treatments-xml/data/01/30/77/01307731FF989932B2B5FE463FFEFB1C.xml
2024-06-21 12:21:35 +02:00

515 lines
66 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="C9B09E50C8CB2423792E13CD6FA5A1A8" ID-DOI="10.5281/zenodo.213642" ID-GBIF-Dataset="5c1cffb4-b78d-4739-9ec0-39664411f750" ID-ISSN="1175-5326" ID-Zenodo-Dep="213642" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460563468625" checkinUser="plazi" docAuthor="Huang, Chao, Huang, Jian Rong &amp; Ng, Peter K. L." docDate="2012" docId="01307731FF989932B2B5FE463FFEFB1C" docLanguage="en" docName="zt03588p063.pdf" docOrigin="Zootaxa 3588" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nanhaipotamon zhuhaiense Huang, Huang &amp; Ng, 2012, n. sp." docType="treatment" docVersion="8" lastPageNumber="62" masterDocId="FD090F49FF999935B222FF933B4CFFD0" masterDocTitle="A new species of Nanhaipotamon Bott, 1968 (Crustacea: Decapoda: Brachyura: Potamidae) from Zhuhai, Guangdong Province, China" masterLastPageNumber="63" masterPageNumber="55" pageNumber="56" updateTime="1698320599326" updateUser="plazi">
<mods:mods id="5E1FC474935A1991403C2C6E48B93AA1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A67881843F0F09BE83DAFE5C0214731D">
<mods:title id="C8AF12CBBC417F6EE4292B6B1339DDCF">A new species of Nanhaipotamon Bott, 1968 (Crustacea: Decapoda: Brachyura: Potamidae) from Zhuhai, Guangdong Province, China</mods:title>
</mods:titleInfo>
<mods:name id="07D0FA7EBB4A960DE86ED099172161A7" type="personal">
<mods:role id="1233875BAF7776293980CB8D07E0E26F">
<mods:roleTerm id="68CEE8FE90B3728394B0CF418D12A9D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="22365F7B1F819017F0B6A46749541322">Huang, Chao</mods:namePart>
</mods:name>
<mods:name id="6F893E1A4B2E8B31C1A58DDC7DA0FECE" type="personal">
<mods:role id="DEE873D7CA8466B59532FB3D0B423E68">
<mods:roleTerm id="A23E48335847C56689C9FAA98875C0B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3349D28CE6AE3769FC9749151F8EDD52">Huang, Jian Rong</mods:namePart>
</mods:name>
<mods:name id="9C809FE9ED1D96AF8EA6765D58BD8E13" type="personal">
<mods:role id="8AAFA6F30886B35EC896280BE35E873B">
<mods:roleTerm id="2C5A123F0A26D3B159A6321A4B374637">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AD4999339B7DD4D90C0F31040519DC8B">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CDDC181ED1B10CCF0DDD8CEF65C172F1">text</mods:typeOfResource>
<mods:relatedItem id="AEF1D84A65C967351ACE51D4E4F03EF2" type="host">
<mods:titleInfo id="6C848129CA5F99AC68EA14DC8617EB7F">
<mods:title id="8A60FBDD02BEB638F6CF33655F12433E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8774AAC6F292D49BFFEE145CFCC5419C">
<mods:date id="25B136C9D89105CFE829B8E4D49D8DDC">2012</mods:date>
<mods:detail id="1581DC7BC9CE46EBE599D575094BC9AB" type="volume">
<mods:number id="BD3693794E95B3340C75E0EE88370AF7">3588</mods:number>
</mods:detail>
<mods:extent id="C610A27EB50BC71C26430345EC7B4F0E" unit="page">
<mods:start id="F1A3330548E8206EEAE291C7A36CB343">55</mods:start>
<mods:end id="7BCF3F976305C9F2C71ED31E8605C59F">63</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F68EC8092E9CCB3C59C6198E79AF4E61">journal article</mods:classification>
<mods:identifier id="1E3D8F3F157267783580DE19B23A8949" type="DOI">10.5281/zenodo.213642</mods:identifier>
<mods:identifier id="9E40C64FFF47306542D76B7ADE17E3F3" type="GBIF-Dataset">5c1cffb4-b78d-4739-9ec0-39664411f750</mods:identifier>
<mods:identifier id="1F05F105F43AD4D7533AD436DD03FCC9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3FDD00313D145C6312E9631BA0D3CC48" type="Zenodo-Dep">213642</mods:identifier>
</mods:mods>
<treatment id="01307731FF989932B2B5FE463FFEFB1C" ID-DOI="http://doi.org/10.5281/zenodo.6180148" ID-GBIF-Taxon="119557176" ID-Zenodo-Dep="6180148" LSID="urn:lsid:plazi:treatment:01307731FF989932B2B5FE463FFEFB1C" httpUri="http://treatment.plazi.org/id/01307731FF989932B2B5FE463FFEFB1C" lastPageId="7" lastPageNumber="62" pageId="1" pageNumber="56">
<subSubSection id="C18395ACFF989934B2B5FE463A3CFDDF" pageId="1" pageNumber="56" type="nomenclature">
<paragraph id="8926C627FF989934B2B5FE463971FE3F" blockId="1.[151,573,469,527]" box="[151,573,469,495]" pageId="1" pageNumber="56">
<heading id="D26E714BFF989934B2B5FE463971FE3F" bold="true" box="[151,573,469,495]" fontSize="11" level="1" pageId="1" pageNumber="56" reason="1">
<emphasis id="BBED1A35FF989934B2B5FE463971FE3F" bold="true" box="[151,573,469,495]" pageId="1" pageNumber="56">
<taxonomicName id="4E99BDA4FF989934B2B5FE463AA0FE3F" ID-CoL="45LV4" box="[151,492,469,495]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF989934B2B5FE463AA0FE3F" bold="true" box="[151,492,469,495]" italics="true" pageId="1" pageNumber="56">Nanhaipotamon zhuhaiense</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A0DEA74EFF989934B3D8FE463971FE3F" box="[506,573,469,495]" pageId="1" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8926C627FF989934B2B5FE643A3CFDDF" blockId="1.[151,573,469,527]" box="[151,368,503,527]" pageId="1" pageNumber="56">
(
<figureCitation id="11A2DAA2FF989934B2BDFE643BA5FDDF" box="[159,233,503,527]" captionStart="FIGURE 1" captionStartId="2.[151,250,1833,1856]" captionTargetBox="[151,1435,220,1812]" captionTargetId="figure@2.[151,1436,193,1812]" captionTargetPageId="2" captionText="FIGURE 1. Colours in life. A, Nanhaipotamon guangdongense Dai, 1997, male (36.2 × 28.4 mm) (SYSU 001003); B, Nanhaipotamon zhuhaiense n. sp., male holotype (38.5 × 30.0 mm) (SYSU 001001), Zhuhai." httpUri="https://zenodo.org/record/213643/files/figure.png" pageId="1" pageNumber="56">Figs. 1</figureCitation>
B, 2, 3, 5D)
</paragraph>
</subSubSection>
<subSubSection id="C18395ACFF989934B2B5FDAC398DFC84" pageId="1" pageNumber="56" type="materials_examined">
<paragraph id="8926C627FF989934B2B5FDAC396FFD4F" blockId="1.[151,1437,575,1968]" pageId="1" pageNumber="56">
<emphasis id="BBED1A35FF989934B2B5FDAC3A32FD88" bold="true" box="[151,382,575,600]" pageId="1" pageNumber="56">Material examined</emphasis>
.
<typeStatus id="56227885FF989934B3B2FDAC3AB1FD87" box="[400,509,575,599]" pageId="1" pageNumber="56" type="holotype">Holotype</typeStatus>
: male (38.5 × 30.0 mm) (
<collectionCode id="EF885EE2FF989934B116FDAC3831FD87" LSID="urn:lsid:biocol.org:col:13728" box="[820,893,575,599]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="1" pageNumber="56">SYSU</collectionCode>
001001), Zhuhai, Guangdong,
<collectingCountry id="F18E86B7FF989934B6D2FDAC3E7AFD87" box="[1264,1334,575,599]" name="China" pageId="1" pageNumber="56">China</collectingCountry>
, coll. C. Huang,
<date id="FD27E0E7FF989934B2D4FDF63A3DFDAC" box="[246,369,612,637]" pageId="1" pageNumber="56" value="2012-05">May 2012</date>
.
