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<document ID-CLB-Dataset="297105" ID-DOI="10.5281/zenodo.7316535" ID-GBIF-Dataset="c7b986d2-d221-4c24-ae0a-037b5d4d8887" ID-ISBN="0-8018-8221-4" checkinTime="1670905166440" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="3B224005A4CB3B1F688393C52960B6C8" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Geomyidae.859-870.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press" docTitle="Thomomys (Megascapheus) townsendii" docType="treatment" docVersion="3" id="B149FDE7D3909CFBF93D8FFBC8EF1D48" lastPageNumber="869" masterDocId="B149FDE7D3909CFBF93D8FFBC8EF1D48" masterDocTitle="Order Rodentia - Family Geomyidae" masterLastPageNumber="870" masterPageNumber="859" pageNumber="869" updateTime="1716775291689" updateUser="ExternalLinkService">
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<mods:title id="F72A0FE98376AE586F773BB1DD27CC3F">Order Rodentia - Family Geomyidae</mods:title>
</mods:titleInfo>
<mods:name id="E8D6AC02AF068A1DCFC1C6FD1DACE152" type="personal">
<mods:role id="B295C468DBE8E02E0F165D08F5BDDAA5">
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<mods:namePart id="D01624259E5614AB7DAD48E663AEA2FE">Wilson, Don E.</mods:namePart>
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<mods:namePart id="BFBC2A5B30FF7FB678A275EED1D511D4">Reeder, DeeAnn</mods:namePart>
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<mods:typeOfResource id="397994C6B8BD6FD896CAD0A5B813EBE0">text</mods:typeOfResource>
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<mods:dateIssued id="DDB6E5D47C54C5AA7FE7953E324F1DFE">2005</mods:dateIssued>
<mods:publisher id="BEEE2F27304A071757C6AC86D14A3CC7">The Johns Hopkins University Press</mods:publisher>
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<mods:titleInfo id="501A583C06FBB7FE810DC7C6AC8F3FA1">
<mods:title id="4310ACFB2ADCB4FC7E8BE8436328F30C">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2</mods:title>
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<mods:identifier id="975B714980956BA4FD900431DC7E0CD2" type="DOI">10.5281/zenodo.7316535</mods:identifier>
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<treatment id="3B224005A4CB3B1F688393C52960B6C8" ID-GBIF-Taxon="231543085" LSID="urn:lsid:plazi:treatment:3B224005A4CB3B1F688393C52960B6C8" httpUri="http://treatment.plazi.org/id/3B224005A4CB3B1F688393C52960B6C8" lastPageNumber="869" pageNumber="869" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=12800373">
<subSubSection id="7E94C70DC1C2673D6278436D70F81412" pageNumber="869" type="nomenclature">
<paragraph id="49FF818ECCF1EA707AB0985210F49131" pageNumber="869">
<taxonomicName id="7D14B9BE398D5A9B47AB2258112EC967" ID-CoL="7X2SP" authority="(Bachman 1839)" baseAuthorityName="Bachman" baseAuthorityYear="1839" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="869" phylum="Chordata" rank="species" species="townsendii" subGenus="Megascapheus">
<emphasis id="827D839993D0834665D08008BF71DCAF" italics="true" pageNumber="869">Thomomys (Megascapheus) townsendii</emphasis>
(Bachman 1839)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="0813FE2C6F19ACD86F797F87EA3A490B" pageNumber="869" type="reference_group">
<paragraph id="80F71A73E2F830D4238DAA7524B008D5" pageNumber="869">
<treatmentCitation id="48B1E21A1C50D585CBB775224903F9B4" author="null" authority="(Bachman 1839)" baseAuthorityName="Bachman" baseAuthorityYear="1839" class="Mammalia" family="Geomyidae" genus="Thomomys" journal="J. Acad. Nat. Sci. Philadelphia" kingdom="Animalia" order="Rodentia" page="105" pageNumber="869" phylum="Chordata" rank="species" species="townsendii" subGenus="Megascapheus" title="Thomomys (Megascapheus) townsendii" volume="8" year="-1">
