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113 lines
14 KiB
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<document id="08C4A866328A1CCD758D94C3F175384F" ID-DOI="http://dx.doi.org/10.3897/zookeys.577.7254" ID-GBIF-Dataset="bad453df-1230-42fb-98f6-30fdcaa45f51" ID-PMC="PMC4829886" ID-Pensoft-Pub="1313-2970-577-125" ID-PubMed="27110191" ID-ZooBank="327F242512D544E28CD6E5BC91773077" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-577-125" ModsDocOrigin="ZooKeys 577" ModsDocTitle="Description of two new species of the Exocelinabroschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1459885117108" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo & Balke, Michael" docDate="2016" docId="93340D770ECFD2C5DCE11D18C3A75B8D" docLanguage="en" docName="ZooKeys 577: 125-148" docOrigin="ZooKeys 577" docSource="http://dx.doi.org/10.3897/zookeys.577.7254" docTitle="Exocelina marinae Shaverdo, Sagata & Balke 2005" docType="treatment" docVersion="5" lastPageNumber="130" masterDocId="FFECB328490DFFA1BA62B2070F4EC00F" masterDocTitle="Description of two new species of the Exocelinabroschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="148" masterPageNumber="125" pageNumber="130" updateTime="1701316882021" updateUser="plazi">
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<mods:title id="C02B0E55270D14C2A039D26F0AB02D56">Description of two new species of the Exocelinabroschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:namePart id="2A069339E9E8E91CB15B7E621B65D55D">Shaverdo, Helena</mods:namePart>
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<mods:namePart id="2967F86AA8F46CC6A6FA214D4262CB11">Sagata, Katayo</mods:namePart>
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<mods:name id="64AD35753FCFF3665E00EC60DF8B7589" type="personal">
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<mods:roleTerm id="EE8627CCA8DA4F5D18936EF1EF859AC1">Author</mods:roleTerm>
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<mods:namePart id="87FB544BFC8B6E95682072F8617CFD89">Balke, Michael</mods:namePart>
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<mods:typeOfResource id="A750C15C76F2AEED1CE88FE06A805EB2">text</mods:typeOfResource>
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<mods:title id="304E299DED9594B52AFA35CF993BBD3F">ZooKeys</mods:title>
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<mods:part id="BD4BD408D42727A68FD91FA18E927E6F">
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<mods:date id="1A5DCC4D2CCB5AF0D0BEC5D36B12B96B">2016</mods:date>
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<mods:detail id="59A69B1109A6A21DEB7899A84965F5A4" type="volume">
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<mods:number id="FD73C253198E7D7B4DA7344F6AFAD093">577</mods:number>
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<mods:start id="9CE7DA742307BE9FAA35370B55D344BC">125</mods:start>
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<mods:end id="542A716FBC6F7365C312060319937D10">148</mods:end>
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<mods:url id="39B9F92B5FFDA81E8E65F6446FC7BF69">http://dx.doi.org/10.3897/zookeys.577.7254</mods:url>
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<mods:classification id="3EB015341A298F14EFB9B13FE19C5357">journal article</mods:classification>
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<mods:identifier id="A6D5822FD89231F06075A051A3C6F2C5" type="DOI">http://dx.doi.org/10.3897/zookeys.577.7254</mods:identifier>
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<mods:identifier id="CA63254DF62461513EA7921621DBB6D0" type="Pensoft-Pub">1313-2970-577-125</mods:identifier>
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<mods:identifier id="F3C31CC5F0141D99D10DCE0956CA0605" type="ZooBank">327F242512D544E28CD6E5BC91773077</mods:identifier>
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<treatment id="93340D770ECFD2C5DCE11D18C3A75B8D" ID-GBIF-Taxon="127880283" LSID="urn:lsid:plazi:treatment:93340D770ECFD2C5DCE11D18C3A75B8D" httpUri="http://treatment.plazi.org/id/93340D770ECFD2C5DCE11D18C3A75B8D" lastPageNumber="130" pageId="5" pageNumber="130">
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<subSubSection id="54FD985A091590FDFC3273258088FE2C" pageId="5" pageNumber="130" type="multiple">
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<paragraph id="BA4A3C9556C5782D2924C24F6DC1269B" pageId="5" pageNumber="130">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="989AD55CAA1CAA82E81A09FAB4C429A1" pageId="5" pageNumber="130" type="nomenclature">
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<paragraph id="D4154F08566AFB5E3EA26501CFCF53EC" pageId="5" pageNumber="130">
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3.
