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<document id="CD141490D1C36F39892D37A027311990" ID-DOI="10.5281/zenodo.189527" ID-GBIF-Dataset="7673d267-e2cc-4e23-89dd-26662a841fae" ID-ISSN="1175-5326" ID-Zenodo-Dep="189527" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461102917121" checkinUser="plazi" docAuthor="Giłka, Wojciech &amp; Paasivirta, Lauri" docDate="2009" docId="7F378787FFB9A569FF64BAB3550AFC65" docLanguage="en" docName="zt02197p042.pdf" docOrigin="Zootaxa 2197" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tanytarsus brundini Lindeberg 1963" docType="treatment" docVersion="7" lastPageNumber="38" masterDocId="830EFFFFFFBDA56EFFF3BE715779FFF6" masterDocTitle="Evaluation of diagnostic characters of the Tanytarsus chinyensis group (Diptera: Chironomidae), with description of a new species from Lapland" masterLastPageNumber="42" masterPageNumber="31" pageNumber="35" updateTime="1698591998081" updateUser="plazi">
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<mods:title id="214CAF21CB641C9D2FBE07E279AEC665">Evaluation of diagnostic characters of the Tanytarsus chinyensis group (Diptera: Chironomidae), with description of a new species from Lapland</mods:title>
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<mods:namePart id="78F35347BD085250C4DC1251F469BB71">Giłka, Wojciech</mods:namePart>
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<mods:namePart id="2E58589B6663CDFAB89ED8E4FAD81089">Paasivirta, Lauri</mods:namePart>
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<mods:date id="C6A8B9917AD40E2979AD736317BAC9D4">2009</mods:date>
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<treatment id="7F378787FFB9A569FF64BAB3550AFC65" ID-DOI="http://doi.org/10.5281/zenodo.6222777" ID-GBIF-Taxon="119622349" ID-Zenodo-Dep="6222777" LSID="urn:lsid:plazi:treatment:7F378787FFB9A569FF64BAB3550AFC65" httpUri="http://treatment.plazi.org/id/7F378787FFB9A569FF64BAB3550AFC65" lastPageId="7" lastPageNumber="38" pageId="4" pageNumber="35">
<subSubSection id="BF84651AFFB9A56AFF64BAB356C8FB09" pageId="4" pageNumber="35" type="nomenclature">
<paragraph id="F7213691FFB9A56AFF64BAB35526FB2A" blockId="4.[151,607,1218,1279]" box="[151,607,1218,1244]" pageId="4" pageNumber="35">
<heading id="AC6981FDFFB9A56AFF64BAB35526FB2A" bold="true" box="[151,607,1218,1244]" fontSize="11" level="1" pageId="4" pageNumber="35" reason="1">
<taxonomicName id="309E4D12FFB9A56AFF64BAB35526FB2A" ID-CoL="8FCVP" authority="Lindeberg, 1963" authorityName="Lindeberg" authorityYear="1963" box="[151,607,1218,1244]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="brundini">
<emphasis id="C5EAEA83FFB9A56AFF64BAB35526FB2A" bold="true" box="[151,607,1218,1244]" pageId="4" pageNumber="35">
<emphasis id="C5EAEA83FFB9A56AFF64BAB356F5FB2A" bold="true" box="[151,396,1218,1244]" italics="true" pageId="4" pageNumber="35">Tanytarsus brundini</emphasis>
<bibRefCitation id="930F4B60FFB9A56AFE60BAB35526FB2A" author="Lindeberg" box="[403,607,1218,1244]" pageId="4" pageNumber="42" refString="Lindeberg, B. (1963) Taxonomy, biology and biometry of Tanytarsus curticornis Kieff. and T. brundini n. sp. (Dipt., Chironomidae). Annales Entomologici Fennici, 29, 118 - 130." type="journal article" year="1963">Lindeberg, 1963</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F7213691FFB9A56AFF64BA9456C8FB09" blockId="4.[151,607,1218,1279]" box="[151,433,1253,1279]" pageId="4" pageNumber="35">
