376 lines
43 KiB
XML
376 lines
43 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1040.66209" ID-GBIF-Dataset="48cd698d-e51b-4bb2-bd4d-1794c8c5955f" ID-PMC="PMC8178291" ID-Pensoft-Pub="1313-2970-1040-167" ID-Pensoft-UUID="1F71B6B55BCB516A825666EA04FC0431" ID-PubMed="34135659" ID-ZooBank="910903D2476A439FAC73292D5B9A18C1" ModsDocID="1313-2970-1040-167" checkinTime="1622233190126" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Srisonchai, Ruttapon & Panha, Somsak" docDate="2021" docId="B4DFAF1674885A80AFDE56D716EED2C4" docLanguage="en" docName="ZooKeys 1040: 167-185" docOrigin="ZooKeys 1040" docPubDate="2021-05-28" docSource="http://dx.doi.org/10.3897/zookeys.1040.66209" docTitle="Tylopus sutchariti Likhitrakarn & Golovatch & Srisonchai & Panha 2021, sp. nov." docType="treatment" docUuid="5385A3A0-C129-41F3-9A8C-C3910EF9178C" docUuidSource="ZooBank" docVersion="4" id="1F71B6B55BCB516A825666EA04FC0431" lastPageNumber="167" masterDocId="1F71B6B55BCB516A825666EA04FC0431" masterDocTitle="Two new species of the millipede genus Tylopus Jeekel, 1968 from Shan State, Myanmar (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="185" masterPageNumber="167" pageNumber="167" updateTime="1668150329396" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of the millipede genus Tylopus Jeekel, 1968 from Shan State, Myanmar (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation>Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai, 50290, Thailand & Biodiversity and Utilization Research Center of Maejo University, Maejo University, Chiang Mai, 50290, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation>Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Srisonchai, Ruttapon</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7142-0999</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, 40002, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-05-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1040</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>167</mods:start>
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<mods:end>185</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1040.66209</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1040.66209</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1040-167</mods:identifier>
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<mods:identifier type="ZooBank">910903D2476A439FAC73292D5B9A18C1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">1F71B6B55BCB516A825666EA04FC0431</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182408860" LSID="urn:lsid:zoobank.org:act:5385A3A0-C129-41F3-9A8C-C3910EF9178C" httpUri="http://treatment.plazi.org/id/B4DFAF1674885A80AFDE56D716EED2C4" lastPageNumber="167" pageId="0" pageNumber="167">
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<subSubSection pageId="0" pageNumber="167" type="nomenclature">
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<paragraph pageId="0" pageNumber="167">
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<taxonomicName LSID="http://zoobank.org/5385A3A0-C129-41F3-9A8C-C3910EF9178C" authority="Likhitrakarn & Golovatch & Srisonchai & Panha, 2021" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus sutchariti" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="sutchariti" status="sp. nov.">Tylopus sutchariti</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="167">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">Figs 1B</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">, 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, sublateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure6" httpUri="https://binary.pensoft.net/fig/549199" pageId="0" pageNumber="167">, 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and triangular process of femorite, l apicolateral lobe of femorite, m dorsoapical lobe of femorite, sl tip of solenomere, sph solenophore, z short and simple process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure7" httpUri="https://binary.pensoft.net/fig/549200" pageId="0" pageNumber="167">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="167" type="materials_examined">
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<paragraph pageId="0" pageNumber="167">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="167">
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<materialsCitation collectingDate="2016-09-21" collectorName="R. Srisonchai" country="Myanmar" elevation="1204" latitude="20.754417" location="Montawa Cave" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" specimenCount="♂" stateProvince="Shan State" typeStatus="Holotype">
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<emphasis bold="true" pageId="0" pageNumber="167">
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<emphasis italics="true" pageId="0" pageNumber="167">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="Myanmar">Myanmar</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="167">
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<collectingRegion country="Myanmar" name="Shan">Shan State</collectingRegion>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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;
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<collectingMunicipality>Taunggyi District</collectingMunicipality>
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, near
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<location LSID="urn:lsid:plazi:treatment:B4DFAF1674885A80AFDE56D716EED2C4:F14960500377E41C184A68EC84C37107" country="Myanmar" latitude="20.754417" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" name="Montawa Cave" stateProvince="Shan State">Montawa Cave</location>
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; elev.