<taxonomicName id="3008850621E0BB9C46D67E4C0AE36CD2" ID-CoL="7X2SP" authority="(Bachman 1839)" baseAuthorityName="Bachman" baseAuthorityYear="1839" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="869" phylum="Chordata" rank="species" species="townsendii" subGenus="Megascapheus">
<emphasis id="8437906FD01E4D4B26E7C922630FC7D3" italics="true" pageNumber="869">Thomomys (Megascapheus) townsendii</emphasis>
(Bachman 1839)
</taxonomicName>
,
<bibCitation id="5CDA4C443A00B1F26BC157AFC8E6309D" author="null" journal="J. Acad. Nat. Sci. Philadelphia" pageNumber="869" pagination="105" title="Thomomys (Megascapheus) townsendii" volume="8" year="-1">J. Acad. Nat. Sci. Philadelphia, 8: 105</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="612F0D556DC3C745294E02C4926A1A78" pageNumber="869" type="materials_examined">
<paragraph id="C4FB16DA5423802265AE18A72C071E6E" pageNumber="869">
<emphasis id="D134A07F836448424BA1FD75C8BE8ECE" inLineHeading="true" pageNumber="869">Type Locality:</emphasis>
<materialsCitation id="2B1D09BA1D7E383333E5CE3799281C8D" ID-GBIF-Occurrence="4867332324" country="United States of America" pageNumber="869" stateProvince="Idaho">
<collectingCountry id="4809343CEB36F8C5748054199666206A" name="United States of America">USA</collectingCountry>
, erroneously given as &quot;Columbia River&quot;, but restricted to
<collectingRegion id="2D87924D88CC8EBFC1D252F45A8FED3A" country="United States of America" name="Idaho">Idaho</collectingRegion>
, Canyon Co., near Nampa, by Bailey (1915).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7AE6051E0E8052AA275422AD484D24F5" pageNumber="869" type="vernacular_names">
<paragraph id="575FA6F18802E36246882193475FC973" pageNumber="869">
<emphasis id="BC99BD4890D22502A7DD0A79CF9194AA" inLineHeading="true" pageNumber="869">Vernacular Names:</emphasis>
<vernacularName id="7B847B80823DC8CBA915EF616795F911" language="eng" pageNumber="869">Townsend's Pocket Gopher</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="00CE4DD3568ED7FC660A97C1287A66CE" pageNumber="869" type="child_taxa">
<paragraph id="89B29D53BEAFAC2D021B6DAEF3F80F97" pageNumber="869">
<emphasis id="5D6B67A40F076921B653E5344C59B0B7" inLineHeading="true" pageNumber="869">Subspecies:</emphasis>
:
</paragraph>
<paragraph id="EA6DD12535FE77B9226C05258497E496" pageNumber="869">
Subspecies
<taxonomicName id="E7FA23561F51E9FFB0B122C955D95C9D" authority="(Bachman 1839)" baseAuthorityName="Bachman" baseAuthorityYear="1839" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="869" phylum="Chordata" rank="subSpecies" species="townsendii" subGenus="Megascapheus" subSpecies="townsendii">
<emphasis id="EAD33D65E5143DF50CD9126ABA1CEB72" italics="true" pageNumber="869">Thomomys (Megascapheus) townsendii</emphasis>
subsp.
<emphasis id="DF2EB50221BBF443CC766D0276ECB844" italics="true" pageNumber="869">townsendii</emphasis>
(Bachman 1839)
</taxonomicName>
</paragraph>
<paragraph id="7C28CF00881CC0BD9F6F04B10FAB9E27" pageNumber="869">
Subspecies
<taxonomicName id="057C8FB599124AA3B403A08C855FC4E9" authority="(Merriam 1897)" baseAuthorityName="Merriam" baseAuthorityYear="1897" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="869" phylum="Chordata" rank="subSpecies" species="townsendii" subGenus="Megascapheus" subSpecies="nevadensis">
<emphasis id="1D99B52B5365EA4A47388E8D24E86766" italics="true" pageNumber="869">Thomomys (Megascapheus) townsendii</emphasis>
subsp.
