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treatments-xml/data/AB/27/87/AB2787FCFFF6FFE7FF1AFE00FE32B3A4.xml
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<document id="23562C6F7BC85CFAF4DF4A0183BF3FC5" ID-DOI="10.25221/fee.470.2" ID-ISSN="2713-2196" ID-Zenodo-Dep="7616385" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1675789169336" checkinUser="felipe" docAuthor="Wang, C. - B., He, T. - L., He, L. &amp; Zhou, C." docDate="2023" docId="AB2787FCFFF6FFE7FF1AFE00FE32B3A4" docLanguage="en" docName="FarEastEntomol.470.8-22.pdf" docOrigin="Far Eastern Entomologist 470" docSource="http://dx.doi.org/10.25221/fee.470.2" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.11:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="11" docTitle="Lucanus niu Wang, He, He et Zhou 2023, sp. n." docType="treatment" docVersion="5" lastPageNumber="16" masterDocId="571EFF84FFF4FFEFFFF9FFECFFAFB27A" masterDocTitle="Two new species of Lucanus Scopoli, 1763 from Yunnan, Southwest China (Coleoptera, Lucanidae, Lucaninae)" masterLastPageNumber="22" masterPageNumber="8" pageNumber="10" updateTime="1712594434517" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="9CEF7407BC48983F78FAE807BBF39508" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="61E7C4F6DB65375F1BC7A429A6A260EC">
<mods:title id="68B4074598BF33CDBB0A785451C16D89">Two new species of Lucanus Scopoli, 1763 from Yunnan, Southwest China (Coleoptera, Lucanidae, Lucaninae)</mods:title>
</mods:titleInfo>
<mods:name id="91FB6EDE916FB76AFA8D89953381C1CB" type="personal">
<mods:role id="DC8411BA5A51081EA8CC808C5134E723">
<mods:roleTerm id="93E36978672B09EACCEE9A6EC9BC5BEE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8795E7F8F74499A545E839CFB35B2D44">Wang, C. - B.</mods:namePart>
</mods:name>
<mods:name id="8E955FCF003D5DCD3CEEABA5C0BB8253" type="personal">
<mods:role id="59A682AD983C2E190F9C2A4CED6D5A80">
<mods:roleTerm id="DCDA211BB0BBDDAEC06BE57C8653DEDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="671528170B8F07286E64FB6D3E976EBB">He, T. - L.</mods:namePart>
</mods:name>
<mods:name id="D492928033FB74E1CF1FDCA5A637FE78" type="personal">
<mods:role id="27A0A552EDD6E318288B89264347769A">
<mods:roleTerm id="392AD727FD2A24EDD1FF72FAC362810C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="20C3B47C60C52857D69C975436FF9911">He, L.</mods:namePart>
</mods:name>
<mods:name id="6A31EBAA1AA9F07AECFE8CC10C904203" type="personal">
<mods:role id="2CC66CA4694E610C9ABADA298DAC003C">
<mods:roleTerm id="9F0931D0E52BA0B50439E78AF4AF0E94">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF5ED89A4DF9EB8D9BDD3EB9CD7CD25B">Zhou, C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C0CBA9A1F395EA299D869F9B3860BEC1">text</mods:typeOfResource>
<mods:relatedItem id="490111868E8E04D705ECA08ED23DB00B" type="host">
<mods:titleInfo id="B342D4CD0CA9B6077F5D30FB7974C609">
<mods:title id="59D47A594DFD91EE8FC75959E3406CAE">Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part id="1E1081660F6B94EA2E380103F97290C6">
<mods:date id="E546F91747894D39C82857B344BFF60D">2023</mods:date>
<mods:detail id="06FAF8F20B54A004A941BD8C3C1A434D" type="pubDate">
<mods:number id="E7454AEBA8393CCFAA71A9F22DC1A027">2023-02-02</mods:number>
</mods:detail>
<mods:detail id="CE4D50C901444DCC07791EB09AF6AFB6" type="volume">
<mods:number id="6C21D014658D96B63C59AA154B904A38">470</mods:number>
</mods:detail>
<mods:extent id="5E74D8CE0635FF14D632D81D12E4975C" unit="page">
<mods:start id="7C076CA9EA78781D358437BF3BA311F4">8</mods:start>
<mods:end id="9FAAE57CC0BC499B59F7C6FCD4B08D14">22</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="2B315B2A33991481F1F4AD4BA51989F5">
<mods:url id="80D172407A40451112D4A97061A60D13">http://dx.doi.org/10.25221/fee.470.2</mods:url>
</mods:location>
<mods:classification id="3FA60DEDB3BE41CC30677CF0DAAFA85E">journal article</mods:classification>
<mods:identifier id="266E59313D025F8A2506ECA03D2F9F89" type="DOI">10.25221/fee.470.2</mods:identifier>
<mods:identifier id="1DAF7C5799A986675D95A2FD2E3EE2B1" type="ISSN">2713-2196</mods:identifier>
<mods:identifier id="10CE1D13B7E47E7187075E607C721AF8" type="Zenodo-Dep">7616385</mods:identifier>
</mods:mods>
<treatment id="AB2787FCFFF6FFE7FF1AFE00FE32B3A4" LSID="urn:lsid:plazi:treatment:AB2787FCFFF6FFE7FF1AFE00FE32B3A4" httpUri="http://treatment.plazi.org/id/AB2787FCFFF6FFE7FF1AFE00FE32B3A4" lastPageId="8" lastPageNumber="16" pageId="2" pageNumber="10">
<subSubSection id="6B946561FFF6FFEDFF1AFE00FB83B059" pageId="2" pageNumber="10" type="nomenclature">
<paragraph id="233136EAFFF6FFEDFF1AFE00FD22B07B" blockId="2.[227,653,492,513]" box="[227,653,492,513]" pageId="2" pageNumber="10">
<heading id="78798186FFF6FFEDFF1AFE00FD22B07B" box="[227,653,492,513]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="10" reason="2">
<emphasis id="11FAEAF8FFF6FFEDFF1AFE00FD22B07B" bold="true" box="[227,653,492,513]" pageId="2" pageNumber="10">
<taxonomicName id="E48E4D69FFF6FFEDFF1AFE00FDE3B07B" ID-CoL="B6JR9" authority="Wang, He, He et Zhou" authorityName="Wang, He, He et Zhou" authorityYear="2023" box="[227,588,492,513]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="10" phylum="Arthropoda" rank="species" species="niu" status="sp. nov.">
<emphasis id="11FAEAF8FFF6FFEDFF1AFE00FECEB07B" bold="true" box="[227,353,492,513]" italics="true" pageId="2" pageNumber="10">Lucanus niu</emphasis>
Wang, He, He et Zhou
</taxonomicName>
,
<taxonomicNameLabel id="0AC95783FFF6FFEDFDAFFE00FD22B07B" box="[598,653,492,513]" pageId="2" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="233136EAFFF6FFEDFF1AFDE1FB83B059" blockId="2.[227,1068,525,547]" box="[227,1068,525,547]" pageId="2" pageNumber="10">
https://zoobank.org/NomenclaturalActs/
<uuid id="57280C3FFFF6FFEDFD94FDE1FB83B059" box="[621,1068,525,547]" pageId="2" pageNumber="10">EE5B2E96-8327-48CC-A51E-476209B24E2B</uuid>
</paragraph>
</subSubSection>
<subSubSection id="6B946561FFF6FFEDFF1AFDC2FE99B03E" box="[227,310,558,580]" pageId="2" pageNumber="10" type="description">
<paragraph id="233136EAFFF6FFEDFF1AFDC2FE99B03E" blockId="2.[227,310,558,580]" box="[227,310,558,580]" pageId="2" pageNumber="10">
<figureCitation id="BBB52A6FFFF6FFEDFF1AFDC2FEB1B03E" box="[227,286,558,580]" captionStart="Fig" captionStartId="3.[265,300,1556,1578]" captionTargetBox="[355,1006,271,1539]" captionTargetId="figure-0@3.[319,1036,257,1550]" captionTargetPageId="3" captionText="Fig. 1. Lucanus niu sp. n., male habitus. a–c – large size, holotype; d–f – moderate size, paratype; g–i – small size, paratype; (a, d, g – dorsal view; b, e, h – ventral view; c, f, i – lateral view). Scale bar = 1.0 cm." figureDoi="http://doi.org/10.5281/zenodo.7616387" httpUri="https://zenodo.org/record/7616387/files/figure.png" pageId="2" pageNumber="10">Figs 1</figureCitation>
,
<figureCitation id="BBB52A6FFFF6FFEDFED3FDC3FE99B03E" box="[298,310,559,580]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="2" pageNumber="10">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6B946561FFF6FFEDFEF0FD8BFC15B097" pageId="2" pageNumber="10" type="materials_examined">
<paragraph id="233136EAFFF6FFEDFEF0FD8BFC15B097" blockId="2.[227,1128,614,1595]" pageId="2" pageNumber="10">
<materialsCitation id="93E63CB7FFF6FFEDFEF0FD8BFBC8B006" box="[265,1127,614,637]" country="China" county="Zhenyuan County" location="Puer City" pageId="2" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
MATERIAL EXAMINED.
