captionTargetPageId="7" captionText="FIGURE 3. 112. Pseudomonticlarella varia Smirnova. 14. PMO 227.416, seep 13, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 4). 58. PMO 227.419, seep 7, dorsal, ventral, lateral, and anterior views. 912. PMO 227.418, seep 13, dorsal, ventral, lateral, and anterior views. 1332. Ptilorhynchia mclachlani sp. nov. 1316. PMO 224.876, seep 9, dorsal, ventral, lateral, and anterior views. 1720, PMO 224.879, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5B). 2124. PMO 227.431, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5A). 2528. PMO 224.871, seep 9, dorsal, ventral, lateral, and anterior views. 2932. Holotype, PMO 224.882, seep 9, dorsal, ventral, lateral, and anterior views. Dotted line delimits photographs at the same magnification. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4951754" httpUri="https://zenodo.org/record/4951754/files/figure.png" pageId="8" pageNumber="509">Fig. 3.13.4</figureCitation>
), but observed from the external examination of specimens.
</paragraph>
</subSubSection>
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<paragraph id="CE79DD25625FFFB8968544CFFE3F378B" blockId="7.[151,1436,1756,1993]" pageId="7" pageNumber="508">
<emphasis id="FCB20137625FFFB8968544CFFEC236B1" bold="true" box="[151,278,1756,1778]" pageId="7" pageNumber="508">FIGURE 3.</emphasis>
112.
<taxonomicName id="09C6A6A6625FFFB8977144CFFD3C36B1" authority="Smirnova." authorityName="Smirnova. It" authorityYear="1987" box="[355,744,1756,1778]" class="Rhynchonellata" family="Norellidae" genus="Pseudomonticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="7" pageNumber="508" phylum="Brachiopoda" rank="species" species="varia">
<emphasis id="FCB20137625FFFB8977144CFFDA136B1" box="[355,629,1756,1778]" italics="true" pageId="7" pageNumber="508">Pseudomonticlarella varia</emphasis>
Smirnova.
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14. PMO 227.416, seep 13, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 4). 58. PMO 227.419, seep 7, dorsal, ventral, lateral, and anterior views. 912. PMO 227.418, seep 13, dorsal, ventral, lateral, and anterior views. 1332.
<taxonomicName id="09C6A6A6625FFFB895B8450AFB7B376C" authority="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little, 2014" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[938,1199,1817,1839]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="7" pageNumber="508" phylum="Brachiopoda" rank="species" species="mclachlani" status="sp. nov.">
<emphasis id="FCB20137625FFFB895B8450AFB7B376C" box="[938,1199,1817,1839]" italics="true" pageId="7" pageNumber="508">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
<emphasis id="FCB20137625FFFB892AA450AFAD8376C" bold="true" box="[1208,1292,1817,1839]" pageId="7" pageNumber="508">
<taxonomicNameLabel id="E781BC4C625FFFB892AA450AFAD8376C" box="[1208,1292,1817,1839]" pageId="7" pageNumber="508" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1316. PMO 224.876, seep 9, dorsal, ventral, lateral, and anterior views. 1720, PMO 224.879, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5B). 2124. PMO 227.431, seep 9, dorsal, ventral, lateral, and anterior views. Specimen serially sectioned (Figure 5A). 2528. PMO 224.871, seep 9, dorsal, ventral, lateral, and anterior views. 2932. Holotype, PMO 224.882, seep 9, dorsal, ventral, lateral, and anterior views. Dotted line delimits photographs at the same magnification. Scale bars = 1 cm.
