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<document id="280DB7B94BF881F7D5B413DACCCD2160" ID-CLB-Dataset="39003" ID-DOI="10.11646/zootaxa.4114.4.1" ID-GBIF-Dataset="47034122-1fc6-4b05-9cb8-0bf9749ef4b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="260588" ID-ZooBank="FBE39B58-678B-495F-8050-E24F89F102BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464157130772" checkinUser="plazi" docAuthor="Travouillon, Kenny J." docDate="2016" docId="1B695A18FB6E0F43FF3BA795FA2460CC" docLanguage="en" docName="zootaxa.4114.4.1.pdf.imf" docOrigin="Zootaxa 4114 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Perameles pallescens Thomas 1923, stat. nov." docType="treatment" docVersion="9" lastPageNumber="376" masterDocId="E7502260FB760F5AFFACA622FFBE6034" masterDocTitle="Investigating dental variation in Perameles nasuta Geoffroy, 1804, with morphological evidence to raise P. nasuta pallescens Thomas, 1923 to species rank" masterLastPageNumber="392" masterPageNumber="351" pageNumber="375" updateTime="1698662609867" updateUser="ExternalLinkService">
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<mods:title id="9DCFA84ECFE69405E6C20AFA38D59F52">Investigating dental variation in Perameles nasuta Geoffroy, 1804, with morphological evidence to raise P. nasuta pallescens Thomas, 1923 to species rank</mods:title>
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<mods:namePart id="CEFF61A21A0613323A394BD55F64F3A7">Travouillon, Kenny J.</mods:namePart>
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<mods:date id="DF2464185709FD83B2AA70ECAC082E4B">2016</mods:date>
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<treatment id="1B695A18FB6E0F43FF3BA795FA2460CC" ID-DOI="http://doi.org/10.5281/zenodo.6087197" ID-GBIF-Taxon="120701245" ID-Zenodo-Dep="6087197" LSID="urn:lsid:plazi:treatment:1B695A18FB6E0F43FF3BA795FA2460CC" httpUri="http://treatment.plazi.org/id/1B695A18FB6E0F43FF3BA795FA2460CC" lastPageId="25" lastPageNumber="376" pageId="24" pageNumber="375">
<subSubSection id="DBDAB885FB6E0F42FF3BA795FEA261C7" pageId="24" pageNumber="375" type="nomenclature">
<paragraph id="937FEB0EFB6E0F42FF3BA795FD7D61E6" blockId="24.[151,707,439,499]" box="[151,707,439,466]" pageId="24" pageNumber="375">
<heading id="C8375C62FB6E0F42FF3BA795FD7D61E6" bold="true" box="[151,707,439,466]" fontSize="11" level="1" pageId="24" pageNumber="375" reason="1">
<emphasis id="A1B4371CFB6E0F42FF3BA795FD7D61E6" bold="true" box="[151,707,439,466]" pageId="24" pageNumber="375">
<taxonomicName id="54C0908DFB6E0F42FF3BA795FDF361E6" authority="Thomas, 1923" authorityName="Thomas" authorityYear="1923" box="[151,589,439,466]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="pallescens" status="stat. nov.">
<emphasis id="A1B4371CFB6E0F42FF3BA795FE2661E5" bold="true" box="[151,408,439,465]" italics="true" pageId="24" pageNumber="375">Perameles pallescens</emphasis>
<bibRefCitation id="F75196FFFB6E0F42FE32A79AFDF361E6" author="Thomas" box="[414,589,440,466]" pageId="24" pageNumber="375" refString="Thomas, O. (1923) The Godman Exploration Fund: List of mammals from north Queensland collected by Mr T. V. Sherrin." type="book" year="1923">Thomas, 1923</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="BA878A67FB6E0F42FDF8A79AFD7D61E6" box="[596,707,440,466]" pageId="24" pageNumber="375">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="937FEB0EFB6E0F42FF3BA7F8FEA261C7" blockId="24.[151,707,439,499]" box="[151,284,474,499]" pageId="24" pageNumber="375">
<figureCitation id="0BFBF78BFB6E0F42FF3BA7F8FF5A61C7" box="[151,228,474,499]" captionStart="FIGURE 3" captionStartId="8.[151,250,1756,1778]" captionTargetBox="[156,1430,193,1733]" captionTargetId="figure@8.[156,1431,193,1735]" captionTargetPageId="8" captionText="FIGURE 3. Dorsal and ventral views of the skins of Perameles nasuta pallescens, represented by the holotype (A, B; adult male; BMNH 22.12.18.40) and all other specimens present in the Queensland Museum collections (C, D). Arrow points to bar pattern in the pelage. Scale bar = 10 cm." httpUri="https://zenodo.org/record/260591/files/figure.png" pageId="24" pageNumber="375">Figs. 3</figureCitation>
,