<typeStatus id="56227885FF989934B3A0FDF63ABFFDAD" box="[386,499,613,637]" pageId="1" pageNumber="56" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9F9F0DAEFF989934B026FDF73919FDAC" box="[516,597,612,636]" pageId="1" pageNumber="56" type="male">1 male</specimenCount>
(27.4 ×
<quantity id="4E616BC2FF989934B0EAFDF7387FFDAC" box="[712,819,612,636]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="1" pageNumber="56" unit="mm" value="22.5">22.5 mm</quantity>
) (
<collectionCode id="EF885EE2FF989934B172FDF738D5FDAC" LSID="urn:lsid:biocol.org:col:13728" box="[848,921,612,636]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="1" pageNumber="56">SYSU</collectionCode>
001002),
<specimenCount id="9F9F0DAEFF989934B63AFDF73F25FDAC" box="[1048,1129,612,636]" pageId="1" pageNumber="56" type="male">1 male</specimenCount>
(30.5 ×
<quantity id="4E616BC2FF989934B6FEFDF73E0BFDAC" box="[1244,1351,612,636]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" pageId="1" pageNumber="56" unit="mm" value="25.2">25.2 mm</quantity>
) (
<collectionCode id="EF885EE2FF989934B746FDF73ED0FDAC" box="[1380,1436,612,636]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="56">ZRC</collectionCode>
2012.0792), same data as
<typeStatus id="56227885FF989934B398FD143953FD4F" box="[442,543,647,671]" pageId="1" pageNumber="56" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8926C627FF989934B2E5FD38398DFC84" blockId="1.[151,1437,575,1968]" pageId="1" pageNumber="56">
<emphasis id="BBED1A35FF989934B2E5FD383A9CFD14" bold="true" box="[199,464,683,708]" pageId="1" pageNumber="56">Comparative material</emphasis>
.
<taxonomicName id="4E99BDA4FF989934B3FCFD3F388FFD14" authority="Dai, 1997" authorityName="Dai" authorityYear="1997" box="[478,963,684,708]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="BBED1A35FF989934B3FCFD3F3806FD14" box="[478,842,684,708]" italics="true" pageId="1" pageNumber="56">Nanhaipotamon guangdongense</emphasis>
<bibRefCitation id="ED08BBD6FF989934B172FD3F388FFD14" author="Dai" box="[848,963,684,708]" pageId="1" pageNumber="56" refString="Dai, A. - Y. (1997) A revision of freshwater crabs of the genus Nanhaipotamon Bott, 1968 from China (Crustacea: Decapoda: Brachyura: Potamidae). Raffles Bulletin of Zoology, 45, 209 - 235." type="journal article" year="1997">Dai, 1997</bibRefCitation>
</taxonomicName>
:
<typeStatus id="56227885FF989934B1EFFD3F3F63FD14" box="[973,1071,684,708]" pageId="1" pageNumber="56" type="holotype">holotype</typeStatus>
male (33.2 ×
<quantity id="4E616BC2FF989934B6EEFD3F3E7DFD14" box="[1228,1329,684,708]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="1" pageNumber="56" unit="mm" value="26.4">26.4 mm</quantity>
) (AS-CB 05141), Guangdong Province, gift from Sun Yat-Sen Medical College, no date [photographs examined];
<specimenCount id="9F9F0DAEFF989934B76CFD5C3ED0FD37" box="[1358,1436,719,743]" pageId="1" pageNumber="56" type="male">1 male</specimenCount>
(36.2 ×
<quantity id="4E616BC2FF989934B2CCFD673A1FFCDC" box="[238,339,756,780]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="1" pageNumber="56" unit="mm" value="28.4">28.4 mm</quantity>
) (
<collectionCode id="EF885EE2FF989934B34BFD673AFEFCDC" LSID="urn:lsid:biocol.org:col:13728" box="[361,434,756,780]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="1" pageNumber="56">SYSU</collectionCode>
001003), Zhuhai, Guangdong,
<collectingCountry id="F18E86B7FF989934B133FD67381BFCDC" box="[785,855,756,780]" name="China" pageId="1" pageNumber="56">China</collectingCountry>
, coll. C. Huang,
<date id="FD27E0E7FF989934B630FD673F9CFCDC" box="[1042,1232,756,780]" pageId="1" pageNumber="56" value="2011-02-01">
February 2011,
<specimenCount id="9F9F0DAEFF989934B6E0FD673F9CFCDC" box="[1218,1232,756,780]" pageId="1" pageNumber="56" type="male">1</specimenCount>
</date>
male (30.5 ×
<quantity id="4E616BC2FF989934B749FD673B88FCFF" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="1" pageNumber="56" unit="mm" value="24.3">24.3 mm</quantity>
) (
<collectionCode id="EF885EE2FF989934B2FFFC843A6AFCFF" LSID="urn:lsid:biocol.org:col:13728" box="[221,294,791,815]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="1" pageNumber="56">SYSU</collectionCode>
001004),
<specimenCount id="9F9F0DAEFF989934B3BFFC843ABAFCFF" box="[413,502,791,815]" pageId="1" pageNumber="56" type="male">3 males</specimenCount>
(42.6 ×
<quantity id="4E616BC2FF989934B07FFC84398AFCE0" box="[605,710,791,816]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" pageId="1" pageNumber="56" unit="mm" value="25.6">25.6 mm</quantity>
, 30.7 ×
<quantity id="4E616BC2FF989934B10AFC8438DCFCE0" box="[808,912,791,816]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="1" pageNumber="56" unit="mm" value="25.3">25.3 mm</quantity>
, 22.1 ×
<quantity id="4E616BC2FF989934B1D0FC843F17FCFF" box="[1010,1115,791,815]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="1" pageNumber="56" unit="mm" value="18.1">18.1 mm</quantity>
) (
<collectionCode id="EF885EE2FF989934B651FC843FE7FCFF" box="[1139,1195,791,815]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="56">ZRC</collectionCode>
2012.0795), Zhuhai, Guangdong,
<collectingCountry id="F18E86B7FF989934B305FCAF3A21FC84" box="[295,365,828,852]" name="China" pageId="1" pageNumber="56">China</collectingCountry>
, coll. C. Huang,
<date id="FD27E0E7FF989934B008FCAF39F2FC84" box="[554,702,828,852]" pageId="1" pageNumber="56" value="2012-08">August 2012</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C18395ACFF989934B2E5FCCC3821FBA4" pageId="1" pageNumber="56" type="diagnosis">
<paragraph id="8926C627FF989934B2E5FCCC3821FBA4" blockId="1.[151,1437,575,1968]" pageId="1" pageNumber="56">
<emphasis id="BBED1A35FF989934B2E5FCCC3A76FCA8" bold="true" box="[199,314,863,888]" pageId="1" pageNumber="56">Diagnosis</emphasis>
. Carapace surface generally smooth (
<figureCitation id="11A2DAA2FF989934B0CAFCCC3861FCA7" box="[744,813,863,887]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[259,1322,183,1966]" captionTargetId="figure@3.[259,1323,183,1966]" captionTargetPageId="3" captionText="FIGURE 2. Nanhaipotamon zhuhaiense n. sp., male paratype (30.5 × 25.2 mm) (ZRC), Zhuhai. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/213644/files/figure.png" pageId="1" pageNumber="56">Fig. 2</figureCitation>
A); epigastric, postorbital cristae confluent, sharp (
<figureCitation id="11A2DAA2FF989934B74DFCCC3BE9FC4C" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[259,1322,183,1966]" captionTargetId="figure@3.[259,1323,183,1966]" captionTargetPageId="3" captionText="FIGURE 2. Nanhaipotamon zhuhaiense n. sp., male paratype (30.5 × 25.2 mm) (ZRC), Zhuhai. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/213644/files/figure.png" pageId="1" pageNumber="56">Fig. 2</figureCitation>
A, B); external orbital angle acutely triangular with small but sharp epibranchial tooth (
<figureCitation id="11A2DAA2FF989934B65DFC173F8EFC4C" box="[1151,1218,900,924]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[259,1322,183,1966]" captionTargetId="figure@3.[259,1323,183,1966]" captionTargetPageId="3" captionText="FIGURE 2. Nanhaipotamon zhuhaiense n. sp., male paratype (30.5 × 25.2 mm) (ZRC), Zhuhai. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/213644/files/figure.png" pageId="1" pageNumber="56">Fig. 2</figureCitation>
A, B); anterolateral border cristate, lined with numerous granules
<figureCitation id="11A2DAA2FF989934B0BBFC343996FC6F" box="[665,730,935,959]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[259,1322,183,1966]" captionTargetId="figure@3.[259,1323,183,1966]" captionTargetPageId="3" captionText="FIGURE 2. Nanhaipotamon zhuhaiense n. sp., male paratype (30.5 × 25.2 mm) (ZRC), Zhuhai. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/213644/files/figure.png" pageId="1" pageNumber="56">Fig. 2</figureCitation>
A, B); sub-orbital, subhepatic, pterygostomian regions covered with rounded granules (
<figureCitation id="11A2DAA2FF989934B386FC5F3AA6FC34" box="[420,490,972,996]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[259,1322,183,1966]" captionTargetId="figure@3.[259,1323,183,1966]" captionTargetPageId="3" captionText="FIGURE 2. Nanhaipotamon zhuhaiense n. sp., male paratype (30.5 × 25.2 mm) (ZRC), Zhuhai. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/213644/files/figure.png" pageId="1" pageNumber="56">Fig. 2</figureCitation>
B); male sterno-abdominal cavity reaching to imaginary line joining median part of coxae of chelipeds (
<figureCitation id="11A2DAA2FF989934B3ACFC7C3A96FBD7" box="[398,474,1007,1031]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[259,1322,183,1966]" captionTargetId="figure@3.[259,1323,183,1966]" captionTargetPageId="3" captionText="FIGURE 2. Nanhaipotamon zhuhaiense n. sp., male paratype (30.5 × 25.2 mm) (ZRC), Zhuhai. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/213644/files/figure.png" pageId="1" pageNumber="56">Fig. 2</figureCitation>
C); male abdomen broadly triangular (
<figureCitation id="11A2DAA2FF989934B18DFC7C3F4AFBD7" box="[943,1030,1007,1031]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[259,1322,183,1966]" captionTargetId="figure@3.[259,1323,183,1966]" captionTargetPageId="3" captionText="FIGURE 2. Nanhaipotamon zhuhaiense n. sp., male paratype (30.5 × 25.2 mm) (ZRC), Zhuhai. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/213644/files/figure.png" pageId="1" pageNumber="56">Figs. 2</figureCitation>
C, 3B); G1 slender, inner margin prominently concave, tip of terminal segment reaches beyond tubercle of abdominal locking structure
<emphasis id="BBED1A35FF989934B73BFB863E13FBFC" box="[1305,1375,1045,1068]" italics="true" pageId="1" pageNumber="56">in situ</emphasis>
(
<figureCitation id="11A2DAA2FF989934B74CFB873BE9FB9F" captionStart="FIGURE 3" captionStartId="4.[151,250,1709,1732]" captionTargetBox="[160,1419,214,1642]" captionTargetId="figure@4.[151,1436,193,1688]" captionTargetPageId="4" captionText="FIGURE 3. Nanhaipotamon zhuhaiense n. sp., male holotype (38.5 × 30.0 mm) (SYSU 001001), Zhuhai. A, left third maxilliped; B, male abdomen; C, sterno-abdominal cavity with G 1 in situ; D, left G 1 (ventral view); E, G 1 terminal segment (ventral view); F, left G 2." httpUri="https://zenodo.org/record/213645/files/figure.png" pageId="1" pageNumber="56">Fig. 3</figureCitation>
C), distal part distinctly elongated, bent laterally outwards at angle of about 90° to vertical (
<figureCitation id="11A2DAA2FF989934B68CFBA43FB7FB9F" box="[1198,1275,1079,1103]" captionStart="FIGURE 3" captionStartId="4.[151,250,1709,1732]" captionTargetBox="[160,1419,214,1642]" captionTargetId="figure@4.[151,1436,193,1688]" captionTargetPageId="4" captionText="FIGURE 3. Nanhaipotamon zhuhaiense n. sp., male holotype (38.5 × 30.0 mm) (SYSU 001001), Zhuhai. A, left third maxilliped; B, male abdomen; C, sterno-abdominal cavity with G 1 in situ; D, left G 1 (ventral view); E, G 1 terminal segment (ventral view); F, left G 2." httpUri="https://zenodo.org/record/213645/files/figure.png" pageId="1" pageNumber="56">Figs. 3</figureCitation>
D, E, 5D); G2 basal segment about 2.2 times length of distal segment (
<figureCitation id="11A2DAA2FF989934B128FBCF3801FBA4" box="[778,845,1116,1140]" captionStart="FIGURE 3" captionStartId="4.[151,250,1709,1732]" captionTargetBox="[160,1419,214,1642]" captionTargetId="figure@4.[151,1436,193,1688]" captionTargetPageId="4" captionText="FIGURE 3. Nanhaipotamon zhuhaiense n. sp., male holotype (38.5 × 30.0 mm) (SYSU 001001), Zhuhai. A, left third maxilliped; B, male abdomen; C, sterno-abdominal cavity with G 1 in situ; D, left G 1 (ventral view); E, G 1 terminal segment (ventral view); F, left G 2." httpUri="https://zenodo.org/record/213645/files/figure.png" pageId="1" pageNumber="56">Fig. 3</figureCitation>
F).
</paragraph>
</subSubSection>
<subSubSection id="C18395ACFF989931B2E5FBEC3BB0F806" lastPageId="4" lastPageNumber="59" pageId="1" pageNumber="56" type="description">
<paragraph id="8926C627FF989934B2E5FBEC3870F997" blockId="1.[151,1437,575,1968]" pageId="1" pageNumber="56">
<emphasis id="BBED1A35FF989934B2E5FBEC3A1DFB48" bold="true" box="[199,337,1151,1176]" pageId="1" pageNumber="56">Description</emphasis>
. Carapace surface generally smooth, convex transversely, longitudinally; regions behind front weakly rugose, pitted. Front deflexed; margin sinuous on dorsal view. Epigastric cristae low, separated by narrow gap. Epigastric, postorbital cristae confluent. Postorbital cristae sharp, confluent with epibranchial teeth. Branchial region swollen; cervical grooves shallow but visible, joining obscure H-shaped gastric groove. External orbital angle prominent, sharp, acutely triangular, outer margin gently convex to almost straight, longer than inner margin. Epibranchial tooth small but sharp, clearly demarcated from external orbital tooth by clear notch. Anterolateral border cristate, lined with numerous granules; posterolateral margin almost smooth or with very low oblique striae, sharply converging towards gently convex posterior carapace margin. Orbits large; margins sinuous, cristate. Suborbital, subhepatic, pterygostomian regions covered with rounded granules. Third maxilliped with merus about 1.1 times as broad as long, ischium about 1.5 times as long as broad; merus quadrate, with prominent median depression, covered with flattened granules; ischium subrectangular with distinct median sulcus; exopod reaching to proximal third of merus, flagellum about as long as width of merus. Posterior margin of epistome narrow; median triangular lobe large, lateral margins almost straight.
</paragraph>
<paragraph id="8926C627FF989934B2E5F9C73A99F907" blockId="1.[151,1437,575,1968]" pageId="1" pageNumber="56">Chelipeds unequal, especially in males. Merus trigonal in cross-section; margins crenulated, granulated, otherwise unarmed. Carpus with sharp spine on inner distal angle, with spinule at base; inner dorsal surface with curved row of low granules. Larger chela about 1.3 times as long as high; fixed finger subequal to movable finger, with small gap when closed.</paragraph>
<paragraph id="8926C627FF989934B2E5F9773AC4F8CF" blockId="1.[151,1437,575,1968]" pageId="1" pageNumber="56">Ambulatory legs slender, surfaces smooth, unarmed; last leg with propodus about 2.4 times as long as broad, shorter than dactylus.</paragraph>
<paragraph id="8926C627FF989934B2E5F8BF3A25F87F" blockId="1.[151,1437,575,1968]" pageId="1" pageNumber="56">Thoracic sternum generally smooth or weakly pitted; sternites 1, 2 completely fused to form triangular structure with convex lateral margins; sternites 3, 4 fused without trace of median sutures; male sterno-abdominal cavity reaching to imaginary line joining median part of coxae of cheliped; median longitudinal groove between sternites 7, 8 deep.</paragraph>
<caption id="DDE696AFFF9B9937B2B5F8BA3F07F88E" httpUri="https://zenodo.org/record/213643/files/figure.png" pageId="2" pageNumber="57" targetBox="[151,1435,220,1812]" targetPageId="2">
<paragraph id="8926C627FF9B9937B2B5F8BA3F07F88E" blockId="2.[151,1435,1833,1887]" pageId="2" pageNumber="57">
<emphasis id="BBED1A35FF9B9937B2B5F8BA3A5BF8EF" bold="true" box="[151,279,1833,1856]" pageId="2" pageNumber="57">FIGURE 1.</emphasis>
Colours in life. A,
<taxonomicName id="4E99BDA4FF9B9937B3D0F8B938F9F8EF" authority="Dai, 1997" authorityName="Dai" authorityYear="1997" box="[498,949,1834,1856]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="BBED1A35FF9B9937B3D0F8B93873F890" box="[498,831,1834,1856]" italics="true" pageId="2" pageNumber="57">Nanhaipotamon guangdongense</emphasis>
Dai, 1997
</taxonomicName>
, male (36.2 × 28.4 mm) (SYSU 001003); B,
<taxonomicName id="4E99BDA4FF9B9937B2B5F8DB3AFDF88E" box="[151,433,1864,1886]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9B9937B2B5F8DB3AFDF88E" box="[151,433,1864,1886]" italics="true" pageId="2" pageNumber="57">Nanhaipotamon zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9B9937B395F8DA3ABCF88E" bold="true" box="[439,496,1865,1886]" pageId="2" pageNumber="57">
<taxonomicNameLabel id="A0DEA74EFF9B9937B395F8DA3ABCF88E" box="[439,496,1865,1886]" pageId="2" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype (38.5 × 30.0 mm) (SYSU 001001), Zhuhai.