</paragraph>
</caption>
<caption id="9AB98DAD6250FFB7968544BBFB183759" ID-DOI="http://doi.org/10.5281/zenodo.4951758" ID-Zenodo-Dep="4951758" httpUri="https://zenodo.org/record/4951758/files/figure.png" pageId="8" pageNumber="509" startId="8.[151,250,1704,1726]" targetBox="[162,1434,188,1692]" targetPageId="8">
<paragraph id="CE79DD256250FFB7968544BBFB183759" blockId="8.[151,1437,1704,1818]" pageId="8" pageNumber="509">
<emphasis id="FCB201376250FFB7968544BBFEC736FD" bold="true" box="[151,275,1704,1726]" pageId="8" pageNumber="509">FIGURE 4.</emphasis>
Transverse serial sections of
<taxonomicName id="09C6A6A66250FFB7945644BBFC6B36FD" authority="Smirnova" authorityName="Smirnova. It" authorityYear="1987" box="[580,959,1704,1726]" class="Rhynchonellata" family="Norellidae" genus="Pseudomonticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="8" pageNumber="509" phylum="Brachiopoda" rank="species" species="varia">
<emphasis id="FCB201376250FFB7945644BBFC8736FD" box="[580,851,1704,1726]" italics="true" pageId="8" pageNumber="509">Pseudomonticlarella varia</emphasis>
Smirnova
</taxonomicName>
, seep 13, PMO 227.416. Sections 0.41.1 mm given at higher magnification below dotted black line. Serial sections prepared perpendicular to maximum length of the specimen, numbering of sections starts at 0.0 mm the initial section through the specimen. Dimensions of sectioned specimen: L 7.0+; W 8.1; T 4.1 mm (+ = dimension damaged/specimen broken in that direction). Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="86DC8EAE6251FFB696D54284FBB033E6" pageId="9" pageNumber="510" type="discussion">
<paragraph id="CE79DD256251FFB696D54284FE593103" blockId="9.[151,1437,151,933]" pageId="9" pageNumber="510">
<emphasis id="FCB201376251FFB696D54284FE9830F3" bold="true" box="[199,332,151,176]" inLineHeading="true" pageId="9" pageNumber="510" reason="1">Discussion.</emphasis>
The general outline and profile of the specimens from Spitsbergen are similar to those of specimens figured by
<bibRefCitation id="AA57A0D46251FFB6979C42AFFD923096" author="Smirnova, T. N." box="[398,582,188,213]" pageId="9" pageNumber="510" pagination="39 - 49" refId="ref18461" refString="Smirnova, T. N. (1987) New taxa of late Volgian and early Hauterivian brachiopods of the Russian platform. Paleontological Journal, 21 (1), 39 - 49." type="journal article" year="1987">Smirnova (1987</bibRefCitation>
, pl. 4, figs. 12) from the Russian Platform, in particular that in pl. 4, fig. 1. This specimen appears to be smooth, developing costae at the anterior margin. The Russian Platform specimens reach maximum lengths close to
<quantity id="093E70C06251FFB694014317FD823158" box="[531,598,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="510" unit="mm" value="6.0">6 mm</quantity>
(
<bibRefCitation id="AA57A0D46251FFB694744317FCCC315E" author="Smirnova, T. N." box="[614,792,260,285]" pageId="9" pageNumber="510" pagination="39 - 49" refId="ref18461" refString="Smirnova, T. N. (1987) New taxa of late Volgian and early Hauterivian brachiopods of the Russian platform. Paleontological Journal, 21 (1), 39 - 49." type="journal article" year="1987">Smirnova 1987</bibRefCitation>
, p. 42) so are smaller than the Spitsbergen material that reach
<quantity id="093E70C06251FFB696C84334FECF317C" box="[218,283,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="510" unit="mm" value="8.0">8 mm</quantity>
in length.
