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<document id="9759DC0C1909531809DB2FF50DA9C591" ID-CLB-Dataset="9037" ID-DOI="10.11646/zootaxa.4861.3.5" ID-GBIF-Dataset="9b512d6d-956c-45a3-9265-8ae1167ba019" ID-ISSN="1175-5326" ID-Zenodo-Dep="4416437" ID-ZooBank="2E2FBB1C-80EC-41B7-806A-43B530297FC2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609799385234" checkinUser="plazi" docAuthor="Schawaller, Wolfgang" docDate="2020" docId="B26D87C8DB7C2C14AF99FF4ABBB6FDC4" docLanguage="en" docName="zootaxa.4861.3.5.pdf" docOrigin="Zootaxa 4861 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laena apfelbecki Schuster 1915" docType="treatment" docVersion="9" lastPageNumber="387" masterDocId="4E54FFB0DB7E2C16AF0EFFDDB861FFFA" masterDocTitle="New taxonomic and faunistic data to species of the genus Laena Dejean from Turkey (Coleoptera, Tenebrionidae, Lagriinae), with descriptions of two new species" masterLastPageNumber="398" masterPageNumber="385" pageNumber="387" updateTime="1698852204079" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="639E33C398510EFAF2069D05309018CB">New taxonomic and faunistic data to species of the genus Laena Dejean from Turkey (Coleoptera, Tenebrionidae, Lagriinae), with descriptions of two new species</mods:title>
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<mods:name id="A9CD5F93180A11A5C7573888AA010B44" type="personal">
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<mods:namePart id="3C3949F4F3279A9C2941A0DF92876951">Schawaller, Wolfgang</mods:namePart>
<mods:affiliation id="C36737E3C0D3B9E45011FAAC422D1F26">Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany. &amp; Contributions to Tenebrionidae no. 161. For no. 160 see: Annals of the Ditsong Museum of Natural History Pretoria 9, 2020</mods:affiliation>
</mods:name>
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<mods:title id="78CC1CFDF36ED1E60802D8A50D403E4C">Zootaxa</mods:title>
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<mods:part id="FD9306AA4E89C0AB6488E847A108ED6D">
<mods:date id="138550E87F27836F4387B08C84F5C5AC">2020</mods:date>
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<mods:number id="7EA7990D78485F75F82861608638082A">2020-10-19</mods:number>
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<mods:detail id="AD72F21E9469FE48749D62C91B32D425" type="volume">
<mods:number id="26B41AC976D3A61D333D8AE0EC3BE974">4861</mods:number>
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<mods:classification id="AACAA737B0F17770A2096862FBE486E0">journal article</mods:classification>
<mods:identifier id="99F77BC32BECECDFAFE07BACF43AE33D" type="CLB-Dataset">9037</mods:identifier>
<mods:identifier id="1EB9E1F8763864AFEE594D730F5A9C65" type="DOI">10.11646/zootaxa.4861.3.5</mods:identifier>
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<mods:identifier id="A2655C64FBEEC137D24202BDDB4F8BBA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8BF079FF4F63E4789823276ACCE5EE98" type="Zenodo-Dep">4416437</mods:identifier>
<mods:identifier id="4FE906BD55B5F9030FC80D8C48D5C270" type="ZooBank">2E2FBB1C-80EC-41B7-806A-43B530297FC2</mods:identifier>
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<treatment id="B26D87C8DB7C2C14AF99FF4ABBB6FDC4" ID-DOI="http://doi.org/10.5281/zenodo.4416580" ID-GBIF-Taxon="176439846" ID-Zenodo-Dep="4416580" LSID="urn:lsid:plazi:treatment:B26D87C8DB7C2C14AF99FF4ABBB6FDC4" httpUri="http://treatment.plazi.org/id/B26D87C8DB7C2C14AF99FF4ABBB6FDC4" lastPageNumber="387" pageId="2" pageNumber="387">