<figureCitation id="0BFBF78BFB6E0F42FF5DA7F8FEA261C7" box="[241,284,474,499]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="10.[151,250,1643,1665]" captionStartId-1="11.[151,250,1923,1945]" captionStartId-2="12.[151,250,1191,1213]" captionStartId-3="16.[151,250,1907,1929]" captionTargetBox-0="[151,1436,193,1604]" captionTargetBox-1="[170,1398,203,1876]" captionTargetBox-2="[155,1425,181,1161]" captionTargetBox-3="[196,1370,207,1845]" captionTargetId-0="figure@10.[151,1436,193,1605]" captionTargetId-1="figure@11.[161,1412,193,1891]" captionTargetId-2="figure@12.[155,1432,175,1170]" captionTargetId-3="figure@16.[180,1386,193,1857]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="16" captionText-0="FIGURE 5. Lateral views of the skins of Perameles nasuta pallescens (A, topotype, QM JM 18575), P. n. nasuta (B, QM J 11236), P. gunnii (C, QM JM 16110) and P. bougainville (D, QM J 11207). Arrow points to bar pattern in the pelage. Scale bar = 10 cm." captionText-1="FIGURE 6. Lateral, dorsal and ventral views of the skull and dentary of Perameles nasuta pallescens (A C, holotype, BMNH 22.12. 18.40, male; D F, topotype, mirrored, QM JM 18575, male) and P. n. nasuta (G I, QM JM 20178, female). Arrow points to accessory cusp on canine. Scale bar = 10 mm." captionText-2="FIGURE 7. Occlusal and lingual views of the upper third premolar and molars of Perameles nasuta pallescens (A B, topotype, QM JM 18575) and P. n. nasuta (C D, QM JM 20178; E F, mirrored image, QM J 4906). Arrows point to morphological differences mentioned in the text. Scale bar = 3 mm." captionText-3="FIGURE 8. Occlusal and buccal views of the lower third premolar and molars of Perameles nasuta pallescens (A B, topotype, QM JM 18575; G, magnesium oxide coated m 1, J 6350) and P. n. nasuta (C D, mirrored image, QM JM 20178; E F, QM J 4906; H, magnesium oxide coated m 1, QM J 4906). Arrows point to morphological differences mentioned in the text. Scale bar = 3 mm." httpUri-0="https://zenodo.org/record/260593/files/figure.png" httpUri-1="https://zenodo.org/record/260594/files/figure.png" httpUri-2="https://zenodo.org/record/260595/files/figure.png" httpUri-3="https://zenodo.org/record/260596/files/figure.png" pageId="24" pageNumber="375">58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DBDAB885FB6E0F42FF3BA400FD38620C" box="[151,646,546,568]" pageId="24" pageNumber="375" type="reference_group">
<paragraph id="937FEB0EFB6E0F42FF3BA400FD38620C" blockId="24.[151,646,546,568]" box="[151,646,546,568]" pageId="24" pageNumber="375">
<treatmentCitationGroup id="B3D0CC20FB6E0F42FF3BA400FD38620C" box="[151,646,546,568]" pageId="24" pageNumber="375">
<taxonomicName id="54C0908DFB6E0F42FF3BA400FD38620C" authority="Thomas, 1923: 173" authorityName="Thomas" authorityPageNumber="173" authorityYear="1923" box="[151,646,546,568]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="subSpecies" species="nasuta" subSpecies="pallescens">
<emphasis id="A1B4371CFB6E0F42FF3BA400FE05620C" box="[151,443,546,568]" italics="true" pageId="24" pageNumber="375">Perameles nasuta pallescens</emphasis>
<treatmentCitation id="1261CD1FFB6E0F42FE6EA400FD38620C" author="Thomas" box="[450,646,546,568]" page="173" pageId="24" pageNumber="375" year="1923">
<bibRefCitation id="F75196FFFB6E0F42FE6EA400FDE8620C" author="Thomas" box="[450,598,546,568]" pageId="24" pageNumber="375" refString="Thomas, O. (1923) The Godman Exploration Fund: List of mammals from north Queensland collected by Mr T. V. Sherrin." type="book" year="1923">Thomas, 1923</bibRefCitation>
: 173
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DBDAB885FB6E0F42FF3BA447FD56634E" pageId="24" pageNumber="375" type="materials_examined">
<paragraph id="937FEB0EFB6E0F42FF3BA447FE266296" blockId="24.[151,1437,613,2006]" pageId="24" pageNumber="375">
<emphasis id="A1B4371CFB6E0F42FF3BA447FEBA624A" bold="true" box="[151,260,613,638]" pageId="24" pageNumber="375">
<typeStatus id="4C7B55ACFB6E0F42FF3BA447FEBA624A" box="[151,260,613,638]" pageId="24" pageNumber="375" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectionCode id="F5D173CBFB6E0F42FEB7A444FED16249" box="[283,367,614,637]" pageId="24" pageNumber="375">BMNH</collectionCode>
22.12.18.40, preserved skin and skeleton of adult male from Vine Creek, Ravenshoe, Queensland,
<collectingCountry id="EBD7AB9EFB6E0F42FE85A4ABFE2D6296" box="[297,403,649,674]" name="Australia" pageId="24" pageNumber="375">Australia</collectingCountry>
.