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.204" unit="m" value="1204.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.204" unit="m" value="1204.0">1204 m</elevation>
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</quantity>
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;
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<geoCoordinate degrees="20" direction="north" minutes="45" orientation="latitude" precision="1" seconds="15.9" value="20.754417">20°45'15.9"N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="01" orientation="longitude" precision="1" seconds="03.4" value="97.01762">97°01'03.4"E</geoCoordinate>
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;
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<collectingDate value="2016-09-21">21 Sep. 2016</collectingDate>
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;
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<collectorName>R. Srisonchai</collectorName>
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leg.; CUMZ
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</materialsCitation>
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.
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="R. Srisonchai" country="Myanmar" elevation="1204" latitude="20.754417" location="Shan State" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" specimenCount="1" specimenCount-female="1" stateProvince="Shan State" typeStatus="Paratype">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="167">
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<typeStatus>Paratype</typeStatus>
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</emphasis>
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:
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<collectingCountry name="Myanmar">Myanmar</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="167">
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<collectingRegion country="Myanmar" name="Shan">Shan State</collectingRegion>
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</emphasis>
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•
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<specimenCount type="female">1 ♀</specimenCount>
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; same collection data as holotype; CUMZ
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="167" type="diagnosis">
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<paragraph pageId="0" pageNumber="167">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="167">
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This new species comes to a dead end in couplet 5 in the latest key to
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<taxonomicName authorityName="Jeekel" authorityYear="1968" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="167">Tylopus</emphasis>
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</taxonomicName>
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species (
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<bibRefCitation DOI="https://doi.org/10.5852/ejt.2016.195" author="Likhitrakarn, N" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="167" pagination="1 - 47" refId="B23" refString="Likhitrakarn, N, Golovatch, SI, Panha, S, 2016. The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina. European Journal of Taxonomy 195: 1 - 47, DOI: https://doi.org/10.5852/ejt.2016.195" title="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina." url="https://doi.org/10.5852/ejt.2016.195" volume="195" year="2016">Likhitrakarn et al. 2016</bibRefCitation>
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), but it seems to be particularly similar to the later described and grossly sympatric
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<taxonomicName lsidName="T. brehieri" pageId="0" pageNumber="167" rank="species" species="brehieri">
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<emphasis italics="true" pageId="0" pageNumber="167">T. brehieri</emphasis>
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</taxonomicName>
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Golovatch, VandenSpiegel & Semenyuk, 2016, especially in its gonopod conformation. Both species compared come from Shan State, Myanmar (
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<bibRefCitation author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="335 - 354" publicationUrl="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" refId="B16" refString="Golovatch, SI, VandenSpiegel, D, Semenyuk, II, 2016. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXI. Arthropoda Selecta 25 (4): 335 - 354, https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXI." url="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" volume="25" year="2016">Golovatch et al. 2016</bibRefCitation>
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), but
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<taxonomicName lsidName="T. sutchariti" pageId="0" pageNumber="167" rank="species" species="sutchariti">
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<emphasis italics="true" pageId="0" pageNumber="167">T. sutchariti</emphasis>
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</taxonomicName>
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sp. nov. differs in the presence of a small and triangular gonopod process
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<emphasis bold="true" pageId="0" pageNumber="167">h</emphasis>
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(vs. absent), and the large and subtrapeziform apicolateral lobe (
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<emphasis bold="true" pageId="0" pageNumber="167">l</emphasis>
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) with a smooth apical margin (Figs
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, sublateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure6" httpUri="https://binary.pensoft.net/fig/549199" pageId="0" pageNumber="167">6B, D</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and triangular process of femorite, l apicolateral lobe of femorite, m dorsoapical lobe of femorite, sl tip of solenomere, sph solenophore, z short and simple process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure7" httpUri="https://binary.pensoft.net/fig/549200" pageId="0" pageNumber="167">7B</figureCitation>
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) (vs. a subtriangular
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<emphasis bold="true" pageId="0" pageNumber="167">l</emphasis>
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with an apically rugose and denticulate margin), as well as the pleurosternal carinae being complete crests with a caudal tooth clearly visible until segments 16 (♂) or 13 (♀), thereafter missing (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5B, D, E</figureCitation>
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) (vs. visible until segment 10), while the sternal lobe between ♂ coxae 4 is deeply notched (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5H, I</figureCitation>
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) (vs. prominent and subquadrate).