<subSubSection id="72DE6555DB7C2C14AF99FF4ABA43FF4B" box="[151,546,151,178]" pageId="2" pageNumber="387" type="nomenclature">
<paragraph id="3A7B36DEDB7C2C14AF99FF4ABA43FF4B" blockId="2.[151,546,151,214]" box="[151,546,151,178]" pageId="2" pageNumber="387">
<heading id="613381B2DB7C2C14AF99FF4ABA43FF4B" bold="true" box="[151,546,151,178]" fontSize="11" level="1" pageId="2" pageNumber="387" reason="1">
<taxonomicName id="FDC44D5DDB7C2C14AF99FF4ABA43FF4B" authority="Schuster, 1915" authorityName="Schuster" authorityYear="1915" box="[151,546,151,178]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="apfelbecki">
<emphasis id="08B0EACCDB7C2C14AF99FF4ABA43FF4B" bold="true" box="[151,546,151,178]" pageId="2" pageNumber="387">
<emphasis id="08B0EACCDB7C2C14AF99FF4AB906FF48" bold="true" box="[151,359,151,178]" italics="true" pageId="2" pageNumber="387">Laena apfelbecki</emphasis>
Schuster, 1915
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="72DE6555DB7C2C14AF99FF66B92BFF2C" box="[151,330,187,214]" pageId="2" pageNumber="387" type="description">
<paragraph id="3A7B36DEDB7C2C14AF99FF66B92BFF2C" blockId="2.[151,546,151,214]" box="[151,330,187,214]" pageId="2" pageNumber="387">
(
<figureCitation id="A2FF2A5BDB7C2C14AF91FF66B88EFF2F" box="[159,239,187,213]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1800,1824]" captionTargetBox="[170,1422,395,1753]" captionTargetId="figure-113@5.[151,1436,381,1777]" captionTargetPageId="5" captionText="FIGURES 1–8. Dorsal view of Laena species from Turkey (and few from Greece and Bulgaria). 1 L. ganglbaueri, Istanbul, SMNS. 2 L. apfelbecki, Greece Évia, SMNS. 3 L. weisei, Greece Sapka, SMNS. 4 L. kraatzi, Sinop, SMNS. 5 L. osmanlis, Bursa, SMNS. 6 L. assingi sp. nov., paratype SMNS. 7 L. quadricollis, Trabzon, SMNS. 8 L. theana, Isparta, SMNS. Scale line: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4416441" httpUri="https://zenodo.org/record/4416441/files/figure.png" pageId="2" pageNumber="387">Figs. 2</figureCitation>
,
<figureCitation id="A2FF2A5BDB7C2C14AFF5FF66B923FF2C" box="[251,322,187,214]" captionStart="FIGURES 22–46" captionStartId="7.[151,264,1694,1718]" captionTargetBox="[191,1396,184,1665]" captionTargetId="figure-16@7.[190,1397,181,1670]" captionTargetPageId="7" captionText="FIGURES 22–46. Right posterior male tibia of Laena species from Turkey and Greece (Figs. 22–25), and apicale of aedeagus of Laena species from Turkey and few from Greece (Figs. 26–46). 22 L. apfelbecki, Greece Évia, SMNS. 23 L. bozdagica sp. nov., holotype Izmir, SMNS. 24 L. kraatzi, Greece Chios, SMNS. 25 L. lindbergi, Bingöl, SMNS. 26 L. deplanata, Izmir, SMNS. 27 L. ferruginea, Greece Corfu, SMNS. 28 L. ferruginea, Antalya, SMNS (det. Kaszab as L. liliputana syn. nov.). 29 L. wanensis, Nur Daglari, SMNS. 30 L. bozdagica sp. nov., paratype Izmir, SMNS. 31 L. merkli, Hakkari, CPE. 32 L. obscuricornis, Meselik, SMNS. 33 L. byzantina, Artvin, SMNS. 34 L. korbi, Konya, SMNS. 35 L. lindbergi, Bitlis, SMNS. 36 L. apfelbecki, Greece Evia, SMNS. 37 L. pici, Antakya, SMNS. 38 L. ganglbaueri, Alem Dagh, SMNS. 39 L. weisei, Greece Sapka, SMNS. 40 L. theana, Erfelek, SMNS. 41 L. assingi sp. nov., holotype Fethiye, CPE. 42 L. graeca, Bolu, SMNS. 43 L. osmanlis, Bursa, SMNS. 44 L. quadricollis, Rize, SMNS. 45 L. quadricollis, Krasnodar Maikop, SMNS. 46 L. kraatzi, Alem Dagh, SMNS. Scale line: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4416445" httpUri="https://zenodo.org/record/4416445/files/figure.png" pageId="2" pageNumber="387">22, 36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72DE6555DB7C2C14AF99FEDEB9E1FEBB" pageId="2" pageNumber="387" type="materials_examined">
<paragraph id="3A7B36DEDB7C2C14AF99FEDEB9E1FEBB" blockId="2.[151,1437,259,574]" pageId="2" pageNumber="387">
<emphasis id="08B0EACCDB7C2C14AF99FEDEB9FCFEE7" bold="true" box="[151,413,259,285]" pageId="2" pageNumber="387">Examined specimens:</emphasis>
<materialsCitation id="8AAC3C83DB7C2C14AEA8FEDEB91DFEBB" ID-GBIF-Occurrence="3012394452" collectingDate="1995-06-05" collectionCode="SMNS" collectorName="Giachino &amp; Vailati" country="Greece" elevation="850" location="Dirfi" municipality="Platania" pageId="2" pageNumber="387" specimenCount="2">
<collectingCountry id="42D3764EDB7C2C14AEA8FEDEB999FEE7" box="[422,504,259,285]" name="Greece" pageId="2" pageNumber="387">Greece</collectingCountry>
, Nom. Évia,
<location id="3F1B6005DB7C2C14AD9DFEDEBAA8FEE7" LSID="urn:lsid:plazi:treatment:B26D87C8DB7C2C14AF99FF4ABBB6FDC4:3F1B6005DB7C2C14AD9DFEDEBAA8FEE7" box="[659,713,259,285]" country="Greece" municipality="Platania" name="Dirfi" pageId="2" pageNumber="387">Dirfi</location>
above
<collectingMunicipality id="DA1FACA4DB7C2C14AC11FEDEBB1DFEE7" box="[799,892,259,285]" pageId="2" pageNumber="387">Plátania</collectingMunicipality>
,
<collectorName id="97315308DB7C2C14AC87FEDEBB98FEE7" box="[905,1017,259,285]" pageId="2" pageNumber="387">Ag. Sofia</collectorName>
,
<quantity id="FD3C9B3BDB7C2C14AB09FEDEBC32FEE4" box="[1031,1107,259,286]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="387" unit="m" value="850.0">
<elevation id="B1E9D1EDDB7C2C14AB09FEDEBC32FEE4" box="[1031,1107,259,286]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="387" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<date id="4E7A101EDB7C2C14AB6FFEDEBCB4FEE4" box="[1121,1237,259,286]" pageId="2" pageNumber="387" value="1995-06-05">
<collectingDate id="5E3EE9F6DB7C2C14AB6FFEDEBCB4FEE4" box="[1121,1237,259,286]" pageId="2" pageNumber="387" value="1995-06-05">5.VI.1995</collectingDate>
</date>
, leg.
<collectorName id="97315308DB7C2C14AA18FEDEBD1CFEE7" box="[1302,1405,259,285]" pageId="2" pageNumber="387">Giachino</collectorName>
&amp;
<collectorName id="97315308DB7C2C14AF99FEFAB882FEBB" box="[151,227,295,321]" pageId="2" pageNumber="387">Vailati</collectorName>
,
<specimenCount id="2CC2FD57DB7C2C14AFE0FEFAB944FEB8" box="[238,293,295,322]" pageId="2" pageNumber="387" type="generic">2 ex.</specimenCount>
<collectionCode id="5CD5AE1BDB7C2C14AE22FEFAB91DFEBB" box="[300,380,295,321]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="387">SMNS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72DE6555DB7C2C14AFC9FE96BBB6FDC4" pageId="2" pageNumber="387" type="discussion">
<paragraph id="3A7B36DEDB7C2C14AFC9FE96B961FDE3" blockId="2.[151,1437,259,574]" pageId="2" pageNumber="387">
<emphasis id="08B0EACCDB7C2C14AFC9FE96B95CFE9F" bold="true" box="[199,317,331,357]" pageId="2" pageNumber="387">Remarks:</emphasis>
This species was listed in the Catalogue of Palaearctic
<taxonomicName id="FDC44D5DDB7C2C14ACA3FE96BC49FE9F" box="[941,1064,331,357]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="2" pageNumber="387" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
(