</paragraph>
<paragraph id="937FEB0EFB6E0F42FF6BA48FFE2662DE" blockId="24.[151,1437,613,2006]" pageId="24" pageNumber="375">
<emphasis id="A1B4371CFB6E0F42FF6BA48FFE8A62F2" bold="true" box="[199,308,685,710]" pageId="24" pageNumber="375">Topotype</emphasis>
. QM JM18575, preserved skin and skeleton of adult male from Mt Father Clancy, Ravenshoe, Queensland,
<collectingCountry id="EBD7AB9EFB6E0F42FE85A4F3FE2D62DE" box="[297,403,721,746]" name="Australia" pageId="24" pageNumber="375">Australia</collectingCountry>
.
</paragraph>
<paragraph id="937FEB0EFB6E0F42FF6BA4D7FD56634E" blockId="24.[151,1437,613,2006]" pageId="24" pageNumber="375">
<emphasis id="A1B4371CFB6E0F42FF6BA4D7FE42633A" bold="true" box="[199,508,757,782]" pageId="24" pageNumber="375">Other material examined</emphasis>
. From the Museum of Natural History, London:
<collectionCode id="F5D173CBFB6E0F42FB91A4D4FB2F6339" box="[1085,1169,758,781]" pageId="24" pageNumber="375">BMNH</collectionCode>
22.12.18.41. From the Australian Museum: AM M15841. From the Queensland Museum: QM J6350, J9421,
<accessionNumber id="8C9376EDFB6E0F42FBD4A53BFB756306" box="[1144,1227,793,818]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J10816" pageId="24" pageNumber="375" type="EnaNcbi">J10816</accessionNumber>
,
<accessionNumber id="8C9376EDFB6E0F42FB75A538FA926306" box="[1241,1324,794,818]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J14308" pageId="24" pageNumber="375" type="EnaNcbi">J14308</accessionNumber>
, JM6561, JM6683, JM11389, JM18575, JM20172, JM20173, JM20174, JM20239, JM20240, JM20241, JM20242, JM20243, JM20244, JM20245, JM20246, JM20247.
</paragraph>
</subSubSection>
<subSubSection id="DBDAB885FB6E0F42FF6BA5A7FE7565A2" pageId="24" pageNumber="375" type="diagnosis">
<paragraph id="937FEB0EFB6E0F42FF6BA5A7FBDA6442" blockId="24.[151,1437,613,2006]" pageId="24" pageNumber="375">
<emphasis id="A1B4371CFB6E0F42FF6BA5A7FE8463AA" bold="true" box="[199,314,901,926]" pageId="24" pageNumber="375">Diagnosis</emphasis>
.