<typeStatus id="FC358848FFF6FFEDFDE0FD8BFDDDB007" box="[537,626,615,637]" pageId="2" pageNumber="10" type="holotype">Holotype</typeStatus>
<emphasis id="11FAEAF8FFF6FFEDFD80FD8BFD2AB006" bold="true" box="[633,645,615,636]" pageId="2" pageNumber="10">–</emphasis>
<specimenCount id="3588FD63FFF6FFEDFD75FD8BFD0FB006" box="[652,672,615,636]" pageId="2" pageNumber="10" type="male">♂</specimenCount>
,
<emphasis id="11FAEAF8FFF6FFEDFD52FD8AFD45B006" bold="true" box="[683,746,614,636]" pageId="2" pageNumber="10">
<collectingCountry id="5B99767AFFF6FFEDFD52FD8AFD45B006" box="[683,746,614,636]" name="China" pageId="2" pageNumber="10">China</collectingCountry>
</emphasis>
:
<collectingRegion id="E14AF808FFF6FFEDFD0EFD8BFCE9B006" box="[759,838,615,636]" country="China" name="Yunnan" pageId="2" pageNumber="10">Yunnan</collectingRegion>
,
<location id="26516031FFF6FFEDFCA9FD8BFC00B007" LSID="urn:lsid:plazi:treatment:AB2787FCFFF6FFE7FF1AFE00FE32B3A4:26516031FFF6FFEDFCA9FD8BFC00B007" box="[848,943,615,637]" country="China" county="Zhenyuan County" name="Puer City" pageId="2" pageNumber="10" stateProvince="Yunnan">Puer City</location>
,
<collectingCounty id="CA504E66FFF6FFEDFC40FD8BFBC8B006" box="[953,1127,615,637]" pageId="2" pageNumber="10">Zhenyuan County</collectingCounty>
</materialsCitation>
[
<materialsCitation id="93E63CB7FFF6FFEDFF12FD92FBCDB0E2" box="[235,1122,638,665]" collectingDate="2020-06-12" collectionCode="MAHU" collectorName="Shao-You Huang" elevation="2000" pageId="2" pageNumber="10" specimenCount="10" specimenCount-male="10" typeStatus="paratype">
普洱市镇沅县],
<date id="5730102AFFF6FFEDFE96FD6FFE71B0E2" box="[367,478,643,664]" pageId="2" pageNumber="10" value="2020-06-12">
<collectingDate id="4774E9C2FFF6FFEDFE96FD6FFE71B0E2" box="[367,478,643,664]" pageId="2" pageNumber="10" value="2020-06-12">12.VI 2020</collectingDate>
</date>
,
<quantity id="E4769B0FFFF6FFEDFE11FD6FFD9CB0E2" box="[488,563,643,664]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="10" unit="m" value="2000.0">
<elevation id="A8A3D1D9FFF6FFEDFE11FD6FFD9CB0E2" box="[488,563,643,664]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="10" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<collectorName id="8E7B533CFFF6FFEDFDC4FD6FFD4BB0E2" box="[573,740,643,665]" pageId="2" pageNumber="10">Shao-You Huang</collectorName>
leg. (
<collectionCode id="459FAE2FFFF6FFEDFCE3FD6FFCC6B0E2" box="[794,873,643,664]" pageId="2" pageNumber="10">MAHU</collectionCode>
).
<typeStatus id="FC358848FFF6FFEDFC80FD6FFC78B0E2" box="[889,983,643,664]" pageId="2" pageNumber="10" type="paratype">Paratypes</typeStatus>
(totally
<specimenCount id="3588FD63FFF6FFEDFBD1FD6FFBF4B0E2" box="[1064,1115,643,664]" pageId="2" pageNumber="10" type="male">10 ♂</specimenCount>
)
</materialsCitation>
:
<materialsCitation id="93E63CB7FFF6FFEDFF1AFD73FBCBB0CE" box="[227,1124,671,693]" collectionCode="CCZC, CTLH" country="China" county="Zhenyuan County" location="Puer City" pageId="2" pageNumber="10" specimenCount="4" specimenCount-male="4" stateProvince="Yunnan" typeStatus="holotype">
the same data as holotype,
<specimenCount id="3588FD63FFF6FFEDFE09FD73FDA1B0CE" box="[496,526,671,692]" pageId="2" pageNumber="10" type="male">1♂</specimenCount>
(
<collectionCode id="459FAE2FFFF6FFEDFDE5FD73FDF1B0CE" box="[540,606,671,692]" pageId="2" pageNumber="10">CCZC</collectionCode>
) and
<specimenCount id="3588FD63FFF6FFEDFD6EFD73FD1AB0CE" box="[663,693,671,692]" pageId="2" pageNumber="10" type="male">3♂</specimenCount>
(
<collectionCode id="459FAE2FFFF6FFEDFD3DFD73FCA7B0CE" box="[708,776,671,692]" pageId="2" pageNumber="10">CTLH</collectionCode>
); the same data as
<typeStatus id="FC358848FFF6FFEDFC3CFD73FBB6B0CF" box="[965,1049,671,693]" pageId="2" pageNumber="10" type="holotype">holotype</typeStatus>
except
</materialsCitation>
:
<date id="5730102AFFF6FFEDFF1AFD57FEFAB0AA" box="[227,341,699,720]" pageId="2" pageNumber="10" value="2020-06-24">
24.
<collectionCode id="459FAE2FFFF6FFEDFF06FD57FEB5B0AA" box="[255,282,699,720]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="10">VI</collectionCode>
<materialsCitation id="93E63CB7FFF6FFEDFEDAFD57FEFAB0AA" box="[291,341,699,720]" collectionCode="CTLH, CYCX" collectorName="Yun-Chuan Xu" pageId="2" pageNumber="10" specimenCount="2" specimenCount-male="2">2020</materialsCitation>
</date>
,
<specimenCount id="3588FD63FFF6FFEDFE9BFD57FE2FB0AA" box="[354,384,699,720]" pageId="2" pageNumber="10" type="male">1♂</specimenCount>
(
<collectionCode id="459FAE2FFFF6FFEDFE76FD57FE7EB0AA" box="[399,465,699,720]" pageId="2" pageNumber="10">CTLH</collectionCode>
) and
<specimenCount id="3588FD63FFF6FFEDFDF2FD57FD85B0AA" box="[523,554,699,720]" pageId="2" pageNumber="10" type="male">1♂</specimenCount>
(
<collectionCode id="459FAE2FFFF6FFEDFDC0FD57FD2EB0AA" box="[569,641,699,720]" pageId="2" pageNumber="10">CYCX</collectionCode>
),
<collectorName id="8E7B533CFFF6FFEDFD6DFD57FC86B0AB" box="[660,809,699,721]" pageId="2" pageNumber="10">Yun-Chuan Xu</collectorName>
leg.;
<materialsCitation id="93E63CB7FFF6FFEDFC9CFD57FED6B097" country="China" county="Zhenyuan County" elevation="2000" location="Puer City" pageId="2" pageNumber="10" specimenCount="1" stateProvince="Yunnan">
the same data as holotype except:
<quantity id="E4769B0FFFF6FFEDFED7FD34FED6B097" box="[302,377,728,749]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="10" unit="m" value="2000.0">
<elevation id="A8A3D1D9FFF6FFEDFED7FD34FED6B097" box="[302,377,728,749]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="10" unit="m" value="2000.0">2000 m</elevation>
</quantity>
</materialsCitation>
,
<date id="5730102AFFF6FFEDFE7AFD3BFE49B097" box="[387,486,727,749]" pageId="2" pageNumber="10" value="2021-06-05">
5.