<bibRefCitation id="5E554B2FDB7C2C14AB38FE96BD69FE9F" author="Ando, K. &amp; Bouchard, P. &amp; Lillig, M. &amp; Lobl, I. &amp; Masumoto, K. &amp; Merkl, O. &amp; Schawaller, W." box="[1078,1288,331,357]" pageId="2" pageNumber="387" refId="ref10605" refString="Ando, K., Bouchard, P., Lillig, M., Lobl, I., Masumoto, K., Merkl, O. &amp; Schawaller, W. (2008) Tenebrionidae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Apollo Books, Stenstrup, pp. 105 - 352." type="book" year="2008">
Ando
<emphasis id="08B0EACCDB7C2C14AB70FE91BCD7FE9F" box="[1150,1206,331,357]" italics="true" pageId="2" pageNumber="387">et al.</emphasis>
(2008)
</bibRefCitation>
from
<collectingCountry id="42D3764EDB7C2C14AA43FE96BDFDFE9F" box="[1357,1436,331,357]" name="Greece" pageId="2" pageNumber="387">Greece</collectingCountry>
and also from
<collectingCountry id="42D3764EDB7C2C14AE39FEB2B9E8FE73" box="[311,393,367,393]" name="Turkey" pageId="2" pageNumber="387">Turkey</collectingCountry>
. The single Turkish record is based on
<bibRefCitation id="5E554B2FDB7C2C14AC34FEADBC25FE73" author="Ferrer, J. &amp; Soldati, L." box="[826,1092,367,394]" pageId="2" pageNumber="387" pagination="53 - 92" refId="ref10680" refString="Ferrer, J. &amp; Soldati, L. (1999) Contribution a l'etude des Tenebrionidae de Turquie (Insecta, Coleoptera). Entomofauna, 20, 53 - 92." type="journal article" year="1999">Ferrer &amp; Soldati (1999)</bibRefCitation>
, considered herein as misidentification. Also
<bibRefCitation id="5E554B2FDB7C2C14AE4BFE4EB996FE54" author="Schuster, A." box="[325,503,403,430]" pageId="2" pageNumber="387" pagination="495 - 629" refId="ref10978" refString="Schuster, A. (1916) Monographie der Coleopterengattung Laena Latreille. Verhandlungen der zoologisch-botanischen Gesellschaft in Wien, 66, 495 - 629." type="journal article" year="1916">Schuster (1916)</bibRefCitation>
did not know this species from
<collectingCountry id="42D3764EDB7C2C14AC50FE4EBBD1FE57" box="[862,944,403,429]" name="Turkey" pageId="2" pageNumber="387">Turkey</collectingCountry>
.
<taxonomicName id="FDC44D5DDB7C2C14ACB4FE49BC98FE57" authority="Reitter, 1891" authorityName="Reitter" authorityYear="1891" box="[954,1273,403,429]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="08B0EACCDB7C2C14ACB4FE49BC00FE57" box="[954,1121,403,429]" italics="true" pageId="2" pageNumber="387">L. ganglbaueri</emphasis>
Reitter, 1891
</taxonomicName>
from northern
<collectingCountry id="42D3764EDB7C2C14AF99FE6AB886FE2B" box="[151,231,439,465]" name="Turkey" pageId="2" pageNumber="387">Turkey</collectingCountry>
is similar in external morphology, but the femora are without distinct teeth, and the apicale of the aedeagus is longer (compare
<figureCitation id="A2FF2A5BDB7C2C14AE78FE06BA62FE0F" box="[374,515,475,501]" captionStart="FIGURES 22–46" captionStartId="7.[151,264,1694,1718]" captionTargetBox="[191,1396,184,1665]" captionTargetId="figure-16@7.[190,1397,181,1670]" captionTargetPageId="7" captionText="FIGURES 22–46. Right posterior male tibia of Laena species from Turkey and Greece (Figs. 22–25), and apicale of aedeagus of Laena species from Turkey and few from Greece (Figs. 26–46). 22 L. apfelbecki, Greece Évia, SMNS. 23 L. bozdagica sp. nov., holotype Izmir, SMNS. 24 L. kraatzi, Greece Chios, SMNS. 25 L. lindbergi, Bingöl, SMNS. 26 L. deplanata, Izmir, SMNS. 27 L. ferruginea, Greece Corfu, SMNS. 28 L. ferruginea, Antalya, SMNS (det. Kaszab as L. liliputana syn. nov.). 29 L. wanensis, Nur Daglari, SMNS. 30 L. bozdagica sp. nov., paratype Izmir, SMNS. 31 L. merkli, Hakkari, CPE. 32 L. obscuricornis, Meselik, SMNS. 33 L. byzantina, Artvin, SMNS. 34 L. korbi, Konya, SMNS. 35 L. lindbergi, Bitlis, SMNS. 36 L. apfelbecki, Greece Evia, SMNS. 37 L. pici, Antakya, SMNS. 38 L. ganglbaueri, Alem Dagh, SMNS. 39 L. weisei, Greece Sapka, SMNS. 40 L. theana, Erfelek, SMNS. 41 L. assingi sp. nov., holotype Fethiye, CPE. 42 L. graeca, Bolu, SMNS. 43 L. osmanlis, Bursa, SMNS. 44 L. quadricollis, Rize, SMNS. 45 L. quadricollis, Krasnodar Maikop, SMNS. 46 L. kraatzi, Alem Dagh, SMNS. Scale line: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4416445" httpUri="https://zenodo.org/record/4416445/files/figure.png" pageId="2" pageNumber="387">Figs. 36, 38</figureCitation>
). In
<taxonomicName id="FDC44D5DDB7C2C14AD37FE01BAADFE0F" authorityName="Schuster" authorityYear="1915" box="[569,716,475,501]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="apfelbecki">
<emphasis id="08B0EACCDB7C2C14AD37FE01BAADFE0F" box="[569,716,475,501]" italics="true" pageId="2" pageNumber="387">L. apfelbecki</emphasis>
</taxonomicName>
the male posterior tibia is distinctly swollen at interior margin (
<figureCitation id="A2FF2A5BDB7C2C14AFAEFE22B895FDE3" box="[160,244,511,537]" captionStart="FIGURES 22–46" captionStartId="7.[151,264,1694,1718]" captionTargetBox="[191,1396,184,1665]" captionTargetId="figure-16@7.[190,1397,181,1670]" captionTargetPageId="7" captionText="FIGURES 22–46. Right posterior male tibia of Laena species from Turkey and Greece (Figs. 22–25), and apicale of aedeagus of Laena species from Turkey and few from Greece (Figs. 26–46). 22 L. apfelbecki, Greece Évia, SMNS. 23 L. bozdagica sp. nov., holotype Izmir, SMNS. 24 L. kraatzi, Greece Chios, SMNS. 25 L. lindbergi, Bingöl, SMNS. 26 L. deplanata, Izmir, SMNS. 27 L. ferruginea, Greece Corfu, SMNS. 28 L. ferruginea, Antalya, SMNS (det. Kaszab as L. liliputana syn. nov.). 29 L. wanensis, Nur Daglari, SMNS. 30 L. bozdagica sp. nov., paratype Izmir, SMNS. 31 L. merkli, Hakkari, CPE. 32 L. obscuricornis, Meselik, SMNS. 33 L. byzantina, Artvin, SMNS. 34 L. korbi, Konya, SMNS. 35 L. lindbergi, Bitlis, SMNS. 36 L. apfelbecki, Greece Evia, SMNS. 37 L. pici, Antakya, SMNS. 38 L. ganglbaueri, Alem Dagh, SMNS. 39 L. weisei, Greece Sapka, SMNS. 40 L. theana, Erfelek, SMNS. 41 L. assingi sp. nov., holotype Fethiye, CPE. 42 L. graeca, Bolu, SMNS. 43 L. osmanlis, Bursa, SMNS. 44 L. quadricollis, Rize, SMNS. 45 L. quadricollis, Krasnodar Maikop, SMNS. 46 L. kraatzi, Alem Dagh, SMNS. Scale line: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4416445" httpUri="https://zenodo.org/record/4416445/files/figure.png" pageId="2" pageNumber="387">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="3A7B36DEDB7C2C14AFC9FDFEBBB6FDC4" blockId="2.[151,1437,259,574]" box="[199,983,547,574]" pageId="2" pageNumber="387">
<emphasis id="08B0EACCDB7C2C14AFC9FDFEB9D7FDC7" bold="true" box="[199,438,547,573]" pageId="2" pageNumber="387">Published localities:</emphasis>
Parnass, Euboea (both
<collectingCountry id="42D3764EDB7C2C14ADB0FDFEBB6EFDC7" box="[702,783,547,573]" name="Greece" pageId="2" pageNumber="387">Greece</collectingCountry>
) (
<bibRefCitation id="5E554B2FDB7C2C14AC2BFDFEBBABFDC4" author="Schuster, A." box="[805,970,547,574]" pageId="2" pageNumber="387" pagination="495 - 629" refId="ref10978" refString="Schuster, A. (1916) Monographie der Coleopterengattung Laena Latreille. Verhandlungen der zoologisch-botanischen Gesellschaft in Wien, 66, 495 - 629." type="journal article" year="1916">Schuster 1916</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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