<taxonomicName id="54C0908DFB6E0F42FEE6A5A7FD8063A9" box="[330,574,901,925]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="A1B4371CFB6E0F42FEE6A5A7FD8063A9" box="[330,574,901,925]" italics="true" pageId="24" pageNumber="375">Perameles pallescens</emphasis>
</taxonomicName>
differs from
<taxonomicName id="54C0908DFB6E0F42FD7BA5A4FC8163A9" box="[727,831,902,925]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="A1B4371CFB6E0F42FD7BA5A4FC8163A9" box="[727,831,902,925]" italics="true" pageId="24" pageNumber="375">P. nasuta</emphasis>
</taxonomicName>
in having the following features: sagittal crest either very small or absent in males; smaller and narrower posterolingual cusp and shelf on P3; StA larger and bladed, ascending the anterior flank of preparacrista on M1; StB shorter than StC on M1; metaconule smaller on M13, with postmetaconule crista ending more lingually; StA more lingually positioned on M3, resulting in a less concave buccal margin of the tooth; StB either absent or present as small conical cusp on M4; protocone smaller on M4 with poorly developed preprotocrista and postprotocrista; postparacrista not connected to metacone on M4; a poorly developed paracristid on m1, not connected to the protoconid; a smaller talonid on m4.
</paragraph>
<paragraph id="937FEB0EFB6E0F42FF6BA2A0FE7565A2" blockId="24.[151,1437,613,2006]" pageId="24" pageNumber="375">
<taxonomicName id="54C0908DFB6E0F42FF6BA2A0FEE664AD" box="[199,344,1153,1177]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="A1B4371CFB6E0F42FF6BA2A0FEE664AD" box="[199,344,1153,1177]" italics="true" pageId="24" pageNumber="375">P. pallescens</emphasis>
</taxonomicName>
differs from
<taxonomicName id="54C0908DFB6E0F42FE42A2A0FDEC64AD" box="[494,594,1154,1177]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="gunnii">
<emphasis id="A1B4371CFB6E0F42FE42A2A0FDEC64AD" box="[494,594,1154,1177]" italics="true" pageId="24" pageNumber="375">P. gunnii</emphasis>
</taxonomicName>
in having the following features: fewer bars in its pelage on its rump; less well-developed orbital rim; no ridges on the parietal; smaller lambdoidal sesamoid; smaller bullae; no accessory palatal fenestrae; smaller incisive and maxillopalatine fenestrae; shallower antorbital fossa;
<date id="E77ECDCEFB6E0F42FB34A2EBFB7264D6" box="[1176,1228,1225,1250]" pageId="24" pageNumber="375">I45</date>
not premolar-like; obvious sexual dimorphism in canines; main cusp of P3 less conical, and less well-developed lingual shelf; metaconule less well-developed on M3; StA larger and bladed, ascending the anterior flank of preparacrista on M1; StD connected to metaconule by a distinct crest on M1; minute anterior cingulum present on M2; less welldeveloped StB and protocone on M4; a poorly developed paracristid on m1, not connected to the protoconid; trigonid of m4 much wider.
</paragraph>
</subSubSection>
<subSubSection id="DBDAB885FB6E0F42FF6BA380FE1E675E" pageId="24" pageNumber="375" type="description">
<paragraph id="937FEB0EFB6E0F42FF6BA380FEA6665A" blockId="24.[151,1437,613,2006]" pageId="24" pageNumber="375">
<taxonomicName id="54C0908DFB6E0F42FF6BA380FEE5658D" box="[199,347,1441,1465]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="A1B4371CFB6E0F42FF6BA380FEE5658D" box="[199,347,1441,1465]" italics="true" pageId="24" pageNumber="375">P. pallescens</emphasis>
</taxonomicName>
differs from
<taxonomicName id="54C0908DFB6E0F42FE55A380FD1D658D" box="[505,675,1441,1465]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="bougainville">
<emphasis id="A1B4371CFB6E0F42FE55A380FD1D658D" box="[505,675,1441,1465]" italics="true" pageId="24" pageNumber="375">P. bougainville</emphasis>
</taxonomicName>
in the following features: larger in overall size; fewer bars in its pelage on its rump; no accessory palatal or palatine fenestrae; smaller maxillopalatine fenestrae; taller postglenoid process; shallower antorbital fossa; lingual cusp and lingual shelf on P3 slightly larger in size; StA larger and bladed, ascending the anterior flank of preparacrista on M1; StD connected to metaconule by a distinct crest on M1; minute anterior cingulum present on M2; a poorly developed paracristid on m1, not connected to the protoconid.