</paragraph>
<paragraph id="CE79DD256251FFB696D5435FFDEC325B" blockId="9.[151,1437,151,933]" pageId="9" pageNumber="510">
Smirnova provided serial sections of
<specimenCount id="D8C016AC6251FFB6946F435FFCFC3126" box="[637,808,332,357]" pageId="9" pageNumber="510" type="generic">two specimens</specimenCount>
of
<taxonomicName id="09C6A6A66251FFB69547435FFB573127" authorityName="Smirnova. It" authorityYear="1987" box="[853,1155,332,357]" class="Rhynchonellata" family="Norellidae" genus="Pseudomonticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="varia">
<emphasis id="FCB201376251FFB69547435FFB573127" box="[853,1155,332,357]" italics="true" pageId="9" pageNumber="510">Pseudomonticlarella varia</emphasis>
</taxonomicName>
(1987, fig. 1ab). Both series of sections show a septalium and crura that arch ventrally anteriorly. These are features that are shared with the Spitsbergen specimen (
<figureCitation id="56FDC1A06251FFB697D84387FDC631EE" box="[458,530,404,429]" captionStart="FIGURE 4" captionStartId="8.[151,250,1704,1726]" captionTargetBox="[162,1434,188,1692]" captionTargetId="figure-173@8.[149,1434,184,1692]" captionTargetPageId="8" captionText="FIGURE 4. Transverse serial sections of Pseudomonticlarella varia Smirnova, seep 13, PMO 227.416. Sections 0.41.1 mm given at higher magnification below dotted black line. Serial sections prepared perpendicular to maximum length of the specimen, numbering of sections starts at 0.0 mm the initial section through the specimen. Dimensions of sectioned specimen: L 7.0+; W 8.1; T 4.1 mm (+ = dimension damaged/specimen broken in that direction). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4951758" httpUri="https://zenodo.org/record/4951758/files/figure.png" pageId="9" pageNumber="510">Fig. 4</figureCitation>
). The internal structures (from the initial section through the brachiopod) were traced for
<quantity id="093E70C06251FFB6971E43A4FEA4318C" box="[268,368,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="4.12" pageId="9" pageNumber="510" unit="mm" value="4.12">4.12 mm</quantity>
in specimen No. 139/584 (
<bibRefCitation id="AA57A0D46251FFB694B643A4FC813193" author="Smirnova, T. N." box="[676,853,439,464]" pageId="9" pageNumber="510" pagination="39 - 49" refId="ref18461" refString="Smirnova, T. N. (1987) New taxa of late Volgian and early Hauterivian brachiopods of the Russian platform. Paleontological Journal, 21 (1), 39 - 49." type="journal article" year="1987">Smirnova 1987</bibRefCitation>
, fig. 1a) and for
<quantity id="093E70C06251FFB6920643A4FBAC318C" box="[1044,1144,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="9" pageNumber="510" unit="mm" value="1.45">1.45 mm</quantity>
in specimen No. 139/590 (fig. 1b). In the Spitsbergen specimen the internal structures were traced to section
<quantity id="093E70C06251FFB6925743CFFB4F31B0" box="[1093,1179,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="510" unit="mm" value="1.2">1.2 mm</quantity>
(i.e.,
<quantity id="093E70C06251FFB692CF43CFFAE031B0" box="[1245,1332,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="510" unit="mm" value="1.2">1.2 mm</quantity>
from the initial section through the specimen).
</paragraph>
<paragraph id="CE79DD256251FFB696D54037FB3C3318" blockId="9.[151,1437,151,933]" pageId="9" pageNumber="510">
It was thought originally that these diminutive specimens could be juveniles of the form referred herein to
<taxonomicName id="09C6A6A66251FFB696854054FE623223" authority="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little, 2014" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[151,438,583,608]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="mclachlani" status="sp. nov.">
<emphasis id="FCB201376251FFB696854054FE623223" box="[151,438,583,608]" italics="true" pageId="9" pageNumber="510">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
<emphasis id="FCB201376251FFB697AF4054FDCD3223" bold="true" box="[445,537,583,608]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="E781BC4C6251FFB697AF4054FDCD3223" box="[445,537,583,608]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
However, posteriorly the ventral valve of
<taxonomicName id="09C6A6A66251FFB695EF4054FAFD321C" authorityName="Smirnova. It" authorityYear="1987" box="[1021,1321,583,608]" class="Rhynchonellata" family="Norellidae" genus="Pseudomonticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="varia">
