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<document id="8E94A7D36CB196AD9DFFBC19E58CB247" ID-DOI="10.5169/seals-985865" ID-ISSN="1662-8500" ID-Zenodo-Dep="8072390" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" approvalRequired="1" approvalRequired_for_originalDoi="1" checkinTime="1638113320454" checkinUser="carolina" docAuthor="Freitag, Anne &amp; Cherix, Daniel" docDate="2019" docId="03C14B04FFA2D515F6966412FDEF3741" docLanguage="fr" docName="EntomoHelv.12.99-110.pdf.imf" docOrigin="Entomo Helvetica 12" docStyle="DocumentStyle:219A02D731FC19F8B159E43A72E20048.2:EntomoHelv.2012-.journal_article" docStyleId="219A02D731FC19F8B159E43A72E20048" docStyleName="EntomoHelv.2012-.journal_article" docStyleVersion="2" docTitle="Tapinoma Forster 1850" docType="treatment" docVersion="2" lastPageNumber="101" masterDocId="FFF8337CFFA3D516F6066219FFF73658" masterDocTitle="Tapinoma magnum Mayr, 1861, une nouvelle espèce de fourmi introduite en Suisse (Hymenoptera, Formicidae)" masterLastPageNumber="110" masterPageNumber="99" pageNumber="100" updateTime="1687479536007" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="0DBD5250F7AFBA77D7F8247DF3F4240A">Tapinoma magnum Mayr, 1861, une nouvelle espèce de fourmi introduite en Suisse (Hymenoptera, Formicidae)</mods:title>
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<mods:namePart id="BDE1774CB36682211FF174F99180E0E2">Freitag, Anne</mods:namePart>
<mods:affiliation id="D27CEAC47798DD20C460B1AAF51957CA">MusÉe cantonal de zoologie, Pl. de la Riponne 6, CH- 1005 Lausanne;</mods:affiliation>
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<mods:namePart id="53B3043BA9EEC8CA55B85C41769851AE">Cherix, Daniel</mods:namePart>
<mods:affiliation id="A59121149E2577C72DCB473F980B375D">DÉpartement d’Ecologie et d’Evolution, Biophore, UniversitÉ de Lausanne, CH- 1015 Lausanne;</mods:affiliation>
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<mods:identifier id="EA66036BBD03A3C104FD5CEA1FB4F828" type="DOI">10.5169/seals-985865</mods:identifier>
<mods:identifier id="04FA2B7D9089277312E9529323AC6F50" type="ISSN">1662-8500</mods:identifier>
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<taxonomicName id="4C688191FFA2D517F6966412FDFF307D" ID-CoL="7SPC" ID-ENA="29038" authorityName="Forster" authorityYear="1850" box="[144,520,1547,1573]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="B91C2600FFA2D517F6966412FE8E307D" bold="true" box="[144,377,1547,1573]" italics="true" pageId="1" pageNumber="100">Tapinoma magnum</emphasis>
Mayr, 1861
</taxonomicName>
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(
<taxonomicName id="4C688191FFA2D517F4116412FD50307D" box="[535,679,1547,1573]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
, Dolichoderinae)
</paragraph>
<paragraph id="8BD7FA12FFA2D514F6966437FD5E3704" blockId="1.[144,1127,1547,1676]" lastBlockId="2.[151,1134,152,519]" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100">
<taxonomicName id="4C688191FFA2D517F6966437FE91301F" authorityName="Forster" authorityYear="1850" box="[144,358,1582,1607]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="B91C2600FFA2D517F6966437FE91301F" box="[144,358,1582,1607]" italics="true" pageId="1" pageNumber="100">Tapinoma magnum</emphasis>
</taxonomicName>
appartient au complexe d’espèces connu sous le nom
<taxonomicName id="4C688191FFA2D517F5C16437FEA13032" authority="(Nylander, 1856)" baseAuthorityName="Nylander" baseAuthorityYear="1856" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="nigerrimum">
<emphasis id="B91C2600FFA2D517F5C16437FB91301F" box="[967,1126,1582,1607]" italics="true" pageId="1" pageNumber="100">T. nigerrimum</emphasis>
(Nylander, 1856)
</taxonomicName>