</paragraph>
<paragraph id="937FEB0EFB6E0F42FF6BA058FEA66682" blockId="24.[151,1437,613,2006]" pageId="24" pageNumber="375">
<taxonomicName id="54C0908DFB6E0F42FF6BA058FEE366A5" box="[199,349,1657,1681]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="A1B4371CFB6E0F42FF6BA058FEE366A5" box="[199,349,1657,1681]" italics="true" pageId="24" pageNumber="375">P. pallescens</emphasis>
</taxonomicName>
differs from
<taxonomicName id="54C0908DFB6E0F42FDAFA058FDD366A5" box="[515,621,1657,1681]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="sobbei">
<emphasis id="A1B4371CFB6E0F42FDAFA058FDD366A5" box="[515,621,1657,1681]" italics="true" pageId="24" pageNumber="375">P. sobbei</emphasis>
</taxonomicName>
having a poorly developed paracristid on m1, not connected to the protoconid.
</paragraph>
<paragraph id="937FEB0EFB6E0F42FF6BA0E0FBA46716" blockId="24.[151,1437,613,2006]" pageId="24" pageNumber="375">
<taxonomicName id="54C0908DFB6E0F42FF6BA0E0FEE666ED" box="[199,344,1729,1753]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="A1B4371CFB6E0F42FF6BA0E0FEE666ED" box="[199,344,1729,1753]" italics="true" pageId="24" pageNumber="375">P. pallescens</emphasis>
</taxonomicName>
differs from
<taxonomicName id="54C0908DFB6E0F42FE42A0E0FDCF66ED" box="[494,625,1729,1753]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="bowensis">
<emphasis id="A1B4371CFB6E0F42FE42A0E0FDCF66ED" box="[494,625,1729,1753]" italics="true" pageId="24" pageNumber="375">P. bowensis</emphasis>
</taxonomicName>
in the following features: all dental dimensions larger; main cusp of P3 less conical and narrower; anterior cingulum on M2 much smaller; anterior cingulum on M3 much smaller (if present); a poorly developed paracristid on m1, not connected to the protoconid.
</paragraph>
<paragraph id="937FEB0EFB6E0F42FF6BA10CFE1E675E" blockId="24.[151,1437,613,2006]" pageId="24" pageNumber="375">
<taxonomicName id="54C0908DFB6E0F42FF6BA10CFEE36771" box="[199,349,1837,1861]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="pallescens">
<emphasis id="A1B4371CFB6E0F42FF6BA10CFEE36771" box="[199,349,1837,1861]" italics="true" pageId="24" pageNumber="375">P. pallescens</emphasis>
</taxonomicName>
differs from
<taxonomicName id="54C0908DFB6E0F42FDAEA10CFD736771" box="[514,717,1837,1861]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="24" pageNumber="375" phylum="Chordata" rank="species" species="allinghamensis">
<emphasis id="A1B4371CFB6E0F42FDAEA10CFD736771" box="[514,717,1837,1861]" italics="true" pageId="24" pageNumber="375">P. allinghamensis</emphasis>
</taxonomicName>
in a much smaller anterior cingulum (when present); and a buccally displaced StA.
</paragraph>
</subSubSection>
<subSubSection id="DBDAB885FB6E0F43FF6BA157FC7060E1" lastPageId="25" lastPageNumber="376" pageId="24" pageNumber="375" type="discussion">
<paragraph id="937FEB0EFB6E0F42FF6BA157FE1A67E1" blockId="24.[151,1437,613,2006]" pageId="24" pageNumber="375">
<emphasis id="A1B4371CFB6E0F42FF6BA157FE8A67BA" bold="true" box="[199,308,1909,1934]" pageId="24" pageNumber="375">Remarks</emphasis>
. A new topotype is established here, to facilitate research and specimen identification in
<collectingCountry id="EBD7AB9EFB6E0F42FA99A157FA2267BA" box="[1333,1436,1909,1934]" name="Australia" pageId="24" pageNumber="375">Australia</collectingCountry>
(from a Queensland Museum specimen), considering that the
<typeStatus id="4C7B55ACFB6E0F42FCE0A1BBFC106786" box="[844,942,1945,1970]" pageId="24" pageNumber="375" type="holotype">holotype</typeStatus>
is located in the British Museum of Natural History in London,
<collectingCountry id="EBD7AB9EFB6E0F42FED8A19CFE1E67E1" box="[372,416,1982,2005]" name="United Kingdom" pageId="24" pageNumber="375">UK</collectingCountry>
.