<collectionCode id="459FAE2FFFF6FFEDFE6BFD3BFE00B097" box="[402,431,727,749]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="10">VI</collectionCode>
<materialsCitation id="93E63CB7FFF6FFEDFE4DFD34FE49B097" box="[436,486,728,749]" collectionCode="CTLH, MYNU" country="China" pageId="2" pageNumber="10" specimenCount="4" specimenCount-male="4" stateProvince="Yunnan">2021</materialsCitation>
</date>
,
<specimenCount id="3588FD63FFF6FFEDFE09FD34FDA1B097" box="[496,526,728,749]" pageId="2" pageNumber="10" type="male">3♂</specimenCount>
(
<collectionCode id="459FAE2FFFF6FFEDFDE2FD34FDF4B097" box="[539,603,728,749]" pageId="2" pageNumber="10">CTLH</collectionCode>
) and
<specimenCount id="3588FD63FFF6FFEDFD6BFD34FD1FB097" box="[658,688,728,749]" pageId="2" pageNumber="10" type="male">1♂</specimenCount>
(
<collectionCode id="459FAE2FFFF6FFEDFD45FD34FCA5B097" box="[700,778,728,749]" pageId="2" pageNumber="10">MYNU</collectionCode>
), local people leg.
</paragraph>
</subSubSection>
<subSubSection id="6B946561FFF6FFE7FEF0FD18FBF8B315" lastPageId="8" lastPageNumber="16" pageId="2" pageNumber="10" type="description">
<paragraph id="233136EAFFF6FFEDFEF0FD18FDFFB138" blockId="2.[227,1128,614,1595]" pageId="2" pageNumber="10">
<materialsCitation id="93E63CB7FFF6FFEDFEF0FD18FBC8B15C" collectionCode="MALE" pageId="2" pageNumber="10" specimenCount="1" typeStatus="holotype">
DESCRIPTION.
<typeStatus id="FC358848FFF6FFEDFE4BFD18FD9EB173" box="[434,561,756,777]" pageId="2" pageNumber="10" type="holotype">HOLOTYPE</typeStatus>
<collectionCode id="459FAE2FFFF6FFEDFDC1FD18FDD0B173" box="[568,639,756,777]" pageId="2" pageNumber="10">MALE</collectionCode>
. Large size, body
<quantity id="E4769B0FFFF6FFEDFCCBFD1FFC27B173" box="[818,904,755,777]" metricMagnitude="-2" metricUnit="m" metricValue="5.74" pageId="2" pageNumber="10" unit="mm" value="57.4">57.4 mm</quantity>
long. Lengths of body parts (mm): head (9.3), mandible (21.2), pronotum (8.3), elytra (23.0); width (mm): head
</materialsCitation>
(14.8), pronotum (13.6), elytra (16.6).
</paragraph>
<paragraph id="233136EAFFF6FFEDFEF0FCA4FE8EB1B5" blockId="2.[227,1128,614,1595]" pageId="2" pageNumber="10">
Habitus (
<figureCitation id="BBB52A6FFFF6FFEDFE99FCA4FE12B127" box="[352,445,840,862]" captionStart="Fig" captionStartId="3.[265,300,1556,1578]" captionTargetBox="[355,1006,271,1539]" captionTargetId="figure-0@3.[319,1036,257,1550]" captionTargetPageId="3" captionText="Fig. 1. Lucanus niu sp. n., male habitus. a–c – large size, holotype; d–f – moderate size, paratype; g–i – small size, paratype; (a, d, g – dorsal view; b, e, h – ventral view; c, f, i – lateral view). Scale bar = 1.0 cm." figureDoi="http://doi.org/10.5281/zenodo.7616387" httpUri="https://zenodo.org/record/7616387/files/figure.png" pageId="2" pageNumber="10">Fig. 1a–c</figureCitation>
). Color mostly reddish brown to blackish brown; elytra reddish brown; femora with reddish stripes on ventral surfaces; tibiae with obscure reddish stripes on ventral surfaces; protibiae tinged with reddish on dorsal surfaces. Body mostly clothed with short, sparse, recumbent, yellowish pubescence, similar on metaventrite, much finer and denser on elytra.
</paragraph>
<paragraph id="233136EAFFF6FFEDFEF0FC39FBE6B441" blockId="2.[227,1128,614,1595]" pageId="2" pageNumber="10">Head 1.6 times as wide as long, widest at anterolateral angles; surfaces matt, finely punctate, intervals finely and irregularly granulose. Frons relatively long, fan-like, weakly concave, fused with clypeolabrum. Clypeal ridge strong, medially interrupted, laterally projecting as two ridges. Labrum slenderly subtriangular, rounded at apex. Anterolateral angles acute, strongly protruding laterally, more prominent than eyes. Ocular canthus protruding, shorter than half eye’s diameter. Postocular margin almost oblique, gradually narrowed posteriorly. Posterior margin widely concave. Anterior ridge distinctly and oblongly elevated, substraight at apical margin. Lateral ridges rather strongly elevated, broad, widely subrounded at apical margins. Mandible long and slender, about 2.3 times as long as head; in dorsal view, mandible curved in basal 1/4 and apical 1/5, and almost straight in between; in lateral view, mandible strongly curved in basal 3/5 and almost straight in apical 2/5; surfaces finely punctate, intervals matt and finely granulose at inner side, feebly sheen and microreticulate at outer side. Inner mandibular margin with unarmed carina at base, preceded by diastema, 7–8 small denticles, major inner tooth, and 2–3 distal small denticles. Major inner tooth situated at apical 1/3, strong, digitiform, shorter than mandibular width, tilting forwards and upwards with apex rounded and slightly higher than mandible. Apical fork with upper tooth bent upwards and inwards; lower tooth markedly shorter, conical, slightly directed downwards and inwards. Antennal club 4-segmented; antennomere VII with much thinner lamella, sharply pointing inwards, only pubescent on apical surface; antennomeres VIII–X with strong, long, pubescent lamellae. Mentum wide, inverted trapezoidal; anterior margin almost straight and with obliquely truncated corners; surface matt, finely punctate, intervals finely and irregularly granulose, and transversely depressed near anterior margin.</paragraph>
<caption id="77F16662FFF7FFECFEF0F9F8FD8DB41B" ID-DOI="http://doi.org/10.5281/zenodo.7616387" ID-Zenodo-Dep="7616387" httpUri="https://zenodo.org/record/7616387/files/figure.png" pageId="3" pageNumber="11" startId="3.[265,300,1556,1578]" subCaptionStartIDs="3.[582,653,1556,1578]" subCaptionStarts="habitus. a–" targetBox="[355,1006,271,1539]" targetPageId="3">
<paragraph id="233136EAFFF7FFECFEF0F9F8FD8DB41B" blockId="3.[227,1127,1556,1633]" pageId="3" pageNumber="11">
Fig. 1.
<taxonomicName id="E48E4D69FFF7FFECFEA9F9F9FE68B450" authority="Wang &amp; He &amp; He &amp; Zhou, 2023" authorityName="Wang &amp; He &amp; He &amp; Zhou" authorityYear="2023" box="[336,455,1557,1578]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="11" phylum="Arthropoda" rank="species" species="niu" status="sp. nov.">
<emphasis id="11FAEAF8FFF7FFECFEA9F9F9FE68B450" box="[336,455,1557,1578]" italics="true" pageId="3" pageNumber="11">Lucanus niu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AC95783FFF7FFECFE37F9F9FDACB450" box="[462,515,1557,1578]" pageId="3" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
, male habitus. a–c – large size, holotype; d–f – moderate size,
<typeStatus id="FC358848FFF7FFECFF1AF9DCFE96B43F" box="[227,313,1584,1605]" pageId="3" pageNumber="11" type="paratype">paratype</typeStatus>
; g–i – small size,
<typeStatus id="FC358848FFF7FFECFE09F9DCFDE9B43F" box="[496,582,1584,1605]" pageId="3" pageNumber="11" type="paratype">paratype</typeStatus>
; (a, d, g – dorsal view; b, e, h – ventral view; c, f, i – lateral view). Scale bar = 1.0 cm.
</paragraph>
</caption>
<caption id="77F16662FFF0FFEBFEF0FB42FB9BB683" ID-DOI="http://doi.org/10.5281/zenodo.7616389" ID-Zenodo-Dep="7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="4" pageNumber="12" startId="4.[265,300,1198,1220]" targetBox="[323,1012,266,1159]" targetPageId="4">
<paragraph id="233136EAFFF0FFEBFEF0FB42FB9BB683" blockId="4.[227,1126,1198,1274]" pageId="4" pageNumber="12">
Fig. 2.
