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<document id="9A478B580D838474FB32932B6A7A7A59" ID-CLB-Dataset="31199" ID-DOI="10.11646/zootaxa.4363.2.6" ID-GBIF-Dataset="f248e89f-ca42-421b-aab5-334a2281af89" ID-ISSN="1175-5326" ID-Zenodo-Dep="1098838" ID-ZooBank="4DF8A3AC-01FA-41D1-93CE-63E23FC7728E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1512980399631" checkinUser="plazi" docAuthor="Arguez, Katherine M., Moore, Matthew R. &amp; Branham, Marc A." docDate="2017" docId="039C878AFFD1FFC10D92FF28FA320ED5" docLanguage="en" docName="zootaxa.4363.2.6.pdf" docOrigin="Zootaxa 4363 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phileurini Burmeister 1847" docType="key" docVersion="8" masterDocId="FFA5FFF2FFD6FFC60D05FFB0FFA90B43" masterDocTitle="Description of the larva and pupa of Phileurus truncatus (Palisot de Beauvois, 1806) (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) with a key to described third instars of New World Phileurini" updateTime="1698508758872" updateUser="ExternalLinkService">
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<mods:title id="42C24A5427D14E37A3070F8933B86553">Description of the larva and pupa of Phileurus truncatus (Palisot de Beauvois, 1806) (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) with a key to described third instars of New World Phileurini</mods:title>
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<mods:namePart id="9BD987FC3696A64535FB08BA4D2D5EC0">Arguez, Katherine M.</mods:namePart>
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<mods:namePart id="F691220B09530CED3226023E7B8054CF">Moore, Matthew R.</mods:namePart>
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<mods:date id="22F9EF0A06D77E6EDC5E670FAC23C550">2017</mods:date>
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<treatment id="039C878AFFD1FFC10D92FF28FA320ED5" ID-DOI="http://doi.org/10.5281/zenodo.6046117" ID-GBIF-Taxon="137665558" ID-Zenodo-Dep="6046117" LSID="urn:lsid:plazi:treatment:039C878AFFD1FFC10D92FF28FA320ED5" httpUri="http://treatment.plazi.org/id/039C878AFFD1FFC10D92FF28FA320ED5" pageId="7">
<subSubSection id="C32F6517FFD1FFC10D92FF28FB170BB4" pageId="7" type="nomenclature">
<paragraph id="8B8A369CFFD1FFC10D92FF28FD5D0BF1" blockId="7.[151,756,152,178]" box="[151,756,152,178]" pageId="7">
<heading id="D0C281F0FFD1FFC10D92FF28FD5D0BF1" bold="true" box="[151,756,152,178]" fontSize="11" level="1" pageId="7" reason="1">
<emphasis id="B941EA8EFFD1FFC10D92FF28FD5D0BF1" bold="true" box="[151,756,152,178]" pageId="7">
Key to the third instars of New World
<taxonomicName id="4C354D1FFFD1FFC10F72FF28FD5D0BF1" ID-CoL="8RYTV" authorityName="Burmeister" authorityYear="1847" box="[631,756,152,178]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="tribe" tribe="Phileurini">Phileurini</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FF6EFB170BB4" blockId="7.[151,1214,222,247]" box="[151,1214,222,247]" pageId="7">
Modified from
<bibRefCitation id="EFA44B6DFFD1FFC10C41FF6EFDF10BB4" author="Neita" box="[324,600,222,247]" pageId="7" refString="Neita Moreno, J. C. &amp; Ratcliffe B. C. (2011) Immature stages of Homophileurus tricuspis Prell (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) with notes on natural history and distribution in Colombia. Zootaxa, 2915, 20 - 28." type="journal article" year="2011">Neita &amp; Ratcliffe (2011)</bibRefCitation>
,
<bibRefCitation id="EFA44B6DFFD1FFC10F61FF6EFC390BB4" author="Ratcliffe" box="[612,912,222,247]" pageId="7" refString="Ratcliffe, B. C. &amp; Skelley, P. E. (2011) Description of the larva of Homophileurus integer (Burmeister, 1847) (Coleoptera: Scarabaeidae: Dynastinae: Phileurini), with notes on biology and a key to the known larvae of New World Phileurini. The Coleopterists Bulletin, 65, 297 - 304. https: // doi. org / 10.1649 / 072.065.0311" type="journal article" year="2011">Ratcliffe &amp; Skelley (2011)</bibRefCitation>
,
<bibRefCitation id="EFA44B6DFFD1FFC10E99FF6EFB120BB4" author="Ibarra-Polesel" box="[924,1211,222,247]" pageId="7" refString="Ibarra-Polesel, M. G., Valle N. G., Neita-Moreno J. C. &amp; Damborsky, M. P. (2017) Description of the pupa and redescription of the third instar of Phileurus valgus (Olivier) (Coleoptera: Scarabaeidae: Dynastinae: Phileurini). Zootaxa, 4290, 571 - 580. https: // doi. org / 10.11646 / zootaxa. 4290.3.10" type="journal article" year="2017">
Ibarra-Polesel
<emphasis id="B941EA8EFFD1FFC10945FF6FFBD80BB4" box="[1088,1137,222,247]" italics="true" pageId="7">et al</emphasis>
. 2017
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32F6517FFD1FFC10D92FE96FA320ED5" pageId="7" type="key">