<emphasis id="3DFF8F15D11FC0D145A1BBDF95AC50F6" italics="true" pageNumber="869">nevadensis</emphasis>
(
<bibRefCitation id="165DF72A0D368A39957685DA9D28FF3E" author="Merriam, C. H." refId="ref238343" refString="Merriam, C. H. 1897. The generic names Ictis, Arctogale, and Arctogalidia. Science, 5: 302." year="1897">Merriam 1897</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="0949991BEB2DE5576782DDEAD11E8903" pageNumber="869" type="distribution">
<paragraph id="E4947F99F6E251FAF46FB8C3718A0F63" pageNumber="869">
<emphasis id="FDA476B11B45C1555DA61041908BBBB8" inLineHeading="true" pageNumber="869">Distribution:</emphasis>
Snake River Valley of
<collectingRegion id="028B398760A15794CB17175A3BE7EA48" country="United States of America" name="Idaho">Idaho</collectingRegion>
south and west to SE
<collectingRegion id="A475AFE9F16ADEEFFA27C0E9201D6CB5" country="United States of America" name="Oregon">Oregon</collectingRegion>
, NE
<collectingRegion id="0117EC15CFD8833C23219E79B60D19B3" country="United States of America" name="California">California</collectingRegion>
, and N
<collectingRegion id="94C62093BA06EFC1FABBA82DE87F12E6" country="United States of America" name="Nevada">Nevada</collectingRegion>
(
<collectingCountry id="CA084B42774D1E7CAF2CFD0C54EECBFC" name="United States of America">USA</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="73A1107A3A4589B52F4B63B29205F826" pageNumber="869" type="conservation">
<paragraph id="3BF43B1E0FA7CFBA4A5957FC1CD1853B" pageNumber="869">
<emphasis id="658E2FC54D559C549CC3FDB7A0FE58A7" inLineHeading="true" pageNumber="869">Conservation:</emphasis>
<collectionCode id="A30EE0F4D547499D1E9DE24C8F804FB1">IUCN</collectionCode>
Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="95348E760EE4275609798815418981A9" pageNumber="869" type="discussion">
<paragraph id="95917E63987AD512F63A1898F5F0BF36" pageNumber="869">
<emphasis id="9E356D3AF77DFACC0169FAE03DFC1D47" inLineHeading="true" pageNumber="869">Discussion:</emphasis>
Subgenus
<taxonomicName id="11CF8B4190C6F6CF239493B82CFDE2A2" authorityName="Elliot" authorityYear="1903" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="subGenus" subGenus="Megascapheus">
<emphasis id="84C6F378C59C7AE8B6610B23D0A47FDC" italics="true" pageNumber="869">Megascapheus</emphasis>
</taxonomicName>
. Revised by
<bibRefCitation id="B072B1B5097BEF62446D440C06779BC7" author="Davis, W. B." refId="ref74963" refString="Davis, W. B. 1937. Variations in Townsend pocket gophers. Journal of Mammalogy, 18: 145 - 158." year="1937">Davis (1937)</bibRefCitation>
and
<bibRefCitation id="2F1CD512BB96007D90B995A0AA861735" author="Rogers, M. A." refId="ref309374" refString="Rogers, M. A. 1991 a. Evolutionary differentiation within the northern Great Basin pocket gopher, Thomomys townsendii. I. Morphological variation. Great Basin Naturalist, 51: 109 - 126." year="1991">
Rogers (1991
<emphasis id="63F776D60121AF17B2E4091044FD7B93" italics="true" pageNumber="869">a</emphasis>
</bibRefCitation>
,
<emphasis id="64175F40A7B0E40194B24829D2773CF0" italics="true" pageNumber="869">b</emphasis>
), who recognized only two valid subspecies (
<taxonomicName id="A386E4D5D43DEE1CB6C83E376E337F7E" baseAuthorityName="Bachman" baseAuthorityYear="1839" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="3951161082C6DF1F325E85EAEBED88A8" italics="true" pageNumber="869">townsendii</emphasis>
</taxonomicName>
and
<emphasis id="9BE092545FAC84BD16F0AD2DA74F108E" italics="true" pageNumber="869">nevadensis</emphasis>
). Considered a distinct species by
<bibRefCitation id="D9A46121D079BF2AE047553BF04C46DE" author="Thaeler, C. S., Jr." refId="ref350523" refString="Thaeler, C. S., Jr. 1968 b. An analysis of three hybrid populations of pocket gophers (genus Thomomys). Evolution, 22: 543 - 555." year="1968">
Thaeler (1968
<emphasis id="76DAE872BEB1B96CF3E6F5B499DB71FF" italics="true" pageNumber="869">b</emphasis>
)
</bibRefCitation>
,
<bibRefCitation id="F83ADA235971392A62E75003844332E7" author="Patton, J. L. &amp; M. F. Smith &amp; R. D. Price &amp; R. A. Hellenthal" refId="ref276497" refString="Patton, J. L., M. F. Smith, R. D. Price, and R. A. Hellenthal. 1984. Genetics of hybridization between the pocket gophers Thomomys bottae and Thomomys townsendii in northeastern California. Great Basin Naturalist, 44: 431 - 440." year="1984">Patton et al. (1984)</bibRefCitation>
,
<bibRefCitation id="E51EC66609B386AA07B4F9C5C18E41D3" author="Patton, J. L. &amp; M. F. Smith" refId="ref276106" refString="Patton, J. L., and M. F. Smith. 1989. Genetic structure and the genetic and morphologic divergence among pocket gopher species (genus Thomomys). Pp. 284 - 304, in Speciation and Its Consequences (D. Otte and J. A. Endler, eds.). Sinauer Associates Incorporated, Sunderland, MA, 679 pp." year="1989">Patton and Smith (1989</bibRefCitation>
, 1994), and
<bibRefCitation id="14FB5FB2414A7A41507F23712F6DD770" author="Rogers, M. A." refId="ref309374" refString="Rogers, M. A. 1991 a. Evolutionary differentiation within the northern Great Basin pocket gopher, Thomomys townsendii. I. Morphological variation. Great Basin Naturalist, 51: 109 - 126." year="1991">
Rogers (1991
<emphasis id="CA592E6EDFD90749C1AA1DC90F3333DB" italics="true" pageNumber="869">a</emphasis>
</bibRefCitation>
,
<emphasis id="4616099EF55A64E79289735047CE0378" italics="true" pageNumber="869">b</emphasis>
), despite limited hybridization with
<taxonomicName id="6DCB6355259057733E8623428CA3AF58" baseAuthorityName="Eydoux and Gervais" baseAuthorityYear="1836" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="bottae">
<emphasis id="71C865982E8EB8C8B0C200C7B9824119" italics="true" pageNumber="869">bottae</emphasis>
</taxonomicName>
in NE
<collectingRegion id="C296FB980FB1BE35E137D50B77894D43" country="United States of America" name="California">California</collectingRegion>
.
<bibRefCitation id="5E9CDAB1D05A73A8FA6C70ADC65A5C51" author="Hall, E. R." refId="ref137310" refString="Hall, E. R. 1981. The mammals of North America. Second ed. John Wiley and Sons, New York, 1: 1 - 600 + 90, 2: 601 - 1181 + 90." year="1981">Hall (1981:469</bibRefCitation>
, 495) reviewed Thaeler's evidence and included
<taxonomicName id="FC54604DD5BCCD2C356B71552D1AD9E3" baseAuthorityName="Bachman" baseAuthorityYear="1839" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="D0E1279E403B59AC7C3E7A0E2331A5C2" italics="true" pageNumber="869">townsendii</emphasis>
</taxonomicName>
in
<taxonomicName id="811A432C414A77DCE5704D479DC45D01" authority="(sensu Hall)" baseAuthorityName="Hall" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="umbrinus">
<emphasis id="CF6D2CD9094ED919A0FB73EDA2D36D99" italics="true" pageNumber="869">umbrinus</emphasis>
(
<emphasis id="6E2D8526F579DAF8AB5208B5E363FB0D" italics="true" pageNumber="869">sensu</emphasis>
Hall)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>