</paragraph>
<paragraph id="937FEB0EFB6F0F43FF6BA6B5FC7060E1" blockId="25.[151,1436,151,248]" pageId="25" pageNumber="376">
<emphasis id="A1B4371CFB6F0F43FF6BA6B5FE9D6084" bold="true" box="[199,291,151,176]" pageId="25" pageNumber="376">Habitat</emphasis>
. Rainforest, wet sclerophyll forests, woodlands, swamps and farmlands (
<bibRefCitation id="F75196FFFB6F0F43FBFAA6B5FB426084" author="Harrison" box="[1110,1276,151,176]" pageId="25" pageNumber="376" refString="Harrison, J. L. (1962) Mammals of Innisfail. 1. Species and distribution. Australian Journal of Zoology, 10 (1), 45 - 83. http: // dx. doi. org / 10.1071 / zo 9620045" type="journal article" year="1962">Harrison 1962</bibRefCitation>
;
<bibRefCitation id="F75196FFFB6F0F43FAABA6B5FF6D60E0" author="Gordon" pageId="25" pageNumber="376" refString="Gordon, G., Hall, L. S. &amp; Atherton, R. G. (1990) Status of bandicoots in Queensland. In: Seebeck, J. H., Wallis, R. L., Brown, P. R. &amp; Kemper, C. M. (Eds.), Bandicoots and bilbies. Surrey Beatty and Sons, Sydney, pp. 37 - 42." type="book chapter" year="1990">
Gordon
<emphasis id="A1B4371CFB6F0F43FAC8A6BBFA226084" box="[1380,1436,152,176]" italics="true" pageId="25" pageNumber="376">et al.</emphasis>
1990
</bibRefCitation>
;
<bibRefCitation id="F75196FFFB6F0F43FF71A69EFE4F60E0" author="Williams" box="[221,497,188,213]" pageId="25" pageNumber="376" refString="Williams, S. E. &amp; Marsh, H. (1998) Changes in small mammal assemblage structure across a rain forest / open forest ecotone. Journal of Tropical Ecology, 14, 187 - 198. http: // dx. doi. org / 10.1017 / s 0266467498000157" type="journal article" year="1998">Williams &amp; Marsh 1998</bibRefCitation>
;
<bibRefCitation id="F75196FFFB6F0F43FE50A69FFD1C60E0" author="Bateman" box="[508,674,188,212]" pageId="25" pageNumber="376" refString="Bateman, B. (2010) Beyond simple means: integrating extreme events and biotic interactions in species distribution models. Conservation implications for the Northern Bettong (Bettongia tropica) under climate change. PhD Thesis, James Cook University, Townsville, 225 pp." type="book" year="2010">Bateman 2010</bibRefCitation>
;
<bibRefCitation id="F75196FFFB6F0F43FD02A69EFC1260E0" author="Kanowski" box="[686,940,188,213]" pageId="25" pageNumber="376" refString="Kanowski, J., Middleton, J., Jensen, R. &amp; Legge, S. (2010 a) Camera trap survey of Mount Zero - Taravale Sanctuary, 2008 - 2009. Australian Wildlife Conservancy, Subiaco, 4 pp." type="book chapter" year="2010" yearSuffix="a">
Kanowski
<emphasis id="A1B4371CFB6F0F43FC8AA69FFCE060E0" box="[806,862,188,212]" italics="true" pageId="25" pageNumber="376">et al.</emphasis>
2010a
</bibRefCitation>
,
<bibRefCitation id="F75196FFFB6F0F43FC1BA69EFC7860E1" author="Kanowski" box="[951,966,188,213]" pageId="25" pageNumber="376" refString="Kanowski, J., Mulder, E. &amp; Jensen, R. (2010 b) Mount Zero Taravale Survey September 2010. Australian Wildlife Conservancy, Subiaco, 4 pp." type="book" year="2010" yearSuffix="b">b</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DBDAB885FB6F0F43FF6BA6FDFA2460CC" box="[199,1434,223,248]" pageId="25" pageNumber="376" type="distribution">
<paragraph id="937FEB0EFB6F0F43FF6BA6FDFA2460CC" blockId="25.[151,1436,151,248]" box="[199,1434,223,248]" pageId="25" pageNumber="376">
<emphasis id="A1B4371CFB6F0F43FF6BA6FDFEE760CC" bold="true" box="[199,345,223,248]" pageId="25" pageNumber="376">Distribution</emphasis>
. From the Townsville region to Cape York Peninsula, on the east coast of Queensland,
<collectingCountry id="EBD7AB9EFB6F0F43FA87A6FDFA2B60CC" box="[1323,1429,223,248]" name="Australia" pageId="25" pageNumber="376">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>