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<document id="19A26568BE50AA0DC961BD231055B6D6" ID-DOI="10.1080/00222933.2018.1435833" ID-ISSN="1464-5262" ID-Zenodo-Dep="5174451" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628558511936" checkinUser="felipe" docAuthor="Wagner, Philipp, Greenbaum, Eli, Bauer, Aaron M., Kusamba, Chifundera &amp; Leaché, Adam D." docDate="2018" docId="03BFD058FFB8FF8E14268BF7BA02FD1B" docLanguage="en" docName="JNatHist.52.13-16.771-817.pdf" docOrigin="Journal of Natural History 52 (13 - 16)" docSource="http://dx.doi.org/10.1080/00222933.2018.1435833" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Acanthocercus kiwuensis" docType="treatment" docVersion="3" lastPageNumber="807" masterDocId="FF86A820FF9BFFAB154A8901BE4DFF9E" masterDocTitle="Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae)" masterLastPageNumber="817" masterPageNumber="771" pageNumber="805" updateTime="1699046512504" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="354C345639C2EFFE6C499791BECEF22D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1E3740254546807F55350E4C2CD0120A">Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae)</mods:title>
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<mods:namePart id="993D0F7C042812B01CE535699A54EE0E">Wagner, Philipp</mods:namePart>
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<mods:namePart id="3E45938D0A73D704291B92F2C694FFD6">Greenbaum, Eli</mods:namePart>
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<mods:namePart id="75312920032B836AAD2CA4514403E3C9">Bauer, Aaron M.</mods:namePart>
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<mods:namePart id="242E208DDE07C8555F044A0B9D205705">Kusamba, Chifundera</mods:namePart>
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<mods:namePart id="A8C0C2FD58830CCBB6D19F1D152F6290">Leaché, Adam D.</mods:namePart>
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<mods:title id="1DCB20A91C009B7C4BAB29644DF6078E">Journal of Natural History</mods:title>
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<mods:date id="3217B662F9A224672DD26E9534860E54">2018</mods:date>
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<mods:number id="6E9B2824F0DBC0D1CD42E2FC5277EC70">2018-03-19</mods:number>
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<mods:detail id="F41DB59D017E5FEA9D99AF253511DA79" type="volume">
<mods:number id="AD30BEE70784DAA7E19FB3CADE0FD4B6">52</mods:number>
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<mods:number id="B6F28692868B59FF1A829C9F86A48B26">13 - 16</mods:number>
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<mods:start id="B86E54F6C36384C2BEC62479BBA2CE6A">771</mods:start>
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<mods:url id="4B77BD7DAA6CECC59DCAE646C0BFB01D">http://dx.doi.org/10.1080/00222933.2018.1435833</mods:url>
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<mods:identifier id="EABB7665F94AA21AD2EC416BABF89B80" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="6D64A2B3D2905F0CEB336B4B31C459CF" type="Zenodo-Dep">5174451</mods:identifier>
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<treatment id="03BFD058FFB8FF8E14268BF7BA02FD1B" LSID="urn:lsid:plazi:treatment:03BFD058FFB8FF8E14268BF7BA02FD1B" httpUri="http://treatment.plazi.org/id/03BFD058FFB8FF8E14268BF7BA02FD1B" lastPageId="37" lastPageNumber="807" pageId="35" pageNumber="805">
<subSubSection id="C30C32C5FFB8FF8814268BF7BDFBFC8E" box="[364,950,758,784]" pageId="35" pageNumber="805" type="nomenclature">
<paragraph id="8BA9614EFFB8FF8814268BF7BDFBFC8E" blockId="35.[364,950,758,819]" box="[364,950,758,784]" pageId="35" pageNumber="805">
<heading id="D0E1D622FFB8FF8814268BF7BDFBFC8E" box="[364,950,758,784]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="805" reason="5">