<emphasis id="FCB201376251FFB695EF4054FAFD321C" box="[1021,1321,583,608]" italics="true" pageId="9" pageNumber="510">Pseudomonticlarella varia</emphasis>
</taxonomicName>
lacks any developing or incipient costae, which are often present posteriorly on specimens of
<taxonomicName id="09C6A6A66251FFB69266407FFA4032C6" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[1140,1428,620,645]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376251FFB69266407FFA4032C6" box="[1140,1428,620,645]" italics="true" pageId="9" pageNumber="510">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
. Also, posteriorly, the dorsal valve of
<taxonomicName id="09C6A6A66251FFB69459409CFCAD32E4" authorityName="Smirnova. It" authorityYear="1987" box="[587,889,655,680]" class="Rhynchonellata" family="Norellidae" genus="Pseudomonticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="varia">
<emphasis id="FCB201376251FFB69459409CFCAD32E4" box="[587,889,655,680]" italics="true" pageId="9" pageNumber="510">Pseudomonticlarella varia</emphasis>
</taxonomicName>
is weakly sulcate (possibly enhanced by some decortication). This sulcate condition does not appear to characterize specimens of
<taxonomicName id="09C6A6A66251FFB6924740A7FAA0328E" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[1109,1396,692,717]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376251FFB6924740A7FAA0328E" box="[1109,1396,692,717]" italics="true" pageId="9" pageNumber="510">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
. In addition,
<taxonomicName id="09C6A6A66251FFB6971340C4FDCB32B3" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[257,543,727,752]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376251FFB6971340C4FDCB32B3" box="[257,543,727,752]" italics="true" pageId="9" pageNumber="510">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
and
<taxonomicName id="09C6A6A66251FFB6944440C4FC5532AC" authorityName="Smirnova. It" authorityYear="1987" box="[598,897,727,752]" class="Rhynchonellata" family="Norellidae" genus="Pseudomonticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="varia">
<emphasis id="FCB201376251FFB6944440C4FC5532AC" box="[598,897,727,752]" italics="true" pageId="9" pageNumber="510">Pseudomonticlarella varia</emphasis>
</taxonomicName>
are not co-occuring. Obviously they should cooccur if they are conspecific (although this could reflect a collecting failure). The specimens of
<taxonomicName id="09C6A6A66251FFB69685410CFE1F3374" authorityName="Smirnova. It" authorityYear="1987" box="[151,459,799,824]" class="Rhynchonellata" family="Norellidae" genus="Pseudomonticlarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="varia">
<emphasis id="FCB201376251FFB69685410CFE1F3374" box="[151,459,799,824]" italics="true" pageId="9" pageNumber="510">Pseudomonticlarella varia</emphasis>
</taxonomicName>
all occur with similar maximum dimensions and there are currently no other intermediate-sized forms between these and larger-sized species such as
<taxonomicName id="09C6A6A66251FFB695D64157FB35331E" authorityName="Sandy &amp; Hryniewicz &amp; Hammer &amp; Nakrem &amp; Little" authorityYear="2014" box="[964,1249,836,861]" class="Rhynchonellata" family="Rhynchonellidae" genus="Ptilorhynchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="9" pageNumber="510" phylum="Brachiopoda" rank="species" species="mclachlani">
<emphasis id="FCB201376251FFB695D64157FB35331E" box="[964,1249,836,861]" italics="true" pageId="9" pageNumber="510">Ptilorhynchia mclachlani</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CE79DD256251FFB696D54174FBB033E6" blockId="9.[151,1437,151,933]" pageId="9" pageNumber="510">
<emphasis id="FCB201376251FFB696D54174FD1C333C" box="[199,712,871,896]" inLineHeading="true" pageId="9" pageNumber="510" reason="1">
<emphasis id="FCB201376251FFB696D54174FD1533C3" bold="true" box="[199,705,871,896]" pageId="9" pageNumber="510">Stratigraphic and geographic distribution</emphasis>
.
</emphasis>
Upper VolgianLower (?Upper) Ryazanian of Spitsbergen and originally described from the Upper Volgian of the Russian Platform (
<bibRefCitation id="AA57A0D46251FFB695B4419FFB8C33E6" author="Smirnova, T. N." box="[934,1112,908,933]" pageId="9" pageNumber="510" pagination="39 - 49" refId="ref18461" refString="Smirnova, T. N. (1987) New taxa of late Volgian and early Hauterivian brachiopods of the Russian platform. Paleontological Journal, 21 (1), 39 - 49." type="journal article" year="1987">Smirnova 1987</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>