</paragraph>
</caption>
<paragraph id="8926C627FF9B9937B2E5F81838E5F83B" blockId="2.[151,1436,1931,2027]" pageId="2" pageNumber="57">Male abdomen broadly triangular; somites 36 progressively broader longitudinally, lateral margins almost straight to gently convex; somite 6 about 2.1 times as broad as long; telson about 1.2 times as broad as long, slightly concave proximally but otherwise gently convex, tip rounded.</paragraph>
<caption id="DDE696AFFF9A9936B2B5F8283F67F820" httpUri="https://zenodo.org/record/213644/files/figure.png" pageId="3" pageNumber="58" targetBox="[259,1322,183,1966]" targetPageId="3">
<paragraph id="8926C627FF9A9936B2B5F8283F67F820" blockId="3.[151,1435,1979,2032]" pageId="3" pageNumber="58">
<emphasis id="BBED1A35FF9A9936B2B5F8283A59F801" bold="true" box="[151,277,1979,2002]" pageId="3" pageNumber="58">FIGURE 2.</emphasis>
<taxonomicName id="4E99BDA4FF9A9936B33FF82F3976F802" box="[285,570,1980,2002]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9A9936B33FF82F3976F802" box="[285,570,1980,2002]" italics="true" pageId="3" pageNumber="58">Nanhaipotamon zhuhaiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0DEA74EFF9A9936B060F82F3931F801" box="[578,637,1980,2001]" pageId="3" pageNumber="58" rank="species">
<emphasis id="BBED1A35FF9A9936B060F82F3931F801" bold="true" box="[578,637,1980,2001]" pageId="3" pageNumber="58">n. sp.</emphasis>
</taxonomicNameLabel>
, male paratype (30.5 × 25.2 mm) (ZRC), Zhuhai. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen.
</paragraph>
</caption>
<caption id="DDE696AFFF9D9931B2B5F93E3AD6F8D0" httpUri="https://zenodo.org/record/213645/files/figure.png" pageId="4" pageNumber="59" targetBox="[160,1419,214,1642]" targetPageId="4">
<paragraph id="8926C627FF9D9931B2B5F93E3AD6F8D0" blockId="4.[151,1436,1709,1793]" pageId="4" pageNumber="59">
<emphasis id="BBED1A35FF9D9931B2B5F93E3A5BF913" bold="true" box="[151,279,1709,1732]" pageId="4" pageNumber="59">FIGURE 3.</emphasis>
<taxonomicName id="4E99BDA4FF9D9931B301F93D390EF914" box="[291,578,1710,1732]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9D9931B301F93D390EF914" box="[291,578,1710,1732]" italics="true" pageId="4" pageNumber="59">Nanhaipotamon zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9D9931B06EF93D39C6F913" bold="true" box="[588,650,1710,1731]" pageId="4" pageNumber="59">
<taxonomicNameLabel id="A0DEA74EFF9D9931B06EF93D39C6F913" box="[588,650,1710,1731]" pageId="4" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype (38.5 × 30.0 mm) (SYSU 001001), Zhuhai. A, left third maxilliped; B, male abdomen; C, sterno-abdominal cavity with G1 in situ; D, left G1 (ventral view); E, G1 terminal segment (ventral view); F, left G2.
</paragraph>
</caption>
<paragraph id="8926C627FF9D9931B2E5F8BE3BB0F806" blockId="4.[151,1436,1837,2006]" pageId="4" pageNumber="59">G1 slender, inner (sternal) margin prominently concave, tip of terminal segment reaches beyond tubercle of abdominal locking structure in situ; subterminal segment about 3.1 times length of terminal segment; inner-distal angle of terminal segment angular but edge rounded, anterior margin sinuous; distal part distinctly elongated, bent laterally outwards at angle of about 90° to vertical. G2 basal segment about 2.2 times length of flagelliform distal segment.</paragraph>
</subSubSection>
<caption id="DDE696AFFF9C9930B2B5F82F3E54F821" httpUri="https://zenodo.org/record/213646/files/figure.png" pageId="5" pageNumber="60" targetBox="[258,1328,193,1958]" targetPageId="5">
<paragraph id="8926C627FF9C9930B2B5F82F3E54F821" blockId="5.[151,1436,1980,2033]" pageId="5" pageNumber="60">
<emphasis id="BBED1A35FF9C9930B2B5F82F3A59F801" bold="true" box="[151,277,1980,2003]" pageId="5" pageNumber="60">FIGURE 4.</emphasis>
<taxonomicName id="4E99BDA4FF9C9930B33DF82F3990F802" authority="Dai, 1997" authorityName="Dai" authorityYear="1997" box="[287,732,1980,2002]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="BBED1A35FF9C9930B33DF82F3926F802" box="[287,618,1980,2002]" italics="true" pageId="5" pageNumber="60">Nanhaipotamon guangdongense</emphasis>
Dai, 1997
</taxonomicName>
, male holotype (33.2 × 26.4 mm) (AS-CB 05141), Guangdong. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen.
</paragraph>
</caption>
<caption id="DDE696AFFF9F9933B2B5FB9739D0FB88" httpUri="https://zenodo.org/record/213647/files/figure.png" pageId="6" pageNumber="61" targetBox="[151,1435,193,1006]" targetPageId="6">
<paragraph id="8926C627FF9F9933B2B5FB9739D0FB88" blockId="6.[151,1436,1028,1112]" pageId="6" pageNumber="61">
<emphasis id="BBED1A35FF9F9933B2B5FB973A5FFBCA" bold="true" box="[151,275,1028,1051]" pageId="6" pageNumber="61">FIGURE 5.</emphasis>
Left G1s (ventral views). AC,
<taxonomicName id="4E99BDA4FF9F9933B07EFB963F5EFBCA" authority="Dai, 1997" authorityName="Dai" authorityYear="1997" box="[604,1042,1028,1051]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="BBED1A35FF9F9933B07EFB9638E8FBCB" box="[604,932,1029,1051]" italics="true" pageId="6" pageNumber="61">Nanhaipotamon guangdongense</emphasis>
Dai, 1997
</taxonomicName>
; D,
<taxonomicName id="4E99BDA4FF9F9933B61EFB963E19FBCB" box="[1084,1365,1029,1051]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9F9933B61EFB963E19FBCB" box="[1084,1365,1029,1051]" italics="true" pageId="6" pageNumber="61">Nanhaipotamon zhuhaiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0DEA74EFF9F9933B77EFB963EDAFBCA" box="[1372,1430,1029,1050]" pageId="6" pageNumber="61" rank="species">
<emphasis id="BBED1A35FF9F9933B77EFB963EDAFBCA" bold="true" box="[1372,1430,1029,1050]" pageId="6" pageNumber="61">n. sp.</emphasis>
</taxonomicNameLabel>
. A, holotype male (33.2 × 26.4 mm) (AS-CB 05141), Guangdong; B, male (30.7 × 25.3 mm) (ZRC); C, male (42.6 × 25.6 mm) (ZRC); D, paratype male (30.5 × 25.2 mm) (ZRC).
</paragraph>
</caption>
<subSubSection id="C18395ACFF9F9933B2E5FB163925FA48" pageId="6" pageNumber="61" type="etymology">
<paragraph id="8926C627FF9F9933B2E5FB163ED7FB4C" blockId="6.[151,1436,1156,2009]" box="[199,1435,1156,1181]" pageId="6" pageNumber="61">
<emphasis id="BBED1A35FF9F9933B2E5FB163A08FB4D" bold="true" box="[199,324,1157,1181]" pageId="6" pageNumber="61">Etymology</emphasis>
.