qui comprend quatre espèces distinctes à distribution essentiellement méditerranéenne (
<bibRefCitation id="EFF987E3FFA2D517F7A46468FD8530D3" author="Seifert" box="[418,626,1649,1676]" etAl="et al." firstAuthor="Seifert" pageId="1" pageNumber="100" pagination="123 - 144" refId="ref6591" refString="Seifert B., d'Eustacchio D., Kaufmann B., Centorame M., Lorite P. &amp; Modica M. V. 2017. Four species within the supercolonial ants of the Tapinoma nigerrimum complex revealed by integrative taxonomy (Hymenoptera: Formicidae). Myrmecological News 24: 123 - 144." type="journal article" year="2017">Seifert et al. 2017</bibRefCitation>
). Jusqu’à la révision de
<bibRefCitation id="EFF987E3FFA2D517F5856468FB9530D3" author="Seifert" box="[899,1122,1649,1676]" etAl="et al." firstAuthor="Seifert" pageId="1" pageNumber="100" pagination="123 - 144" refId="ref6591" refString="Seifert B., d'Eustacchio D., Kaufmann B., Centorame M., Lorite P. &amp; Modica M. V. 2017. Four species within the supercolonial ants of the Tapinoma nigerrimum complex revealed by integrative taxonomy (Hymenoptera: Formicidae). Myrmecological News 24: 123 - 144." type="journal article" year="2017">Seifert et al. (2017)</bibRefCitation>
, les différentes espèces étaient très souvent confondues et la littérature antérieure à 2017 traitant de
<taxonomicName id="4C688191FFA1D514F75262A2FE0C368C" baseAuthorityName="Nylander" baseAuthorityYear="1856" box="[340,507,187,212]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="nigerrimum">
<emphasis id="B91C2600FFA1D514F75262A2FE0C368C" box="[340,507,187,212]" italics="true" pageId="2" pageNumber="101">T. nigerrimum</emphasis>
</taxonomicName>
concerne potentiellement l’une ou l’autre des espèces de ce groupe. Celles-ci sont difficiles à distinguer entre elles (
<bibRefCitation id="EFF987E3FFA1D514F55E62C5FBD236AE" author="Seifert" box="[856,1061,220,247]" etAl="et al." firstAuthor="Seifert" pageId="2" pageNumber="101" pagination="123 - 144" refId="ref6591" refString="Seifert B., d'Eustacchio D., Kaufmann B., Centorame M., Lorite P. &amp; Modica M. V. 2017. Four species within the supercolonial ants of the Tapinoma nigerrimum complex revealed by integrative taxonomy (Hymenoptera: Formicidae). Myrmecological News 24: 123 - 144." type="journal article" year="2017">Seifert et al. 2017</bibRefCitation>
), mais elles se diffÉrencient des autres
<taxonomicName id="4C688191FFA1D514F41762E6FD733740" authorityName="Forster" authorityYear="1850" box="[529,644,255,280]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="B91C2600FFA1D514F41762E6FD733740" box="[529,644,255,280]" italics="true" pageId="2" pageNumber="101">Tapinoma</emphasis>
</taxonomicName>
europÉennes entre autres par un très fort polymorphisme des ouvrières (
<figureCitation id="1353E697FFA1D514F4036339FDBF3762" box="[517,584,288,314]" captionStart="Fig" captionStartId="2.[659,692,1600,1622]" captionTargetBox="[659,1134,546,1579]" captionTargetId="figure-163@2.[656,1137,543,1579]" captionTargetPageId="2" captionText="Fig. 1. Ouvrières de Tapinoma magnum, montrant le fort polymorphisme chez cette espèce; Cully, août 2017. (Photo Marion Podolak)" figureDoi="http://doi.org/10.5281/zenodo.8072392" httpUri="https://zenodo.org/record/8072392/files/figure.png" pageId="2" pageNumber="101">Fig. 1</figureCitation>
), les plus grands individus possÉdant une tête dont la largeur dÉpasse
<quantity id="4C9057F7FFA1D514F7A1635BFE093705" box="[423,510,322,349]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="101" unit="mm" value="0.9">0.9 mm</quantity>
(