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</paragraph>
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<caption doi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="167">
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<emphasis bold="true" pageId="0" pageNumber="167">Figure 5.</emphasis>
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<taxonomicName authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus sutchariti" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="sutchariti">
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<emphasis italics="true" pageId="0" pageNumber="167">Tylopus sutchariti</emphasis>
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</taxonomicName>
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sp. nov., ♂ holotype (CUMZ)
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<emphasis bold="true" pageId="0" pageNumber="167">A, B</emphasis>
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anterior part of body, dorsal and lateral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="167">C, D</emphasis>
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segments 10 and 11, dorsal and lateral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="167">E-G</emphasis>
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posterior part of body, lateral, subdorsal and subventral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="167">H, I</emphasis>
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sternal cones between coxae 4, subcaudal and sublateral views, respectively.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="167" type="description">
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<paragraph pageId="0" pageNumber="167">Description.</paragraph>
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<paragraph pageId="0" pageNumber="167">Length 24.1 (♂) or 22.5 mm (♀), width of midbody pro- and metazona 1.8 and 2.7 mm (♂) or 2.2 and 2.6 mm (♀), respectively.</paragraph>
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<paragraph pageId="0" pageNumber="167">
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Colouration of live animals dark brown (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">1B</figureCitation>
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); venter and legs brown; colouration of alcohol material after two years of preservation blackish, calluses of paraterga yellowish brown, head and antennae dark brown, venter and legs light yellowish, increasingly darker brown distally (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="167">
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Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae short (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">1B</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5B</figureCitation>
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), reaching body segment 3 (♂) or surpassing body segment 2 (♀) when stretched dorsally. In width, head <segment 3 <4 <5 <collum <segment 2 <6-17 (♂, ♀); thereafter body gently and gradually tapering. Collum with three transverse rows of strong setae: 3+3 anterior, 1+1 intermediate, and 3+3 posterior; a small lateral incision near midway; caudal corner of paraterga rounded, slightly declined ventrad, not surpassing rear tergal margin (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
Tegument rather smooth and shining, prozonae very finely shagreened, metaterga smooth and finely rugulose; surface below paraterga finely microgranulate (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A-F</figureCitation>
|
|
). Postcollum metaterga with two transverse rows of rather long setae: 2+2 in anterior and 2(3)+2(3) in posterior row, the latter often abraded, but then readily traceable as insertion points. Tergal setae long, strong, slender, about 1/3 metatergal length. Axial line visible only on metazonae.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
Paraterga strongly developed (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A-F</figureCitation>
|
|
), especially well so in ♂, set high, at upper 1/3 of midbody height, slightly upturned, but remaining below dorsum; anterior edge well-developed, mostly regularly rounded and narrowly bordered, continuous with callus; caudal corner narrowly rounded to fully pointed, extending increasingly past rear tergal margin, especially well so on segments 15-19; on segments 16-19, tips strongly curved mesad, posterior edge slightly oblique (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A, C, F</figureCitation>
|
|
); paraterga very thin blunt blades in lateral view, a little thicker only on pore-bearing segments (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5D</figureCitation>
|
|
). Calluses on paraterga delimited by a sulcus both dorsally and ventrally. Paraterga 2 broad, horizontal, anterior edge angular, lateral edge with three evident incisions, one in anterior 1/3, middle one at midway, caudal incision near tip; anterior incision particularly evident. Paraterga 3 and 4 with two small incisions at lateral edge (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A</figureCitation>
|
|
), one in anterior 1/3, the other at midway, anterior one also particularly evident. Following segments each with lateral edge showing an evident incision near front 1/3 (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5C</figureCitation>
|
|
). Ozopores evident, lateral, lying in an ovoid groove at about 1/3 metatergal length in front of posterior edge of metaterga (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5D</figureCitation>
|
|
). Transverse sulcus usually distinct (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A, C, F</figureCitation>
|
|
), complete and visible on metaterga 5-18, deep, narrow, reaching bases of paraterga, line-shaped, clearly beaded at bottom. Stricture between pro- and metazonae wide, deep, clearly ribbed at bottom down to base of paraterga (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A, C, F</figureCitation>
|
|
). Pleurosternal carinae complete crests on segment 2-4 (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5B</figureCitation>
|
|
), with anteriorly bulged crests and a sharp denticle caudally on segments 5-8 (♂, ♀), thereafter increasingly reduced and broken, remaining only a small sharp caudal tooth until segment 16 (♂) or 13 (♀), thereafter missing (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5B, D, E</figureCitation>
|
|
). Epiproct (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5E-G</figureCitation>
|
|
) conical, flattened dorsoventrally, subtruncate, with two evident apical papillae directed caudally, both pointed at tip; pre-apical papillae evident, lying close to tip. Hypoproct roundly subtrapeziform (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5G</figureCitation>
|
|
), small setiferous knobs at caudal edge well-separated and evident.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
Sterna densely setose, without modifications (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5G</figureCitation>
|
|
); cross-impressions shallow; a deeply notched sternal lobe between ♂ coxae 4 (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5H, I</figureCitation>
|
|
). Legs long and slender, midbody ones ca 1.4-1.5 (♂) or 1.1-1.2 (♀) as long as body height; ♂ legs of segments 4-17 with prefemora distinctly swollen laterally; ♂ legs of segments 2-16 each with femur, postfemur, tibia and tarsus with particularly dense setae and carrying ventral microgranulations (Fig.