<taxonomicName id="4E99BDA4FF9F9933B373FB1639CEFB4D" box="[337,642,1157,1181]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="zhuhaiense">
<emphasis id="BBED1A35FF9F9933B373FB1639CEFB4D" box="[337,642,1157,1181]" italics="true" pageId="6" pageNumber="61">Nanhaipotamon zhuhaiense</emphasis>
</taxonomicName>
is named after the
<typeStatus id="56227885FF9F9933B171FB1638CEFB4D" box="[851,898,1157,1181]" pageId="6" pageNumber="61">type</typeStatus>
locality, Zhuhai, in Guangdong Province,
<collectingCountry id="F18E86B7FF9F9933B770FB173EDBFB4C" box="[1362,1431,1156,1180]" name="China" pageId="6" pageNumber="61">China</collectingCountry>
.
</paragraph>
<paragraph id="8926C627FF9F9933B2E5FB3B3F99FB10" blockId="6.[151,1436,1156,2009]" box="[199,1237,1192,1217]" pageId="6" pageNumber="61">
<emphasis id="BBED1A35FF9F9933B2E5FB3B3A6CFB11" bold="true" box="[199,288,1192,1217]" pageId="6" pageNumber="61">Colour.</emphasis>
Carapace and ambulatory legs are bluish-white, chelipeds generally white (
<figureCitation id="11A2DAA2FF9F9933B652FB3B3FE3FB10" box="[1136,1199,1192,1216]" captionStart="FIGURE 1" captionStartId="2.[151,250,1833,1856]" captionTargetBox="[151,1435,220,1812]" captionTargetId="figure@2.[151,1436,193,1812]" captionTargetPageId="2" captionText="FIGURE 1. Colours in life. A, Nanhaipotamon guangdongense Dai, 1997, male (36.2 × 28.4 mm) (SYSU 001003); B, Nanhaipotamon zhuhaiense n. sp., male holotype (38.5 × 30.0 mm) (SYSU 001001), Zhuhai." httpUri="https://zenodo.org/record/213643/files/figure.png" pageId="6" pageNumber="61">Fig. 1</figureCitation>
B).
</paragraph>
<paragraph id="8926C627FF9F9933B2E5FB5E3925FA48" blockId="6.[151,1436,1156,2009]" pageId="6" pageNumber="61">
<emphasis id="BBED1A35FF9F9933B2E5FB5E3A68FB35" bold="true" box="[199,292,1229,1253]" pageId="6" pageNumber="61">Ecology</emphasis>
. Like
<taxonomicName id="4E99BDA4FF9F9933B34CFB5D390CFB35" box="[366,576,1229,1253]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="BBED1A35FF9F9933B34CFB5D390CFB35" box="[366,576,1229,1253]" italics="true" pageId="6" pageNumber="61">N. guangdongense</emphasis>
</taxonomicName>
, the new species inhabits mud burrows near the banks of densely vegetated creeks. The burrows can be as deep as one meter and always have some water at the bottom. Crabs reside in their burrows during the day and are relatively inactive, but are particularly active during hot, humid nights. They are omnivorous, eating dead leaves, earthworms, freshwater shrimps, other crabs, etc. Brooding females have been observed in February. Cannibalism has been observed in captive specimens and may also occur in the wild; large specimens are generally more aggressive.
</paragraph>
</subSubSection>
<subSubSection id="C18395ACFF9F9932B2E5FA363FFEFB1C" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61" type="discussion">
<paragraph id="8926C627FF9F9933B2E5FA363AB5F9F8" blockId="6.[151,1436,1156,2009]" pageId="6" pageNumber="61">
<emphasis id="BBED1A35FF9F9933B2E5FA363A78FA6D" bold="true" box="[199,308,1445,1469]" pageId="6" pageNumber="61">Remarks</emphasis>
.
<taxonomicName id="4E99BDA4FF9F9933B361FA363931FA6D" box="[323,637,1445,1469]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9F9933B361FA363931FA6D" box="[323,637,1445,1469]" italics="true" pageId="6" pageNumber="61">Nanhaipotamon zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9F9933B0A7FA36398AFA6D" bold="true" box="[645,710,1445,1469]" pageId="6" pageNumber="61">
<taxonomicNameLabel id="A0DEA74EFF9F9933B0A7FA36398AFA6D" box="[645,710,1445,1469]" pageId="6" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can most easily be distinguished from
<taxonomicName id="4E99BDA4FF9F9933B6AAFA353E01FA6D" box="[1160,1357,1445,1469]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guandongense">
<emphasis id="BBED1A35FF9F9933B6AAFA353E01FA6D" box="[1160,1357,1445,1469]" italics="true" pageId="6" pageNumber="61">N. guandongense</emphasis>
</taxonomicName>
by the live coloration of its carapace, the former being bluish-white (
<figureCitation id="11A2DAA2FF9F9933B179FA5B38D0FA30" box="[859,924,1480,1504]" captionStart="FIGURE 1" captionStartId="2.[151,250,1833,1856]" captionTargetBox="[151,1435,220,1812]" captionTargetId="figure@2.[151,1436,193,1812]" captionTargetPageId="2" captionText="FIGURE 1. Colours in life. A, Nanhaipotamon guangdongense Dai, 1997, male (36.2 × 28.4 mm) (SYSU 001003); B, Nanhaipotamon zhuhaiense n. sp., male holotype (38.5 × 30.0 mm) (SYSU 001001), Zhuhai." httpUri="https://zenodo.org/record/213643/files/figure.png" pageId="6" pageNumber="61">Fig. 1</figureCitation>
B) with the latter generally brown (
<figureCitation id="11A2DAA2FF9F9933B711FA5B3E38FA30" box="[1331,1396,1480,1504]" captionStart="FIGURE 1" captionStartId="2.[151,250,1833,1856]" captionTargetBox="[151,1435,220,1812]" captionTargetId="figure@2.[151,1436,193,1812]" captionTargetPageId="2" captionText="FIGURE 1. Colours in life. A, Nanhaipotamon guangdongense Dai, 1997, male (36.2 × 28.4 mm) (SYSU 001003); B, Nanhaipotamon zhuhaiense n. sp., male holotype (38.5 × 30.0 mm) (SYSU 001001), Zhuhai." httpUri="https://zenodo.org/record/213643/files/figure.png" pageId="6" pageNumber="61">Fig. 1</figureCitation>
A). Although the colour of
<taxonomicName id="4E99BDA4FF9F9933B38EFA7D393EF9D5" box="[428,626,1517,1541]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guandongense">
<emphasis id="BBED1A35FF9F9933B38EFA7D393EF9D5" box="[428,626,1517,1541]" italics="true" pageId="6" pageNumber="61">N. guandongense</emphasis>
</taxonomicName>
varies slightly from yellowish to orangish-brown, it never attains the colours of
<taxonomicName id="4E99BDA4FF9F9933B32CF9813AE7F9F9" box="[270,427,1553,1577]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9F9933B32CF9813AE7F9F9" box="[270,427,1553,1577]" italics="true" pageId="6" pageNumber="61">N. zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9F9933B391F9823ABDF9F9" bold="true" box="[435,497,1553,1577]" pageId="6" pageNumber="61">
<taxonomicNameLabel id="A0DEA74EFF9F9933B391F9823ABDF9F9" box="[435,497,1553,1577]" pageId="6" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
<paragraph id="8926C627FF9F9933B2E5F9A63969F808" blockId="6.[151,1436,1156,2009]" pageId="6" pageNumber="61">
As with most
<taxonomicName id="4E99BDA4FF9F9933B348F9A63953F99D" box="[362,543,1589,1613]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="BBED1A35FF9F9933B348F9A63953F99D" box="[362,543,1589,1613]" italics="true" pageId="6" pageNumber="61">Nanhaipotamon</emphasis>
</taxonomicName>
species, the carapace, chelipeds and ambulatory legs are extremely similar in form and we could discern no major differences between
<taxonomicName id="4E99BDA4FF9F9933B111F9C938B5F9A1" box="[819,1017,1625,1649]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guandongense">
<emphasis id="BBED1A35FF9F9933B111F9C938B5F9A1" box="[819,1017,1625,1649]" italics="true" pageId="6" pageNumber="61">N. guandongense</emphasis>
</taxonomicName>
and
<taxonomicName id="4E99BDA4FF9F9933B616F9C93F98F9A1" box="[1076,1236,1625,1649]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9F9933B616F9C93F98F9A1" box="[1076,1236,1625,1649]" italics="true" pageId="6" pageNumber="61">N. zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9F9933B6FFF9CA3E53F9A1" bold="true" box="[1245,1311,1625,1649]" pageId="6" pageNumber="61">
<taxonomicNameLabel id="A0DEA74EFF9F9933B6FFF9CA3E53F9A1" box="[1245,1311,1625,1649]" pageId="6" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. The male abdomen of
<taxonomicName id="4E99BDA4FF9F9933B305F9ED3AA0F945" box="[295,492,1661,1685]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guandongense">
<emphasis id="BBED1A35FF9F9933B305F9ED3AA0F945" box="[295,492,1661,1685]" italics="true" pageId="6" pageNumber="61">N. guandongense</emphasis>
</taxonomicName>
is relatively wider, especially somite 6 (
<figureCitation id="11A2DAA2FF9F9933B19DF9EF3F48F944" box="[959,1028,1660,1684]" captionStart="FIGURE 4" captionStartId="5.[151,250,1980,2003]" captionTargetBox="[258,1328,193,1958]" captionTargetId="figure@5.[258,1329,193,1959]" captionTargetPageId="5" captionText="FIGURE 4. Nanhaipotamon guangdongense Dai, 1997, male holotype (33.2 × 26.4 mm) (AS-CB 05141), Guangdong. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/213646/files/figure.png" pageId="6" pageNumber="61">Fig. 4</figureCitation>
C), with that of
<taxonomicName id="4E99BDA4FF9F9933B69CF9ED3E11F945" box="[1214,1373,1661,1685]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9F9933B69CF9ED3E11F945" box="[1214,1373,1661,1685]" italics="true" pageId="6" pageNumber="61">N. zhuhaiense</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A0DEA74EFF9F9933B74FF9EE3BA7F968" pageId="6" pageNumber="61" rank="species">new species</taxonomicNameLabel>
, relatively narrower (
<figureCitation id="11A2DAA2FF9F9933B3F8F9333951F968" box="[474,541,1696,1720]" captionStart="FIGURE 2" captionStartId="3.[151,250,1979,2002]" captionTargetBox="[259,1322,183,1966]" captionTargetId="figure@3.[259,1323,183,1966]" captionTargetPageId="3" captionText="FIGURE 2. Nanhaipotamon zhuhaiense n. sp., male paratype (30.5 × 25.2 mm) (ZRC), Zhuhai. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/213644/files/figure.png" pageId="6" pageNumber="61">Fig. 2</figureCitation>
C) (Table 1). The male abdomen of
<taxonomicName id="4E99BDA4FF9F9933B18CF9313F3CF969" box="[942,1136,1697,1721]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guandongense">
<emphasis id="BBED1A35FF9F9933B18CF9313F3CF969" box="[942,1136,1697,1721]" italics="true" pageId="6" pageNumber="61">N. guandongense</emphasis>
</taxonomicName>
(see
<bibRefCitation id="ED08BBD6FF9F9933B689F9333E5AF968" author="Dai" box="[1195,1302,1696,1720]" pageId="6" pageNumber="61" refString="Dai, A. - Y. (1997) A revision of freshwater crabs of the genus Nanhaipotamon Bott, 1968 from China (Crustacea: Decapoda: Brachyura: Potamidae). Raffles Bulletin of Zoology, 45, 209 - 235." type="journal article" year="1997">Dai 1997</bibRefCitation>
: Fig. 9-2) is not accurate as it shows a relatively narrower structure. The male abdomen of the
<typeStatus id="56227885FF9F9933B68CF9573E5CF90C" box="[1198,1296,1732,1756]" pageId="6" pageNumber="61" type="holotype">holotype</typeStatus>
is actually proportionately wider (
<figureCitation id="11A2DAA2FF9F9933B380F97B3AA7F8D0" box="[418,491,1768,1792]" captionStart="FIGURE 4" captionStartId="5.[151,250,1980,2003]" captionTargetBox="[258,1328,193,1958]" captionTargetId="figure@5.[258,1329,193,1959]" captionTargetPageId="5" captionText="FIGURE 4. Nanhaipotamon guangdongense Dai, 1997, male holotype (33.2 × 26.4 mm) (AS-CB 05141), Guangdong. A, dorsal overall view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/213646/files/figure.png" pageId="6" pageNumber="61">Fig. 4</figureCitation>
C). The most significant difference is in the structure of the G1. While they are superficially similar, that of
<taxonomicName id="4E99BDA4FF9F9933B3F7F89D39D4F8F5" box="[469,664,1805,1829]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guandongense">
<emphasis id="BBED1A35FF9F9933B3F7F89D39D4F8F5" box="[469,664,1805,1829]" italics="true" pageId="6" pageNumber="61">N. guandongense</emphasis>
</taxonomicName>
is more curved, with the inner distal margin distinctly concave (
<figureCitation id="11A2DAA2FF9F9933B74CF89F3BE8F898" captionStart="FIGURE 5" captionStartId="6.[151,250,1028,1051]" captionTargetBox="[151,1435,193,1006]" captionTargetId="figure@6.[151,1436,193,1007]" captionTargetPageId="6" captionText="FIGURE 5. Left G 1 s (ventral views). A C, Nanhaipotamon guangdongense Dai, 1997; D, Nanhaipotamon zhuhaiense n. sp.. A, holotype male (33.2 × 26.4 mm) (AS-CB 05141), Guangdong; B, male (30.7 × 25.3 mm) (ZRC); C, male (42.6 × 25.6 mm) (ZRC); D, paratype male (30.5 × 25.2 mm) (ZRC)." httpUri="https://zenodo.org/record/213647/files/figure.png" pageId="6" pageNumber="61">Fig. 5</figureCitation>
AC), while that of
<taxonomicName id="4E99BDA4FF9F9933B3A5F8A13969F899" box="[391,549,1841,1865]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9F9933B3A5F8A13969F899" box="[391,549,1841,1865]" italics="true" pageId="6" pageNumber="61">N. zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9F9933B00EF8A23920F899" bold="true" box="[556,620,1841,1865]" pageId="6" pageNumber="61">
<taxonomicNameLabel id="A0DEA74EFF9F9933B00EF8A23920F899" box="[556,620,1841,1865]" pageId="6" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, is relatively straight (
<figureCitation id="11A2DAA2FF9F9933B14BF8A338E9F898" box="[873,933,1840,1864]" captionStart="FIGURE 5" captionStartId="6.[151,250,1028,1051]" captionTargetBox="[151,1435,193,1006]" captionTargetId="figure@6.[151,1436,193,1007]" captionTargetPageId="6" captionText="FIGURE 5. Left G 1 s (ventral views). A C, Nanhaipotamon guangdongense Dai, 1997; D, Nanhaipotamon zhuhaiense n. sp.. A, holotype male (33.2 × 26.4 mm) (AS-CB 05141), Guangdong; B, male (30.7 × 25.3 mm) (ZRC); C, male (42.6 × 25.6 mm) (ZRC); D, paratype male (30.5 × 25.2 mm) (ZRC)." httpUri="https://zenodo.org/record/213647/files/figure.png" pageId="6" pageNumber="61">Fig 5</figureCitation>
D). The form of the terminal segment is also different; in
<taxonomicName id="4E99BDA4FF9F9933B300F8C53AF3F8BD" box="[290,447,1877,1901]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9F9933B300F8C53AF3F8BD" box="[290,447,1877,1901]" italics="true" pageId="6" pageNumber="61">N. zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9F9933B3E4F8C63948F8BD" bold="true" box="[454,516,1877,1901]" pageId="6" pageNumber="61">
<taxonomicNameLabel id="A0DEA74EFF9F9933B3E4F8C63948F8BD" box="[454,516,1877,1901]" pageId="6" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
the distal part is distinctively more elongated and is bent laterally at almost 90° to the vertical (
<figureCitation id="11A2DAA2FF9F9933B300F8EB3A22F840" box="[290,366,1912,1936]" captionStart="FIGURE 3" captionStartId="4.[151,250,1709,1732]" captionTargetBox="[160,1419,214,1642]" captionTargetId="figure@4.[151,1436,193,1688]" captionTargetPageId="4" captionText="FIGURE 3. Nanhaipotamon zhuhaiense n. sp., male holotype (38.5 × 30.0 mm) (SYSU 001001), Zhuhai. A, left third maxilliped; B, male abdomen; C, sterno-abdominal cavity with G 1 in situ; D, left G 1 (ventral view); E, G 1 terminal segment (ventral view); F, left G 2." httpUri="https://zenodo.org/record/213645/files/figure.png" pageId="6" pageNumber="61">Figs. 3</figureCitation>
D, E, 5D). In
<taxonomicName id="4E99BDA4FF9F9933B024F8E93984F841" box="[518,712,1913,1937]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guandongense">
<emphasis id="BBED1A35FF9F9933B024F8E93984F841" box="[518,712,1913,1937]" italics="true" pageId="6" pageNumber="61">N. guandongense</emphasis>
</taxonomicName>
, the distal part is shorter and bent obliquely outwards at an angle of about 45° to the vertical (
<figureCitation id="11A2DAA2FF9F9933B3F1F80F3958F864" box="[467,532,1948,1972]" captionStart="FIGURE 5" captionStartId="6.[151,250,1028,1051]" captionTargetBox="[151,1435,193,1006]" captionTargetId="figure@6.[151,1436,193,1007]" captionTargetPageId="6" captionText="FIGURE 5. Left G 1 s (ventral views). A C, Nanhaipotamon guangdongense Dai, 1997; D, Nanhaipotamon zhuhaiense n. sp.. A, holotype male (33.2 × 26.4 mm) (AS-CB 05141), Guangdong; B, male (30.7 × 25.3 mm) (ZRC); C, male (42.6 × 25.6 mm) (ZRC); D, paratype male (30.5 × 25.2 mm) (ZRC)." httpUri="https://zenodo.org/record/213647/files/figure.png" pageId="6" pageNumber="61">Fig. 5</figureCitation>