<taxonomicName id="E48E4D69FFF0FFEBFEAAFB43FE63B6BE" authority="Wang &amp; He &amp; He &amp; Zhou, 2023" authorityName="Wang &amp; He &amp; He &amp; Zhou" authorityYear="2023" box="[339,460,1199,1220]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="12" phylum="Arthropoda" rank="species" species="niu" status="sp. nov.">
<emphasis id="11FAEAF8FFF0FFEBFEAAFB43FE63B6BE" box="[339,460,1199,1220]" italics="true" pageId="4" pageNumber="12">Lucanus niu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AC95783FFF0FFEBFE2CFB42FDA4B6B9" box="[469,523,1198,1219]" pageId="4" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
, male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="233136EAFFF0FFEBFEF0FAF7FB98B7FB" blockId="4.[227,1127,1306,1598]" pageId="4" pageNumber="12">Pronotum 1.6 times as wide as long, widest at middle. Anterior corners weakly acute; posterior corners obtuse. Surface feebly sheen, finely punctate, intervals finely and irregularly granulose, and with median longitudinal depression rather shallow. Prosternal apophysis strong, distinctly convex ventrally in middle of posterior part, and truncated at posterior margin.</paragraph>
<paragraph id="233136EAFFF0FFEBFEF0FA6AFD5BB7CD" blockId="4.[227,1127,1306,1598]" pageId="4" pageNumber="12">Scutellum linguiform, about 1.8 times as wide as long, rounded at apical margin. Surface feebly sheen, finely punctate, intervals microreticulate.</paragraph>
<paragraph id="233136EAFFF0FFEBFEF0FA50FE7BB472" blockId="4.[227,1127,1306,1598]" pageId="4" pageNumber="12">Elytra oval, 1.4 times as long as wide, widest around basal 4/9, 2.8 times as long as and 1.2 times as wide as pronotum. Humeri rounded. Surface sheen, finely and densely punctate, intervals microreticulate.</paragraph>
<paragraph id="233136EAFFF0FFE9FEF0F9E2FC0AB706" blockId="4.[227,1127,1306,1598]" lastBlockId="6.[227,1127,1327,1625]" lastPageId="6" lastPageNumber="14" pageId="4" pageNumber="12">Legs slender. Protibia with 5 (left, basal one tiny) or 4 (right, basal one tiny) teeth along outer margin; apex bifurcate with branches narrowly rounded at tips. Except apical spurs and spines, mesotibia with 3 (right) or 4 (left, basal one tiny) small lateral spines and metatibia with 3 lateral spines (basal one tiny). Surfaces finely punctate, intervals matt and finely granulose on femora and tibiae, feebly sheen and microreticulate on tarsi.</paragraph>
<caption id="77F16662FFF1FFEAFEF0F9FAFDC2B418" ID-DOI="http://doi.org/10.5281/zenodo.7616391" ID-Zenodo-Dep="7616391" httpUri="https://zenodo.org/record/7616391/files/figure.png" pageId="5" pageNumber="13" startId="5.[265,300,1558,1580]" targetBox="[351,1002,269,1546]" targetPageId="5">
<paragraph id="233136EAFFF1FFEAFEF0F9FAFDC2B418" blockId="5.[227,1127,1558,1635]" pageId="5" pageNumber="13">
Fig. 3. Male habitus of
<emphasis id="11FAEAF8FFF1FFEAFE05F9FAFD37B451" box="[508,664,1558,1579]" italics="true" pageId="5" pageNumber="13">
<taxonomicName id="E48E4D69FFF1FFEAFE05F9FAFDE2B451" authorityName="SCOPOLI" authorityYear="1763" box="[508,589,1558,1579]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="13" phylum="Arthropoda" rank="genus">Lucanus</taxonomicName>
vitalisi
</emphasis>
Pouillaude, 1913 from Yunnan, China. a–c – large size; d–f – moderate size; g–i – small size; (a, d, g – dorsal view; b, e, h – ventral view; c, f, I – lateral view). Scale bar = 1.0 cm.
</paragraph>
</caption>
<caption id="77F16662FFF2FFE9FEF0FB49FE47B777" ID-DOI="http://doi.org/10.5281/zenodo.7616393" ID-Zenodo-Dep="7616393" httpUri="https://zenodo.org/record/7616393/files/figure.png" pageId="6" pageNumber="14" startId="6.[265,300,1189,1211]" targetBox="[360,1002,271,1163]" targetPageId="6">
<paragraph id="233136EAFFF2FFE9FEF0FB49FE47B777" blockId="6.[227,1127,1189,1294]" pageId="6" pageNumber="14">
Fig. 4. Male genitalia of
<emphasis id="11FAEAF8FFF2FFE9FDF4FB4AFD05B6C0" box="[525,682,1189,1211]" italics="true" pageId="6" pageNumber="14">
<taxonomicName id="E48E4D69FFF2FFE9FDF4FB4AFDF1B6C1" authorityName="SCOPOLI" authorityYear="1763" box="[525,606,1190,1211]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="14" phylum="Arthropoda" rank="genus">Lucanus</taxonomicName>
vitalisi
</emphasis>
Pouillaude, 1913 from Yunnan, China. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h – dorsal view; b, c, f, g – ventral view; e, I – lateral view). Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="233136EAFFF2FFE8FEF0FA6EFD72B7C0" blockId="6.[227,1127,1327,1625]" lastBlockId="7.[227,1127,1389,1604]" lastPageId="7" lastPageNumber="15" pageId="6" pageNumber="14">
Male genitalia. Abdominal tergite VIII (
<figureCitation id="BBB52A6FFFF2FFE9FD60FA6EFD4FB7ED" box="[665,736,1410,1432]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="6" pageNumber="14">Fig. 2a</figureCitation>
) with subround lateral angles and with longitudinal, thin, membranous stripe in middle of basal 5/9; sternite VIII (
<figureCitation id="BBB52A6FFFF2FFE9FC18FA71FB84B7C9" box="[993,1067,1437,1459]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="6" pageNumber="14">Fig. 2b</figureCitation>
) with transverse, semielliptical, membranous area in middle of basal half. Abdominal sternite IX (
<figureCitation id="BBB52A6FFFF2FFE9FF13FA39FE80B790" box="[234,303,1493,1515]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="6" pageNumber="14">Fig. 2c</figureCitation>
) with apical expansion relatively slender, and with long, membranous, median stripe in apical 5/8; basal stalk not extended laterally. Aedeagus (
<figureCitation id="BBB52A6FFFF2FFE9FCE4FA1CFCF0B47C" box="[797,863,1520,1542]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="6" pageNumber="14">Fig. 2f</figureCitation>
) about 2.3 times as long as wide in ventral view. Basal piece (
<figureCitation id="BBB52A6FFFF2FFE9FDDFF9E0FDD1B458" box="[550,638,1548,1570]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="6" pageNumber="14">Fig. 2f–h</figureCitation>
) constricted in basal part, about 1.8 times as long as parameres, with a pair of sclerotized dorsal plates (
<figureCitation id="BBB52A6FFFF2FFE9FD2DF9CBFCB1B447" box="[724,798,1575,1597]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="6" pageNumber="14">Fig. 2h</figureCitation>
); ventral plate (
<figureCitation id="BBB52A6FFFF2FFE9FC43F9CBFBAFB446" box="[954,1024,1575,1597]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="6" pageNumber="14">
Fig.
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</figureCitation>
) bifurcate at apex of distal end, with relatively deep emargination. Paramere with basal process (
<figureCitation id="BBB52A6FFFF2FFE9FC0EF9AFFBF4B423" box="[1015,1115,1603,1625]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="6" pageNumber="14">
Fig.
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–h
</figureCitation>
); apex strongly upturned in lateral view (
<figureCitation id="BBB52A6FFFF3FFE8FD93FA81FD01B7F9" box="[618,686,1389,1411]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="7" pageNumber="15">Fig. 2i</figureCitation>
). Penis (
<figureCitation id="BBB52A6FFFF3FFE8FCFCFA81FCCEB7F9" box="[773,865,1389,1411]" captionStart="Fig" captionStartId="4.[265,300,1198,1220]" captionTargetBox="[323,1012,266,1159]" captionTargetId="figure-2@4.[302,1032,257,1166]" captionTargetPageId="4" captionText="Fig. 2. Lucanus niu sp. n., male genitalia of holotype. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h: – dorsal view; b, c, f, g: – ventral view; e, i: l– ateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616389" httpUri="https://zenodo.org/record/7616389/files/figure.png" pageId="7" pageNumber="15">Fig. 2f–h</figureCitation>
) moderately wide, weakly expanded near apex, 0.2 times shorter than parameres. Flagellum relatively short, about 2.2 times as long as parameres, apex distinctly enlarged.