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and triangular process of femorite, l apicolateral lobe of femorite, m dorsoapical lobe of femorite, sl tip of solenomere, sph solenophore, z short and simple process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure7" httpUri="https://binary.pensoft.net/fig/549200" pageId="0" pageNumber="167">7C</figureCitation>
|
|
), tarsal brushes absent.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
Gonopods (Figs
|
|
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, sublateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure6" httpUri="https://binary.pensoft.net/fig/549199" pageId="0" pageNumber="167">6</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and triangular process of femorite, l apicolateral lobe of femorite, m dorsoapical lobe of femorite, sl tip of solenomere, sph solenophore, z short and simple process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure7" httpUri="https://binary.pensoft.net/fig/549200" pageId="0" pageNumber="167">7A, B</figureCitation>
|
|
) complex; coxa slightly curved caudad, sparsely setose distoventrally. Prefemorite as usual, densely setose, about 1/2 as long as femorite +
|
|
<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
|
part. Femorite rather stout, suberect, expanded distad, showing a distinct mesal groove/hollow (
|
|
<emphasis bold="true" pageId="0" pageNumber="167">g</emphasis>
|
|
) and a prominent, rounded, dorso-apical lobe (
|
|
<emphasis bold="true" pageId="0" pageNumber="167">m</emphasis>
|
|
), apicolateral lobe (
|
|
<emphasis bold="true" pageId="0" pageNumber="167">l</emphasis>
|
|
) large, subtrapeziform, with a smooth apical margin, mostly delimited at base by a transverse sulcus, with process
|
|
<emphasis bold="true" pageId="0" pageNumber="167">h</emphasis>
|
|
short and triangular; process (
|
|
<emphasis bold="true" pageId="0" pageNumber="167">z</emphasis>
|
|
) rather short and simple, narrowly rounded at tip. Solenophore (
|
|
<emphasis bold="true" pageId="0" pageNumber="167">sph</emphasis>
|
|
) typically coiled, lamellar, expanded apically into a subtruncate lobe, almost fully sheathing a similarly long, flagelliform solenomere, with only its tip (
|
|
<emphasis bold="true" pageId="0" pageNumber="167">sl</emphasis>
|
|
) being exposed.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1040.66209.figure6" httpUri="https://binary.pensoft.net/fig/549199" pageId="0" pageNumber="167" start="Figure 6" startId="F6">
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">Figure 6.</emphasis>
|
|
<taxonomicName authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus sutchariti" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="sutchariti">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus sutchariti</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., ♂ holotype (CUMZ), left gonopod
|
|
<emphasis bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
|
mesal and lateral views, respectively
|
|
<emphasis bold="true" pageId="0" pageNumber="167">C-F</emphasis>
|
|
distal part, submesal, sublateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="167" type="name">
|
|
<paragraph pageId="0" pageNumber="167">Name.</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">To honour Dr. Chirasak Sutcharit, Professor at the Department of Biology of the Chulalongkorn University, Bangkok, who participated in collecting the type series.</paragraph>
|
|
<caption doi="10.3897/zookeys.1040.66209.figure7" httpUri="https://binary.pensoft.net/fig/549200" pageId="0" pageNumber="167" start="Figure 7" startId="F7">
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">Figure 7.</emphasis>
|
|
<taxonomicName authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus sutchariti" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="sutchariti">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus sutchariti</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., ♂ holotype (CUMZ)
|
|
<emphasis bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
|
left gonopod, mesal and lateral views, respectively
|
|
<emphasis bold="true" pageId="0" pageNumber="167">C</emphasis>
|
|
leg of segment 10. Abbreviations:
|
|
<emphasis bold="true" pageId="0" pageNumber="167">g</emphasis>
|
|
mesal groove/hollow of femorite,
|
|
<emphasis bold="true" pageId="0" pageNumber="167">h</emphasis>
|
|
short and triangular process of femorite,
|
|
<emphasis bold="true" pageId="0" pageNumber="167">l</emphasis>
|
|
apicolateral lobe of femorite,
|
|
<emphasis bold="true" pageId="0" pageNumber="167">m</emphasis>
|
|
dorsoapical lobe of femorite,
|
|
<emphasis bold="true" pageId="0" pageNumber="167">sl</emphasis>
|
|