AC). These differences are consistent and are beyond the variation observed or known for
<taxonomicName id="4E99BDA4FF9F9933B330F8523A8BF809" box="[274,455,1985,2009]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="BBED1A35FF9F9933B330F8523A8BF809" box="[274,455,1985,2009]" italics="true" pageId="6" pageNumber="61">Nanhaipotamon</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="8926C627FF9E9932B2B5FF0B38BAFB54" blockId="7.[151,1253,152,176]" lastBlockId="7.[151,1437,664,1228]" pageId="7" pageNumber="62">
<emphasis id="BBED1A35FF9E9932B2B5FF0B3A4BFF60" bold="true" box="[151,263,152,176]" pageId="7" pageNumber="62">TABLE I.</emphasis>
Morphological differences among
<taxonomicName id="4E99BDA4FF9E9932B05CFF0A38E4FF60" box="[638,936,153,176]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9E9932B05CFF0A38E4FF60" box="[638,936,153,176]" italics="true" pageId="7" pageNumber="62">Nanhaipotamon zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9E9932B18CFF0B38A6FF60" bold="true" box="[942,1002,152,176]" pageId="7" pageNumber="62">
<taxonomicNameLabel id="A0DEA74EFF9E9932B18CFF0B38A6FF60" box="[942,1002,152,176]" pageId="7" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="BBED1A35FF9E9932B63CFF0A394CFD60" italics="true" pageId="7" pageNumber="62">
<taxonomicName id="4E99BDA4FF9E9932B63CFF0A3FA9FF60" box="[1054,1253,153,176]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="guangdongense">N. guangdongense</taxonomicName>
<taxonomicName id="4E99BDA4FF9E9932B2E5FD0B394CFD60" box="[199,512,664,688]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">Nanhaipotamon zhuhaiense</taxonomicName>
</emphasis>
<emphasis id="BBED1A35FF9E9932B02AFD0B3904FD60" bold="true" box="[520,584,664,688]" pageId="7" pageNumber="62">
<taxonomicNameLabel id="A0DEA74EFF9E9932B02AFD0B3904FD60" box="[520,584,664,688]" pageId="7" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has been found only in small numbers from just one site in Zhuhai, where they are sympatric with
<taxonomicName id="4E99BDA4FF9E9932B381FD2E393FFD04" box="[419,627,700,724]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="BBED1A35FF9E9932B381FD2E393FFD04" box="[419,627,700,724]" italics="true" pageId="7" pageNumber="62">N. guangdongense</emphasis>
</taxonomicName>
. Their population does not appear to be large, and considering their very limited range, we had concerns about their conservation status and therefore restricted our collection of samples. Observations of several more adult and juvenile specimens of
<taxonomicName id="4E99BDA4FF9E9932B153FC963F5EFCCC" box="[881,1042,772,796]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9E9932B153FC963F5EFCCC" box="[881,1042,772,796]" italics="true" pageId="7" pageNumber="62">N. zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9E9932B63FFC973F13FCCC" bold="true" box="[1053,1119,772,796]" pageId="7" pageNumber="62">
<taxonomicNameLabel id="A0DEA74EFF9E9932B63FFC973F13FCCC" box="[1053,1119,772,796]" pageId="7" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in the field and laboratory nevertheless confirm the validity of the colour differences observed with
<taxonomicName id="4E99BDA4FF9E9932B1C0FCBA3FF9FC90" box="[994,1205,808,832]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="BBED1A35FF9E9932B1C0FCBA3FF9FC90" box="[994,1205,808,832]" italics="true" pageId="7" pageNumber="62">N. guangdongense</emphasis>
</taxonomicName>
. Although there are reports, based on the basis of their distinctive coloration, that it is present in other areas around of Zhuhai, we have not been able to find them. In all the other areas surveyed around Zhuhai, the only species present is
<taxonomicName id="4E99BDA4FF9E9932B7A0FCE23A0BFC7C" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="BBED1A35FF9E9932B7A0FCE23A0BFC7C" italics="true" pageId="7" pageNumber="62">N. guangdongense</emphasis>
</taxonomicName>
. This is only the second case of sympatric occurrence of two species in the genus, the other being
<taxonomicName id="4E99BDA4FF9E9932B2B5FC2A3AE0FC00" authority="Dai, 1997" authorityName="Dai" authorityYear="1997" box="[151,428,952,976]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="hepingense">
<emphasis id="BBED1A35FF9E9932B2B5FC2A3A79FC00" box="[151,309,952,976]" italics="true" pageId="7" pageNumber="62">N. hepingense</emphasis>
<bibRefCitation id="ED08BBD6FF9E9932B319FC2B3AE0FC00" author="Dai" box="[315,428,952,976]" pageId="7" pageNumber="62" refString="Dai, A. - Y. (1997) A revision of freshwater crabs of the genus Nanhaipotamon Bott, 1968 from China (Crustacea: Decapoda: Brachyura: Potamidae). Raffles Bulletin of Zoology, 45, 209 - 235." type="journal article" year="1997">Dai, 1997</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4E99BDA4FF9E9932B3C4FC2A39A3FC00" authority="Dai, 1997" authorityName="Dai" authorityYear="1997" box="[486,751,952,976]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="pinghense">
<emphasis id="BBED1A35FF9E9932B3C4FC2A3934FC00" box="[486,632,952,976]" italics="true" pageId="7" pageNumber="62">N. pinghense</emphasis>
<bibRefCitation id="ED08BBD6FF9E9932B05CFC2B39A3FC00" author="Dai" box="[638,751,952,976]" pageId="7" pageNumber="62" refString="Dai, A. - Y. (1997) A revision of freshwater crabs of the genus Nanhaipotamon Bott, 1968 from China (Crustacea: Decapoda: Brachyura: Potamidae). Raffles Bulletin of Zoology, 45, 209 - 235." type="journal article" year="1997">Dai, 1997</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="ED08BBD6FF9E9932B0D8FC2B38FAFC00" author="Shih" box="[762,950,952,976]" pageId="7" pageNumber="62" refString="Shih, H. - T., Zhou X. - M., Chen G. - X., Chien I-C. &amp; Ng, P. K. L. (2011) Recent vicariant and dispersal events affecting the phylogeny and biogeography of East Asian freshwater crab genus Nanhaipotamon (Decapoda: Potamidae). Molecular Phylogenetics and Evolution, 58, 427 - 438." type="journal article" year="2011">
Shih
<emphasis id="BBED1A35FF9E9932B116FC2A3820FC00" box="[820,876,952,976]" italics="true" pageId="7" pageNumber="62">et al.</emphasis>
(2011
</bibRefCitation>
: 437) indicated, however, that there is good evidence that these two species (both described on the basis of only one male each from the same location and collected on the same date) are synonymous. In the case of
<taxonomicName id="4E99BDA4FF9E9932B162FB923F5EFBC8" box="[832,1042,1024,1048]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="BBED1A35FF9E9932B162FB923F5EFBC8" box="[832,1042,1024,1048]" italics="true" pageId="7" pageNumber="62">N. guangdongense</emphasis>
</taxonomicName>
and
<taxonomicName id="4E99BDA4FF9E9932B669FB923FA5FBC8" box="[1099,1257,1024,1048]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9E9932B669FB923FA5FBC8" box="[1099,1257,1024,1048]" italics="true" pageId="7" pageNumber="62">N. zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9E9932B6D0FB933E7EFBC8" bold="true" box="[1266,1330,1024,1048]" pageId="7" pageNumber="62">
<taxonomicNameLabel id="A0DEA74EFF9E9932B6D0FB933E7EFBC8" box="[1266,1330,1024,1048]" pageId="7" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
the most likely explanation is that the two species were originally allopatric and human activities (land development and moving of agricultural plants and animals) causing part of their ranges to overlap. More thorough surveys will need to be done to determine the precise and original distributions of both species.