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<taxonomicName id="371095983724B80AF25A4268D663CA6B" ID-CoL="3DLQK" authority="Shaverdo, Sagata & Balke, 2005" authorityName="Shaverdo, Sagata & Balke" authorityYear="2005" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina marinae" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="marinae">Exocelina marinae (Shaverdo, Sagata & Balke, 2005)</taxonomicName>
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Figs 5, 10
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</paragraph>
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</subSubSection>
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<subSubSection id="DFD01A07C3FAB2F4DCA10E83B5156332" pageId="5" pageNumber="130" type="reference_group">
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<paragraph id="E880939F3FC1C67F380737D23DF7630A" pageId="5" pageNumber="130">
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<taxonomicName id="D35D46D69A7C2DC33B4C350BF1504D04" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes marinae" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="marinae">Papuadytes marinae</taxonomicName>
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Shaverdo, Sagata & Balke 2005: 272.
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</paragraph>
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<paragraph id="D88E403E86139DB78CDC31F318E700D5" pageId="5" pageNumber="130">
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<taxonomicName id="D5A88D8D71C0614F2B55CB9A9C39F6E7" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina marinae" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="marinae">Exocelina marinae</taxonomicName>
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(Shaverdo, Sagata & Balke, 2005):
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<bibRefCitation id="1F876400849E19A8FF976E24758339E6" author="Nilsson, AN" journalOrPublisher="Latissimus" pageId="13" pageNumber="138" pagination="33 - 34" title="Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998." volume="23" year="2007">Nilsson 2007</bibRefCitation>
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: 34 (comb. n.);
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<bibRefCitation id="BBF2A595431E9D1544B326A6AADCCC9E" author="Toussaint, EFA" journalOrPublisher="Nature Communications" pageId="13" pageNumber="138" pagination="1 - 10" title="The towering orogeny of New Guinea as a trigger for arthropod megadiversity." url="10.1038/ncomms5001" volume="1" year="2014">Toussaint et al. 2014</bibRefCitation>
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: Supplementary figs 1-4, tab. 2 (
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<taxonomicName id="46BCCB53710E8DBF7BDF1DD46F63E7E4" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina marinae" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="marinae">Exocelina marinae</taxonomicName>
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MB1291).
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</paragraph>
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</subSubSection>
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<subSubSection id="5E6BC2A8E6EE37513DB6D1735501BFC9" pageId="5" pageNumber="130" type="type locality">
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<paragraph id="03A72A8076DADC1443AA056E278A5151" pageId="5" pageNumber="130">Type locality.</paragraph>
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<paragraph id="54E51EDAA7222D680214C10180AB3CA3" pageId="5" pageNumber="130">Papua New Guinea: Sandaun Province, trail from Telefomin to Eliptamin.</paragraph>
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</subSubSection>
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<subSubSection id="8F2C02BD1372B324E32C63BCD99B75CE" pageId="5" pageNumber="130" type="type material">
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<paragraph id="55AA20270B43955F38FBF1172A985DCD" pageId="5" pageNumber="130">Type material studied.</paragraph>
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<paragraph id="BB17728B5AB8D32E208612C5203594B6" pageId="5" pageNumber="130">
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<typeStatus id="ADF4423A4F98E593CE53967C7B3B0AB0" pageId="5" pageNumber="130">Holotype</typeStatus>
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: male "Papua N. G.: Sandaun Prov. Telefomin, 16.- 17.5.1998 trail to Eliptamin 1700-1800 m; leg. Riedel", "Holotypus
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<taxonomicName id="2490B5C9275643AA41422C2E681A648A" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes marinae" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="marinae">Papuadytes marinae</taxonomicName>
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sp.n. des. H. Shaverdo, K. Sagata & M. Balke, 2005" (NHMW).
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</paragraph>
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</subSubSection>
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<subSubSection id="9F8DB6F78A4FDB32B97915D7F9C36C81" pageId="5" pageNumber="130" type="materials_examined">
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<paragraph id="E0CF6A94D927A161DE353F46B4CAD8D4" pageId="5" pageNumber="130">Additional material.</paragraph>
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<paragraph id="AA655FCFB5FEA57F11E243E13C0B725F" pageId="5" pageNumber="130">