<treatmentCitationGroup id="AB064660FFB8FF8814268BF7BDFBFC8E" box="[364,950,758,784]" pageId="35" pageNumber="805">
<treatmentCitation id="0AB7475FFFB8FF8814268BF7BDFBFC8E" author="Klausewitz W" box="[364,950,758,784]" page="167" pageId="35" pageNumber="805" year="1957">
<taxonomicName id="4C161ACDFFB8FF8814268BF7BDFBFC8E" authority="(Klausewitz, 1957: 167)" baseAuthorityName="Klausewitz" baseAuthorityPageNumber="167" baseAuthorityYear="1957" box="[364,950,758,784]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B962BD5CFFB8FF8814268BF7BCD2FC8E" bold="true" box="[364,671,758,784]" italics="true" pageId="35" pageNumber="805">Acanthocercus kiwuensis</emphasis>
(
<bibRefCitation id="EF871CBFFFB8FF8817E48BF6BDFCFC8E" author="Klausewitz W" box="[686,945,759,784]" pageId="35" pageNumber="805" pagination="157 - 174" refId="ref20422" refString="Klausewitz W. 1957. Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. Die Unterarten von Agama atricollis [Eidonomic studies about the species complex of Agama cyanogaster and Agama atricollis. The subspecies of Agama atricollis]. Senckenb Biol. 38: 157 - 174." type="journal article" year="1957">Klausewitz, 1957: 167</bibRefCitation>
)
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30C32C5FFB8FF88171B8A1BBCCDFC24" pageId="35" pageNumber="805" type="description">
<paragraph id="8BA9614EFFB8FF88171B8A1BBC9FFCAD" blockId="35.[364,950,758,819]" box="[593,722,794,819]" pageId="35" pageNumber="805">
(
<figureCitation id="132D7DCBFFB8FF8817138A1BBC87FCAD" box="[601,714,794,819]" captionStart="Figure 15" captionStartId="36.[160,225,1253,1276]" captionTargetBox="[178,1138,142,1222]" captionTargetId="figure-183@36.[178,1138,142,1222]" captionTargetPageId="36" captionText="Figure 15. Holotype (ZMB 23906) of Acanthocercus kiwuensis, from Kissenji, Rwanda." figureDoi="http://doi.org/10.5281/zenodo.5174483" httpUri="https://zenodo.org/record/5174483/files/figure.png" pageId="35" pageNumber="805">Figure 15</figureCitation>
)
</paragraph>
<paragraph id="8BA9614EFFB8FF8815EA8A5CBCCDFC24" blockId="35.[160,1156,861,954]" pageId="35" pageNumber="805">
1957
<taxonomicName id="4C161ACDFFB8FF8815AA8A5CBDBCFCEB" authority="Klausewitz, Eidonomische Untersuchungen" authorityName="Klausewitz, Eidonomische Untersuchungen" authorityYear="1957" box="[224,1009,861,885]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="subSpecies" species="atricollis" subSpecies="kiwuensis">
<emphasis id="B962BD5CFFB8FF8815AA8A5CBC4CFCEB" box="[224,513,861,885]" italics="true" pageId="35" pageNumber="805">Agama atricollis kiwuensis</emphasis>
Klausewitz, Eidonomische Untersuchungen
</taxonomicName>
über die Rassenkreise
<taxonomicName id="4C161ACDFFB8FF8814088A81BC5FFC06" authorityName="Ruppell" authorityYear="1835" box="[322,530,896,920]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="cyanogaster">
<emphasis id="B962BD5CFFB8FF8814088A81BC5FFC06" box="[322,530,896,920]" italics="true" pageId="35" pageNumber="805">Agama cyanogaster</emphasis>
</taxonomicName>
und
<taxonomicName id="4C161ACDFFB8FF88170E8A81BCA7FC06" authorityName="Smith, Illustrations" authorityYear="1849" box="[580,746,896,920]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFFB8FF88170E8A81BCA7FC06" box="[580,746,896,920]" italics="true" pageId="35" pageNumber="805">Agama atricollis</emphasis>
</taxonomicName>
. 2. Die Unterarten von
<taxonomicName id="4C161ACDFFB8FF8816928A81BA33FC06" authorityName="Smith, Illustrations" authorityYear="1849" box="[984,1150,896,920]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFFB8FF8816928A81BA33FC06" box="[984,1150,896,920]" italics="true" pageId="35" pageNumber="805">Agama atricollis</emphasis>
</taxonomicName>
. Senckenbergiana biologica 38: 157–174.