</paragraph>
<paragraph id="8926C627FF9E9932B287FF423FEFFD84" pageId="7" pageNumber="62">
<table id="FB993487FF9E66CAB287FF423EC4FD84" box="[165,1416,209,596]" gridcols="3" gridrows="7" pageId="7" pageNumber="62">
<tr id="37A9C465FF9E66CAB287FF423EC4FF38" box="[165,1416,209,232]" gridrow="0" pageId="7" pageNumber="62">
<th id="7478AD19FF9E66CAB287FF4239D5FF38" box="[165,665,209,232]" gridcol="0" gridrow="0" pageId="7" pageNumber="62">Character</th>
<th id="7478AD19FF9E66CAB097FF423F48FF38" box="[693,1028,209,232]" gridcol="1" gridrow="0" pageId="7" pageNumber="62">
<taxonomicName id="4E99BDA4FF9E9932B097FF40380EFF37" box="[693,834,209,232]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9E9932B097FF40380EFF37" box="[693,834,209,232]" italics="true" pageId="7" pageNumber="62">N. zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9E9932B16BFF4138CEFF37" bold="true" box="[841,898,210,231]" pageId="7" pageNumber="62">
<taxonomicNameLabel id="A0DEA74EFF9E9932B16BFF4138CEFF37" box="[841,898,210,231]" pageId="7" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
<th id="7478AD19FF9E66CAB602FF423EC4FF38" box="[1056,1416,209,232]" gridcol="2" gridrow="0" pageId="7" pageNumber="62">
<taxonomicName id="4E99BDA4FF9E9932B602FF403E77FF37" authority="Dai, 1997" authorityName="Dai" authorityYear="1997" box="[1056,1339,209,232]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="guandongense">
<emphasis id="BBED1A35FF9E9932B602FF403F9CFF37" box="[1056,1232,209,232]" italics="true" pageId="7" pageNumber="62">N. guandongense</emphasis>
Dai, 1997
</taxonomicName>
</th>
</tr>
<tr id="37A9C465FF9E66CAB287FF6E3EC4FEE5" box="[165,1416,253,309]" gridrow="1" pageId="7" pageNumber="62">
<th id="7478AD19FF9E66CAB287FF6E39D5FEE5" box="[165,665,253,309]" gridcol="0" gridrow="1" pageId="7" pageNumber="62">Ratio of lengths of G1 subterminal to terminal segments</th>
<td id="7478AD19FF9E66CAB097FF6E3F48FEE5" box="[693,1028,253,309]" gridcol="1" gridrow="1" pageId="7" pageNumber="62">3.1 (Figs. 3D, E, 5D)</td>
<td id="7478AD19FF9E66CAB602FF6E3EC4FEE5" box="[1056,1416,253,309]" gridcol="2" gridrow="1" pageId="7" pageNumber="62">2.7 (Fig. 5AC)</td>
</tr>
<tr id="37A9C465FF9E66CAB287FED93EC4FE52" box="[165,1416,330,386]" gridrow="2" pageId="7" pageNumber="62">
<th id="7478AD19FF9E66CAB287FED939D5FE52" box="[165,665,330,386]" gridcol="0" gridrow="2" pageId="7" pageNumber="62">Tip of G1 in relation to tubercle of male abdominal locking structure, in situ</th>
<td id="7478AD19FF9E66CAB097FED93F48FE52" box="[693,1028,330,386]" gridcol="1" gridrow="2" pageId="7" pageNumber="62">reaches outwards, beyond tubercle (Fig. 3C)</td>
<td id="7478AD19FF9E66CAB602FED93EC4FE52" box="[1056,1416,330,386]" gridcol="2" gridrow="2" pageId="7" pageNumber="62">does not reach tubercle</td>
</tr>
<tr id="37A9C465FF9E66CAB287FE0B3EC4FE7E" box="[165,1416,408,430]" gridrow="3" pageId="7" pageNumber="62">
<th id="7478AD19FF9E66CAB287FE0B39D5FE7E" box="[165,665,408,430]" gridcol="0" gridrow="3" pageId="7" pageNumber="62">Inner margin of G1 terminal segment</th>
<td id="7478AD19FF9E66CAB097FE0B3F48FE7E" box="[693,1028,408,430]" gridcol="1" gridrow="3" pageId="7" pageNumber="62">gently concave (Figs. 3D, E, 5D)</td>
<td id="7478AD19FF9E66CAB602FE0B3EC4FE7E" box="[1056,1416,408,430]" gridcol="2" gridrow="3" pageId="7" pageNumber="62">prominently concave (Fig. 5AC)</td>
</tr>
<tr id="37A9C465FF9E66CAB287FE573EC4FE2B" box="[165,1416,452,507]" gridrow="4" pageId="7" pageNumber="62">
<th id="7478AD19FF9E66CAB287FE5739D5FE2B" box="[165,665,452,507]" gridcol="0" gridrow="4" pageId="7" pageNumber="62">Distal part of G1 terminal segment, in situ</th>
<td id="7478AD19FF9E66CAB097FE573F48FE2B" box="[693,1028,452,507]" gridcol="1" gridrow="4" pageId="7" pageNumber="62">directed obliquely outwards, pointed towards coxa of cheliped</td>
<td id="7478AD19FF9E66CAB602FE573EC4FE2B" box="[1056,1416,452,507]" gridcol="2" gridrow="4" pageId="7" pageNumber="62">directed laterally outwards, pointed towards coxa of first ambulatory leg</td>
</tr>
<tr id="37A9C465FF9E66CAB287FD823EC4FDF8" box="[165,1416,529,552]" gridrow="5" pageId="7" pageNumber="62">
<th id="7478AD19FF9E66CAB287FD8239D5FDF8" box="[165,665,529,552]" gridcol="0" gridrow="5" pageId="7" pageNumber="62">Ratio of lengths of G2 distal and basal segments</th>
<td id="7478AD19FF9E66CAB097FD823F48FDF8" box="[693,1028,529,552]" gridcol="1" gridrow="5" pageId="7" pageNumber="62">2.2 (Fig. 3F)</td>
<td id="7478AD19FF9E66CAB602FD823EC4FDF8" box="[1056,1416,529,552]" gridcol="2" gridrow="5" pageId="7" pageNumber="62">1.8 (Dai 1997: fig. 96)</td>
</tr>
<tr id="37A9C465FF9E66CAB287FDAE3EC4FD84" box="[165,1416,573,596]" gridrow="6" pageId="7" pageNumber="62">
<th id="7478AD19FF9E66CAB287FDAE39D5FD84" box="[165,665,573,596]" gridcol="0" gridrow="6" pageId="7" pageNumber="62">Width to length ratio of male abdominal somite 6</th>
<td id="7478AD19FF9E66CAB097FDAE3F48FD84" box="[693,1028,573,596]" gridcol="1" gridrow="6" pageId="7" pageNumber="62">2.1 (Figs. 2C, 3B)</td>
<td id="7478AD19FF9E66CAB602FDAE3EC4FD84" box="[1056,1416,573,596]" gridcol="2" gridrow="6" pageId="7" pageNumber="62">2.3 (Fig. 4C)</td>
</tr>
</table>
</paragraph>
<paragraph id="8926C627FF9E9932B2E5FB023FFEFB1C" blockId="7.[151,1437,664,1228]" pageId="7" pageNumber="62">
At the one site where both
<taxonomicName id="4E99BDA4FF9E9932B3DCFB02399CFB78" box="[510,720,1168,1192]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="guangdongense">
<emphasis id="BBED1A35FF9E9932B3DCFB02399CFB78" box="[510,720,1168,1192]" italics="true" pageId="7" pageNumber="62">N. guangdongense</emphasis>
</taxonomicName>
and
<taxonomicName id="4E99BDA4FF9E9932B128FB0238E5FB78" box="[778,937,1168,1192]" class="Malacostraca" family="Potamidae" genus="Nanhaipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="zhuhaiense" status="sp. nov.">
<emphasis id="BBED1A35FF9E9932B128FB0238E5FB78" box="[778,937,1168,1192]" italics="true" pageId="7" pageNumber="62">N. zhuhaiense</emphasis>
</taxonomicName>
<emphasis id="BBED1A35FF9E9932B193FB0338BDFB78" bold="true" box="[945,1009,1168,1192]" pageId="7" pageNumber="62">
<taxonomicNameLabel id="A0DEA74EFF9E9932B193FB0338BDFB78" box="[945,1009,1168,1192]" pageId="7" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are present, the former species is the dominant one. The burrows of these two species have even been found adjacent to each other.
</paragraph>
</subSubSection>
</treatment>
</document>