</paragraph>
<caption id="77F16662FFF3FFE8FF1AFEEBFDA0B342" ID-Table-UUID="77F16662FFF3FFE8FF1AFEEBFDA0B342" httpUri="http://table.plazi.org/id/77F16662FFF3FFE8FF1AFEEBFDA0B342" pageId="7" pageNumber="15" startId="7.[227,282,263,285]" targetBox="[227,1123,350,1351]" targetIsTable="true" targetPageId="7">
<paragraph id="233136EAFFF3FFE8FF1AFEEBFDA0B342" blockId="7.[227,1127,263,313]" pageId="7" pageNumber="15">
Table 1. Selected important morphological differences between males of
<taxonomicName id="E48E4D69FFF3FFE8FC4BFEE4FB85B366" authority="Wang &amp; He &amp; He &amp; Zhou, 2023" authorityName="Wang &amp; He &amp; He &amp; Zhou" authorityYear="2023" box="[946,1066,263,285]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="15" phylum="Arthropoda" rank="species" species="niu" status="sp. nov.">
<emphasis id="11FAEAF8FFF3FFE8FC4BFEE4FB85B366" box="[946,1066,263,285]" italics="true" pageId="7" pageNumber="15">Lucanus niu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AC95783FFF3FFE8FBC8FEEBFBC8B366" box="[1073,1127,263,284]" pageId="7" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
and
<emphasis id="11FAEAF8FFF3FFE8FEF5FECFFEC9B342" box="[268,358,291,312]" italics="true" pageId="7" pageNumber="15">L. vitalisi</emphasis>
Pouillaude, 1913
</paragraph>
</caption>
<paragraph id="233136EAFFF3FFE8FF1AFEB2FC67B73D" pageId="7" pageNumber="15">
<table id="518EC44AFFF30010FF1AFEB2FBCCB73D" box="[227,1123,350,1351]" gridcols="3" gridrows="36" pageId="7" pageNumber="15">
<tr id="9DBE34A8FFF30010FF1AFEB2FBCCB30E" box="[227,1123,350,372]" gridrow="0" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFEB2FED1B30E" box="[227,382,350,372]" gridcol="0" gridrow="0" pageId="7" pageNumber="15">
<emphasis id="11FAEAF8FFF3FFE8FF1AFEB2FEF9B30E" bold="true" box="[227,342,350,372]" pageId="7" pageNumber="15">Characters</emphasis>
</th>
<th id="DE6F5DD4FFF30010FE62FEB2FD58B30E" box="[411,759,350,372]" gridcol="1" gridrow="0" pageId="7" pageNumber="15">
<emphasis id="11FAEAF8FFF3FFE8FE14FEB3FD08B30E" bold="true" box="[493,679,350,372]" pageId="7" pageNumber="15">
<taxonomicName id="E48E4D69FFF3FFE8FE14FEB3FDC5B309" authority="Wang &amp; He &amp; He &amp; Zhou, 2023" authorityName="Wang &amp; He &amp; He &amp; Zhou" authorityYear="2023" box="[493,618,350,372]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="15" phylum="Arthropoda" rank="species" species="niu" status="sp. nov.">
<emphasis id="11FAEAF8FFF3FFE8FE14FEB3FDC5B309" bold="true" box="[493,618,350,372]" italics="true" pageId="7" pageNumber="15">Lucanus niu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AC95783FFF3FFE8FD88FEB3FD08B30E" box="[625,679,351,372]" pageId="7" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<th id="DE6F5DD4FFF30010FCECFEB2FBCCB30E" box="[789,1123,350,372]" gridcol="2" gridrow="0" pageId="7" pageNumber="15">
<emphasis id="11FAEAF8FFF3FFE8FC94FEB3FBA4B30E" bold="true" box="[877,1035,350,372]" italics="true" pageId="7" pageNumber="15">
<taxonomicName id="E48E4D69FFF3FFE8FC94FEB3FC6BB30E" authorityName="SCOPOLI" authorityYear="1763" box="[877,964,351,372]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="15" phylum="Arthropoda" rank="genus">Lucanus</taxonomicName>
vitalisi
</emphasis>
</th>
</tr>
<tr id="9DBE34A8FFF30010FF1AFE97FBCCB3EB" box="[227,1123,379,401]" gridrow="1" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFE97FED1B3EB" box="[227,382,379,401]" gridcol="0" gridrow="1" pageId="7" pageNumber="15">Body length</th>
<td id="DE6F5DD4FFF30010FE62FE97FD58B3EB" box="[411,759,379,401]" gridcol="1" gridrow="1" pageId="7" pageNumber="15">large-sized male shorter than 60</td>
<td id="DE6F5DD4FFF30010FCECFE97FBCCB3EB" box="[789,1123,379,401]" gridcol="2" gridrow="1" pageId="7" pageNumber="15">large-sized male generally longer</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFE7BFBCCB3D7" box="[227,1123,407,429]" gridrow="2" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FE7BFD58B3D7" box="[411,759,407,429]" gridcol="1" gridrow="2" pageId="7" pageNumber="15">mm (Fig. 1a–c)</td>
<td id="DE6F5DD4FFF30010FCECFE7BFBCCB3D7" box="[789,1123,407,429]" gridcol="2" gridrow="2" pageId="7" pageNumber="15">than 70 mm (Fig. 3a–c)</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFE5FFBCCB3B3" box="[227,1123,435,457]" gridrow="3" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFE5FFED1B3B3" box="[227,382,435,457]" gridcol="0" gridrow="3" pageId="7" pageNumber="15">Color of femora</th>
<td id="DE6F5DD4FFF30010FE62FE5FFD58B3B3" box="[411,759,435,457]" gridcol="1" gridrow="3" pageId="7" pageNumber="15">reddish stripes distinct on ventral</td>
<td id="DE6F5DD4FFF30010FCECFE5FFBCCB3B3" box="[789,1123,435,457]" gridcol="2" gridrow="3" pageId="7" pageNumber="15">reddish stripes distinct on ventral</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFE23FBCCB39F" box="[227,1123,463,485]" gridrow="4" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FE23FD58B39F" box="[411,759,463,485]" gridcol="1" gridrow="4" pageId="7" pageNumber="15">surfaces in large-sized male (Fig.</td>
<td id="DE6F5DD4FFF30010FCECFE23FBCCB39F" box="[789,1123,463,485]" gridcol="2" gridrow="4" pageId="7" pageNumber="15">surfaces from large-sized to small-</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFE07FBCCB07B" box="[227,1123,491,513]" gridrow="5" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FE07FD58B07B" box="[411,759,491,513]" gridcol="1" gridrow="5" pageId="7" pageNumber="15">1b), but obscure in moderate-sized</td>
<td id="DE6F5DD4FFF30010FCECFE07FBCCB07B" box="[789,1123,491,513]" gridcol="2" gridrow="5" pageId="7" pageNumber="15">sized males (Fig. 3b, e, h)</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFDEAFBCCB066" box="[227,1123,518,540]" gridrow="6" pageId="7" pageNumber="15" rowspan-0="1" rowspan-2="1">
<td id="DE6F5DD4FFF30010FE62FDEAFD58B066" box="[411,759,518,540]" gridcol="1" gridrow="6" pageId="7" pageNumber="15">and small-sized males (Fig. 1e, h)</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFDCFFBCCB043" box="[227,1123,547,569]" gridrow="7" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFDCFFED1B043" box="[227,382,547,569]" gridcol="0" gridrow="7" pageId="7" pageNumber="15">Color of tibiae</th>
<td id="DE6F5DD4FFF30010FE62FDCFFD58B043" box="[411,759,547,569]" gridcol="1" gridrow="7" pageId="7" pageNumber="15">reddish stripes obscure on ventral</td>
<td id="DE6F5DD4FFF30010FCECFDCFFBCCB043" box="[789,1123,547,569]" gridcol="2" gridrow="7" pageId="7" pageNumber="15">reddish stripes distinct on ventral</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFDD3FBCCB02F" box="[227,1123,575,597]" gridrow="8" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FDD3FD58B02F" box="[411,759,575,597]" gridcol="1" gridrow="8" pageId="7" pageNumber="15">surfaces (Fig. 1b, e, h), and absent</td>
<td id="DE6F5DD4FFF30010FCECFDD3FBCCB02F" box="[789,1123,575,597]" gridcol="2" gridrow="8" pageId="7" pageNumber="15">surfaces (Fig. 3b, e, h), and obscure</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFDB6FBCCB00A" box="[227,1123,602,624]" gridrow="9" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FDB6FD58B00A" box="[411,759,602,624]" gridcol="1" gridrow="9" pageId="7" pageNumber="15">on dorsal surfaces (protibiae tinged</td>