<keyStep id="30C12E39FFD1FFC10D92FE96FA320A1B" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FE96FA320A78" blockId="7.[151,1436,294,1430]" box="[151,1435,294,315]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FE96FA320A78" box="[151,1435,294,315]" pageId="7">1. Anterior frontal setae absent............................................................................. 2</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FEF3FA320A1B" blockId="7.[151,1436,294,1430]" box="[151,1435,323,344]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FEF3FA320A1B" box="[151,1435,323,344]" pageId="7">- Anterior frontal setae present............................................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FED1FA320AF3" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FED1FA350AD0" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FED1FA350AD0" pageId="7">
2. Lateral margins of labrum broadly rounded, not angulate (
<collectingCountry id="F322760CFFD1FFC10E08FED1FCD90A35" box="[781,880,353,374]" name="Argentina" pageId="7">Argentina</collectingCountry>
, southern
<collectingCountry id="F322760CFFD1FFC10ED6FED1FBB90A35" box="[979,1040,353,374]" name="Brazil" pageId="7">Brazil</collectingCountry>
,
<collectingRegion id="49F1F87EFFD1FFC1091CFED1FBDA0A35" box="[1049,1139,353,374]" country="Paraguay" name="Asuncion" pageId="7">
<collectingCountry id="F322760CFFD1FFC1091CFED1FBDA0A35" box="[1049,1139,353,374]" name="Paraguay" pageId="7">Paraguay</collectingCountry>
</collectingRegion>
,
<collectingCountry id="F322760CFFD1FFC10979FED1FB780A35" box="[1148,1233,353,374]" name="Uruguay" pageId="7">Uruguay</collectingCountry>
)...................................................................................
<taxonomicName id="4C354D1FFFD1FFC1090AFECEFA350AD0" authority="(Mannerheim, 1829)" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[1039,1436,382,403]" class="Insecta" family="Dynastidae" genus="Trioplus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="B941EA8EFFD1FFC1090AFECEFB660AD0" box="[1039,1231,382,403]" italics="true" pageId="7">Trioplus cylindricus</emphasis>
(Mannerheim, 1829)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FE2BFA320AF3" blockId="7.[151,1436,294,1430]" box="[151,1435,411,432]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FE2BFA320AF3" box="[151,1435,411,432]" pageId="7">- At least one lateral margin of labrum angulate............................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FE09FA32094B" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FE09FA350AA8" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FE09FA350AA8" pageId="7">
3. Left and right lateral margins of labrum angulate. Maxilla with a row of six conical, stridulatory teeth (
<collectingCountry id="F322760CFFD1FFC109BFFE09FAB90A8D" box="[1210,1296,441,462]" name="Uruguay" pageId="7">Uruguay</collectingCountry>
,
<collectingCountry id="F322760CFFD1FFC1081CFE09FAFC0A8D" box="[1305,1365,441,462]" name="Brazil" pageId="7">Brazil</collectingCountry>
)....................................................................
<taxonomicName id="4C354D1FFFD1FFC10EE1FE66FA350AA8" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[996,1436,470,491]" class="Insecta" family="Dynastidae" genus="Archophileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="B941EA8EFFD1FFC10EE1FE66FB480AA8" box="[996,1249,470,491]" italics="true" pageId="7">Archophileurus fimbriatus</emphasis>
(Burmeister, 1847)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FE43FA32094B" blockId="7.[151,1436,294,1430]" box="[151,1435,499,520]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FE43FA32094B" box="[151,1435,499,520]" pageId="7">- Left lateral margin of labrum angulate. Maxilla with 610 stridulatory teeth....................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FDA1FA350900" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FDA1FA320965" blockId="7.[151,1436,294,1430]" box="[151,1435,529,550]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FDA1FA320965" box="[151,1435,529,550]" pageId="7">4. Terminal antennomere with 5 dorsal sensory spots........................................................... 5</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FD9EFA350900" blockId="7.[151,1436,294,1430]" box="[151,1436,558,579]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FD9EFA350900" box="[151,1436,558,579]" pageId="7">- Terminal antennomere with less than 5 dorsal sensory spots....................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FDFBFA3509FA" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FDFBFA35093D" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FDFBFA35093D" pageId="7">
5. Abdominal spiracle I smaller than spiracles IIVII. Maxilla with row of 67 truncate, stridulatory teeth (
<collectingCountry id="F322760CFFD1FFC109CCFDFBFA870923" box="[1225,1326,587,608]" name="Colombia" pageId="7">Colombia</collectingCountry>
).........................................................................
