194 lines
16 KiB
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194 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.816.29738" ID-GBIF-Dataset="31bce27d-7094-467b-95ae-009a27e00cf5" ID-PMC="PMC6341056" ID-Pensoft-Pub="1313-2970-816-1" ID-PubMed="30686917" ID-ZBK="51CEEF2E1E1040A8A6731140426ED5A7" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-816-1" ModsDocOrigin="ZooKeys 816" ModsDocTitle="A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)" checkinTime="1547561011487" checkinUser="pensoft" docAuthor="Hunting, Wesley & Yang, Man-Miao" docDate="2019" docId="9CE4A03AEEA685F0A81DD7845072417D" docLanguage="en" docName="ZooKeys 816: 1-164" docOrigin="ZooKeys 816" docSource="http://dx.doi.org/10.3897/zookeys.816.29738" docTitle="Coptodera (Coptoderina) japonica Bates" docType="treatment" docVersion="4" lastPageNumber="65" masterDocId="AA26FF84157BFFAE490E7F133744FF98" masterDocTitle="A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)" masterLastPageNumber="164" masterPageNumber="1" pageNumber="60" updateTime="1668166739332" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunting, Wesley</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yang, Man-Miao</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>816</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>164</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.816.29738</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.816.29738</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-816-1</mods:identifier>
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<mods:identifier type="ZBK">51CEEF2E1E1040A8A6731140426ED5A7</mods:identifier>
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<mods:identifier type="ZooBank">51CEEF2E1E1040A8A6731140426ED5A7</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="154125979" LSID="urn:lsid:plazi:treatment:9CE4A03AEEA685F0A81DD7845072417D" httpUri="http://treatment.plazi.org/id/9CE4A03AEEA685F0A81DD7845072417D" lastPageId="64" lastPageNumber="65" pageId="59" pageNumber="60">
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<subSubSection pageId="59" pageNumber="60" type="nomenclature">
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<paragraph pageId="59" pageNumber="60">
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<taxonomicName authority="Bates" class="Insecta" family="Carabidae" genus="Coptodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodera (Coptoderina) japonica" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="japonica" subGenus="Coptoderina">Coptodera (Coptoderina) japonica Bates</taxonomicName>
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Figs 41, 42, 43
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 57C, 59
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</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="reference_group">
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<paragraph pageId="60" pageNumber="61">
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<taxonomicName class="Insecta" family="Carabidae" genus="Coptodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodera japonica" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="japonica">
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<pageBreakToken pageId="60" pageNumber="61" start="start">Coptodera</pageBreakToken>
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japonica
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</taxonomicName>
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Bates, 1883: 281;
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<bibRefCitation pageId="60" pageNumber="61">Csiki 1932</bibRefCitation>
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: 1371;
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<bibRefCitation pageId="60" pageNumber="61">
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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1963
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</bibRefCitation>
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: 348;
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae). Second ed. Germany" pageId="152" pageNumber="153" title="Systematic List of Extant Ground Beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae & Carabidae incl." year="2005">Lorenz 2005</bibRefCitation>
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: 458;
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<bibRefCitation author="Park, J K" journalOrPublisher="The Entomological review of Japan" pageId="153" pageNumber="154" url="https://doi.org/10.1111/j.1748-5967.2011.00324.x" year="2011">Park et al. 2011</bibRefCitation>
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: 103.