tip of solenomere,
|
|
<emphasis bold="true" pageId="0" pageNumber="167">sph</emphasis>
|
|
solenophore,
|
|
<emphasis bold="true" pageId="0" pageNumber="167">z</emphasis>
|
|
short and simple process of femorite. Scale bars: 0.5 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.1040.66209.figure8" httpUri="https://binary.pensoft.net/fig/549201" pageId="0" pageNumber="167" start="Figure 8" startId="F8">
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">Figure 8.</emphasis>
|
|
Gonopod structure of
|
|
<taxonomicName authorityName="Golovatch & Enghoff" authorityYear="1993" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus rugosus" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="rugosus">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus rugosus</emphasis>
|
|
</taxonomicName>
|
|
Golovatch & Enghoff, 1993, ♂ holotype, left gonopod
|
|
<emphasis bold="true" pageId="0" pageNumber="167">A-C</emphasis>
|
|
mesal, lateral and dorsal views, respectively. Abbreviations:
|
|
<emphasis bold="true" pageId="0" pageNumber="167">h</emphasis>
|
|
strong hook-shaped process of femorite,
|
|
<emphasis bold="true" pageId="0" pageNumber="167">l</emphasis>
|
|
apicolateral lobe of femorite,
|
|
<emphasis bold="true" pageId="0" pageNumber="167">z</emphasis>
|
|
serrate process of femorite. Scale bar: 0.5 mm (after
|
|
<bibRefCitation author="Golovatch, SI" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="167" pagination="85 - 125" refId="B14" refString="Golovatch, SI, Enghoff, H, 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125" title="Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="19" year="1993">Golovatch and Enghoff 1993</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="167" type="remarks">
|
|
<paragraph pageId="0" pageNumber="167">Remark.</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
Both new species described here have been found to occur syntopically (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distributions of Tylopus species recorded from Myanmar. Pink circle: T. brehieri Golovatch, VandenSpiegel & Semenyuk, 2016; Red Asterisk: T. punctus Likhitrakarn, Golovatch & Panha, 2016; Yellow inverted triangle: T. monticola sp. nov.; Blue square: T. sutchariti sp. nov. and T. monticola sp. nov.; Orange diamond: T. silvestris (Pocock, 1895); Green triangle: T. doriae (Pocock, 1895)." figureDoi="10.3897/zookeys.1040.66209.figure9" httpUri="https://binary.pensoft.net/fig/549202" pageId="0" pageNumber="167">9</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1040.66209.figure9" httpUri="https://binary.pensoft.net/fig/549202" pageId="0" pageNumber="167" start="Figure 9" startId="F9">
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">Figure 9.</emphasis>
|
|
Distributions of
|
|
<taxonomicName authorityName="Jeekel" authorityYear="1968" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus</emphasis>
|
|
</taxonomicName>
|
|
species recorded from Myanmar. Pink circle:
|
|
<taxonomicName lsidName="T. brehieri" pageId="0" pageNumber="167" rank="species" species="brehieri">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">T. brehieri</emphasis>
|
|
</taxonomicName>
|
|
Golovatch, VandenSpiegel & Semenyuk, 2016; Red Asterisk:
|
|
<taxonomicName lsidName="T. punctus" pageId="0" pageNumber="167" rank="species" species="punctus">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">T. punctus</emphasis>
|
|
</taxonomicName>
|
|
Likhitrakarn, Golovatch & Panha, 2016; Yellow inverted triangle:
|
|
<taxonomicName lsidName="T. monticola" pageId="0" pageNumber="167" rank="species" species="monticola">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">T. monticola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.; Blue square:
|
|
<taxonomicName lsidName="T. sutchariti" pageId="0" pageNumber="167" rank="species" species="sutchariti">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">T. sutchariti</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName lsidName="T. monticola" pageId="0" pageNumber="167" rank="species" species="monticola">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">T. monticola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.; Orange diamond:
|
|
<taxonomicName lsidName="T. silvestris" pageId="0" pageNumber="167" rank="species" species="silvestris">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">T. silvestris</emphasis>
|
|
</taxonomicName>
|
|
(Pocock, 1895); Green triangle:
|
|
<taxonomicName lsidName="T. doriae" pageId="0" pageNumber="167" rank="species" species="doriae">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">T. doriae</emphasis>
|
|
</taxonomicName>
|
|
(Pocock, 1895).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |