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<document id="52B8297DB4360C31840BD25570FF2049" ID-DOI="10.11646/zootaxa.4020.3.9" ID-GBIF-Dataset="ff6b61bc-bb45-4436-9d88-8f1c678daf06" ID-ISSN="1175-5326" ID-Zenodo-Dep="238547" ID-ZooBank="A254E350-F0C8-495D-A003-43EAF1512B0F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461271511584" checkinUser="plazi" docAuthor="De Matos-Pita, Susana S. &amp; Ramil, Fran" docDate="2015" docId="002487F22011FFD8FF142DC3FADCFD2B" docLanguage="en" docName="zt04020p587.pdf" docOrigin="Zootaxa 4020 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trypaea oblonga Le Loeuff &amp; Intes 1974, n. comb." docType="treatment" docVersion="7" lastPageNumber="578" masterDocId="FC1DFF8A2014FFDFFF83282EFFCCFFE6" masterDocTitle="Additions to thalassinidean fauna (Crustacea: Decapoda) off Mauritania (NW Africa) with the description of a new genus and a new species" masterLastPageNumber="587" masterPageNumber="571" pageNumber="576" updateTime="1698611913965" updateUser="plazi">
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<mods:title id="281B6CE1D467B47C593AB4EC34F61916">Additions to thalassinidean fauna (Crustacea: Decapoda) off Mauritania (NW Africa) with the description of a new genus and a new species</mods:title>
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<mods:namePart id="21DD0E4280ED4790DC2134175AE40840">De Matos-Pita, Susana S.</mods:namePart>
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<mods:namePart id="24F0F0CBBAA2E73D5710F267105A2AED">Ramil, Fran</mods:namePart>
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<treatment id="002487F22011FFD8FF142DC3FADCFD2B" ID-DOI="http://doi.org/10.5281/zenodo.6108163" ID-GBIF-Taxon="119638872" ID-Zenodo-Dep="6108163" LSID="urn:lsid:plazi:treatment:002487F22011FFD8FF142DC3FADCFD2B" httpUri="http://treatment.plazi.org/id/002487F22011FFD8FF142DC3FADCFD2B" lastPageId="7" lastPageNumber="578" pageId="5" pageNumber="576">
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<heading id="D37A81882011FFDAFF142DC3FCD4F9E1" bold="true" box="[151,792,1517,1543]" fontSize="11" level="1" pageId="5" pageNumber="576" reason="1">
<emphasis id="BAF9EAF62011FFDAFF142DC3FCD4F9E1" bold="true" box="[151,792,1517,1543]" pageId="5" pageNumber="576">
<taxonomicName id="4F8D4D672011FFDAFF142DC3FD6AF9E1" ID-CoL="66175202-cd95-4167-bc8c-3e4bd427279a" authority="Le Loeuff &amp; Intes, 1974" authorityName="Le Loeuff &amp; Intes" authorityYear="1974" box="[151,678,1517,1543]" class="Malacostraca" family="Callianassidae" genus="Trypaea" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="oblonga" status="comb. nov.">
<emphasis id="BAF9EAF62011FFDAFF142DC3FEAEF9E1" bold="true" box="[151,354,1517,1543]" italics="true" pageId="5" pageNumber="576">Trypaea oblonga</emphasis>
(
<bibRefCitation id="EC1C4B152011FFDAFEF12DC3FD52F9E1" author="Le" box="[370,670,1517,1543]" pageId="5" pageNumber="576" refString="Le Loeuff, P. &amp; Intes, A. (1974) Les Thalassinidea (Crustacea Decapoda) du Golfe de Guinee: systematique - ecologie. Cahiers ORSTOM, serie Oceanographie, 12 (1), 17 - 69." type="journal article" year="1974">Le Loeuff &amp; Intès, 1974</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A1CA578D2011FFDAFD2E2DC3FCD4F9E1" box="[685,792,1517,1543]" pageId="5" pageNumber="576">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="883236E42011FFDAFF142E21FEE0F9CE" blockId="5.[151,792,1517,1576]" box="[151,300,1551,1576]" pageId="5" pageNumber="576">
(
<figureCitation id="10B62A612011FFDAFF1C2E21FF20F9CE" box="[159,236,1551,1576]" captionStart="FIGURE 2" captionStartId="6.[151,250,1091,1113]" captionTargetBox="[157,1428,412,1061]" captionTargetId="figure@6.[150,1436,405,1067]" captionTargetPageId="6" captionText="FIGURE 2. Trypaea oblonga (Le Loeuff &amp; Intès, 1974) n. comb. Male MU 215: A) Left pleopod 4; B) Left pleopod 4 appendix interna detail. Female MUDR 26: C) Left pleopod 1; D) Left pleopod 2, mesial view. Scale bars: A, 2 mm; B, 0.2 mm; C, D, 1 mm." httpUri="https://zenodo.org/record/238549/files/figure.png" pageId="5" pageNumber="576">Figs. 2</figureCitation>
A–D)
</paragraph>
</subSubSection>