<td id="DE6F5DD4FFF30010FCECFDB6FBCCB00A" box="[789,1123,602,624]" gridcol="2" gridrow="9" pageId="7" pageNumber="15">on dorsal surfaces (Fig. 3a, d, j)</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFD9AFBCCB0F6" box="[227,1123,630,652]" gridrow="10" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFD9AFBCCB0F6" box="[227,1123,630,652]" colspan="3" colspanRight="2" gridcol="0" gridrow="10" pageId="7" pageNumber="15">with reddish) (Fig. 1a, d, j)</th>
</tr>
<tr id="9DBE34A8FFF30010FF1AFD7FFBCCB0D3" box="[227,1123,659,681]" gridrow="11" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFD7FFED1B0D3" box="[227,382,659,681]" gridcol="0" gridrow="11" pageId="7" pageNumber="15">Mandibles</th>
<td id="DE6F5DD4FFF30010FE62FD7FFD58B0D3" box="[411,759,659,681]" gridcol="1" gridrow="11" pageId="7" pageNumber="15">about 2.3 times as long as head in</td>
<td id="DE6F5DD4FFF30010FCECFD7FFBCCB0D3" box="[789,1123,659,681]" gridcol="2" gridrow="11" pageId="7" pageNumber="15">about 3.0 times as long as head in</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFD43FBCCB0BF" box="[227,1123,687,709]" gridrow="12" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FD43FD58B0BF" box="[411,759,687,709]" gridcol="1" gridrow="12" pageId="7" pageNumber="15">large-sized male (Fig. 1a–c); inner</td>
<td id="DE6F5DD4FFF30010FCECFD43FBCCB0BF" box="[789,1123,687,709]" gridcol="2" gridrow="12" pageId="7" pageNumber="15">large-sized male (Fig. 3a–c); inner</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFD26FBCCB09A" box="[227,1123,714,736]" gridrow="13" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FD26FD58B09A" box="[411,759,714,736]" gridcol="1" gridrow="13" pageId="7" pageNumber="15">mandibular margin with unarmed</td>
<td id="DE6F5DD4FFF30010FCECFD26FBCCB09A" box="[789,1123,714,736]" gridcol="2" gridrow="13" pageId="7" pageNumber="15">mandibular margin with unarmed</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFD0AFBCCB086" box="[227,1123,742,764]" gridrow="14" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FD0AFD58B086" box="[411,759,742,764]" gridcol="1" gridrow="14" pageId="7" pageNumber="15">carina at base, preceded by diastema,</td>
<td id="DE6F5DD4FFF30010FCECFD0AFBCCB086" box="[789,1123,742,764]" gridcol="2" gridrow="14" pageId="7" pageNumber="15">carina at base, preceded by diaste-</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFCEDFBCCB16D" box="[227,1123,769,791]" gridrow="15" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FCEDFD58B16D" box="[411,759,769,791]" gridcol="1" gridrow="15" pageId="7" pageNumber="15">7–8 small denticles, major inner</td>
<td id="DE6F5DD4FFF30010FCECFCEDFBCCB16D" box="[789,1123,769,791]" gridcol="2" gridrow="15" pageId="7" pageNumber="15">ma, 17–18 small denticles, major</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFCF1FBCCB149" box="[227,1123,797,819]" gridrow="16" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FCF1FD58B149" box="[411,759,797,819]" gridcol="1" gridrow="16" pageId="7" pageNumber="15">tooth, and 2–3 distal small denticles</td>
<td id="DE6F5DD4FFF30010FCECFCF1FBCCB149" box="[789,1123,797,819]" gridcol="2" gridrow="16" pageId="7" pageNumber="15">inner tooth, and 7–8 distal small</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFCD4FBCCB134" box="[227,1123,824,846]" gridrow="17" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FCD4FD58B134" box="[411,759,824,846]" gridcol="1" gridrow="17" pageId="7" pageNumber="15">in large-sized male (Fig. 1a, b);</td>
<td id="DE6F5DD4FFF30010FCECFCD4FBCCB134" box="[789,1123,824,846]" gridcol="2" gridrow="17" pageId="7" pageNumber="15">denticles in large-sized male (Fig.</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFCB8FBCCB110" box="[227,1123,852,874]" gridrow="18" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FCB8FD58B110" box="[411,759,852,874]" gridcol="1" gridrow="18" pageId="7" pageNumber="15">major inner tooth relatively more</td>
<td id="DE6F5DD4FFF30010FCECFCB8FBCCB110" box="[789,1123,852,874]" gridcol="2" gridrow="18" pageId="7" pageNumber="15">3a, b); major inner tooth relatively</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFC9CFBCCB1FC" box="[227,1123,880,902]" gridrow="19" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FC9CFD58B1FC" box="[411,759,880,902]" gridcol="1" gridrow="19" pageId="7" pageNumber="15">prominent in large-sized male (Fig.</td>
<td id="DE6F5DD4FFF30010FCECFC9CFBCCB1FC" box="[789,1123,880,902]" gridcol="2" gridrow="19" pageId="7" pageNumber="15">less prominent in large-sized male</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFC67FBCCB1DB" box="[227,1123,907,929]" gridrow="20" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FC67FD58B1DB" box="[411,759,907,929]" gridcol="1" gridrow="20" pageId="7" pageNumber="15">1a, b); apical part thick and strong</td>
<td id="DE6F5DD4FFF30010FCECFC67FBCCB1DB" box="[789,1123,907,929]" gridcol="2" gridrow="20" pageId="7" pageNumber="15">(Fig. 3a, b); apical part much more</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFC4BFBCCB1C7" box="[227,1123,935,957]" gridrow="21" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FC4BFD58B1C7" box="[411,759,935,957]" gridcol="1" gridrow="21" pageId="7" pageNumber="15">in moderate-sized and small-sized</td>
<td id="DE6F5DD4FFF30010FCECFC4BFBCCB1C7" box="[789,1123,935,957]" gridcol="2" gridrow="21" pageId="7" pageNumber="15">slender in moderate-sized and</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFC2FFBCCB1A3" box="[227,1123,963,985]" gridrow="22" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FC2FFD58B1A3" box="[411,759,963,985]" gridcol="1" gridrow="22" pageId="7" pageNumber="15">males (Fig. 1d, e, g, h)</td>
<td id="DE6F5DD4FFF30010FCECFC2FFBCCB1A3" box="[789,1123,963,985]" gridcol="2" gridrow="22" pageId="7" pageNumber="15">small-sized males (Fig. 3d, e, g, h)</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFC33FBCCB18F" box="[227,1123,991,1013]" gridrow="23" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFC33FED1B18F" box="[227,382,991,1013]" gridcol="0" gridrow="23" pageId="7" pageNumber="15">Head: anterior</th>
<td id="DE6F5DD4FFF30010FE62FC33FD58B18F" box="[411,759,991,1013]" gridcol="1" gridrow="23" pageId="7" pageNumber="15">narrower; substraight at apical</td>
<td id="DE6F5DD4FFF30010FCECFC33FBCCB18F" box="[789,1123,991,1013]" gridcol="2" gridrow="23" pageId="7" pageNumber="15">broader; rounded at apical margin</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFC17FBCCB66B" box="[227,1123,1019,1041]" gridrow="24" pageId="7" pageNumber="15" rowspan-2="1">
<th id="DE6F5DD4FFF30010FF1AFC17FED1B66B" box="[227,382,1019,1041]" gridcol="0" gridrow="24" pageId="7" pageNumber="15">ridge</th>
<td id="DE6F5DD4FFF30010FE62FC17FD58B66B" box="[411,759,1019,1041]" gridcol="1" gridrow="24" pageId="7" pageNumber="15">margin</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFBF4FBCCB654" box="[227,1123,1048,1070]" gridrow="25" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFBF4FED1B654" box="[227,382,1048,1070]" gridcol="0" gridrow="25" pageId="7" pageNumber="15">Abdominal</th>
<td id="DE6F5DD4FFF30010FE62FBF4FD58B654" box="[411,759,1048,1070]" gridcol="1" gridrow="25" pageId="7" pageNumber="15">membranous area semielliptical</td>
<td id="DE6F5DD4FFF30010FCECFBF4FBCCB654" box="[789,1123,1048,1070]" gridcol="2" gridrow="25" pageId="7" pageNumber="15">membranous area trapezoidal</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFBDFFBCCB633" box="[227,1123,1075,1097]" gridrow="26" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFBDFFED1B633" box="[227,382,1075,1097]" gridcol="0" gridrow="26" pageId="7" pageNumber="15">sternite VIII</th>
<td id="DE6F5DD4FFF30010FE62FBDFFD58B633" box="[411,759,1075,1097]" gridcol="1" gridrow="26" pageId="7" pageNumber="15">(Fig. 2b)</td>
<td id="DE6F5DD4FFF30010FCECFBDFFBCCB633" box="[789,1123,1075,1097]" gridcol="2" gridrow="26" pageId="7" pageNumber="15">(Fig. 4b)</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFBBCFBCCB61C" box="[227,1123,1104,1126]" gridrow="27" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFBBCFED1B61C" box="[227,382,1104,1126]" gridcol="0" gridrow="27" pageId="7" pageNumber="15">Abdominal</th>
<td id="DE6F5DD4FFF30010FE62FBBCFD58B61C" box="[411,759,1104,1126]" gridcol="1" gridrow="27" pageId="7" pageNumber="15">apical expansion relatively slender,</td>
<td id="DE6F5DD4FFF30010FCECFBBCFBCCB61C" box="[789,1123,1104,1126]" gridcol="2" gridrow="27" pageId="7" pageNumber="15">apical expansion relatively broad,</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFB80FBCCB6F8" box="[227,1123,1132,1154]" gridrow="28" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFB80FED1B6F8" box="[227,382,1132,1154]" gridcol="0" gridrow="28" pageId="7" pageNumber="15">sternite IX</th>
<td id="DE6F5DD4FFF30010FE62FB80FD58B6F8" box="[411,759,1132,1154]" gridcol="1" gridrow="28" pageId="7" pageNumber="15">and with long, membranous,</td>
<td id="DE6F5DD4FFF30010FCECFB80FBCCB6F8" box="[789,1123,1132,1154]" gridcol="2" gridrow="28" pageId="7" pageNumber="15">
and with short, membranous,
<tr id="9DBE34A8FFF30010FF1AFB6BFBCCB6E4" box="[227,1123,1159,1182]" gridrow="29" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FB6BFD58B6E4" box="[411,759,1159,1182]" gridcol="1" gridrow="29" pageId="7" pageNumber="15">memedian stripe in apical 5/8 (Fig. 2c)</td>
<td id="DE6F5DD4FFF30010FCECFB6BFBCCB6E4" box="[789,1123,1159,1182]" gridcol="2" gridrow="29" pageId="7" pageNumber="15">dian stripe in apical 1/3 (Fig. 4c)</td>
</tr>
</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFB49FBCCB6C1" box="[227,1123,1189,1211]" gridrow="30" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFB49FED1B6C1" box="[227,382,1189,1211]" gridcol="0" gridrow="30" pageId="7" pageNumber="15">Basal piece</th>
<td id="DE6F5DD4FFF30010FE62FB49FD58B6C1" box="[411,759,1189,1211]" gridcol="1" gridrow="30" pageId="7" pageNumber="15">apex of distal end with relatively</td>
<td id="DE6F5DD4FFF30010FCECFB49FBCCB6C1" box="[789,1123,1189,1211]" gridcol="2" gridrow="30" pageId="7" pageNumber="15">apex of distal end with relatively</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFB2CFBCCB6AC" box="[227,1123,1216,1238]" gridrow="31" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FB2CFD58B6AC" box="[411,759,1216,1238]" gridcol="1" gridrow="31" pageId="7" pageNumber="15">deep emargination (Fig. 2g)</td>
<td id="DE6F5DD4FFF30010FCECFB2CFBCCB6AC" box="[789,1123,1216,1238]" gridcol="2" gridrow="31" pageId="7" pageNumber="15">shallow emargination (Fig. 4g)</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFB31FBCCB689" box="[227,1123,1245,1267]" gridrow="32" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFB31FED1B689" box="[227,382,1245,1267]" gridcol="0" gridrow="32" pageId="7" pageNumber="15">Parameres</th>
<td id="DE6F5DD4FFF30010FE62FB31FD58B689" box="[411,759,1245,1267]" gridcol="1" gridrow="32" pageId="7" pageNumber="15">apex strongly upturned in lateral</td>
<td id="DE6F5DD4FFF30010FCECFB31FBCCB689" box="[789,1123,1245,1267]" gridcol="2" gridrow="32" pageId="7" pageNumber="15">with apex less strongly upturned</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFB15FBCCB775" box="[227,1123,1273,1295]" gridrow="33" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FB15FD58B775" box="[411,759,1273,1295]" gridcol="1" gridrow="33" pageId="7" pageNumber="15">view (Fig. 2i)</td>
<td id="DE6F5DD4FFF30010FCECFB15FBCCB775" box="[789,1123,1273,1295]" gridcol="2" gridrow="33" pageId="7" pageNumber="15">in lateral view (Fig. 4i)</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFAFAFBCCB756" box="[227,1123,1302,1324]" gridrow="34" pageId="7" pageNumber="15">
<th id="DE6F5DD4FFF30010FF1AFAFAFED1B756" box="[227,382,1302,1324]" gridcol="0" gridrow="34" pageId="7" pageNumber="15">Penis</th>
<td id="DE6F5DD4FFF30010FE62FAFAFD58B756" box="[411,759,1302,1324]" gridcol="1" gridrow="34" pageId="7" pageNumber="15">moderately wide, weakly expanded</td>
<td id="DE6F5DD4FFF30010FCECFAFAFBCCB756" box="[789,1123,1302,1324]" gridcol="2" gridrow="34" pageId="7" pageNumber="15">rather wide, strongly expanded</td>
</tr>
<tr id="9DBE34A8FFF30010FF1AFADDFBCCB73D" box="[227,1123,1329,1351]" gridrow="35" pageId="7" pageNumber="15" rowspan-0="1">
<td id="DE6F5DD4FFF30010FE62FADDFD58B73D" box="[411,759,1329,1351]" gridcol="1" gridrow="35" pageId="7" pageNumber="15">near apex (Fig. 2f)</td>
<td id="DE6F5DD4FFF30010FCECFADDFBCCB73D" box="[789,1123,1329,1351]" gridcol="2" gridrow="35" pageId="7" pageNumber="15">near apex (Fig. 4f)</td>
</tr>
</table>
</paragraph>
<paragraph id="233136EAFFF3FFE8FEF0FA2CFBE4B43E" blockId="7.[227,1127,1389,1604]" pageId="7" pageNumber="15">
<collectionCode id="459FAE2FFFF3FFE8FEF0FA2CFEE2B7AF" box="[265,333,1472,1493]" pageId="7" pageNumber="15">MALE</collectionCode>
<typeStatus id="FC358848FFF3FFE8FEA1FA2CFE4AB7AF" box="[344,485,1472,1493]" pageId="7" pageNumber="15" type="paratype">PARATYPES</typeStatus>
. Body 43.5–58.0 mm long. All male
<typeStatus id="FC358848FFF3FFE8FC93FA2CFC69B7AF" box="[874,966,1472,1493]" pageId="7" pageNumber="15" type="paratype">paratypes</typeStatus>
consistent with
<typeStatus id="FC358848FFF3FFE8FF1AFA37FE98B78B" box="[227,311,1499,1521]" pageId="7" pageNumber="15" type="holotype">holotype</typeStatus>
except for usual allometric differences (
<figureCitation id="BBB52A6FFFF3FFE8FD3EFA37FC8CB78B" box="[711,803,1499,1521]" captionStart="Fig" captionStartId="3.[265,300,1556,1578]" captionTargetBox="[355,1006,271,1539]" captionTargetId="figure-0@3.[319,1036,257,1550]" captionTargetPageId="3" captionText="Fig. 1. Lucanus niu sp. n., male habitus. a–c – large size, holotype; d–f – moderate size, paratype; g–i – small size, paratype; (a, d, g – dorsal view; b, e, h – ventral view; c, f, i – lateral view). Scale bar = 1.0 cm." figureDoi="http://doi.org/10.5281/zenodo.7616387" httpUri="https://zenodo.org/record/7616387/files/figure.png" pageId="7" pageNumber="15">Fig. 1d–i</figureCitation>
). Large-sized male like
<typeStatus id="FC358848FFF3FFE8FBEAFA37FBC8B78B" box="[1043,1127,1499,1521]" pageId="7" pageNumber="15" type="holotype">holotype</typeStatus>
with more pronounced morphological characters, while in small-sized male, mandible strongly shorter, apical fork of mandible less opened to vestigial, number of inner teeth fewer, major inner tooth ill-developed, clypeolabrum shorter, anterior and lateral ridges of head weaker.
</paragraph>
<paragraph id="233136EAFFFCFFE7FEF0FEEBFE60B367" blockId="8.[227,1127,263,478]" box="[265,463,263,285]" pageId="8" pageNumber="16">
<pageTitle id="6311EE8DFFFCFFE7FEF0FEEBFE60B367" box="[265,463,263,285]" pageId="8" pageNumber="16">FEMALE. Unkown.</pageTitle>
</paragraph>
<paragraph id="233136EAFFFCFFE7FEF0FECFFBF8B315" blockId="8.[227,1127,263,478]" pageId="8" pageNumber="16">
<title id="518ADDDCFFFCFFE7FEF0FECFFDF6B32E" jopScore="0.12855114" pageId="8" pageNumber="16">
DIFFERENTIAL DIAGNOSIS. This new species should be assigned to the
<emphasis id="11FAEAF8FFFCFFE7FBECFECFFE94B329" italics="true" pageId="8" pageNumber="16">Lucanus laminifer</emphasis>
group (
<emphasis id="11FAEAF8FFFCFFE7FE71FED3FE12B32E" box="[392,445,319,340]" italics="true" pageId="8" pageNumber="16">sensu</emphasis>
<bibRefCitation id="471F4B1BFFFCFFE7FE3DFED3FD37B329" author="Huang, H. &amp; Chen, C. - C." box="[452,664,318,340]" pageId="8" pageNumber="16" refId="ref6248" refString="Huang, H. &amp; Chen, C. - C. 2010. Stag beetles of China I. Formosa Ecological Company, Taipei. 288 pp." type="book" year="2010">
Huang &amp; Chen,
<year id="572677B9FFFCFFE7FD9DFED2FD37B329" box="[612,664,318,339]" pageId="8" pageNumber="16">2010</year>
</bibRefCitation>
</title>
).
<journalOrPublisher id="F94F3C26FFFCFFE7FD50FED3FCFCB32E" box="[681,851,318,340]" jopScore="0.06666667" pageId="8" pageNumber="16" type="_" typeScore="4">It most resembles</journalOrPublisher>
<emphasis id="11FAEAF8FFFCFFE7FCA3FED3FC5CB329" box="[858,1011,318,340]" italics="true" pageId="8" pageNumber="16">
<author id="FB33DE0FFFFCFFE7FCA3FED3FC04B32E" box="[858,939,319,340]" fns="*" inm="true" npo="*" pageId="8" pageNumber="16">Lucanus</author>
vitalisi
</emphasis>
Pouillaude,
<year id="572677B9FFFCFFE7FF1AFEB6FEBDB315" box="[227,274,346,367]" pageId="8" pageNumber="16">1913</year>
(
<figureCitation id="BBB52A6FFFFCFFE7FEE7FEB6FEF5B315" box="[286,346,346,368]" captionStart="Fig" captionStartId="5.[265,300,1558,1580]" captionTargetBox="[351,1002,269,1546]" captionTargetId="figure-0@5.[327,1026,257,1553]" captionTargetPageId="5" captionText="Fig. 3. Male habitus of Lucanus vitalisi Pouillaude, 1913 from Yunnan, China. a–c – large size; d–f – moderate size; g–i – small size; (a, d, g – dorsal view; b, e, h – ventral view; c, f, I – lateral view). Scale bar = 1.0 cm." figureDoi="http://doi.org/10.5281/zenodo.7616391" httpUri="https://zenodo.org/record/7616391/files/figure.png" pageId="8" pageNumber="16">
<author id="FB33DE0FFFFCFFE7FEE7FEB6FEE7B30A" box="[286,328,346,368]" fns="*" inm="true" npo="*" pageId="8" pageNumber="16">Figs</author>
3
</figureCitation>
,
<figureCitation id="BBB52A6FFFFCFFE7FE9DFEB6FEDDB315" box="[356,370,346,367]" captionStart="Fig" captionStartId="6.[265,300,1189,1211]" captionTargetBox="[360,1002,271,1163]" captionTargetId="figure-0@6.[343,1012,257,1167]" captionTargetPageId="6" captionText="Fig. 4. Male genitalia of Lucanus vitalisi Pouillaude, 1913 from Yunnan, China. a – abdominal tergite VIII; b – abdominal sternite VIII; c–e – abdominal segment IX; f–i – aedeagus (penis removed in g); (a, d, h – dorsal view; b, c, f, g – ventral view; e, I – lateral view). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7616393" httpUri="https://zenodo.org/record/7616393/files/figure.png" pageId="8" pageNumber="16">4</figureCitation>
),
<title id="518ADDDCFFFCFFE7FE7AFEB6FBFCB315" box="[387,1107,346,368]" pageId="8" pageNumber="16">
but can be distinguished by the combination of characters shown in
<tableCitation id="6E0C0351FFFCFFE7FBF5FEB6FBFCB315" box="[1036,1107,346,368]" captionStart="Table 1" captionStartId="7.[227,282,263,285]" captionTargetBox="[227,1123,350,1351]" captionTargetId="graphics-27@7.[211,1137,343,1352]" captionTargetPageId="7" captionText="Table 1. Selected important morphological differences between males of Lucanus niu sp. n. and L. vitalisi Pouillaude, 1913" httpUri="http://table.plazi.org/id/77F16662FFF3FFE8FF1AFEEBFDA0B342" pageId="8" pageNumber="16" tableUuid="77F16662FFF3FFE8FF1AFEEBFDA0B342">Table 1</tableCitation>
</title>
.
</paragraph>
</subSubSection>
<subSubSection id="6B946561FFFCFFE7FEF0FE99FDF0B3F0" box="[265,607,373,395]" pageId="8" pageNumber="16" type="distribution">
<paragraph id="233136EAFFFCFFE7FEF0FE99FDF0B3F0" blockId="8.[227,1127,263,478]" box="[265,607,373,395]" pageId="8" pageNumber="16">
DISTRIBUTION.
<collectingCountry id="5B99767AFFFCFFE7FE44FE99FE59B3F1" box="[445,502,373,395]" name="China" pageId="8" pageNumber="16">China</collectingCountry>
(
<collectingRegion id="E14AF808FFFCFFE7FDFAFE99FDFBB3F0" box="[515,596,373,394]" country="China" name="Yunnan" pageId="8" pageNumber="16">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="6B946561FFFCFFE7FEF0FE7DFE32B3A4" pageId="8" pageNumber="16" type="etymology">
<paragraph id="233136EAFFFCFFE7FEF0FE7DFE32B3A4" blockId="8.[227,1127,263,478]" pageId="8" pageNumber="16">
<title id="518ADDDCFFFCFFE7FEF0FE7DFD77B3B8" jopScore="0.008381943" pageId="8" pageNumber="16">ETYMOLOGY. The specific epithet is from the Chinese Pinyin “niú [牛]”, which means ox, indicating this species was identified as new in</title>
<year id="572677B9FFFCFFE7FD19FE41FCBDB3B8" box="[736,786,429,450]" pageId="8" pageNumber="16">2021</year>
,
<journalOrPublisher id="F94F3C26FFFCFFE7FCE7FE40FC7FB3B8" box="[798,976,428,450]" jopScore="0.1534303" pageId="8" pageNumber="16" type="_" typeScore="4">the year of the ox</journalOrPublisher>
.
<author id="FB33DE0FFFFCFFE7FC22FE40FBAFB3B8" box="[987,1024,428,450]" fns="*" inm="true" npo="*" pageId="8" pageNumber="16">The</author>
name is a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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