</paragraph>
</subSubSection>
<subSubSection id="C30C32C5FFB8FF8815EA8AE3BAC9FB83" box="[160,1156,994,1053]" pageId="35" pageNumber="805" type="materials_examined">
<paragraph id="8BA9614EFFB8FF8815EA8AE3BF46FC65" blockId="35.[160,1156,994,1053]" box="[160,267,994,1019]" pageId="35" pageNumber="805">
<emphasis id="B962BD5CFFB8FF8815EA8AE3BF46FC65" bold="true" box="[160,267,994,1019]" italics="true" pageId="35" pageNumber="805">
<typeStatus id="54ADDFECFFB8FF8815EA8AE3BF46FC65" box="[160,267,994,1019]" pageId="35" pageNumber="805" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA9614EFFB8FF8815EA8D04BAC9FB83" blockId="35.[160,1156,994,1053]" box="[160,1156,1029,1053]" pageId="35" pageNumber="805">
<materialsCitation id="3B7E6B13FFB8FF8815EA8D04BAC9FB83" accessionNumber="ZMB 23906" box="[160,1156,1029,1053]" collectionCode="ZMB" country="Rwanda" county="East Africa." location="Kissenji" municipality="Lake Kivu'" pageId="35" pageNumber="805" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED07F98BFFB8FF8815EA8D04BE9EFB83" box="[160,211,1029,1053]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="35" pageNumber="805" type="Museum">ZMB</collectionCode>
23906, from ‘
<location id="8EC93795FFB8FF88143A8D04BF8EFB83" LSID="urn:lsid:plazi:treatment:03BFD058FFB8FF8E14268BF7BA02FD1B:8EC93795FFB8FF88143A8D04BF8EFB83" box="[368,451,1029,1053]" country="Rwanda" county="East Africa." municipality="Lake Kivu'" name="Kissenji" pageId="35" pageNumber="805">Kissenji</location>
[=Gisenyi] north-east shore of
<collectingMunicipality id="6BCDFB34FFB8FF8816698D04BDDEFB83" box="[803,915,1029,1053]" pageId="35" pageNumber="805">Lake Kivu’</collectingMunicipality>
,
<collectingCountry id="F30121DEFFB8FF8816EB8D04BDB0FB83" box="[929,1021,1029,1053]" name="Rwanda" pageId="35" pageNumber="805">Rwanda</collectingCountry>
,
<collectingCounty id="62C819C2FFB8FF8811428D04BAC9FB83" box="[1032,1156,1029,1053]" pageId="35" pageNumber="805">East Africa.</collectingCounty>
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</paragraph>
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<subSubSection id="C30C32C5FFB8FF8E15EA8D44BF90FE90" lastPageId="37" lastPageNumber="807" pageId="35" pageNumber="805" type="description">
<paragraph id="8BA9614EFFB8FF8815EA8D44BF6BFBC0" blockId="35.[160,1156,1093,1395]" box="[160,294,1093,1118]" pageId="35" pageNumber="805">
<emphasis id="B962BD5CFFB8FF8815EA8D44BF6BFBC0" bold="true" box="[160,294,1093,1118]" italics="true" pageId="35" pageNumber="805">Description</emphasis>
</paragraph>
<paragraph id="8BA9614EFFB8FF8815EA8D69BACEFAED" blockId="35.[160,1156,1093,1395]" pageId="35" pageNumber="805">
A small species of the genus (SVL 87.3–130.3, x = 116.1, n = 15) with a total length usually about
<quantity id="4CEECCABFFB8FF88140C8D8ABFE5FB3D" box="[326,424,1163,1187]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="805" unit="mm" value="300.0">300 mm</quantity>
, tail more than half again longer than the SVL in adult males. Head narrow (particularly in males), occipital scale lacking. Nasal scale not convex, smooth, below the canthus rostralis, often not visible from above. Ear openings as large as eyes, with the tympanum conspicuous. No nuchal crest. Dorsal scalation heterogeneous, matrix scales keeled and mucronate, irregularly intermixed with enlarged keeled and mucronate scales of two sizes, but not as distinct as in