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</paragraph>
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<paragraph pageId="60" pageNumber="61">
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<taxonomicName class="Insecta" family="Carabidae" genus="Coptodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodera formosana" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="formosana">Coptodera formosana</taxonomicName>
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Dupuis, 1912: 328;
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<bibRefCitation pageId="60" pageNumber="61">
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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1963
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</bibRefCitation>
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: 347.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="type material">
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<paragraph pageId="60" pageNumber="61">Types and other material examined.</paragraph>
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<paragraph pageId="60" pageNumber="61">
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Cotype (male) labeled: "Japan./G. Lewis./1910-320";
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<normalizedToken originalValue="“Cotype”">"Cotype"</normalizedToken>
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[circular, ringed with green]"; "Ex Coll./Brit. Mus."; "
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<taxonomicName class="Insecta" family="Carabidae" genus="Coptodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodera" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">Coptodera</taxonomicName>
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/
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<taxonomicName lsidName="japonica" pageId="60" pageNumber="61" rank="species" species="japonica">japonica</taxonomicName>
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/cotype Bates/H. E. Andrewes det.";
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<normalizedToken originalValue="“Syntypus”">"Syntypus"</normalizedToken>
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[rectangular, red paper]. Eight specimens of
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<taxonomicName lsidName="C. japonica" pageId="60" pageNumber="61" rank="species" species="japonica">C. japonica</taxonomicName>
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, three males and five females. For further details see EH Strickland Virtual Entomology Museum Database.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="type locality">
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<paragraph pageId="60" pageNumber="61">Type locality.</paragraph>
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<paragraph pageId="60" pageNumber="61">Japan: Kyushu (Kiushiu).</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="diagnosis">
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<paragraph pageId="60" pageNumber="61">Diagnosis.</paragraph>
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<paragraph pageId="60" pageNumber="61">
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Specimens of this species are easily distinguished from other Taiwanese
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<taxonomicName class="Insecta" family="Carabidae" genus="Coptodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodera" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">Coptodera</taxonomicName>
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by a combination of elytra with a seta in the apical 1/3 of stria 2 and black lateral margins.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="63" lastPageNumber="64" pageId="60" pageNumber="61" type="description">
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<paragraph pageId="60" pageNumber="61">Description.</paragraph>
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<paragraph pageId="60" pageNumber="61">OBL 9.16 - 10.33 mm. Length (n = three males, five females): head 1.00 - 1.12, pronotum 1.36 - 1.58, elytra 5.50 - 6.33, metepisternum 1.32 - 1.44 mm; width: head 2.00 - 2.20, pronotum 2.40 - 2.76, elytra 4.16 - 4.50, metepisternum 0.80 - 0.84 mm.</paragraph>
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<paragraph pageId="60" pageNumber="61">Body proportions. HW/HL 1.93 - 2.12; PWM/PL 1.64 - 1.82; EL/EW 1.30 - 1.42; ML/MW 1.57 - 1.67.</paragraph>
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<paragraph lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
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Color. Fig. 41. Dorsum of head piceous, area nearest to antennal socket brunneous to rufo-piceous, clypeus black with outer margin brunneous to rufo-brunneous, labrum, antennae, and palpi brunneous to rufo-brunneous; disc of pronotum rufo-piceous to piceous, lateral margins rufo-brunneous, only slightly lighter then disc; elytral disc black, with four testaceous maculae, two anterior and two posterior, anterior macula large, from interval 3 to 7 in most specimens, closest to base of elytra in
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<pageBreakToken pageId="61" pageNumber="62" start="start">interval</pageBreakToken>
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5 and 6, closest to apex of elytra in interval 4, posterior macula, from interval 2-8 (some from interval 1), closest to base of elytra in intervals 4 and 5, closest to apex of elytra in interval 4; margins of elytra rufo-brunneous to piceous; ventral surface rufo-brunneous to piceous; legs with trochanter and femora and tarsi brunneous to rufo-brunneous, tibia rufo-brunneous to piceous on dorsal surface.
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</paragraph>
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<caption pageId="61" pageNumber="62">
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<paragraph pageId="61" pageNumber="62">
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Figure 41. Dorsal habitus and color pattern of
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<taxonomicName lsidName="C. (Coptoderina) japonica" pageId="61" pageNumber="62" rank="species" species="japonica" subGenus="Coptoderina">C. (Coptoderina) japonica</taxonomicName>
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Bates. (OBL 10.24 mm).
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</paragraph>
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</caption>
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<paragraph pageId="61" pageNumber="62">
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Microsculpture. Dorsum of head with microsculpture somewhat granulate and isodiametric, easily visible at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification; pronotum somewhat granulate, isodiametric to somewhat transverse mesh pattern easily visible at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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; elytral intervals with transverse sculpticells, center of striae with isodiametric sculpticells a few cells wide; ventral surface of head with microsculpture transverse, faintly visible at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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; prosternum, proepipleuron, mesepisternum and metepisternum with sculpticells forming a shallow transverse mesh.
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</paragraph>
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<paragraph pageId="61" pageNumber="62">
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Macrosculpture and pilosity. Dorsum of head and clypeus smooth, with relatively dense, fine, scattered setigerous punctures, setae hardly visible at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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; pronotum with relatively dense, fine and scattered setigerous punctures, visible in side view at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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, disc shallowly rugulose; elytra with intervals broadly rounded, single row of fine setigerous punctures in the center of each interval, these punctures larger than additional scattered setigerous punctures throughout disc, striae with single row of fine setigerous punctures hardly visible in lateral view at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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; ventrally, thoracic and abdominal sclerites with scattered setigerous punctures throughout.
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</paragraph>
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<paragraph pageId="62" pageNumber="63">
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<pageBreakToken pageId="62" pageNumber="63" start="start">Fixed</pageBreakToken>
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setae. Elytra with two setae in apical 1/3 of stria 2, one seta near base in stria 3.