<bibRefCitation id="EFF987E3FFA1D514F40B635BFD6B3704" author="Seifert" box="[525,668,322,348]" firstAuthor="Seifert" pageId="2" pageNumber="101" pagination="139 - 147" refId="ref6543" refString="Seifert B. 2012. Clarifying naming and identification of the outdoor species of the ant genus Tapinoma Forster, 1850 (Hymenoptera: Formicidae) in Europe north of the Mediterranean region with description of a new species. Myrmecological News 16: 139 - 147." type="journal article" year="2012">Seifert 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8BD7FA12FFA1D514F6D2637CFDC13233" blockId="2.[151,1134,152,519]" lastBlockId="2.[151,627,526,1676]" pageId="2" pageNumber="101">
<taxonomicName id="4C688191FFA1D514F6D2637CFE423726" authorityName="Forster" authorityYear="1850" box="[212,437,357,382]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="B91C2600FFA1D514F6D2637CFE423726" box="[212,437,357,382]" italics="true" pageId="2" pageNumber="101">Tapinoma magnum</emphasis>
</taxonomicName>
est prÉsente sur une grande partie du pourtour mÉditerranéen, en Afrique du Nord, en
<collectingCountry id="F37FBA82FFA1D514F7EC639FFDDF37F8" box="[490,552,390,416]" name="Italy" pageId="2" pageNumber="101">Italie</collectingCountry>
, en
<collectingCountry id="F37FBA82FFA1D514F452639FFD6D37F8" box="[596,666,390,416]" name="France" pageId="2" pageNumber="101">Corse</collectingCountry>
, en Sardaigne et dans le Sud de la
<collectingCountry id="F37FBA82FFA1D514F226639EFB9937F9" box="[1056,1134,391,417]" name="France" pageId="2" pageNumber="101">France</collectingCountry>
(
<bibRefCitation id="EFF987E3FFA1D514F69963B1FE9C379A" author="Seifert" box="[159,363,424,451]" etAl="et al." firstAuthor="Seifert" pageId="2" pageNumber="101" pagination="123 - 144" refId="ref6591" refString="Seifert B., d'Eustacchio D., Kaufmann B., Centorame M., Lorite P. &amp; Modica M. V. 2017. Four species within the supercolonial ants of the Tapinoma nigerrimum complex revealed by integrative taxonomy (Hymenoptera: Formicidae). Myrmecological News 24: 123 - 144." type="journal article" year="2017">Seifert et al. 2017</bibRefCitation>
). Depuis les années 2000, elle est observée plus au nord en Europe, dans des zones urbaines ou suburbaines où elle est devenue envahissante (
<bibRefCitation id="EFF987E3FFA1D514F5E563D3FF23345E" author="Seifert" etAl="et al." firstAuthor="Seifert" pageId="2" pageNumber="101" pagination="123 - 144" refId="ref6591" refString="Seifert B., d'Eustacchio D., Kaufmann B., Centorame M., Lorite P. &amp; Modica M. V. 2017. Four species within the supercolonial ants of the Tapinoma nigerrimum complex revealed by integrative taxonomy (Hymenoptera: Formicidae). Myrmecological News 24: 123 - 144." type="journal article" year="2017">Seifert et al. 2017</bibRefCitation>
).
<bibRefCitation id="EFF987E3FFA1D514F6EA63F5FDB2345E" author="Lenoir &amp; Galkowski" box="[236,581,492,518]" firstAuthor="Lenoir" pageId="2" pageNumber="101" pagination="449 - 453" refId="ref6085" refString="Lenoir L. &amp; Galkowski C. 2017. Sur la prEsence d'une fourmi envahissante (Tapinoma magnum) dans le Sud-Ouest de la France. Bulletin Societe Linneenne Bordeaux 45 (4): 449 - 453." type="journal article" year="2017">Lenoir &amp; Galkowski (2017)</bibRefCitation>
l’ont observée en 2008 dans la banlieue de Bordeaux, puis en 2017 dans deux villes des Pyrénées-Atlantiques.
<bibRefCitation id="EFF987E3FFA1D514F7DE6029FD853412" author="Heller" box="[472,626,560,586]" firstAuthor="Heller" pageId="2" pageNumber="101" pagination="273 - 281" refId="ref5910" refString="Heller G. 2011. Pheidole pallidula (Nylander, 1849) und Tapinoma nigerrimum (Nylander, 1886) (Hymenoptera Formicidae), etablierte Neozoa in Rheinland-Pfalz. Mainzer naturwissenschaftliches Archiv 48: 273 - 281." type="journal article" year="2011">Heller (2011)</bibRefCitation>
la signale en
<collectingCountry id="F37FBA82FFA1D514F723604BFE683434" box="[293,415,594,620]" name="Germany" pageId="2" pageNumber="101">Allemagne</collectingCountry>
dans le Rheinland-Pfalz dès 2009. En 2013, elle est découverte aux
<collectingCountry id="F37FBA82FFA1D514F71F608EFE7434E9" box="[281,387,663,689]" name="Netherlands" pageId="2" pageNumber="101">Pays-Bas</collectingCountry>
à Wageningen par
<bibRefCitation id="EFF987E3FFA1D514F69160A1FEB9348B" author="Noordijk" box="[151,334,696,723]" firstAuthor="Noordijk" pageId="2" pageNumber="101" pagination="86 - 93" refId="ref6274" refString="Noordijk A. 2016. Leefwijze van Tapinoma nigerrimum (Hymenoptera: Formicidae), een nieuwe exotische mier in Nederland. Entomologische Berichten 76 (3): 86 - 93." type="journal article" year="2016">Noordijk (2016)</bibRefCitation>
, puis en 2014 en
<collectingCountry id="F37FBA82FFA1D514F40A60A1FD84348A" box="[524,627,696,722]" name="Belgium" pageId="2" pageNumber="101">Belgique</collectingCountry>
à Ostende par
<bibRefCitation id="EFF987E3FFA1D514F74860C3FD9834AD" author="Dekoninck" box="[334,623,730,757]" etAl="et al." firstAuthor="Dekoninck" pageId="2" pageNumber="101" pagination="206 - 209" refId="ref5739" refString="Dekoninck W., Parmentier T. &amp; Seifert B. 2015. First records of a supercolonial species of the Tapinoma nigerrimum c omplex in Belgium (Hymenoptera: Formicidae). Bulletin de la societe royale belge d'Entomologie 151: 206 - 209." type="journal article" year="2015">Dekoninck et al. (2015)</bibRefCitation>
. Tous ces sites se caractÉrisent par la prÉsence de l’espèce dans des milieux fortement anthropisés tels que parking de centre commercial, quartiers résidentiels, trottoirs pavés, bords de route. L’expansion de
<taxonomicName id="4C688191FFA1D514F72361BEFE5B3598" authorityName="Forster" authorityYear="1850" box="[293,428,935,960]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="B91C2600FFA1D514F72361BEFE5B3598" box="[293,428,935,960]" italics="true" pageId="2" pageNumber="101">T. magnum</emphasis>
</taxonomicName>
vers le nord de l’Europe est sans doute due à du transport passif, par exemple avec des plantes, buissons ou arbres en pot d’origine méditerranÉenne (
<bibRefCitation id="EFF987E3FFA1D514F7426636FE283211" author="Heller" box="[324,479,1071,1097]" firstAuthor="Heller" pageId="2" pageNumber="101" pagination="273 - 281" refId="ref5910" refString="Heller G. 2011. Pheidole pallidula (Nylander, 1849) und Tapinoma nigerrimum (Nylander, 1886) (Hymenoptera Formicidae), etablierte Neozoa in Rheinland-Pfalz. Mainzer naturwissenschaftliches Archiv 48: 273 - 281." type="journal article" year="2011">Heller 2011</bibRefCitation>
,
<bibRefCitation id="EFF987E3FFA1D514F7F16636FEA53233" author="Lenoir &amp; Galkowski" firstAuthor="Lenoir" pageId="2" pageNumber="101" pagination="449 - 453" refId="ref6085" refString="Lenoir L. &amp; Galkowski C. 2017. Sur la prEsence d'une fourmi envahissante (Tapinoma magnum) dans le Sud-Ouest de la France. Bulletin Societe Linneenne Bordeaux 45 (4): 449 - 453." type="journal article" year="2017">Lenoir &amp; Galkowski 2017</bibRefCitation>
,
<bibRefCitation id="EFF987E3FFA1D514F75B6648FDDD3233" author="Seifert" box="[349,554,1105,1131]" etAl="et al." firstAuthor="Seifert" pageId="2" pageNumber="101" pagination="123 - 144" refId="ref6591" refString="Seifert B., d'Eustacchio D., Kaufmann B., Centorame M., Lorite P. &amp; Modica M. V. 2017. Four species within the supercolonial ants of the Tapinoma nigerrimum complex revealed by integrative taxonomy (Hymenoptera: Formicidae). Myrmecological News 24: 123 - 144." type="journal article" year="2017">Seifert et al. 2017</bibRefCitation>
).
</paragraph>
<paragraph id="8BD7FA12FFA1D515F6D2666AFDEF3741" blockId="2.[151,627,526,1676]" lastBlockId="3.[144,1127,152,281]" lastPageId="3" lastPageNumber="102" pageId="2" pageNumber="101">
<taxonomicName id="4C688191FFA1D514F6D2666AFE4A32D4" authorityName="Forster" authorityYear="1850" box="[212,445,1139,1164]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="B91C2600FFA1D514F6D2666AFE4A32D4" box="[212,445,1139,1164]" italics="true" pageId="2" pageNumber="101">Tapinoma magnum</emphasis>
</taxonomicName>
est fortement polygyne et souvent supercoloniale. Au sein du groupe
<taxonomicName id="4C688191FFA1D514F74166A1FE383289" baseAuthorityName="Nylander" baseAuthorityYear="1856" box="[327,463,1208,1233]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="nigerrimum">
<emphasis id="B91C2600FFA1D514F74166A1FE383289" box="[327,463,1208,1233]" italics="true" pageId="2" pageNumber="101">nigerrimum</emphasis>
</taxonomicName>
, c’est l’espèce avec le plus fort potentiel envahissant (
<bibRefCitation id="EFF987E3FFA1D514F69966E2FE99334D" author="Seifert" box="[159,366,1275,1301]" etAl="et al." firstAuthor="Seifert" pageId="2" pageNumber="101" pagination="123 - 144" refId="ref6591" refString="Seifert B., d'Eustacchio D., Kaufmann B., Centorame M., Lorite P. &amp; Modica M. V. 2017. Four species within the supercolonial ants of the Tapinoma nigerrimum complex revealed by integrative taxonomy (Hymenoptera: Formicidae). Myrmecological News 24: 123 - 144." type="journal article" year="2017">Seifert et al. 2017</bibRefCitation>
). En rÉgion mÉditerranÉenne,
<taxonomicName id="4C688191FFA1D514F6FD6707FE88336F" authorityName="Forster" authorityYear="1850" box="[251,383,1310,1335]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="B91C2600FFA1D514F6FD6707FE88336F" box="[251,383,1310,1335]" italics="true" pageId="2" pageNumber="101">T. magnum</emphasis>
</taxonomicName>
est particulièrement abondante dans les milieux ouverts instables ou dégradés, avec une forte influence anthropique et un faible développement de la strate arborée. Elle forme des nids souterrains, souvent très Étendus et pouvant atteindre
<quantity id="4C9057F7FFA1D514F79667F0FE4B305C" box="[400,444,1513,1540]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="101" unit="m" value="1.0">1 m</quantity>
de profondeur, les entrÉes des galeries se signalant par la prÉsence de petits cônes de particules de sol excavÉes (
<bibRefCitation id="EFF987E3FFA1D514F7336456FDF13031" author="Seifert" box="[309,518,1615,1642]" etAl="et al." firstAuthor="Seifert" pageId="2" pageNumber="101" pagination="123 - 144" refId="ref6591" refString="Seifert B., d'Eustacchio D., Kaufmann B., Centorame M., Lorite P. &amp; Modica M. V. 2017. Four species within the supercolonial ants of the Tapinoma nigerrimum complex revealed by integrative taxonomy (Hymenoptera: Formicidae). Myrmecological News 24: 123 - 144." type="journal article" year="2017">Seifert et al. 2017</bibRefCitation>
). Les ouvrières fourragent en longues colonnes souvent très denses, longeant volontiers les bords de trottoirs, murs et murets. Elles se nourrissent de liquides sucrés, tels que miellat d’homoptères, nectar de plantes ou jus sucré sur des fruits, et chassent des petits invertébrés morts ou vivants (
<bibRefCitation id="EFF987E3FFA0D515F5D862C5FF3E3740" author="Cerda X. &amp; Retana J. &amp; Bosch J. &amp; Alsina A." journalOrPublisher="Acta Oecologica" pageId="3" pageNumber="102" pagination="419 - 429" part="10" refId="ref5696" refString="Cerda X., Retana J., Bosch J. &amp; Alsina A. 1989. Exploitation of food resources by the ant Tapinoma nigerrimum (Hymenoptera: Formicidae). Acta Oecologica 10 (4): 419 - 429." title="Exploitation of food resources by the ant Tapinoma nigerrimum (Hymenoptera: Formicidae)" type="journal article" year="1989">Cerda et al. 1989</bibRefCitation>
,
<bibRefCitation id="EFF987E3FFA0D515F6D262E7FEAE3740" author="Heller" box="[212,345,254,280]" firstAuthor="Heller" pageId="3" pageNumber="102" pagination="273 - 281" refId="ref5910" refString="Heller G. 2011. Pheidole pallidula (Nylander, 1849) und Tapinoma nigerrimum (Nylander, 1886) (Hymenoptera Formicidae), etablierte Neozoa in Rheinland-Pfalz. Mainzer naturwissenschaftliches Archiv 48: 273 - 281." type="journal article" year="2011">Heller 2011</bibRefCitation>
,
<bibRefCitation id="EFF987E3FFA0D515F76262E7FDFA3741" author="Noordijk" box="[356,525,254,281]" firstAuthor="Noordijk" pageId="3" pageNumber="102" pagination="86 - 93" refId="ref6274" refString="Noordijk A. 2016. Leefwijze van Tapinoma nigerrimum (Hymenoptera: Formicidae), een nieuwe exotische mier in Nederland. Entomologische Berichten 76 (3): 86 - 93." type="journal article" year="2016">Noordijk 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
</emphasis>
</treatment>
</document>
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