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Sandaun: 4 males, 6 females "Papua New Guinea: Sandaun, Mianmin, 670m 20.x.2008, 4.53.292S 141.34.118E, Ibalim (PNG 191)", two males with additional green labels "DNA M.Balke 3733" and "DNA M.Balke 3734" (NHMW, ZSM). 2 females "Papua New Guinea: Sandaun, Mianmin (river), 990m, 23.x.2008, 4.54.570S 141.35.490E, Ibalim (PNG 192)", one of them with an additional green label "DNA M.Balke 3737" (ZSM). 1 male, 1 female "Papua New Guinea: Sandaun, Mianmin (pool), 990m, 23.x.2008, 4.54.570S 141.35.490E, Ibalim (PNG 193)", the female with an additional green label "M.Balke 3777 DNA" (ZSM). 2 males, 2 females "Papua New Guinea: Sandaun, Mianmin (river), 1080m, 24.x.2008, 04.55.780S 141.38.185E, Ibalim (PNG 195)", one male and one female with additional green labels "DNA M.Balke 3742" and "DNA M.Balke 3741", respectively (NHMW, ZSM). 1 male, 5 females "Papua New Guinea: Sandaun, Mianmin (pool), 1080m, 24.x.2008, 04.55.780S 141.38.185E, Ibalim (PNG 196)" (NHMW, ZSM). 1 female "Papua New Guinea: Sandaun, Mianmin (river), 700m, 21.x.2008, 04.52.858S 141.31.706E, Ibalim (PNG 197)" (ZSM). 1 female "Papua New Guinea: Sandaun, Mianmin area,>1000m, 26.xii.2009, Ibalim & Pius (PNG233)" (ZSM). 1 male, 5 females "Papua New Guinea: Sandaun, Mianmin area,>1000m, 23.12.2009, Ibalim & Pius (PNG234)", the male with an additional label "DNA M.Balke 4934" (NHMW, ZSM). 8 males, 7 females "Papua New Guinea: Sandaun, Mianmin area,>600m, 13.i.2010, Ibalim & Pius (PNG235)", one male and one female with additional labels "DNA M.Balke 4938" and "DNA M.Balke 4931", respectively (NHMW, ZSM). 1 male "Papua New Guinea: Sandaun, Mianmin area,>600m, 13.i.2010, Ibalim & Pius (PNG236)" (ZSM). 1 male, 1 female "Papua New Guinea: Sandaun, Mianmin area,>700m, 14.i.2010,
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<geoCoordinate id="C961B00F6D0E5272604A3CF5727C1B2C" direction="south" orientation="latitude" precision="1" value="-0.08181667">0 4 54.540S</geoCoordinate>
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<geoCoordinate id="D86478F9BA0845E340BFD5321ADF12E3" direction="east" orientation="longitude" precision="1" value="141.61589">141 36.953E</geoCoordinate>
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, Ibalim & Pius (PNG238)", the female with an additional label "DNA M.Balke 4933" (ZSM).
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</paragraph>
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<paragraph id="329C30824A397646F63E13B8150E6099" pageId="5" pageNumber="130">Hela: 1 male "Papua New Guinea: Southern Highlands, Tari to Koroba, 1600m, 15.v.2006, 05.46.500S 142.50.000E, Balke (PNG 65)", "DNA M.Balke 1291" [green] (ZSM).</paragraph>
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</subSubSection>
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<subSubSection id="3E2D4A1E2A4779D02E62E7A50423CBBC" pageId="5" pageNumber="130" type="additions to the description">
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<paragraph id="593D71713902C7DF6482BE067412017B" pageId="5" pageNumber="130">Additions to the description</paragraph>
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<paragraph id="E47F16D2E8B3E09A96C5DF46BB64919B" pageId="5" pageNumber="130">
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(original description in
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<bibRefCitation id="C8A3E4FE4B840ECC7EC01EDBAECF3EB5" author="Shaverdo, HV" journalOrPublisher="Aquatic Insects" pageId="13" pageNumber="138" pagination="269 - 280" title="Five new species of the genus Papuadytes Balke, 1998 from New Guinea (Coleoptera: Dytiscidae)." url="10.1080/01650420500290169" volume="27" year="2005">Shaverdo et al. 2005</bibRefCitation>
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, p. 270). Male: Median lobe with apex slightly curved in lateral view and more or less rounded in ventral view. Its ventral sclerite with unequal apical lobes: left lobe very long (very often with broken apex) and right lobe short, relatively narrow (Fig. 10).
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</paragraph>
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<paragraph id="8473B44614DE28C702A760F9961FF18E" pageId="5" pageNumber="130">Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.</paragraph>
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<paragraph id="9F75AD0126FFA69A70B9FBDC053FE1C3" pageId="5" pageNumber="130">Variability: Beetles small to medium-sized: for Mianmin populations: TL-H 3.3-3.55 mm, TL 3.6-3.95 mm, MW 1.75-1.95 mm; for the holotype (Telefomin): TL-H 3.6 mm, TL 4.1 mm, MW 1.9 mm (Fig. 5); for the specimen from Tari-Koroba: TL-H 3.85 mm, TL 4.25 mm, MW 2.15 mm. Also beetles vary in shape of the median lobe and its ventral sclerite as shown in Fig. 10.</paragraph>
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</subSubSection>
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<subSubSection id="3D5928EBFAB0F278A1F642873DCF8B34" pageId="5" pageNumber="130" type="distribution">
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<paragraph id="AF40F8858DFDE85C81307DF6664C0515" pageId="5" pageNumber="130">Distribution.</paragraph>
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<paragraph id="1FD2F15FC714B9D5B36CD9C923D78FBF" pageId="5" pageNumber="130">Papua New Guinea: Sandaun and Hela Provinces (Fig. 14). This species is mainly known from the south of Sandaun Province: Mianmin area. Based on the record from Tari-Koroba, we assume that the species occurs also further southeast.</paragraph>
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</subSubSection>
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</treatment>
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</document> |