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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.899.47070" ID-GBIF-Dataset="2c7e9ce7-8a11-4d64-9a73-40bb2eb487fb" ID-PMC="PMC6926428" ID-Pensoft-Pub="1313-2970-899-109" ID-Pensoft-UUID="E0FA12EBCCD55E3B8BC61796697C69C3" ID-PubMed="31875090" ID-ZooBank="9AA5216D3150475DA165B36EABCB61E2" ModsDocID="1313-2970-899-109" checkinTime="1576248706674" checkinUser="pensoft" docAuthor="Taylor, John D. &amp; Glover, Emily A." docDate="2019" docId="7FBF93FC99455711A759808852ACCB2E" docLanguage="en" docName="ZooKeys 899: 109-140" docOrigin="ZooKeys 899" docSource="http://dx.doi.org/10.3897/zookeys.899.47070" docTitle="Rugalucina angela Melvill 1899" docType="treatment" docVersion="4" id="E0FA12EBCCD55E3B8BC61796697C69C3" lastPageNumber="109" masterDocId="E0FA12EBCCD55E3B8BC61796697C69C3" masterDocTitle="Unloved, paraphyletic or misplaced: new genera and species of small to minute lucinid bivalves and their relationships (Bivalvia, Lucinidae)" masterLastPageNumber="140" masterPageNumber="109" pageNumber="109" updateTime="1668168230035" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Unloved, paraphyletic or misplaced: new genera and species of small to minute lucinid bivalves and their relationships (Bivalvia, Lucinidae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Taylor, John D.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Glover, Emily A.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>899</mods:number>
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<mods:start>109</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.899.47070</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.899.47070</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-899-109</mods:identifier>
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<treatment ID-GBIF-Taxon="160702943" LSID="urn:lsid:plazi:treatment:7FBF93FC99455711A759808852ACCB2E" httpUri="http://treatment.plazi.org/id/7FBF93FC99455711A759808852ACCB2E" lastPageNumber="109" pageId="0" pageNumber="109">
<subSubSection pageId="0" pageNumber="109" type="nomenclature">
<paragraph pageId="0" pageNumber="109">
<taxonomicName LSID="7FBF93FC-9945-5711-A759-808852ACCB2E" authority="Melvill, 1899" authorityName="Melvill" authorityYear="1899" class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela">Rugalucina angela (Melvill, 1899)</taxonomicName>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Rugalucina angela (Melvill, 1899). A-C Syntype of Lucina (Codakia) angela Melvill, 1899 (NHMUK 1899.12.18.20), exterior of left valve and interiors of right and left valves. Gwadur, Pakistan, L 8.1 mm D-F L. (C.) angela syntype (NHMUK 1899.12.18.20), exterior of left valve and interiors of right and left valves, L 6.1 mm G-M Rugalucina angela Ras al Khaimah, Arabian Gulf, (NHMUK 20191071) G exterior SEM of right valve, L 5.0 mm H, I interior SEM of right and left valves, L 7.7 mm J dorsal view, L 5.9 mm K exterior of left valve, L 7.9 mm L, M exterior of left valve, interior of right valve, L 7.6 mm N, O R. angela exterior and interior of left valve, Gulf of Suez (NHMUK 1868.5.29.2), L 13.7 mm P R. angela exterior of right valve and interior of right and left valves, Egypt, 7 km south of Hurgada, H Dekker colln 4569, L 12.8 mm Q interior of left and right valves, Egypt, Port Safaga, H Dekker colln 3263, L 9.4 mm R, S detail of hinge teeth of Q. T R. angela exterior of left valve. Red Sea, Yemen, Orestes Point, N of Midi, H Dekker colln 4553, L 9.8 mm U R. angela exterior of right and interior of left valves, Aden (NHMUK 1963340), L 10.6 mm V R. angela exterior and interior of right valve, Krusadai, India, (NHMUK 1953.1.30.69 - 76), L 8.4 mm." figureDoi="10.3897/zookeys.899.47070.figure2" httpUri="https://binary.pensoft.net/fig/364609" pageId="0" pageNumber="109" tableDoi="10.3897/zookeys.899.47070.figure2">Figs 2</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Internal drawings of left valves of Rugalucina and Pusillolucina species. Not to scale." figureDoi="10.3897/zookeys.899.47070.figure3" httpUri="https://binary.pensoft.net/fig/364610" pageId="0" pageNumber="109" tableDoi="10.3897/zookeys.899.47070.figure3">3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="reference_group">
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Lucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lucina fischeriana" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="fischeriana">Lucina fischeriana</taxonomicName>
Issel, 1869: 83-84, pl. 1, fig. 8 (non
<taxonomicName family="Lucinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="L. fischeriana" order="Lucinida" pageId="0" pageNumber="109" rank="species" species="fischeriana">L. fischeriana</taxonomicName>
<normalizedToken originalValue="dOrbigny">d'Orbigny</normalizedToken>
, 1845: Jurassic fossil).
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Lucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lucina (Codakia) angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela" subGenus="Codakia">Lucina (Codakia) angela</taxonomicName>
Melvill, 1899: 98, pl. 2, fig. 8.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Loripes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loripes fischeriana" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="fischeriana">Loripes fischeriana</taxonomicName>
:
<bibRefCitation author="Lamy, E" journalOrPublisher="Bulletin of Marine Science" pageId="0" pageNumber="109" refId="B29" refString="Lamy, E, 1916. Les Lucines et les Diplodontes de la Mer Rouge (d'apres les materiaux recueillis par M. de Dr Jousseaume). Bulletin du Museum national d'Histoire naturelle, Paris 22: 145-155, 183-190." title="Les Lucines et les Diplodontes de la Mer Rouge (d'apres les materiaux recueillis par M. de Dr Jousseaume). Bulletin du Museum national d'Histoire naturelle, Paris 22: 145 - 155, 183 - 190." year="1916">Lamy 1916</bibRefCitation>
: 151.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pillucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pillucina fischeriana" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="fischeriana">Pillucina fischeriana</taxonomicName>
:
<bibRefCitation author="Oliver, PG" journalOrPublisher="Verlag Christa Hemmen &amp; National Museum of Wales, Wiesbaden &amp; Cardiff" pageId="0" pageNumber="109" refId="B36" refString="Oliver, PG, 1992. Bivalved seashells of the Red Sea. Verlag Christa Hemmen &amp; National Museum of Wales, Wiesbaden &amp; Cardiff" title="Bivalved seashells of the Red Sea." year="1992">Oliver 1992</bibRefCitation>
: 98, pl. 20, fig. 4.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pillucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pillucina fischeriana" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="fischeriana">Pillucina fischeriana</taxonomicName>
:
<bibRefCitation author="Oliver, PG" journalOrPublisher="Verlag Christa Hemmen &amp; National Museum of Wales, Wiesbaden &amp; Cardiff" pageId="0" pageNumber="109" refId="B37" refString="Oliver, PG, 1995. Bivalvia. In: Dance SP (Ed.) Seashells of Eastern Arabia Motivate Publishing, Dubai, 194-281." title="Bivalvia. In: Dance SP (Ed.) Seashells of Eastern Arabia Motivate Publishing, Dubai, 194 - 281." year="1995">Oliver 1995</bibRefCitation>
: 236, fig. 1026.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pillucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pillucina angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela">Pillucina angela</taxonomicName>
:
<bibRefCitation author="Oliver, PG" journalOrPublisher="Verlag Christa Hemmen &amp; National Museum of Wales, Wiesbaden &amp; Cardiff" pageId="0" pageNumber="109" refId="B37" refString="Oliver, PG, 1995. Bivalvia. In: Dance SP (Ed.) Seashells of Eastern Arabia Motivate Publishing, Dubai, 194-281." title="Bivalvia. In: Dance SP (Ed.) Seashells of Eastern Arabia Motivate Publishing, Dubai, 194 - 281." year="1995">Oliver 1995</bibRefCitation>
: 236, fig. 1025.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pillucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pillucina vietnamica" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="vietnamica">Pillucina vietnamica</taxonomicName>
(part):
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.53.2001.1349" author="Glover, EA" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="109" pagination="263 - 292" refId="B18" refString="Glover, EA, Taylor, JD, 2001. Systematic revision of Australian and Indo-Pacific Lucinidae (Mollusca: Bivalvia): Pillucina, Wallucina and descriptions of two new genera and four new species. Records of the Australian Museum 53: 263 - 292" title="Systematic revision of Australian and Indo-Pacific Lucinidae (Mollusca: Bivalvia): Pillucina, Wallucina and descriptions of two new genera and four new species." url="https://doi.org/10.3853/j.0067-1975.53.2001.1349" volume="53" year="2001">Glover and Taylor 2001</bibRefCitation>
: 273.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pillucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pillucina angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela">Pillucina angela</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.53.2001.1349" author="Glover, EA" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="109" pagination="263 - 292" refId="B18" refString="Glover, EA, Taylor, JD, 2001. Systematic revision of Australian and Indo-Pacific Lucinidae (Mollusca: Bivalvia): Pillucina, Wallucina and descriptions of two new genera and four new species. Records of the Australian Museum 53: 263 - 292" title="Systematic revision of Australian and Indo-Pacific Lucinidae (Mollusca: Bivalvia): Pillucina, Wallucina and descriptions of two new genera and four new species." url="https://doi.org/10.3853/j.0067-1975.53.2001.1349" volume="53" year="2001">Glover and Taylor 2001</bibRefCitation>
: 279, figs 9 h, i.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pillucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pillucina angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela">Pillucina angela</taxonomicName>
(part):
<bibRefCitation author="Huber, M" journalOrPublisher="ConchBooks, Harxheim, Germany" pageId="0" pageNumber="109" refId="B24" refString="Huber, M, 2015. Compendium of bivalves 2. ConchBooks, Harxheim, Germany" title="Compendium of bivalves 2." year="2015">Huber 2015</bibRefCitation>
: 422, figs p. 76.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="type material">
<paragraph pageId="0" pageNumber="109">Type material.</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Lucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lucina fischeriana" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis italics="true" pageId="0" pageNumber="109">Lucina fischeriana</emphasis>
</taxonomicName>
5 syntypes (MCG), type locality: Suez, Egypt.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName lsidName="L. angela" pageId="0" pageNumber="109" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">L. angela</emphasis>
</taxonomicName>
two syntypes NHMUK1899.12.18.20-21; L 7.9 mm and 6.1 mm; 1 syntype NMW 1955.158.684.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="type locality">
<paragraph pageId="0" pageNumber="109">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="109">Gwadur, Pakistan, 8 fathoms (15 m).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="description">
<paragraph pageId="0" pageNumber="109">Description.</paragraph>
<paragraph pageId="0" pageNumber="109">Small (L to 15 mm), subcircular, inflated. Colour white, yellow or orange. Waxy appearance. Sculpture of strong diverging radial ribs, broader and more widely spaced to the anterior and posterior. Ribs crossed by fine, closely spaced, commarginal lamellae which curve over ribs producing a roughly scabrous appearance. Central parts of shell generally without ribs. Lunule large, broadly lanceolate, smooth. Ligament internal, obliquely inset. Hinge: RV with single large cardinal tooth and short anterior and posterior lateral teeth, LV with two cardinal teeth, lateral teeth consisting of small sockets. Anterior adductor scar narrow, elongate, detached from pallial line for ca. half of length. Pallial line irregularly lobate, or slightly divided. Inner shell margin coarsely crenulate to anterior and posterior.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="distribution">
<paragraph pageId="0" pageNumber="109">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Red Sea</emphasis>
: Great Bitter Lake (Hoffman et al. 2006), Suez Canal, El Ballah (NHMUK 1950.11.10.1), Suez (NHMUK 1968.5.29.2), Safaga, Dongonab Bay (NHMUK), Oreste Point, Yemen (Dekker colln), Aden (NHMUK 1963340).
<emphasis bold="true" pageId="0" pageNumber="109">Arabian Gulf</emphasis>
: Kuwait (NHMUK), Tarut Bay, Qatar (NHMUK), Abu Dhabi (NHMUK), Ras al Khaimah (NHMUK).
<emphasis bold="true" pageId="0" pageNumber="109">Arabian Sea</emphasis>
: Khor Kalba, Sharjah (NHMUK), Karachi, dredged (NHMUK 1953.1.30.85). Oman: Masirah Island (NHMUK).
<emphasis bold="true" pageId="0" pageNumber="109">Indian Ocean</emphasis>
: South India: Chennai (Madras) (NHMUK 1953.1.30.169-73), Krusadai Island (NHMUK 1953.1.30.110), Kundugal Point, Krusadai Island (NHMUK1953.1.30.175-181), Tuticorin (NHMUK 1953.1.30.99-101). Sri Lanka: Trincomalee (NHMUK 1910.9.28.175-178).
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">Rugalucina angela</emphasis>
</taxonomicName>
(as
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pillucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pillucina vietnamica" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="vietnamica">
<emphasis italics="true" pageId="0" pageNumber="109">Pillucina vietnamica</emphasis>
</taxonomicName>
) is recorded as an invasive species off Israel in the eastern Mediterranean (
<bibRefCitation DOI="https://doi.org/10.3391/bir.2018.7.3.05" author="Steger, J" journalOrPublisher="Bioinvasions Records" pageId="0" pageNumber="109" pagination="245 - 257" refId="B45" refString="Steger, J, Stockinger, M, Ivkic, A, Galil, BS, Albano, PG, 2018. New records of non-indigenous molluscs from the eastern Mediterranean Sea. Bioinvasions Records 7: 245 - 257" title="New records of non-indigenous molluscs from the eastern Mediterranean Sea." url="https://doi.org/10.3391/bir.2018.7.3.05" volume="7" year="2018">Steger et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="remarks">
<paragraph pageId="0" pageNumber="109">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">Rugalucina angela</emphasis>
</taxonomicName>
shows variation in shell morphology between various localities around the Arabian Peninsula and Red Sea. For example, shells from the Arabian Gulf are usually smaller, while those from the northern Red Sea as on Gulf of Suez shores are generally larger, and the marginal crenulations stronger. We regard these differences as ecophenotypic probably associated with the extreme environmental conditions such as the very high salinities experienced in the southern Arabian Gulf, usually more than 40 psu but in shallow lagoons as high as 52-55 psu (
<bibRefCitation DOI="https://doi.org/10.1080/00222938200770421" author="Price, ARG" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="109" pagination="519 - 527" refId="B40" refString="Price, ARG, 1982. Western Arabian Gulf echinoderms in high salinity waters and the occurrence of dwarfism. Journal of Natural History 16: 519 - 527" title="Western Arabian Gulf echinoderms in high salinity waters and the occurrence of dwarfism." url="https://doi.org/10.1080/00222938200770421" volume="16" year="1982">Price 1982</bibRefCitation>
) and approximately 40-46 psu at Safaga in the northern Red Sea (
<bibRefCitation author="Zuschin, M" journalOrPublisher="Naturhistorisches Museum, Vienna" pageId="0" pageNumber="109" refId="B55" refString="Zuschin, M, Oliver, PG, 2003. Bivalves and bivalve habitats in the northern Red Sea. The northern Bay of Safaga (Red Sea, Egypt): an actuopalaeontological approach. VI. Bivalvia. Naturhistorisches Museum, Vienna" title="Bivalves and bivalve habitats in the northern Red Sea. The northern Bay of Safaga (Red Sea, Egypt): an actuopalaeontological approach. VI. Bivalvia." year="2003">Zuschin and Oliver 2003</bibRefCitation>
) and probably higher for the habitats occupied by
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Rugalucina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="109">
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">Rugalucina angela</emphasis>
</taxonomicName>
is closely similar in shell morphology to
<taxonomicName lsidName="R. vietnamica" pageId="0" pageNumber="109" rank="species" species="vietnamica">
<emphasis italics="true" pageId="0" pageNumber="109">R. vietnamica</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="R. munda" pageId="0" pageNumber="109" rank="species" species="munda">
<emphasis italics="true" pageId="0" pageNumber="109">R. munda</emphasis>
</taxonomicName>
. The differences are subtle; externally they share diverging radial ribs that are stronger to anterior and posterior, and the fine commarginal lamellae.
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">Rugalucina angela</emphasis>
</taxonomicName>
has a shorter anterior dorsal area, larger hinge plate and teeth, and a slightly more divergent anterior adductor scar.
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina vietnamica" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="vietnamica">
<emphasis italics="true" pageId="0" pageNumber="109">Rugalucina vietnamica</emphasis>
</taxonomicName>
is higher, with a longer anterior dorsal area.
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina munda" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="munda">
<emphasis italics="true" pageId="0" pageNumber="109">Rugalucina munda</emphasis>
</taxonomicName>
is similar but the radial sculpture is much less pronounced with finer margin denticulations and subdued commarginal sculpture.
</paragraph>
<caption doi="10.3897/zookeys.899.47070.figure2" httpUri="https://binary.pensoft.net/fig/364609" pageId="0" pageNumber="109" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 2.</emphasis>
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">Rugalucina angela</emphasis>
</taxonomicName>
(Melvill, 1899).
<emphasis bold="true" pageId="0" pageNumber="109">
<normalizedToken originalValue="AC">A-C</normalizedToken>
</emphasis>
Syntype of
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Lucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lucina (Codakia) angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela" subGenus="Codakia">Lucina (Codakia) angela</taxonomicName>
Melvill, 1899 (NHMUK 1899.12.18.20), exterior of left valve and interiors of right and left valves. Gwadur, Pakistan, L 8.1mm
<emphasis bold="true" pageId="0" pageNumber="109">
<normalizedToken originalValue="DF">D-F</normalizedToken>
</emphasis>
<taxonomicName lsidName="L. (C.) angela" pageId="0" pageNumber="109" rank="species" species="angela" subGenus="C.">L. (C.) angela</taxonomicName>
syntype (NHMUK 1899.12.18.20), exterior of left valve and interiors of right and left valves, L 6.1 mm
<emphasis bold="true" pageId="0" pageNumber="109">
<normalizedToken originalValue="GM">G-M</normalizedToken>
</emphasis>
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina angela" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">Rugalucina angela</emphasis>
</taxonomicName>
Ras al Khaimah, Arabian Gulf, (NHMUK 20191071)
<emphasis bold="true" pageId="0" pageNumber="109">G</emphasis>
exterior SEM of right valve, L 5.0 mm
<emphasis bold="true" pageId="0" pageNumber="109">H, I</emphasis>
interior SEM of right and left valves, L 7.7 mm
<emphasis bold="true" pageId="0" pageNumber="109">J</emphasis>
dorsal view, L 5.9 mm
<emphasis bold="true" pageId="0" pageNumber="109">K</emphasis>
exterior of left valve, L 7.9 mm
<emphasis bold="true" pageId="0" pageNumber="109">L, M</emphasis>
exterior of left valve, interior of right valve, L 7.6 mm
<emphasis bold="true" pageId="0" pageNumber="109">N, O</emphasis>
<taxonomicName lsidName="R. angela" pageId="0" pageNumber="109" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">R. angela</emphasis>
</taxonomicName>
exterior and interior of left valve, Gulf of Suez (NHMUK1868.5.29.2), L 13.7 mm
<emphasis bold="true" pageId="0" pageNumber="109">P</emphasis>
<taxonomicName lsidName="R. angela" pageId="0" pageNumber="109" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">R. angela</emphasis>
</taxonomicName>
exterior of right valve and interior of right and left valves, Egypt, 7km south of Hurgada, H Dekker colln 4569, L 12.8 mm
<emphasis bold="true" pageId="0" pageNumber="109">Q</emphasis>
interior of left and right valves, Egypt, Port Safaga, H Dekker colln 3263, L 9.4 mm
<emphasis bold="true" pageId="0" pageNumber="109">R, S</emphasis>
detail of hinge teeth of
<emphasis bold="true" pageId="0" pageNumber="109">Q</emphasis>
.
<emphasis bold="true" pageId="0" pageNumber="109">T</emphasis>
<taxonomicName lsidName="R. angela" pageId="0" pageNumber="109" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">R. angela</emphasis>
</taxonomicName>
exterior of left valve. Red Sea, Yemen, Orestes Point, N of Midi, H Dekker colln 4553, L 9.8 mm
<emphasis bold="true" pageId="0" pageNumber="109">U</emphasis>
<taxonomicName lsidName="R. angela" pageId="0" pageNumber="109" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">R. angela</emphasis>
</taxonomicName>
exterior of right and interior of left valves, Aden (NHMUK 1963340), L 10.6 mm
<emphasis bold="true" pageId="0" pageNumber="109">V</emphasis>
<taxonomicName lsidName="R. angela" pageId="0" pageNumber="109" rank="species" species="angela">
<emphasis italics="true" pageId="0" pageNumber="109">R. angela</emphasis>
</taxonomicName>
exterior and interior of right valve, Krusadai, India, (NHMUK 1953.1.30.69-76), L 8.4 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.899.47070.figure3" httpUri="https://binary.pensoft.net/fig/364610" pageId="0" pageNumber="109" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 3.</emphasis>
Internal drawings of left valves of
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Rugalucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rugalucina" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Rugalucina</emphasis>
</taxonomicName>
and
<taxonomicName class="Bivalvia" family="Lucinidae" genus="Pusillolucina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pusillolucina" order="Lucinida" pageId="0" pageNumber="109" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Pusillolucina</emphasis>
</taxonomicName>
species. Not to scale.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>