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</paragraph>
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<paragraph pageId="62" pageNumber="63">Luster. Dorsal and ventral surfaces moderately glossy.</paragraph>
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<paragraph pageId="62" pageNumber="63">Head. Mandibles somewhat curved at apex, relatively long and narrow in form, when measured on outside diameter, visible portion longer than length of labrum; labrum relatively stout and rounded, slightly wider than clypeus at max width, broadly bilobed to almost flat apically.</paragraph>
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<paragraph pageId="62" pageNumber="63">Pronotum. Transverse impression deep; posterior transverse impression deep, median longitudinal impression moderately deep; lateral margins explanate, apico-lateral margins broadly rounded, posterio-lateral margins obtuse.</paragraph>
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<paragraph pageId="62" pageNumber="63">Elytra. Apex slightly sinuate.</paragraph>
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<paragraph pageId="62" pageNumber="63">Legs. Two rows of small squamo-setae on tarsomeres 1-2 of mid-leg; males with two notches apically on ventral side of mid-tibia. Note: one individual with two notches on left mid-tibia and three notches on right mid-tibia has been observed.</paragraph>
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<paragraph lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
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Male genitalia. Figs 42, 43
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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Length 1.86 - 1.92 mm. Ostium catopic, positioned slightly to right when viewed ventrally, phallus cylindrical, slanting right from base of ostium towards apex when viewed ventrally, apical area with short, bluntly rounded apex; endophallus long and narrow, positioned to right of phallus when everted and viewed ventrally, microtrichia dispersed evenly and rather divergently on most
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<pageBreakToken pageId="63" pageNumber="64" start="start">of</pageBreakToken>
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surface, one large and typically more dense microtrichial field (mtf) present on left side of phallus, just before apical constriction, sclerotized ring of spines (esr) at narrowest portion of apical constriction.
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</paragraph>
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<caption pageId="63" pageNumber="64">
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<paragraph pageId="63" pageNumber="64">
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Figure 42. Digital image of male genitalia of
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<taxonomicName lsidName="C. (Coptoderina) japonica" pageId="63" pageNumber="64" rank="species" species="japonica" subGenus="Coptoderina">C. (Coptoderina) japonica</taxonomicName>
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Bates, ventral aspect, endophallus everted. Legend: esr endophallic sclerotized ring.
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</paragraph>
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</caption>
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<caption pageId="63" pageNumber="64">
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<paragraph pageId="63" pageNumber="64">
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Figure 43. Digital images of male genitalia of
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<taxonomicName lsidName="C. (Coptoderina) japonica" pageId="63" pageNumber="64" rank="species" species="japonica" subGenus="Coptoderina">C. (Coptoderina) japonica</taxonomicName>
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Bates. A right lateral aspect B ventral aspect C left lateral aspect.
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</paragraph>
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</caption>
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<paragraph pageId="63" pageNumber="64">Female genitalia. 57C. Width 1.28 mm. One spermatheca (sp1), cylindrical and long, ribbed laterally along length, distinctively curved at base; one spermathecal accessory gland (sg), large and bulbous; spermathecal gland duct (sgd) only slightly longer than length of spermatheca, slightly swollen at apical end, just before gland, attachment site medially on distinctively shaped diverticulum (div) of spermatheca; bursa copulatrix (bc) with distinctive sac at apical end (bs), highly constricted near opening to common oviduct, then expanding out into and oblong mushroom top shaped chamber.</paragraph>
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</subSubSection>
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<subSubSection pageId="63" pageNumber="64" type="habitat">
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<paragraph pageId="63" pageNumber="64">Habitat, habits, and seasonal occurrence.</paragraph>
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<paragraph pageId="63" pageNumber="64">
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The known elevational range of
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<taxonomicName lsidName="C. japonica" pageId="63" pageNumber="64" rank="species" species="japonica">C. japonica</taxonomicName>
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is from 640 to 2770 meters. Adults of this species are found in mixed forest of montane areas. Little is known of the habits of this species but one specimen of this species was collected on shelf fungus. Specimens have been collected from April to October in Taiwan and methods of collecting include u.v. light, malaise trap, and hand collecting.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="64" pageNumber="65" type="distribution">
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<paragraph pageId="64" pageNumber="65">
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<pageBreakToken pageId="64" pageNumber="65" start="start">Geographical</pageBreakToken>
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distribution.
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</paragraph>
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<paragraph pageId="64" pageNumber="65">
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<taxonomicName class="Insecta" family="Carabidae" genus="Coptodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodera japonica" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="japonica">Coptodera japonica</taxonomicName>
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is known from Japan, Korea, and Taiwan. For Taiwan collecting localities see Figure 59.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |