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<document ID-DOI="10.1206/0003-0082(2003)421&lt;0001:ANSOTR&gt;2.0.CO;2" ID-GBIF-Dataset="f4c2f80e-4917-4b5b-9415-0ee6377973ce" ID-ISSN="0003-0082" ID-Zenodo-Dep="4734917" approvalRequired="4" approvalRequired_for_taxonomicNames="4" checkinTime="1620068002453" checkinUser="felipe" docAuthor="VOSS, ROBERT S." docDate="2003" docId="03FAB267FFA3FFDEFF70FC51D884DB3D" docLanguage="en" docName="AmMusNovit.2003.3421.1-47.pdf" docOrigin="American Museum Novitates 3421" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0082%282003%29421%3C0001%3AANSOTR%3E2.0.CO%3B2" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.1:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="1" docTitle="Akodon mollis Thomas 1894" docType="treatment" docVersion="8" lastPageNumber="22" masterDocId="FFC3CA1FFFB6FFCBFF8AFF92DB29DF75" masterDocTitle="A New Species of Thomasomys (Rodentia: Muridae) from Eastern Ecuador, with Remarks on Mammalian Diversity and Biogeography in the Cordillera Oriental" masterLastPageNumber="48" masterPageNumber="1" pageNumber="22" updateTime="1671043263584" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A New Species of Thomasomys (Rodentia: Muridae) from Eastern Ecuador, with Remarks on Mammalian Diversity and Biogeography in the Cordillera Oriental</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>VOSS, ROBERT S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>American Museum Novitates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2003</mods:date>
<mods:detail type="pubDate">
<mods:number>2003-12-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3421</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>48</mods:end>
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<mods:url>http://www.bioone.org/doi/abs/10.1206/0003-0082%282003%29421%3C0001%3AANSOTR%3E2.0.CO%3B2</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1206/0003-0082(2003)421&lt;0001:ANSOTR&gt;2.0.CO;2</mods:identifier>
<mods:identifier type="GBIF-Dataset">f4c2f80e-4917-4b5b-9415-0ee6377973ce</mods:identifier>
<mods:identifier type="ISSN">0003-0082</mods:identifier>
<mods:identifier type="Zenodo-Dep">4734917</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5605917" ID-GBIF-Taxon="189547089" ID-Zenodo-Dep="5605917" LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D" httpUri="http://treatment.plazi.org/id/03FAB267FFA3FFDEFF70FC51D884DB3D" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection pageId="21" pageNumber="22" type="nomenclature">
<paragraph blockId="21.[250,524,963,986]" box="[250,524,963,986]" pageId="21" pageNumber="22">
<heading box="[250,524,963,986]" centered="true" fontSize="36" level="2" pageId="21" pageNumber="22" reason="8">
<taxonomicName authority="Thomas" authorityName="Thomas" authorityYear="1894" box="[250,524,963,986]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis box="[250,420,963,986]" italics="true" pageId="21" pageNumber="22">Akodon mollis</emphasis>
Thomas
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="21.[123,651,1014,1741]" pageId="21" pageNumber="22">
SPECIMENS COLLECTED: 12.5 km W (by road) Papallacta,
<quantity box="[326,432,1044,1067]" metricMagnitude="0" metricUnit="m" metricValue="4.120896" pageId="21" pageNumber="22" unit="ft" value="13.52">13,520 ft</quantity>
(UMMZ 155597); 10.6 km W (by road) Papallacta,
<quantity box="[541,651,1073,1096]" metricMagnitude="0" metricUnit="m" metricValue="3.84048" pageId="21" pageNumber="22" unit="ft" value="12.6">12,600 ft</quantity>
(UMMZ 155596, 155777); 7.5 km W (by road) Papallacta,
<quantity box="[327,433,1132,1155]" metricMagnitude="0" metricUnit="m" metricValue="3.6576" pageId="21" pageNumber="22" unit="ft" value="12.0">12,000 ft</quantity>
(UMMZ 155591 155595); 6.2 km W (by road) Papallacta,
<quantity box="[123,229,1190,1213]" metricMagnitude="0" metricUnit="m" metricValue="3.56616" pageId="21" pageNumber="22" unit="ft" value="11.7">11,700 ft</quantity>
(UMMZ 155586155590, 155778 155780).
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="description">
<paragraph blockId="21.[123,651,1014,1741]" pageId="21" pageNumber="22">
<materialsCitation ID-GBIF-Occurrence="3395013323" collectionCode="OTHER, MATERIAL" collectorName="L. Soderstrom " county="Forty-two" elevation="3" location="Three" municipality="Mt. Antisana" pageId="21" pageNumber="22" specimenCode="AMNH 47156-47164, AMNH 66450 -66491" specimenCount="1">
<collectionCode box="[152,356,1249,1272]" pageId="21" pageNumber="22">OTHER MATERIAL</collectionCode>
:
<location LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFEF2FB73DA95DB8D" box="[376,444,1249,1272]" county="Forty-two" municipality="Mt. Antisana" name="Three" pageId="21" pageNumber="22">Three</location>
additional specimens were collected at
<location LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFE2AFB6CDAD0DA60" box="[416,505,1278,1301]" county="Forty-two" municipality="Mt. Antisana" name="Tablon" pageId="21" pageNumber="22">Tablon</location>
, road to
<location LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFDEEFB6CDBC5DA46" county="Forty-two" municipality="Mt. Antisana" name="Papallacta'" pageId="21" pageNumber="22">Papallacta</location>
with recorded elevations of
<quantity box="[123,325,1337,1360]" metricMagnitude="0" metricUnit="m" metricValue="3.3528" metricValueMax="3.5052" metricValueMin="3.2003999999999997" pageId="21" pageNumber="22" unit="ft" value="11.0" valueMax="11.5" valueMin="10.5">
<elevation box="[123,325,1337,1360]" metricMagnitude="0" metricUnit="m" metricValue="3.3528" metricValueMax="3.5052" metricValueMin="3.2003999999999997" pageId="21" pageNumber="22" unit="ft" value="11.0" valueMax="11.5" valueMin="10.5">10,50011,500 ft</elevation>
</quantity>
[
<quantity box="[340,509,1337,1360]" metricMagnitude="3" metricUnit="m" metricValue="3.3525" metricValueMax="3.505" metricValueMin="3.2" pageId="21" pageNumber="22" unit="m" value="3352.5" valueMax="3505.0" valueMin="3200.0">
<elevation box="[340,509,1337,1360]" metricMagnitude="3" metricUnit="m" metricValue="3.3525" metricValueMax="3.505" metricValueMin="3.2" pageId="21" pageNumber="22" unit="m" value="3352.5" valueMax="3505.0" valueMin="3200.0">32003505 m</elevation>
</quantity>
] by
<collectorName pageId="21" pageNumber="22">L. Söderström</collectorName>
in 1903 and 1913.
<location LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFE62FAC4D96EDA18" box="[488,583,1366,1389]" county="Forty-two" municipality="Mt. Antisana" name="Another" pageId="21" pageNumber="22">Another</location>
<specimenCount pageId="21" pageNumber="22" type="generic">nine specimens</specimenCount>
(
<specimenCode box="[263,528,1396,1419]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH 4715647164</specimenCode>
) taken by the same collector in 1914 are labeled
<collectingMunicipality pageId="21" pageNumber="22">Mt. Antisana</collectingMunicipality>
12000 feet [
<quantity box="[407,510,1454,1477]" metricMagnitude="3" metricUnit="m" metricValue="3.658" pageId="21" pageNumber="22" unit="m" value="3658.0">
<elevation box="[407,510,1454,1477]" metricMagnitude="3" metricUnit="m" metricValue="3.658" pageId="21" pageNumber="22" unit="m" value="3658.0">3658 m</elevation>
</quantity>
].
<collectingCounty box="[535,651,1454,1477]" pageId="21" pageNumber="22">Forty­two</collectingCounty>
more specimens (
<specimenCode box="[356,642,1484,1507]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH 66450 66491</specimenCode>
) were collected by
<collectorName box="[342,506,1513,1536]" pageId="21" pageNumber="22">H.E. Anthony</collectorName>
and
<collectorName pageId="21" pageNumber="22">G.H.H. Tate</collectorName>
between 4115 and
<quantity box="[409,500,1542,1565]" metricMagnitude="3" metricUnit="m" metricValue="4.57" pageId="21" pageNumber="22" unit="m" value="4570.0">
<elevation box="[409,500,1542,1565]" metricMagnitude="3" metricUnit="m" metricValue="4.57" pageId="21" pageNumber="22" unit="m" value="4570.0">4570 m</elevation>
</quantity>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3395013309" collectionCode="AMNH" pageId="21" pageNumber="22" specimenCode="AMNH 67330, 67332, 67334" specimenCount="1">
on Antisana in 1923. Lastly,
<specimenCount box="[341,536,1572,1595]" pageId="21" pageNumber="22" type="generic">three specimens</specimenCount>
(
<specimenCode collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH 67330</specimenCode>
,
<specimenCode box="[215,291,1601,1624]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">67332</specimenCode>
,
<specimenCode box="[306,381,1601,1624]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">67334</specimenCode>
) collected in 1924 by
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3395013324" collectionCode="R" county="Papallacta'" elevation="4" location="Olalla" municipality="Mt. Guamani'" pageId="21" pageNumber="22" specimenCount="1">
<collectionCode box="[123,146,1630,1653]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="22">R</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:03FAB267FFA3FFDEFF70FC51D884DB3D:8E8C55AAFFA3FFDEFF15F9CCDBCFD900" box="[159,230,1630,1653]" county="Papallacta'" municipality="Mt. Guamani'" name="Olalla" pageId="21" pageNumber="22">Olalla</location>
are labeled
<collectingMunicipality box="[390,555,1630,1653]" pageId="21" pageNumber="22">Mt. Guamanı´</collectingMunicipality>
, road to
<collectingCounty box="[123,267,1660,1683]" pageId="21" pageNumber="22">Papallacta</collectingCounty>
with recorded elevations of
<quantity box="[123,327,1689,1712]" metricMagnitude="0" metricUnit="m" metricValue="3.81" metricValueMax="3.9624" metricValueMin="3.6576" pageId="21" pageNumber="22" unit="ft" value="12.5" valueMax="13.0" valueMin="12.0">
<elevation box="[123,327,1689,1712]" metricMagnitude="0" metricUnit="m" metricValue="3.81" metricValueMax="3.9624" metricValueMin="3.6576" pageId="21" pageNumber="22" unit="ft" value="12.5" valueMax="13.0" valueMin="12.0">12,00013,000 ft</elevation>
</quantity>
[
<quantity box="[343,515,1689,1712]" metricMagnitude="3" metricUnit="m" metricValue="3.81" metricValueMax="3.9619999999999997" metricValueMin="3.658" pageId="21" pageNumber="22" unit="m" value="3810.0" valueMax="3962.0" valueMin="3658.0">
<elevation box="[343,515,1689,1712]" metricMagnitude="3" metricUnit="m" metricValue="3.81" metricValueMax="3.9619999999999997" metricValueMin="3.658" pageId="21" pageNumber="22" unit="m" value="3810.0" valueMax="3962.0" valueMin="3658.0">36583962 m</elevation>
</quantity>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="21.[123,651,1014,1741]" lastBlockId="21.[699,1228,193,1096]" pageId="21" pageNumber="22">
TAXONOMY: The Papallacta material closely resembles the
<typeStatus box="[911,961,193,216]" pageId="21" pageNumber="22">type</typeStatus>
of
<taxonomicName authorityName="Thomas" authorityYear="1913" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="subSpecies" species="mollis" subSpecies="altorum">
<emphasis italics="true" pageId="21" pageNumber="22">Akodon mollis altorum</emphasis>
</taxonomicName>
as originally described by
<bibRefCitation author="Thomas, O." pageId="21" pageNumber="22" pagination="404 - 409" refId="ref25296" refString="Thomas, O. 1913. New forms of Akodon and Phyllotis, and a new genus for '' Akodon '' teguina. Annals and Magazine of Natural History 8 (11): 404 - 409." type="journal article" year="1913">Thomas (1913)</bibRefCitation>
based on a specimen collected at
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="21" pageNumber="22" unit="m" value="2600.0">2600 m</quantity>
in the Ecuadorean province of
<collectingRegion box="[1107,1178,281,304]" country="Ecuador" name="Canar" pageId="21" pageNumber="22">Cañar</collectingRegion>
. No substantive analysis of character data, however, is available to support the current hypothesis (
<bibRefCitation author="Cabrera, A." box="[814,987,369,392]" pageId="21" pageNumber="22" pagination="309 - 732" refId="ref21669" refString="Cabrera, A. 1961. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' (Ciencias Zoologicas) 4 (2): 309 - 732." type="journal article" year="1961">Cabrera, 1961</bibRefCitation>
;
<bibRefCitation author="Musser, G. G. &amp; M. D. Carleton" pageId="21" pageNumber="22" pagination="501 - 755" refId="ref23955" refString="Musser, G. G., and M. D. Carleton. 1993. Family Muridae. In D. E. Wilson and D. M. Reeder (editors), Mammal species of the world, 2 nd ed.: 501 - 755. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1993">Musser and Carleton, 1993</bibRefCitation>
) that
<emphasis box="[884,976,399,422]" italics="true" pageId="21" pageNumber="22">altorum</emphasis>
, a highland taxon, is really conspecific with the geographically adjacent lowland forms
<taxonomicName authority="Thomas (1894)" authorityName="Thomas" authorityYear="1894" box="[963,1227,457,480]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis box="[963,1034,457,480]" italics="true" pageId="21" pageNumber="22">mollis</emphasis>
<bibRefCitation author="Thomas, O." box="[1045,1227,457,480]" pageId="21" pageNumber="22" pagination="346 - 366" refId="ref24959" refString="Thomas, O. 1894. Descriptions of some new Neotropical Muridae. Annals and Magazine of Natural History 6 (14): 346 - 366." type="journal article" year="1894">Thomas (1894)</bibRefCitation>
</taxonomicName>
and
<taxonomicName authority="Hershkovitz (1940)" authorityName="Hershkovitz" authorityYear="1940" box="[750,1110,486,510]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="fulvescens">
<emphasis box="[750,869,487,510]" italics="true" pageId="21" pageNumber="22">fulvescens</emphasis>
<bibRefCitation author="Hershkovitz, P." box="[878,1110,486,509]" pageId="21" pageNumber="22" pagination="1 - 3" refId="ref22833" refString="Hershkovitz, P. 1940. Notes on the distribution of the akodont rodent, Akodon mollis, in Ecuador with a description of a new race. Occasional Papers of the Museum of Zoology, University of Michigan 418: 1 - 3." type="journal article" year="1940">Hershkovitz (1940)</bibRefCitation>
</taxonomicName>
. As noted by
<bibRefCitation author="Myers, P. &amp; J. L. Patton" box="[739,1042,516,539]" pageId="21" pageNumber="22" pagination="1 - 35" refId="ref24077" refString="Myers, P., and J. L. Patton. 1989. Akodon of Peru and Bolivia - revision of the fumeus group (Rodentia: Sigmodontinae). Occasional Papers of the Museum of Zoology, University of Michigan 721: 1 - 35." type="journal article" year="1989">Myers and Patton (1989)</bibRefCitation>
,
<taxonomicName authorityName="Thomas" authorityYear="1894" box="[1057,1227,516,539]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis box="[1057,1227,516,539]" italics="true" pageId="21" pageNumber="22">Akodon mollis</emphasis>
</taxonomicName>
as currently recognized has a very large geographic range and exhibits substantial geographic variation. Given their conclusion that
<emphasis box="[699,781,633,656]" italics="true" pageId="21" pageNumber="22">fumeus</emphasis>
<bibRefCitation author="Thomas, O." box="[794,974,633,656]" pageId="21" pageNumber="22" pagination="125 - 143" refId="ref25218" refString="Thomas, O. 1902. On mammals from Cochabamba, Bolivia, and the region north of that place. Annals and Magazine of Natural History 7 (9): 125 - 143." type="journal article" year="1902">Thomas (1902)</bibRefCitation>
—another taxon formerly ranked as a subspecies or synonym of
<taxonomicName authorityName="Thomas" authorityYear="1894" box="[699,770,692,715]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis box="[699,770,692,715]" italics="true" pageId="21" pageNumber="22">mollis</emphasis>
</taxonomicName>
—is a valid species, a comprehensive revision of this complex is long overdue.
</paragraph>
<paragraph blockId="21.[699,1228,193,1096]" pageId="21" pageNumber="22">
<materialsCitation ID-GBIF-Occurrence="3395013329" collectionCode="FIELD, OBSERVATIONS" elevation="3880" pageId="21" pageNumber="22" specimenCount="16">
<collectionCode box="[728,968,750,773]" pageId="21" pageNumber="22">FIELD OBSERVATIONS</collectionCode>
: The
<specimenCount box="[1037,1195,750,773]" pageId="21" pageNumber="22" type="generic">16 specimens</specimenCount>
of
<taxonomicName authorityName="Thomas" authorityYear="1894" box="[699,868,780,803]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mollis">
<emphasis box="[699,868,780,803]" italics="true" pageId="21" pageNumber="22">Akodon mollis</emphasis>
</taxonomicName>
that I collected near Papallacta in 1980 were trapped at elevations ranging from
<quantity box="[764,954,838,861]" metricMagnitude="3" metricUnit="m" metricValue="3.88" metricValueMax="4.16" metricValueMin="3.6" pageId="21" pageNumber="22" unit="m" value="3880.0" valueMax="4160.0" valueMin="3600.0">
<elevation box="[764,954,838,861]" metricMagnitude="3" metricUnit="m" metricValue="3.88" metricValueMax="4.16" metricValueMin="3.6" pageId="21" pageNumber="22" unit="m" value="3880.0" valueMax="4160.0" valueMin="3600.0">3600 to 4160 m</elevation>
</quantity>
. Of these, 13 were taken in the shrubby páramo/forest ecotone, and
<quantity box="[699,751,897,920]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="21" pageNumber="22" unit="in" value="3.0">3 in</quantity>
grassy páramo. All recorded captures were on the ground:
<quantity box="[951,998,926,949]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="21" pageNumber="22" unit="in" value="5.0">5 in</quantity>
tunnels among the bases of tall bunch grass, 5 among wet litter under mossy shrubs,
<quantity box="[962,1012,985,1008]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="21" pageNumber="22" unit="in" value="4.0">4 in</quantity>
runways through mixed bunch grass and bushes,
<quantity box="[1080,1127,1014,1037]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="21" pageNumber="22" unit="in" value="1.0">1 in</quantity>
a rabbit trail beneath low herbaceous cover, and 1 beneath an earth bank
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>