<taxonomicName id="4C354D1FFFD1FFC10EDBFDD9FA35093D" authority="Neita &amp; Ratcliffe, 2010" authorityName="Neita &amp; Ratcliffe" authorityYear="2010" box="[990,1436,617,638]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="elbitae">
<emphasis id="B941EA8EFFD1FFC10EDBFDD9FB1D093D" box="[990,1204,617,638]" italics="true" pageId="7">Hemiphileurus elbitae</emphasis>
Neita &amp; Ratcliffe, 2010
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FD36FA3509FA" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FD36FA3509FA" pageId="7">
- Abdominal spiracles similar in size. Maxilla with row of 710 truncate, stridulatory teeth (
<collectingCountry id="F322760CFFD1FFC10953FD36FAE709D8" box="[1110,1358,646,667]" name="United States of America" pageId="7">United States of America</collectingCountry>
to Pan- ama)..........................................................
<taxonomicName id="4C354D1FFFD1FFC10ECBFD14FA3509FA" authority="(Palisot de Beauvois, 1806)" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1806" box="[974,1436,676,697]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B941EA8EFFD1FFC10ECBFD14FB2409FA" box="[974,1165,676,697]" italics="true" pageId="7">Phileurus truncatus</emphasis>
(Palisot de Beauvois, 1806)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FD71FA35086D" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FD71FA3509B0" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FD71FA3509B0" pageId="7">
6. Terminal segment of antenna with 2 dorsal sensory spots. Inner margin of left mandible, distad of molar area, with short, rounded tooth (Hispaniola)...................................................
<taxonomicName id="4C354D1FFFD1FFC10948FD6EFA3509B0" authority="Kolbe, 1910" authorityName="Kolbe" authorityYear="1910" box="[1101,1436,734,755]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B941EA8EFFD1FFC10948FD6EFAB609B0" box="[1101,1311,734,755]" italics="true" pageId="7">Hemiphileurus dispar</emphasis>
Kolbe, 1910
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FD4CFA35086D" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FD4CFA35086D" pageId="7">
- Terminal segment of antenna with 34 dorsal sensory spots. Inner margin of left mandible, distad of molar area, with promi- nent, triangular tooth (
<collectingCountry id="F322760CFFD1FFC10CB2FCA9FD04086D" box="[439,685,793,814]" name="United States of America" pageId="7">United States of America</collectingCountry>
,
<collectingRegion id="49F1F87EFFD1FFC10FB2FCA9FCAB086D" box="[695,770,793,814]" country="Mexico" name="Mexico" pageId="7">
<collectingCountry id="F322760CFFD1FFC10FB2FCA9FCAB086D" box="[695,770,793,814]" name="Mexico" pageId="7">Mexico</collectingCountry>
</collectingRegion>
).......................
<taxonomicName id="4C354D1FFFD1FFC10922FCA9FA35086D" authority="(LeConte, 1854)" baseAuthorityName="LeConte" baseAuthorityYear="1854" box="[1063,1436,793,814]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="illatus">
<emphasis id="B941EA8EFFD1FFC10922FCA9FB5E086D" box="[1063,1271,793,814]" italics="true" pageId="7">Hemiphileurus illatus</emphasis>
(LeConte, 1854)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FC86FA3C082A" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FC86FA3C0808" blockId="7.[151,1436,294,1430]" box="[151,1429,822,843]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FC86FA3C0808" box="[151,1429,822,843]" pageId="7">7. Protarsal claw longer than mesotarsal and metatarsal claws.....................................................8</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FCE4FA3C082A" blockId="7.[151,1436,294,1430]" box="[151,1429,852,873]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FCE4FA3C082A" box="[151,1429,852,873]" pageId="7">- Protarsal claw similar in size to mesotarsal and metatarsal claws.............................................. 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FCC1FA3C089D" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FCC1FA3508E0" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FCC1FA3508E0" pageId="7">
8. Surface of head coarsely pitted, with dense covering of setae. Stridulatory area of mandible formed by approximately 30 sepa- rated ridges (
<collectingCountry id="F322760CFFD1FFC10C5BFC3EFE3208E0" box="[350,411,910,931]" name="Brazil" pageId="7">Brazil</collectingCountry>
)......................................................