<taxonomicName id="4C161ACDFFB8FF8811478C39BA33FACE" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[1037,1150,1336,1360]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFB8FF8811478C39BA33FACE" box="[1037,1150,1336,1360]" italics="true" pageId="35" pageNumber="805">A. gregorii</emphasis>
</taxonomicName>
. Vertebral scales enlarged, becoming continuously smaller to the flanks, not as distinct as
</paragraph>
<paragraph id="8BA9614EFFB8FF8815EA8CE3BFECF92F" blockId="35.[160,1157,1506,1713]" pageId="35" pageNumber="805">
64 = northern slopes Ngorongoro; 65 = Olduvai Gorge,
<collectingRegion id="49D2AFACFFB8FF8817EA8CE3BC92FA68" box="[672,735,1506,1526]" country="Tanzania" name="Arusha" pageId="35" pageNumber="805">Arusha</collectingRegion>
; 66 = Serengeti, Naabi Gate; 67 = Kakitumba; 68 = Ntaruka; 69 =
<collectingRegion id="49D2AFACFFB8FF8814138CFCBFDCF98F" box="[345,401,1533,1553]" country="Uganda" name="Kibale" pageId="35" pageNumber="805">Kibale</collectingRegion>
; 70 = Mpanga; 71 = Ntaruka?; 72 = Lake Nasho; 73 = Mpanga?; 74 =
<collectingRegion id="49D2AFACFFB8FF8811578CFCBACDF98F" box="[1053,1152,1533,1553]" country="Burundi" name="Bujumbura Mairie" pageId="35" pageNumber="805">Bujumbura</collectingRegion>
; 75 = Digg
<emphasis id="B962BD5CFFB8FF8814498F19BF4AF9B2" box="[259,263,1560,1580]" italics="true" pageId="35" pageNumber="805">’</emphasis>
s Farm nr. Amoressa; 76 = Sanga; 77 = only coordinates; 78 =
<collectingRegion id="49D2AFACFFB8FF88163A8F19BDE1F9B2" box="[880,940,1560,1580]" country="Tanzania" name="Tabora" pageId="35" pageNumber="805">Tabora</collectingRegion>
Udjidji; 79 = Kakoma; 80 = Ukimbu; 81 = Nguru Mountains; 82 = Bagamoyo; 83 = Kilosa; 84 = Karema; 85 = Kakokwe; 86 = Tukuyu; 87 = Rutenganio; 88 = Mwaya; 89 = Matema; 90 = Utengule; 91 = Ilolo; 92 = Uhehe; 93 = Liwale; 94 = Manengue; 95 = Kilwa; 96 = Arba Minch; 97 = Lake Shala; 98 = Mahal Uonz; 99 = Masai
<collectingRegion id="49D2AFACFFB8FF8815EA8F82BE81F909" box="[160,204,1667,1687]" country="Tanzania" name="Mara" pageId="35" pageNumber="805">Mara</collectingRegion>
NP; 100 = Lyanza; 101 = Kikambo; 102 = Sanga; 103 = Metemna; 104 = NE Songea District; 105 =
<collectingRegion id="49D2AFACFFB8FF8811778F82BACDF909" box="[1085,1152,1667,1687]" country="Tanzania" name="Singida" pageId="35" pageNumber="805">Singida</collectingRegion>
; 106 =
<collectingRegion id="49D2AFACFFB8FF8815978F9CBF59F92F" box="[221,276,1693,1713]" country="Mozambique" name="Niassa" pageId="35" pageNumber="805">Niassa</collectingRegion>
Game Reserve.
</paragraph>
<caption id="DF6931C6FFBFFF8F15EA8DE4BDBEFB62" ID-DOI="http://doi.org/10.5281/zenodo.5174483" ID-Zenodo-Dep="5174483" box="[160,1011,1253,1276]" httpUri="https://zenodo.org/record/5174483/files/figure.png" pageId="36" pageNumber="806" startId="36.[160,225,1253,1276]" targetBox="[178,1138,142,1222]" targetPageId="36">
<paragraph id="8BA9614EFFBFFF8F15EA8DE4BDBEFB62" blockId="36.[160,1011,1253,1276]" box="[160,1011,1253,1276]" pageId="36" pageNumber="806">
<emphasis id="B962BD5CFFBFFF8F15EA8DE4BF44FB62" bold="true" box="[160,265,1253,1276]" pageId="36" pageNumber="806">Figure 15.</emphasis>
Holotype (ZMB 23906) of
<taxonomicName id="4C161ACDFFBFFF8F175C8DE4BD4EFB62" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[534,771,1253,1276]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="806" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B962BD5CFFBFFF8F175C8DE4BD4EFB62" box="[534,771,1253,1276]" italics="true" pageId="36" pageNumber="806">Acanthocercus kiwuensis</emphasis>
</taxonomicName>
, from Kissenji, Rwanda.
</paragraph>
</caption>
<paragraph id="8BA9614EFFBFFF8F15EA8C30BDC3F9A2" blockId="36.[160,1156,1329,1700]" pageId="36" pageNumber="806">
in other species. Body scales in 73–96 rows [up to 116 mentioned by (
<bibRefCitation id="EF871CBFFFBFFF8F169E8C30BA31FAD7" author="Sternfeld R" box="[980,1148,1329,1353]" pageId="36" pageNumber="806" refId="ref21426" refString="Sternfeld R. 1912. Wissenschaftliche Ergebnisse der Deutschen Zentral-Afrika Expedition 1907 - 1908, unter Fuhrung A. Friedrichs, Herzogs zu Mecklenburg. Klinkhard und Biermann, Leipzig. IV. Zoologie II. Lfg. Reptilia; p. 197 - 279 [the volume was published in parts from 1911 to 1913 but this part was published in 1912]." type="book" year="1912">Sternfeld 1912</bibRefCitation>
) around midbody (x = 83.6, n = 29) and 55–77 (x = 62.7, n = 15) longitudinal rows along the vertebrae. Ventral scales smooth and small, as large as the medium-sized dorsal scales, in 57–74 (x = 63.9, n = 15) longitudinal rows. Gular scales small, keeled, and exposed, slightly smaller than the ventral scales. Males with two to three continuous rows (x = 2.2, n = 18) and a total number of 16–28 precloacal pores (x = 20.2, n = 18), absent in females. Dorsal scales of the tail base not as distinctly swollen as in e.g.
<taxonomicName id="4C161ACDFFBFFF8F15EA8F25BF5CF9A2" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[160,273,1572,1596]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="806" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFBFFF8F15EA8F25BF5CF9A2" box="[160,273,1572,1596]" italics="true" pageId="36" pageNumber="806">A. gregorii</emphasis>
</taxonomicName>
or
<taxonomicName id="4C161ACDFFBFFF8F14708F25BFFCF9A2" authorityName="Smith, Illustrations" authorityYear="1849" box="[314,433,1572,1596]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="806" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFFBFFF8F14708F25BFFCF9A2" box="[314,433,1572,1596]" italics="true" pageId="36" pageNumber="806">A. atricollis</emphasis>
</taxonomicName>
, but distinct from other scales of the tail.
</paragraph>
<paragraph id="8BA9614EFFBFFF8E158A8F46BF90FE90" blockId="36.[160,1156,1329,1700]" lastBlockId="37.[160,1155,142,270]" lastPageId="37" lastPageNumber="807" pageId="36" pageNumber="806">
According to
<bibRefCitation id="EF871CBFFFBFFF8F14118F46BC6DF9C1" author="Klausewitz W" box="[347,544,1607,1631]" pageId="36" pageNumber="806" pagination="157 - 174" refId="ref20422" refString="Klausewitz W. 1957. Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. Die Unterarten von Agama atricollis [Eidonomic studies about the species complex of Agama cyanogaster and Agama atricollis. The subspecies of Agama atricollis]. Senckenb Biol. 38: 157 - 174." type="journal article" year="1957">Klausewitz (1957)</bibRefCitation>
this species has a very characteristic coloration: adult males with a yellowish green to greenish blue head, throat whitish to blue, body deep blue with some pale blue dots, base and first third of the tail yellow, followed by yellowish green second third and a brownish tip.
<bibRefCitation id="EF871CBFFFBEFF8E17B4898FBDFAFF38" author="Sternfeld R" box="[766,951,142,166]" pageId="37" pageNumber="807" refId="ref21426" refString="Sternfeld R. 1912. Wissenschaftliche Ergebnisse der Deutschen Zentral-Afrika Expedition 1907 - 1908, unter Fuhrung A. Friedrichs, Herzogs zu Mecklenburg. Klinkhard und Biermann, Leipzig. IV. Zoologie II. Lfg. Reptilia; p. 197 - 279 [the volume was published in parts from 1911 to 1913 but this part was published in 1912]." type="book" year="1912">Sternfeld (1912)</bibRefCitation>
mentioned male specimens from ‘Kissenje’ [=Gisenyi] having the entire body blue and the tail uniform olive-yellow. However, the non-nuptial coloration shows a pattern of yellow spots and stripes on the body as well.