<subSubSection id="C097656F2011FFDAFF142E76FB47F94D" pageId="5" pageNumber="576" type="reference_group">
<paragraph id="883236E42011FFDAFF142E76FE58F96A" blockId="5.[151,1436,1624,1707]" pageId="5" pageNumber="576">
<treatmentCitationGroup id="A89D11CA2011FFDAFF142E76FE58F96A" infered="true" pageId="5" pageNumber="576">
<taxonomicName id="4F8D4D672011FFDAFF142E76FD42F988" ID-CoL="512cef42-0090-4368-8cf7-783a6af00f4a" authority="Le Loeuff &amp; Intes, 1974: 38" authorityName="Le Loeuff &amp; Intes" authorityPageNumber="38" authorityYear="1974" box="[151,654,1624,1646]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="oblonga">
<emphasis id="BAF9EAF62011FFDAFF142E76FEA5F988" box="[151,361,1624,1646]" italics="true" pageId="5" pageNumber="576">Callianassa oblonga</emphasis>
<treatmentCitation id="092C10F52011FFDAFEEC2E77FD42F988" author="Le" box="[367,654,1624,1646]" page="38" pageId="5" pageNumber="576" year="1974">
<bibRefCitation id="EC1C4B152011FFDAFEEC2E77FDA5F988" author="Le" box="[367,617,1624,1646]" pageId="5" pageNumber="576" refString="Le Loeuff, P. &amp; Intes, A. (1974) Les Thalassinidea (Crustacea Decapoda) du Golfe de Guinee: systematique - ecologie. Cahiers ORSTOM, serie Oceanographie, 12 (1), 17 - 69." type="journal article" year="1974">Le Loeuff &amp; Intès, 1974</bibRefCitation>
: 38
</treatmentCitation>
</taxonomicName>
, figs. 9a–r.—
<treatmentCitation id="092C10F52011FFDAFC952E76FB47F988" author="Saint" box="[790,1163,1624,1646]" page="55" pageId="5" pageNumber="576" year="1979">
<bibRefCitation id="EC1C4B152011FFDAFC952E76FBA4F988" author="Saint" box="[790,1128,1624,1646]" pageId="5" pageNumber="576" refString="Saint Laurent, M. de &amp; Le Loeuff, P. (1979) Campagnes de la Calypso au large des cotes Atlantiques Africaines (1956 et 1959) (suite) 22. Crustaces Decapodes Thalassinidea. I. Upogebiidae et Callianassidae. Resultats Scientifiques des Campagnes de la Calypso, 11, 29 - 101." type="journal article" year="1979">Saint Laurent &amp; Le Loeuff, 1979</bibRefCitation>
: 55
</treatmentCitation>
.—Sakai, 1999: 18; 2005a: 32, figs. 2E, 3E, 4E.
</treatmentCitationGroup>
</paragraph>
<paragraph id="883236E42011FFDAFF142EBBFB47F94D" blockId="5.[151,1436,1624,1707]" box="[151,1163,1685,1707]" pageId="5" pageNumber="576">
<treatmentCitationGroup id="A89D11CA2011FFDAFF142EBBFB47F94D" box="[151,1163,1685,1707]" pageId="5" pageNumber="576">
<taxonomicName id="4F8D4D672011FFDAFF142EBBFEAEF94D" ID-CoL="92bce01a-d14a-46d9-9d59-fed703372648" box="[151,354,1685,1707]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="BAF9EAF62011FFDAFF142EBBFEAEF94D" box="[151,354,1685,1707]" italics="true" pageId="5" pageNumber="576">Cheramus oblongus</emphasis>
</taxonomicName>
.—
<treatmentCitation id="092C10F52011FFDAFE032EBBFD7DF94D" author="Manning" box="[384,689,1685,1707]" page="780" pageId="5" pageNumber="576" year="1991">
<bibRefCitation id="EC1C4B152011FFDAFE032EBBFD4CF94D" author="Manning" box="[384,640,1685,1707]" pageId="5" pageNumber="576" refString="Manning, R. B. &amp; Felder, D. L. (1991) Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington, 104 (4), 764 - 792." type="journal article" year="1991">Manning &amp; Felder, 1991</bibRefCitation>
: 780
</treatmentCitation>
.—
<treatmentCitation id="092C10F52011FFDAFD4D2EBBFC75F94D" author="Tudge" box="[718,953,1685,1707]" page="145" pageId="5" pageNumber="576" year="2000">
<bibRefCitation id="EC1C4B152011FFDAFD4D2EBBFC40F94D" author="Tudge" box="[718,908,1685,1707]" pageId="5" pageNumber="576" refString="Tudge, C. C., Poore, G. C. B. &amp; Lemaitre, R. (2000) Preliminary phylogenetic analysis of generic relationships within the Callianassidae and Ctenochelidae (Decapoda: Thalassinidea: Callianassoidea). Journal of Crustacean Biology, 20 (Special Number 2), pp. 129 - 149." type="journal article" year="2000">
Tudge
<emphasis id="BAF9EAF62011FFDAFC962EB8FC85F94D" box="[789,841,1685,1707]" italics="true" pageId="5" pageNumber="576">et al.</emphasis>
, 2000
</bibRefCitation>
: 145
</treatmentCitation>
.—
<treatmentCitation id="092C10F52011FFDAFC5A2EBBFB4BF94D" author="Sakai" box="[985,1159,1685,1707]" page="369" pageId="5" pageNumber="576" year="2011">
<bibRefCitation id="EC1C4B152011FFDAFC5A2EBBFB99F94D" author="Sakai" box="[985,1109,1685,1707]" pageId="5" pageNumber="576" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai, 2011</bibRefCitation>
: 369
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C097656F2011FFDAFF142EF9FD97F82F" pageId="5" pageNumber="576" type="materials_examined">
<paragraph id="883236E42011FFDAFF142EF9FD97F82F" blockId="5.[151,1436,1751,1993]" pageId="5" pageNumber="576">
<emphasis id="BAF9EAF62011FFDAFF142EF9FE41F916" bold="true" box="[151,397,1751,1776]" pageId="5" pageNumber="576">Material examined.</emphasis>
Expedition Maurit0911: Stn. MU215, off S Nouakchott (
<collectingCountry id="F09A76742011FFDAFBF42EF9FB39F916" box="[1143,1269,1751,1776]" name="Mauritania" pageId="5" pageNumber="576">Mauritania</collectingCountry>
) (17°29´44″– 17°28´06″N; 16°39´31″–16°40´10″W), 358–364-m depth, muddy fine sand, commercial trawl,
<date id="FC3310242011FFDAFB7A2ED2FAAFF8F2" box="[1273,1379,1788,1812]" pageId="5" pageNumber="576" value="2009-02-02">2.II.2009</date>
; one male TL/CL: 19.09/
<quantity id="4F759B012011FFDAFEFA2F31FE2CF8D1" box="[377,480,1823,1848]" metricMagnitude="-3" metricUnit="m" metricValue="4.52" pageId="5" pageNumber="576" unit="mm" value="4.52">4.52 mm</quantity>
, both pereiopods 1 missing, left P4 merus to dactylus detached and both pleopods 3 missing. Expedition Maurit1011: Stn.
<collectionCode id="EE9CAE212011FFDAFDFA2F6AFD1CF8BB" box="[633,720,1860,1885]" pageId="5" pageNumber="576">MUDR</collectionCode>
26, off S Nouakchott (
<collectingCountry id="F09A76742011FFDAFC7E2F6AFBB7F8BB" box="[1021,1147,1860,1885]" name="Mauritania" pageId="5" pageNumber="576">Mauritania</collectingCountry>
) cold-water coral reef (17°29´42″N, 16°41´04″W), 441-m depth, mud with coral rubble and shell debris, rock dredge,
<date id="FC3310242011FFDAFB582F46FAABF866" box="[1243,1383,1896,1920]" pageId="5" pageNumber="576" value="2010-12-12">12.XII.2010</date>
; one incomplete female, TL/CL: +12.23/
<quantity id="4F759B012011FFDAFDAB2FA2FD5CF842" box="[552,656,1932,1956]" metricMagnitude="-3" metricUnit="m" metricValue="4.94" pageId="5" pageNumber="576" unit="mm" value="4.94">4.94 mm</quantity>
, right pereiopod 1 carpus and chela, both pereiopods 3–5, abdominal segments 3–5 and telson are all missing.
</paragraph>
</subSubSection>
<subSubSection id="C097656F2012FFD9FF4428B9FB22FA69" pageId="6" pageNumber="577" type="description">
<paragraph id="883236E42012FFD9FF4428B9FEB7FE6E" blockId="6.[151,1437,151,392]" pageId="6" pageNumber="577">
<emphasis id="BAF9EAF62012FFD9FF4428B9FE95FF56" bold="true" box="[199,345,151,176]" pageId="6" pageNumber="577">Description.</emphasis>
Our male specimen mostly corresponds to the
<typeStatus id="573688462012FFD9FCFA28B6FC65FF56" box="[889,937,152,176]" pageId="6" pageNumber="577">type</typeStatus>
description provided by
<bibRefCitation id="EC1C4B152012FFD9FB4928B6FF28FF32" pageId="6" pageNumber="577" refString="Le Loeuff, P. &amp; Intes, A. (1974) Les Thalassinidea (Crustacea Decapoda) du Golfe de Guinee: systematique - ecologie. Cahiers ORSTOM, serie Oceanographie, 12 (1), 17 - 69." type="journal article">Le Loeuff &amp; Intès (1974)</bibRefCitation>
. Both specimens present a rudimentary, uniramous and bisegmented pleopod 1 (as shown in
<bibRefCitation id="EC1C4B152012FFD9FA8F2893FEDFFF1E" author="Le" pageId="6" pageNumber="577" refString="Le Loeuff, P. &amp; Intes, A. (1974) Les Thalassinidea (Crustacea Decapoda) du Golfe de Guinee: systematique - ecologie. Cahiers ORSTOM, serie Oceanographie, 12 (1), 17 - 69." type="journal article" year="1974">Le Loeuff &amp; Intès 1974</bibRefCitation>
: fig. 9r). While the
<typeStatus id="573688462012FFD9FE7E28CEFDE1FF1E" box="[509,557,224,248]" pageId="6" pageNumber="577">type</typeStatus>
specimen was described with pleopods 2–5 missing, in our male specimen, although no scar or trace of pleopods can be appreciated on pleomere 2, a conspicuous scar is present in place of pleopod 3; pleopods 4 and 5 are biramous and foliaceous (
<figureCitation id="10B62A612012FFD9FCB32909FCB9FEA6" box="[816,885,295,320]" captionStart="FIGURE 2" captionStartId="6.[151,250,1091,1113]" captionTargetBox="[157,1428,412,1061]" captionTargetId="figure@6.[150,1436,405,1067]" captionTargetPageId="6" captionText="FIGURE 2. Trypaea oblonga (Le Loeuff &amp; Intès, 1974) n. comb. Male MU 215: A) Left pleopod 4; B) Left pleopod 4 appendix interna detail. Female MUDR 26: C) Left pleopod 1; D) Left pleopod 2, mesial view. Scale bars: A, 2 mm; B, 0.2 mm; C, D, 1 mm." httpUri="https://zenodo.org/record/238549/files/figure.png" pageId="6" pageNumber="577">Fig. 2</figureCitation>
A), with an appendix interna projecting from the distal proximal third of the endopod inner border, and with distal margin obliquely truncate and furnished with hooks (
<figureCitation id="10B62A612012FFD9FF6A2941FEFAFE6E" box="[233,310,367,392]" captionStart="FIGURE 2" captionStartId="6.[151,250,1091,1113]" captionTargetBox="[157,1428,412,1061]" captionTargetId="figure@6.[150,1436,405,1067]" captionTargetPageId="6" captionText="FIGURE 2. Trypaea oblonga (Le Loeuff &amp; Intès, 1974) n. comb. Male MU 215: A) Left pleopod 4; B) Left pleopod 4 appendix interna detail. Female MUDR 26: C) Left pleopod 1; D) Left pleopod 2, mesial view. Scale bars: A, 2 mm; B, 0.2 mm; C, D, 1 mm." httpUri="https://zenodo.org/record/238549/files/figure.png" pageId="6" pageNumber="577">Figs. 2</figureCitation>
A, B).
</paragraph>
<caption id="DCF2666C2012FFD9FF142C6DFF14FB71" httpUri="https://zenodo.org/record/238549/files/figure.png" pageId="6" pageNumber="577" targetBox="[157,1428,412,1061]" targetPageId="6">
<paragraph id="883236E42012FFD9FF142C6DFF14FB71" blockId="6.[151,1436,1091,1175]" pageId="6" pageNumber="577">
<emphasis id="BAF9EAF62012FFD9FF142C6DFEC0FBBE" bold="true" box="[151,268,1091,1113]" pageId="6" pageNumber="577">FIGURE 2</emphasis>
.
<taxonomicName id="4F8D4D672012FFD9FE9B2C6AFD02FBBF" authority="Le Loeuff &amp; Intes, 1974" authorityName="Le Loeuff &amp; Intes" authorityYear="1974" box="[280,718,1091,1113]" class="Malacostraca" family="Callianassidae" genus="Trypaea" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="oblonga" status="comb. nov.">
<emphasis id="BAF9EAF62012FFD9FE9B2C6AFE0FFBBF" box="[280,451,1091,1113]" italics="true" pageId="6" pageNumber="577">Trypaea oblonga</emphasis>
(Le Loeuff &amp; Intès, 1974)
</taxonomicName>
<taxonomicNameLabel id="A1CA578D2012FFD9FD502C6AFCE7FBBF" box="[723,811,1091,1113]" pageId="6" pageNumber="577">n. comb.</taxonomicNameLabel>
Male MU215: A) Left pleopod 4; B) Left pleopod 4 appendix interna detail. Female MUDR26: C) Left pleopod 1; D) Left pleopod 2, mesial view. Scale bars: A, 2 mm; B, 0.2 mm; C, D, 1 mm.
</paragraph>
</caption>
<paragraph id="883236E42012FFD9FF442CEAFB21FAC2" blockId="6.[151,1437,1219,2036]" pageId="6" pageNumber="577">
Our incomplete female specimen also corresponds to the original description. Differences observed are those referring to the sex: conspicuous gonopore on both pereiopod 3 coxa, pleopod 1 uniramous and bisegmented (
<figureCitation id="10B62A612012FFD9FAEC2CC9FF6AFAC2" captionStart="FIGURE 2" captionStartId="6.[151,250,1091,1113]" captionTargetBox="[157,1428,412,1061]" captionTargetId="figure@6.[150,1436,405,1067]" captionTargetPageId="6" captionText="FIGURE 2. Trypaea oblonga (Le Loeuff &amp; Intès, 1974) n. comb. Male MU 215: A) Left pleopod 4; B) Left pleopod 4 appendix interna detail. Female MUDR 26: C) Left pleopod 1; D) Left pleopod 2, mesial view. Scale bars: A, 2 mm; B, 0.2 mm; C, D, 1 mm." httpUri="https://zenodo.org/record/238549/files/figure.png" pageId="6" pageNumber="577">Fig. 2</figureCitation>
C), and pleopod 2 slender, biramous, with endopod bearing a terminal appendix interna (
<figureCitation id="10B62A612012FFD9FB052D25FB06FAC2" box="[1158,1226,1291,1316]" captionStart="FIGURE 2" captionStartId="6.[151,250,1091,1113]" captionTargetBox="[157,1428,412,1061]" captionTargetId="figure@6.[150,1436,405,1067]" captionTargetPageId="6" captionText="FIGURE 2. Trypaea oblonga (Le Loeuff &amp; Intès, 1974) n. comb. Male MU 215: A) Left pleopod 4; B) Left pleopod 4 appendix interna detail. Female MUDR 26: C) Left pleopod 1; D) Left pleopod 2, mesial view. Scale bars: A, 2 mm; B, 0.2 mm; C, D, 1 mm." httpUri="https://zenodo.org/record/238549/files/figure.png" pageId="6" pageNumber="577">Fig. 2</figureCitation>
D).
</paragraph>
<paragraph id="883236E42012FFD9FF442D01FB22FA69" blockId="6.[151,1437,1219,2036]" pageId="6" pageNumber="577">
<emphasis id="BAF9EAF62012FFD9FF442D01FEE6FAAE" bold="true" box="[199,298,1327,1352]" pageId="6" pageNumber="577">Habitat.</emphasis>
The
<typeStatus id="573688462012FFD9FEE62D1EFE59FAAE" box="[357,405,1328,1352]" pageId="6" pageNumber="577">type</typeStatus>
specimen was collected at
<quantity id="4F759B012012FFD9FD4D2D1EFCD9FAA1" box="[718,789,1328,1352]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="577" unit="m" value="200.0">200 m</quantity>
with a DRB: drague à dents (72×
<quantity id="4F759B012012FFD9FB162D1EFB2CFAAE" box="[1173,1248,1328,1352]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="577" unit="cm" value="30.0">30 cm</quantity>
); no bottom was specified (
<bibRefCitation id="EC1C4B152012FFD9FE8D2D7AFDE4FA8A" author="Le" box="[270,552,1363,1388]" pageId="6" pageNumber="577" refString="Le Loeuff, P. &amp; Intes, A. (1974) Les Thalassinidea (Crustacea Decapoda) du Golfe de Guinee: systematique - ecologie. Cahiers ORSTOM, serie Oceanographie, 12 (1), 17 - 69." type="journal article" year="1974">Le Loeuff &amp; Intès 1974</bibRefCitation>
). Our specimens were captured off South Nouakchott, at two geographically close stations, on muddy fine sand and muddy coral rubble, at depths between 358–364 and
<specimenCount id="9E8BFD6D2012FFD9FB232D56FB22FA69" box="[1184,1262,1400,1424]" pageId="6" pageNumber="577" type="generic">
<quantity id="4F759B012012FFD9FB232D56FB26FA69" box="[1184,1258,1400,1424]" metricMagnitude="2" metricUnit="m" metricValue="4.41" pageId="6" pageNumber="577" unit="m" value="441.0">441 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C097656F2012FFD9FF442DB5FCB8FA3E" pageId="6" pageNumber="577" type="distribution">
<paragraph id="883236E42012FFD9FF442DB5FCB8FA3E" blockId="6.[151,1437,1219,2036]" pageId="6" pageNumber="577">
<emphasis id="BAF9EAF62012FFD9FF442DB5FEACFA52" bold="true" box="[199,352,1435,1460]" pageId="6" pageNumber="577">Distribution.</emphasis>
Previously reported only from the
<typeStatus id="573688462012FFD9FD7E2DB2FCE1FA52" box="[765,813,1436,1460]" pageId="6" pageNumber="577">type</typeStatus>
locality (
<collectingCountry id="F09A76742012FFD9FC1D2DB2FBEBFA52" box="[926,1063,1436,1460]" name="Ivory Coast" pageId="6" pageNumber="577">Ivory Coast</collectingCountry>
) and now from
<collectingCountry id="F09A76742012FFD9FB6B2DB5FAAAFA52" box="[1256,1382,1435,1460]" name="Mauritania" pageId="6" pageNumber="577">Mauritania</collectingCountry>
, this species seems to be restricted to the West African tropical region.
</paragraph>
</subSubSection>
<subSubSection id="C097656F2012FFD8FF442DCDFADCFD2B" lastPageId="7" lastPageNumber="578" pageId="6" pageNumber="577" type="discussion">
<paragraph id="883236E42012FFD9FF442DCDFC07F96D" blockId="6.[151,1437,1219,2036]" pageId="6" pageNumber="577">
<emphasis id="BAF9EAF62012FFD9FF442DCDFEF7FA1A" bold="true" box="[199,315,1507,1532]" pageId="6" pageNumber="577">Remarks.</emphasis>
To date, the
<typeStatus id="573688462012FFD9FE5F2DCAFDC0FA1A" box="[476,524,1508,1532]" pageId="6" pageNumber="577">type</typeStatus>
specimen (male CL:
<quantity id="4F759B012012FFD9FC8D2DCDFCA9FA1A" box="[782,869,1507,1532]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="577" unit="mm" value="3.5">3.5 mm</quantity>
) from
<collectingCountry id="F09A76742012FFD9FC3A2DCAFB8DFA1A" box="[953,1089,1508,1532]" name="Ivory Coast" pageId="6" pageNumber="577">Ivory Coast</collectingCountry>
(
<quantity id="4F759B012012FFD9FBD72DCAFB51FA1A" box="[1108,1181,1508,1532]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="577" unit="m" value="200.0">200 m</quantity>
) is the only one ever recorded. After its original description as
<taxonomicName id="4F8D4D672012FFD9FDF52E29FCADF9C6" box="[630,865,1543,1568]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="oblonga">
<emphasis id="BAF9EAF62012FFD9FDF52E29FCADF9C6" box="[630,865,1543,1568]" italics="true" pageId="6" pageNumber="577">Callianassa oblonga</emphasis>
</taxonomicName>
, the species was included in the genus
<taxonomicName id="4F8D4D672012FFD9FAAA2E29FED9F9A2" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62012FFD9FAAA2E29FA50F9C6" box="[1321,1436,1543,1568]" italics="true" pageId="6" pageNumber="577">Cheramus</emphasis>
<bibRefCitation id="EC1C4B152012FFD9FF142E02FED9F9A2" author="Bate" box="[151,277,1580,1604]" pageId="6" pageNumber="577" refString="Bate, C. S. (1888) Report on the Crustacea Macrura dredged by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger During the Years 1873 - 76, Zoology, 24, i - xc, 1 - 942, pls. 1 - 157." type="journal article" year="1888">Bate, 1888</bibRefCitation>
</taxonomicName>
as
<taxonomicName id="4F8D4D672012FFD9FEC72E02FE07F9A2" box="[324,459,1579,1604]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="BAF9EAF62012FFD9FEC72E02FE07F9A2" box="[324,459,1579,1604]" italics="true" pageId="6" pageNumber="577">C. oblongus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EC1C4B152012FFD9FE7C2E05FCFEF9A2" author="Manning" box="[511,818,1579,1604]" pageId="6" pageNumber="577" refString="Manning, R. B. &amp; Felder, D. L. (1991) Revision of the American Callianassidae (Crustacea: Decapoda: Thalassinidea). Proceedings of the Biological Society of Washington, 104 (4), 764 - 792." type="journal article" year="1991">Manning &amp; Felder (1991)</bibRefCitation>
, and accepted by
<bibRefCitation id="EC1C4B152012FFD9FB8F2E05FB3EF9A2" box="[1036,1266,1579,1604]" pageId="6" pageNumber="577" refString="Tudge, C. C., Poore, G. C. B. &amp; Lemaitre, R. (2000) Preliminary phylogenetic analysis of generic relationships within the Callianassidae and Ctenochelidae (Decapoda: Thalassinidea: Callianassoidea). Journal of Crustacean Biology, 20 (Special Number 2), pp. 129 - 149." type="journal article">
Tudge
<emphasis id="BAF9EAF62012FFD9FBDD2E02FB50F9A2" box="[1118,1180,1579,1604]" italics="true" pageId="6" pageNumber="577">et al.</emphasis>
(2000)
</bibRefCitation>
in this genus. However, Sakai (2005: 20) was unable to find enough morphological characters that reliably separated both genera; he therefore synonymized the genus
<taxonomicName id="4F8D4D672012FFD9FD0F2E5DFD33F96A" box="[652,767,1651,1676]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62012FFD9FD0F2E5DFD33F96A" box="[652,767,1651,1676]" italics="true" pageId="6" pageNumber="577">Cheramus</emphasis>
</taxonomicName>
with
<taxonomicName id="4F8D4D672012FFD9FCBD2E5DFC08F96A" box="[830,964,1651,1676]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62012FFD9FCBD2E5DFC08F96A" box="[830,964,1651,1676]" italics="true" pageId="6" pageNumber="577">Callianassa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="883236E42012FFD9FF442EB6FE60F812" blockId="6.[151,1437,1219,2036]" pageId="6" pageNumber="577">
On the other hand, a tendency to group
<taxonomicName id="4F8D4D672012FFD9FD2A2EB9FCE3F956" box="[681,815,1687,1712]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62012FFD9FD2A2EB9FCE3F956" box="[681,815,1687,1712]" italics="true" pageId="6" pageNumber="577">Callianassa</emphasis>
</taxonomicName>
species based on pleopod morphology, among other characters, led
<bibRefCitation id="EC1C4B152012FFD9FECB2E95FD25F932" author="Saint" box="[328,745,1723,1748]" pageId="6" pageNumber="577" refString="Saint Laurent, M. de &amp; Le Loeuff, P. (1979) Campagnes de la Calypso au large des cotes Atlantiques Africaines (1956 et 1959) (suite) 22. Crustaces Decapodes Thalassinidea. I. Upogebiidae et Callianassidae. Resultats Scientifiques des Campagnes de la Calypso, 11, 29 - 101." type="journal article" year="1979">
Saint Laurent and Le
<bibRefCitation id="EC1C4B152012FFD9FDCE2E95FD25F932" author="Loeuff" box="[589,745,1723,1748]" pageId="6" pageNumber="577" refString="Saint Laurent, M. de &amp; Le Loeuff, P. (1979) Campagnes de la Calypso au large des cotes Atlantiques Africaines (1956 et 1959) (suite) 22. Crustaces Decapodes Thalassinidea. I. Upogebiidae et Callianassidae. Resultats Scientifiques des Campagnes de la Calypso, 11, 29 - 101." type="journal article" year="1979">Loeuff (1979</bibRefCitation>
</bibRefCitation>
: 55) to recognize three groups: one including species with pleopod 1, but not pleopod 2; a second group of species lacking either pleopod 1 or pleopod 2; and a third group with ‛des pléopodes présents, bien que faiblement développés, sur les deux premiers segments abdominaux du mâle’ (pleopods on male absominal segments 1 and 2 present, although reduced). These authors included only one species,
<taxonomicName id="4F8D4D672012FFD9FF752F65FE13F882" box="[246,479,1867,1892]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="oblonga">
<emphasis id="BAF9EAF62012FFD9FF752F65FE13F882" box="[246,479,1867,1892]" italics="true" pageId="6" pageNumber="577">Callianassa oblonga</emphasis>
</taxonomicName>
, in the last group, although this species was originally described as with ‛pléopode 1 réduit, en forme de doigt de gant à extrémité courbe; les autres pléopodes manquent.’ (pleopod 1 reduced, fingerlike and with tip curved; pleopods 2–5 absent) (
<bibRefCitation id="EC1C4B152012FFD9FD522FBAFBC8F84A" author="Le" box="[721,1028,1939,1964]" pageId="6" pageNumber="577" refString="Le Loeuff, P. &amp; Intes, A. (1974) Les Thalassinidea (Crustacea Decapoda) du Golfe de Guinee: systematique - ecologie. Cahiers ORSTOM, serie Oceanographie, 12 (1), 17 - 69." type="journal article" year="1974">Le Loeuff and Intès 1974</bibRefCitation>
: 40); however, although a reduced pleopod 1 is present (and bisegmented as shown in
<bibRefCitation id="EC1C4B152012FFD9FD5D2F96FBD8F836" box="[734,1044,1975,2000]" pageId="6" pageNumber="577" refString="Le Loeuff, P. &amp; Intes, A. (1974) Les Thalassinidea (Crustacea Decapoda) du Golfe de Guinee: systematique - ecologie. Cahiers ORSTOM, serie Oceanographie, 12 (1), 17 - 69." type="journal article">Le Loeuff and Intès (1974)</bibRefCitation>
: fig. 9r), there is no clear evidence of an existing pleopod 2.
</paragraph>
<paragraph id="883236E42013FFD8FF4428B6FBD9FDDA" blockId="7.[151,1436,151,717]" pageId="7" pageNumber="578">
In his synopsis of the family
<taxonomicName id="4F8D4D672013FFD8FD8E28B9FC80FF56" authority="Sakai (1999)" authorityName="Sakai" authorityYear="1999" box="[525,844,151,176]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="family">Callianassidae, Sakai (1999)</taxonomicName>
first associated
<taxonomicName id="4F8D4D672013FFD8FB8128B6FBB2FF56" box="[1026,1150,151,176]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="oblonga">
<emphasis id="BAF9EAF62013FFD8FB8128B6FBB2FF56" box="[1026,1150,151,176]" italics="true" pageId="7" pageNumber="578">C. oblonga</emphasis>
</taxonomicName>
with
<taxonomicName id="4F8D4D672013FFD8FB3D28B9FA88FF56" box="[1214,1348,151,176]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62013FFD8FB3D28B9FA88FF56" box="[1214,1348,151,176]" italics="true" pageId="7" pageNumber="578">Callianassa</emphasis>
</taxonomicName>
species without pleopod 2 (Sakai 1999: 5). However, in his account of the species (Sakai 1999: 18), he described it as having male pleopods 1 and 2, as did
<bibRefCitation id="EC1C4B152013FFD8FDBA28F1FC05FF1E" box="[569,969,223,248]" pageId="7" pageNumber="578" refString="Saint Laurent, M. de &amp; Le Loeuff, P. (1979) Campagnes de la Calypso au large des cotes Atlantiques Africaines (1956 et 1959) (suite) 22. Crustaces Decapodes Thalassinidea. I. Upogebiidae et Callianassidae. Resultats Scientifiques des Campagnes de la Calypso, 11, 29 - 101." type="journal article">Saint Laurent and Le Loeuff (1979)</bibRefCitation>
. Some years later, with the revision of the
<taxonomicName id="4F8D4D672013FFD8FF14292AFE8BFEFB" box="[151,327,260,285]" class="Malacostraca" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="superFamily" superFamily="Callianassoidea">Callianassoidea</taxonomicName>
of the world,
<bibRefCitation id="EC1C4B152013FFD8FE72292AFD5CFEFB" author="Sakai" box="[497,656,260,285]" pageId="7" pageNumber="578" refString="Sakai, K. (2005 a) Callianassoidea of the world (Decapoda: Thalassinidea). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 4. Leiden, Brill, 286 pp." type="book" year="2005" yearSuffix="a">Sakai (2005a)</bibRefCitation>
arranged the
<taxonomicName id="4F8D4D672013FFD8FCB1292AFC74FEFB" box="[818,952,260,285]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62013FFD8FCB1292AFC74FEFB" box="[818,952,260,285]" italics="true" pageId="7" pageNumber="578">Callianassa</emphasis>
</taxonomicName>
species into four groups according to the morphology of pleopods l and 2, including
<taxonomicName id="4F8D4D672013FFD8FD212906FCD3FEA6" box="[674,799,295,320]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="oblonga">
<emphasis id="BAF9EAF62013FFD8FD212906FCD3FEA6" box="[674,799,295,320]" italics="true" pageId="7" pageNumber="578">C. oblonga</emphasis>
</taxonomicName>
within the group with uniramous and two-segmented pleopod l, and with pleopod 2 absent (Sakai 2005: 25, 32). The same author (
<bibRefCitation id="EC1C4B152013FFD8FC7A2962FB7EFE83" author="Sakai" box="[1017,1202,332,357]" pageId="7" pageNumber="578" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai 2011: 355</bibRefCitation>
) later considered the morphology of male pleopods 1 and 2 as the single valid character for callianassid genera classification, redescribing, among others, the genera
<taxonomicName id="4F8D4D672013FFD8FDED29BAFD38FE4B" box="[622,756,404,429]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62013FFD8FDED29BAFD38FE4B" box="[622,756,404,429]" italics="true" pageId="7" pageNumber="578">Callianassa</emphasis>
</taxonomicName>
(to include species with male pleopod 1 uniramous and bisegmented, and with male pleopod 2 uniramous, in which the protopod rarely protruded short distolaterally),
<taxonomicName id="4F8D4D672013FFD8FF1429F3FF3EFE12" box="[151,242,477,500]" class="Malacostraca" family="Callianassidae" genus="Trypaea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62013FFD8FF1429F3FF3EFE12" box="[151,242,477,500]" italics="true" pageId="7" pageNumber="578">Trypaea</emphasis>
</taxonomicName>
(male pleopod 1 uniramous, uni-, bi- or trisegmented, and male pleopod 2 absent), and
<taxonomicName id="4F8D4D672013FFD8FB6229F2FA98FE13" box="[1249,1364,476,501]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62013FFD8FB6229F2FA98FE13" box="[1249,1364,476,501]" italics="true" pageId="7" pageNumber="578">Cheramus</emphasis>
</taxonomicName>
(male pleopod 1 absent, or uniramous and bisegmented, and with slender, biramous male pleopod 2), and subsequently, returning
<taxonomicName id="4F8D4D672013FFD8FE872A0AFE21FDDB" box="[260,493,548,573]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="oblonga">
<emphasis id="BAF9EAF62013FFD8FE872A0AFE21FDDB" box="[260,493,548,573]" italics="true" pageId="7" pageNumber="578">Callianassa oblonga</emphasis>
</taxonomicName>
to the genus
<taxonomicName id="4F8D4D672013FFD8FD012A0AFD39FDDB" box="[642,757,548,573]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62013FFD8FD012A0AFD39FDDB" box="[642,757,548,573]" italics="true" pageId="7" pageNumber="578">Cheramus</emphasis>
</taxonomicName>
(as
<taxonomicName id="4F8D4D672013FFD8FCA72A0AFBC9FDDB" box="[804,1029,548,573]" class="Malacostraca" family="Callianassidae" genus="Cheramus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="BAF9EAF62013FFD8FCA72A0AFBC9FDDB" box="[804,1029,548,573]" italics="true" pageId="7" pageNumber="578">Cheramus oblongus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="883236E42013FFD8FF442A69FADCFD2B" blockId="7.[151,1436,151,717]" pageId="7" pageNumber="578">
All the foregoing propositions of classification were based on pleopod morphology. However, neither of the only two males ever recorded (the
<typeStatus id="573688462013FFD8FDBB2A43FDA4FD63" box="[568,616,621,645]" pageId="7" pageNumber="578">type</typeStatus>
specimen and the one captured during the Maurit surveys) displayed pleopod 2, and there is no evidence of their existence (at least in our material). In consequence, we propose here the relocation of
<taxonomicName id="4F8D4D672013FFD8FEAF2A9BFE65FD2B" box="[300,425,692,717]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="oblonga">
<emphasis id="BAF9EAF62013FFD8FEAF2A9BFE65FD2B" box="[300,425,692,717]" italics="true" pageId="7" pageNumber="578">C. oblonga</emphasis>
</taxonomicName>
to the genus
<taxonomicName id="4F8D4D672013FFD8FDBD2A9BFD55FD2A" box="[574,665,693,716]" class="Malacostraca" family="Callianassidae" genus="Trypaea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62013FFD8FDBD2A9BFD55FD2A" box="[574,665,693,716]" italics="true" pageId="7" pageNumber="578">Trypaea</emphasis>
</taxonomicName>
as
<taxonomicName id="4F8D4D672013FFD8FD3C2A9BFB6BFD2A" authority="Le Loeuff &amp; Intes, 1974" authorityName="Le Loeuff &amp; Intes" authorityYear="1974" box="[703,1191,692,717]" class="Malacostraca" family="Callianassidae" genus="Trypaea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="oblonga" status="comb. nov.">
<emphasis id="BAF9EAF62013FFD8FD3C2A9BFCB1FD2B" box="[703,893,692,717]" italics="true" pageId="7" pageNumber="578">Trypaea oblonga</emphasis>
(
<bibRefCitation id="EC1C4B152013FFD8FC0F2A9AFB53FD2A" author="Le" box="[908,1183,692,717]" pageId="7" pageNumber="578" refString="Le Loeuff, P. &amp; Intes, A. (1974) Les Thalassinidea (Crustacea Decapoda) du Golfe de Guinee: systematique - ecologie. Cahiers ORSTOM, serie Oceanographie, 12 (1), 17 - 69." type="journal article" year="1974">Le Loeuff &amp; Intès, 1974</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A1CA578D2013FFD8FB2D2A9BFADCFD2B" box="[1198,1296,692,717]" pageId="7" pageNumber="578">n. comb.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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