<taxonomicName id="4C354D1FFFD1FFC10922FC3EFA3508E0" authority="Luderwaldt, 1910" authorityName="Luderwaldt" authorityYear="1910" box="[1063,1436,910,931]" class="Insecta" family="Dynastidae" genus="Actinobolus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="trilobus">
<emphasis id="B941EA8EFFD1FFC10922FC3EFB4308E0" box="[1063,1258,910,931]" italics="true" pageId="7">Actinobolus trilobus</emphasis>
Lüderwaldt, 1910
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FC1CFA3C089D" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FC1CFA3C089D" pageId="7">- Surface of head moderately punctate and sparsely setae. Stridulatory area of mandible formed by 40 or more separated ridges....................................................................................................9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FC56FA3F0F75" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FC56FA3C08B8" blockId="7.[151,1436,294,1430]" box="[151,1429,998,1019]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FC56FA3C08B8" box="[151,1429,998,1019]" pageId="7">9. Left lateral margin of labrum angulate................................................................... 10</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FBB4FA3F0F75" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FBB4FA3F0F75" pageId="7">
- Lateral margins of labrum broadly rounded, not angulate (northeastern South America)................................................................................................
<taxonomicName id="4C354D1FFFD1FFC10EF7FB91FA3F0F75" authority="(Ohaus, 1910)" baseAuthorityName="Ohaus" baseAuthorityYear="1910" box="[1010,1430,1057,1078]" class="Insecta" family="Dynastidae" genus="Homophileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B941EA8EFFD1FFC10EF7FB91FAAF0F75" box="[1010,1286,1057,1078]" italics="true" pageId="7">Homophileurus luederwaldti</emphasis>
(Ohaus, 1910)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FB8EFA3F0F32" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FB8EFA3F0F10" blockId="7.[151,1436,294,1430]" box="[151,1430,1086,1107]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FB8EFA3F0F10" box="[151,1430,1086,1107]" pageId="7">
10. Anterior clypeal setae absent (
<collectingCountry id="F322760CFFD1FFC10F06FB8EFDE80F10" box="[515,577,1086,1107]" name="Brazil" pageId="7">Brazil</collectingCountry>
,
<collectingRegion id="49F1F87EFFD1FFC10F4EFB8EFD0F0F10" box="[587,678,1086,1107]" country="Paraguay" name="Asuncion" pageId="7">
<collectingCountry id="F322760CFFD1FFC10F4EFB8EFD0F0F10" box="[587,678,1086,1107]" name="Paraguay" pageId="7">Paraguay</collectingCountry>
</collectingRegion>
)...........................
<taxonomicName id="4C354D1FFFD1FFC10EFEFB8EFA3F0F10" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1019,1430,1086,1107]" class="Insecta" family="Dynastidae" genus="Homophileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="integer">
<emphasis id="B941EA8EFFD1FFC10EFEFB8EFB730F10" box="[1019,1242,1086,1107]" italics="true" pageId="7">Homophileurus integer</emphasis>
(Burmeister, 1847)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FBECFA3F0F32" blockId="7.[151,1436,294,1430]" box="[151,1430,1116,1137]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FBECFA3F0F32" box="[151,1430,1116,1137]" pageId="7">
- Anterior clypeal setae present (Mexico to Brazil)...............................
<taxonomicName id="4C354D1FFFD1FFC1093DFBECFA3F0F32" authority="Prell, 1914" authorityName="Prell" authorityYear="1914" box="[1080,1430,1116,1137]" class="Insecta" family="Dynastidae" genus="Homophileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="B941EA8EFFD1FFC1093DFBECFA8F0F32" box="[1080,1318,1116,1137]" italics="true" pageId="7">Homophileurus tricuspis</emphasis>
Prell, 1914
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FBC9FA3C0FE8" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FBC9FA3C0FCD" blockId="7.[151,1436,294,1430]" box="[151,1429,1145,1166]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FBC9FA3C0FCD" box="[151,1429,1145,1166]" pageId="7">11. Ocelli absent....................................................................................... 12</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FB26FA3C0FE8" blockId="7.[151,1436,294,1430]" box="[151,1429,1174,1195]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FB26FA3C0FE8" box="[151,1429,1174,1195]" pageId="7">- Ocelli present....................................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FB04FA3F0E62" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FB04FA3F0FA5" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FB04FA3F0FA5" pageId="7">
12. Right lateral margin of labrum rounded, raster and lower anal lobe with short setae alternates with numerous long setae (South America)............................................................
<taxonomicName id="4C354D1FFFD1FFC10914FB61FA3F0FA5" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[1041,1430,1233,1254]" class="Insecta" family="Dynastidae" genus="Archophileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="vervex">
<emphasis id="B941EA8EFFD1FFC10914FB61FB430FA5" box="[1041,1258,1233,1254]" italics="true" pageId="7">Archophileurus vervex</emphasis>
Burmeister, 1847
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FB5EFA3F0E62" blockId="7.[151,1436,294,1430]" pageId="7">
<materialsCitation id="3B5D3CC1FFD1FFC10D92FB5EFA3F0E62" ID-GBIF-Occurrence="1703351499" country="Argentina" location="United States of America" pageId="7" specimenCount="1">
<keyLead id="30C495A9FFD1FFC10D92FB5EFA3F0E62" pageId="7">
- Right lateral margin of labrum angulate, raster and lower anal lobe with short and thin setae (
<location id="8EEA6047FFD1FFC10980FB5EFA290E40" LSID="urn:lsid:plazi:treatment:039C878AFFD1FFC10D92FF28FA320ED5:8EEA6047FFD1FFC10980FB5EFA290E40" box="[1157,1408,1262,1283]" country="Argentina" name="United States of America" pageId="7">United States of America</location>
to
<collectingCountry id="F322760CFFD1FFC10DE4FABCFEEB0E62" box="[225,322,1292,1313]" name="Argentina" pageId="7">Argentina</collectingCountry>
).................................................................
<taxonomicName id="4C354D1FFFD1FFC1095AFABCFA3F0E62" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[1119,1430,1292,1313]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="valgus">
<emphasis id="B941EA8EFFD1FFC1095AFABCFAA80E62" box="[1119,1281,1292,1313]" italics="true" pageId="7">Phileurus valgus</emphasis>
(Olivier, 1789)
</taxonomicName>
</keyLead>
</materialsCitation>
</paragraph>
</keyStep>
<keyStep id="30C12E39FFD1FFC10D92FA99FA320ED5" pageId="7">
<paragraph id="8B8A369CFFD1FFC10D92FA99FA3F0E18" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FA99FA3F0E18" pageId="7">
13. Lateral margins of labrum broadly rounded, not angulate. Laephoba with 11 slender, short setae Maxilla with a row of 7 strid- ulatory teeth (Mexico to Paraguay)...........................................
<taxonomicName id="4C354D1FFFD1FFC1093DFAF6FA3F0E18" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1080,1430,1350,1371]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="didymus">
<emphasis id="B941EA8EFFD1FFC1093DFAF6FB420E18" box="[1080,1259,1350,1371]" italics="true" pageId="7">Phileurus didymus</emphasis>
(Linnaeus, 1758)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8A369CFFD1FFC10D92FAD4FA320ED5" blockId="7.[151,1436,294,1430]" pageId="7">
<keyLead id="30C495A9FFD1FFC10D92FAD4FA320ED5" pageId="7">
- Left and right lateral margins of labrum angulate. Laephoba with 17 slender, short setae. Maxilla with a row of 9 stridulatory teeth (
<collectingRegion id="49F1F87EFFD1FFC10C19FA31FE070ED5" box="[284,430,1409,1430]" country="France" name="Guyane" pageId="7">
<collectingCountry id="F322760CFFD1FFC10C19FA31FE070ED5" box="[284,430,1409,1430]" name="French Guiana" pageId="7">French Guiana</collectingCountry>
</collectingRegion>
,
<collectingCountry id="F322760CFFD1FFC10CB2FA31FE5D0ED5" box="[439,500,1409,1430]" name="Brazil" pageId="7">Brazil</collectingCountry>
,
<collectingCountry id="F322760CFFD1FFC10CFBFA31FDFA0ED5" box="[510,595,1409,1430]" name="Uruguay" pageId="7">Uruguay</collectingCountry>
).........................................
<taxonomicName id="4C354D1FFFD1FFC10955FA31FA320ED5" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[1104,1435,1409,1430]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B941EA8EFFD1FFC10955FA31FB460ED5" box="[1104,1263,1409,1430]" italics="true" pageId="7">Phileurus affinis</emphasis>
Burmeister, 1847
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>