</paragraph>
</subSubSection>
<subSubSection id="C30C32C5FFBEFF8E15EA8848BA02FD1B" pageId="37" pageNumber="807" type="distribution">
<paragraph id="8BA9614EFFBEFF8E15EA8848BF61FEFC" blockId="37.[160,1155,329,492]" box="[160,300,329,354]" pageId="37" pageNumber="807">
<emphasis id="B962BD5CFFBEFF8E15EA8848BF61FEFC" bold="true" box="[160,300,329,354]" italics="true" pageId="37" pageNumber="807">Distribution</emphasis>
</paragraph>
<paragraph id="8BA9614EFFBEFF8E15EA886DBFA7FE72" blockId="37.[160,1155,329,492]" pageId="37" pageNumber="807">
Since its description (
<bibRefCitation id="EF871CBFFFBEFF8E14D2886DBC1CFE1A" author="Klausewitz W" box="[408,593,364,388]" pageId="37" pageNumber="807" pagination="157 - 174" refId="ref20422" refString="Klausewitz W. 1957. Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. Die Unterarten von Agama atricollis [Eidonomic studies about the species complex of Agama cyanogaster and Agama atricollis. The subspecies of Agama atricollis]. Senckenb Biol. 38: 157 - 174." type="journal article" year="1957">Klausewitz 1957</bibRefCitation>
),
<taxonomicName id="4C161ACDFFBEFF8E1722886DBCA2FE1A" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[616,751,364,388]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="807" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B962BD5CFFBEFF8E1722886DBCA2FE1A" box="[616,751,364,388]" italics="true" pageId="37" pageNumber="807">A. kiwuensis</emphasis>
</taxonomicName>
had been considered restricted to the mountainous areas around Lake Kivu (
<collectingCountry id="F30121DEFFBEFF8E1732888EBC9EFE39" box="[632,723,399,423]" name="Rwanda" pageId="37" pageNumber="807">Rwanda</collectingCountry>
and
<collectingCountry id="F30121DEFFBEFF8E1646888EBD0DFE39" box="[780,832,399,423]" name="Democratic Republic of the Congo" pageId="37" pageNumber="807">DRC</collectingCountry>
). Herein, it is recognized as a wide-ranging species from Lake
<collectingRegion id="49D2AFACFFBEFF8E175988B0BC0EFE57" box="[531,579,433,457]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="37" pageNumber="807">Kivu</collectingRegion>
in the south to central
<collectingCountry id="F30121DEFFBEFF8E161C88B0BDE3FE57" box="[854,942,433,457]" name="Uganda" pageId="37" pageNumber="807">Uganda</collectingCountry>
and north-eastern
<collectingCountry id="F30121DEFFBEFF8E15EA88D5BE9DFE72" box="[160,208,468,492]" name="Democratic Republic of the Congo" pageId="37" pageNumber="807">DRC</collectingCountry>
in the north (
<figureCitation id="132D7DCBFFBEFF8E143888D5BF90FE72" box="[370,477,468,492]" captionStart="Figure 16" captionStartId="38.[160,225,1079,1102]" captionTargetBox="[178,1138,142,1048]" captionTargetId="figure-42@38.[178,1138,142,1048]" captionTargetPageId="38" captionText="Figure 16. Distribution of Acanthocercus kiwuensis and A. ugandaensis. The numbers refer to the specific localities." figureDoi="http://doi.org/10.5281/zenodo.5174485" httpUri="https://zenodo.org/record/5174485/files/figure.png" pageId="37" pageNumber="807">Figure 16</figureCitation>
).
</paragraph>
<paragraph id="8BA9614EFFBEFF8E15EA8B27BFD8FDA1" blockId="37.[160,1155,550,645]" box="[160,405,550,575]" pageId="37" pageNumber="807">
<emphasis id="B962BD5CFFBEFF8E15EA8B27BFD8FDA1" bold="true" box="[160,405,550,575]" italics="true" pageId="37" pageNumber="807">Habitat and ecology</emphasis>
</paragraph>
<paragraph id="8BA9614EFFBEFF8E15EA8B4BBA02FD1B" blockId="37.[160,1155,550,645]" pageId="37" pageNumber="807">
<taxonomicName id="4C161ACDFFBEFF8E15EA8B4BBFFDFDFC" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[160,432,586,610]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="807" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B962BD5CFFBEFF8E15EA8B4BBFFDFDFC" box="[160,432,586,610]" italics="true" pageId="37" pageNumber="807">Acanthocercus kiwuensis</emphasis>
</taxonomicName>
has been found on trees in savannah woodland, but also on man-made structures (e.g. houses, old walls) near cities and towns (EG, pers. obs.).
</paragraph>
</subSubSection>
</treatment>
</document>
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