(
<figureCitation id="6FA52A14FFB9A56AFF6CBA945790FB09" box="[159,233,1253,1279]" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1259,1283]" captionTargetBox="[174,1412,196,1223]" captionTargetId="figure@3.[154,1439,158,1256]" captionTargetPageId="3" captionText="FIGURES 1 3. Male wing. 1: Tanytarsus brundini, 2: T. curticornis, 3: T. salmelai sp. n." httpUri="https://zenodo.org/record/189528/files/figure.png" pageId="4" pageNumber="35">Figs 1</figureCitation>
,
<figureCitation id="6FA52A14FFB9A56AFF0BBA94565FFB09" box="[248,294,1253,1279]" captionStart="FIGURES 5 18" captionStartId="6.[151,274,1982,2006]" captionTargetBox="[242,1340,373,1967]" captionTargetId="figure@6.[234,1353,373,1968]" captionTargetPageId="6" captionText="FIGURES 5 18. Variability in hypopygial anal point. 5 9: Tanytarsus brundini, 10 14: T. curticornis, 15 18: T. salmelai sp. n." httpUri="https://zenodo.org/record/189530/files/figure.png" pageId="4" pageNumber="35">59</figureCitation>
,
<figureCitation id="6FA52A14FFB9A56AFEC1BA9456D1FB09" box="[306,424,1253,1279]" captionStart="FIGURES 19 27" captionStartId="8.[151,269,1787,1811]" captionTargetBox="[177,1384,204,1746]" captionTargetId="figure@8.[162,1424,194,1793]" captionTargetPageId="8" captionText="FIGURES 19 27. Hypopygial volsellae. 19 21: superior volsella and digitus, 22 24: median volsella and inner margin of coxite, 25 27: inferior volsella. 19, 22, 25: Tanytarsus brundini; 20, 23, 26: T. curticornis; 21, 24, 27: T. salmelai sp. n." httpUri="https://zenodo.org/record/189531/files/figure.png" pageId="4" pageNumber="35">19, 22, 25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BF84651AFFB9A56AFF64BB5B5402FA96" pageId="4" pageNumber="35" type="reference_group">
<paragraph id="F7213691FFB9A56AFF64BB5B5402FA96" blockId="4.[151,1003,1321,1377]" pageId="4" pageNumber="35">
<treatmentCitationGroup id="D78E11BFFFB9A56AFF64BB5B5402FA96" pageId="4" pageNumber="35">
<taxonomicName id="309E4D12FFB9A56AFF64BB5B55D3FAB6" ID-CoL="8FCVS" authority="Kieffer, 1911" authorityName="Kieffer" authorityYear="1911" box="[151,682,1321,1345]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="curticornis" subGenus="Tanytarsus">
<emphasis id="C5EAEA83FFB9A56AFF64BB5B5672FAB7" box="[151,267,1322,1345]" italics="true" pageId="4" pageNumber="35">Tanytarsus</emphasis>
(
<emphasis id="C5EAEA83FFB9A56AFEE9BB5B56F7FAB7" box="[282,398,1322,1345]" italics="true" pageId="4" pageNumber="35">Tanytarsus</emphasis>
)
<emphasis id="C5EAEA83FFB9A56AFE6EBB5B556BFAB7" box="[413,530,1322,1345]" italics="true" pageId="4" pageNumber="35">curticornis</emphasis>
Kieffer, 1911
</taxonomicName>
:
<treatmentCitation id="763F1080FFB9A56AFD46BB5854FDFAB6" author="Edwards" box="[693,900,1321,1344]" page="415" pageId="4" pageNumber="35" year="1929">
<bibRefCitation id="930F4B60FFB9A56AFD46BB58542BFAB6" author="Edwards" box="[693,850,1321,1344]" pageId="4" pageNumber="41" refString="Edwards, F. W. (1929) British non-biting midges (Diptera, Chironomidae). Transactions of the Entomological Society of London, 77, 279 - 430." type="journal article" year="1929">Edwards 1929</bibRefCitation>
: 415
</treatmentCitation>
(
<emphasis id="C5EAEA83FFB9A56AFC67BB5B54A2FAB7" box="[916,987,1322,1345]" italics="true" pageId="4" pageNumber="35">partim</emphasis>
).
<taxonomicName id="309E4D12FFB9A56AFF64BB3B5525FA96" ID-CoL="8FCVP" authority="Lindeberg, 1963: 127" authorityName="Lindeberg" authorityPageNumber="127" authorityYear="1963" box="[151,604,1353,1377]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="brundini">
<emphasis id="C5EAEA83FFB9A56AFF64BB3B5614FA97" box="[151,365,1354,1377]" italics="true" pageId="4" pageNumber="35">Tanytarsus brundini</emphasis>
<treatmentCitation id="763F1080FFB9A56AFE87BB385525FA96" author="Lindeberg" box="[372,604,1353,1376]" page="127" pageId="4" pageNumber="35" year="1963">
<bibRefCitation id="930F4B60FFB9A56AFE87BB38555EFA96" author="Lindeberg" box="[372,551,1353,1376]" pageId="4" pageNumber="42" refString="Lindeberg, B. (1963) Taxonomy, biology and biometry of Tanytarsus curticornis Kieff. and T. brundini n. sp. (Dipt., Chironomidae). Annales Entomologici Fennici, 29, 118 - 130." type="journal article" year="1963">Lindeberg, 1963</bibRefCitation>
: 127
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="763F1080FFB9A56AFD94BB38540EFA96" author="Reiss" box="[615,887,1353,1376]" page="98" pageId="4" pageNumber="35" year="1971">
<bibRefCitation id="930F4B60FFB9A56AFD94BB385436FA96" author="Reiss" box="[615,847,1353,1376]" pageId="4" pageNumber="42" refString="Reiss, F. &amp; Fittkau, E. J. (1971) Taxonomie und Okologie europaisch verbreiteter Tanytarsus - Arten (Chironomidae, Diptera). Archiv fur Hydrobiologie, Supplement 40, 75 - 200." type="book chapter" year="1971">Reiss &amp; Fittkau 1971</bibRefCitation>
: 98
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BF84651AFFB9A568FF64BBF853D4FF21" lastPageId="6" lastPageNumber="37" pageId="4" pageNumber="35" type="materials_examined">
<paragraph id="F7213691FFB9A56AFF64BBF8526AFA55" blockId="4.[151,1437,1417,2030]" box="[151,1299,1417,1443]" pageId="4" pageNumber="35">
<emphasis id="C5EAEA83FFB9A56AFF64BBF856F2FA55" bold="true" box="[151,395,1417,1443]" pageId="4" pageNumber="35">Material examined.</emphasis>
<collectingCountry id="8F897601FFB9A56AFE61BBF8556EFA55" box="[402,535,1417,1443]" name="Finland" pageId="4" pageNumber="35">FINLAND</collectingCountry>
. Hietanen near Peranka,
<date id="83201051FFB9A56AFCCABBF85490FA55" box="[825,1001,1417,1443]" pageId="4" pageNumber="35" value="2002-08-02">2 August 2002</date>
,
<specimenCount id="E198FD18FFB9A56AFC07BBF85328FA55" box="[1012,1105,1417,1443]" pageId="4" pageNumber="35" type="male">6 males</specimenCount>
, W. Giłka. Inari,
</paragraph>
<paragraph id="F7213691FFB9A56AFF64BBC1556FF92F" blockId="4.[151,1437,1417,2030]" pageId="4" pageNumber="35">
Vuopajanniemi,
<date id="83201051FFB9A56AFEABBBC1568AFA3C" box="[344,499,1456,1482]" pageId="4" pageNumber="35" value="2002-07-28">28 July 2002</date>
,
<specimenCount id="E198FD18FFB9A56AFE0DBBC15529FA3C" box="[510,592,1456,1482]" pageId="4" pageNumber="35" type="male">1 male</specimenCount>
,
<date id="83201051FFB9A56AFDAFBBC15472FA3C" box="[604,779,1456,1482]" pageId="4" pageNumber="35" value="2003-08-04">4 August 2003</date>
,
<specimenCount id="E198FD18FFB9A56AFCE4BBC154FAFA3C" box="[791,899,1456,1482]" pageId="4" pageNumber="35" type="male">12 males</specimenCount>
, W. Giłka. Jänisjärvi near Näätämö,
<date id="83201051FFB9A56AFAC3BBC157ACFA07" pageId="4" pageNumber="35" value="2002-07-26">26 July 2002</date>
,
<specimenCount id="E198FD18FFB9A56AFF13BBA6564BFA07" box="[224,306,1495,1521]" pageId="4" pageNumber="35" type="male">1 male</specimenCount>
, W. Giłka. Korettoja on Utsjoki river,
<date id="83201051FFB9A56AFD01BBA654F5FA07" box="[754,908,1495,1521]" pageId="4" pageNumber="35" value="2003-07-26">26 July 2003</date>
,
<specimenCount id="E198FD18FFB9A56AFC6BBBA65493FA07" box="[920,1002,1495,1521]" pageId="4" pageNumber="35" type="male">1 male</specimenCount>
, W. Giłka. Kotka, Santalahti,
<date id="83201051FFB9A56AFAB4BBA65649F9E1" pageId="4" pageNumber="35" value="2002-08-05">5 August 2002</date>
,
<specimenCount id="E198FD18FFB9A56AFEC9BB8C56EFF9E1" box="[314,406,1533,1559]" pageId="4" pageNumber="35" type="male">6 males</specimenCount>
, W. Giłka. Olhava near Oulu, Bothnian Gulf,
<date id="83201051FFB9A56AFC5ABB8C533BF9E1" box="[937,1090,1533,1559]" pageId="4" pageNumber="35" value="2002-07-13">13 July 2002</date>
,
<specimenCount id="E198FD18FFB9A56AFBBFBB8C53D1F9E1" box="[1100,1192,1533,1559]" pageId="4" pageNumber="35" type="male">4 males</specimenCount>
, W. Giłka. Päijänne,
<date id="83201051FFB9A56AFF64B855564CF9C8" box="[151,309,1572,1598]" pageId="4" pageNumber="35" value="2003-05-27">27 May 2003</date>
,
<specimenCount id="E198FD18FFB9A56AFEB1B85556EDF9C8" box="[322,404,1572,1598]" pageId="4" pageNumber="35" type="male">1 male</specimenCount>
, L. Paasivirta. Supru, Kuosnajärvi,
<date id="83201051FFB9A56AFCC6B85554B6F9C8" box="[821,975,1572,1598]" pageId="4" pageNumber="35" value="2002-07-26">26 July 2002</date>
,
<specimenCount id="E198FD18FFB9A56AFC28B8555341F9C8" box="[987,1080,1572,1598]" pageId="4" pageNumber="35" type="male">6 males</specimenCount>
, W. Giłka. Tervola on Kemijoki river,
<date id="83201051FFB9A56AFEA1B83B5695F992" box="[338,492,1610,1636]" pageId="4" pageNumber="35" value="2002-07-13">13 July 2002</date>
,
<specimenCount id="E198FD18FFB9A56AFE0BB83B5530F992" box="[504,585,1610,1636]" pageId="4" pageNumber="35" type="male">1 male</specimenCount>
,
<date id="83201051FFB9A56AFDA6B83B5596F992" box="[597,751,1610,1636]" pageId="4" pageNumber="35" value="2003-07-22">22 July 2003</date>
,
<specimenCount id="E198FD18FFB9A56AFD08B83B5421F992" box="[763,856,1610,1636]" pageId="4" pageNumber="35" type="male">5 males</specimenCount>
,
<date id="83201051FFB9A56AFC97B83B5486F992" box="[868,1023,1610,1636]" pageId="4" pageNumber="35" value="2006-07-19">19 July 2006</date>
,
<specimenCount id="E198FD18FFB9A56AFBF9B83B531EF992" box="[1034,1127,1610,1636]" pageId="4" pageNumber="35" type="male">4 males</specimenCount>
, W. Giłka. Vesijärvi near Lahti,
<date id="83201051FFB9A56AFF10B8005604F97D" box="[227,381,1649,1675]" pageId="4" pageNumber="35" value="2002-07-11">11 July 2002</date>
,
<specimenCount id="E198FD18FFB9A56AFE7BB80056A3F97D" box="[392,474,1649,1675]" pageId="4" pageNumber="35" type="male">1 male</specimenCount>
, W. Giłka.
<collectingCountry id="8F897601FFB9A56AFDAEB80055A2F97D" box="[605,731,1649,1675]" name="Sweden" pageId="4" pageNumber="35">SWEDEN</collectingCountry>
. Bureå, Skelleftebukten, Bothnian Gulf,
<date id="83201051FFB9A56AFB4BB800521EF97D" box="[1208,1383,1649,1675]" pageId="4" pageNumber="35" value="2003-08-09">9 August 2003</date>
,
<specimenCount id="E198FD18FFB9A56AFA80B80057A7F944" pageId="4" pageNumber="35" type="male">11 males</specimenCount>
, W. Giłka. Ljusnan river,
<quantity id="30669B74FFB9A56AFDF8B8E9552FF944" box="[523,598,1688,1714]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="35" unit="km" value="10.0">10 km</quantity>
E of Sveg,
<date id="83201051FFB9A56AFD2CB8E95400F944" box="[735,889,1688,1714]" pageId="4" pageNumber="35" value="2003-07-19">19 July 2003</date>
,
<specimenCount id="E198FD18FFB9A56AFC76B8E9549BF944" box="[901,994,1688,1714]" pageId="4" pageNumber="35" type="male">2 males</specimenCount>
, W. Giłka. Sikeå, Bothnian Gulf,
<date id="83201051FFB9A56AFA9FB8E95648F92F" pageId="4" pageNumber="35" value="2004-08-20">20 August 2004</date>
,
<specimenCount id="E198FD18FFB9A56AFECFB8CE56E0F92F" box="[316,409,1727,1753]" pageId="4" pageNumber="35" type="male">3 males</specimenCount>
, W. Giłka.
</paragraph>
<paragraph id="F7213691FFB9A56AFF35B89454E6F909" blockId="4.[151,1437,1417,2030]" box="[198,927,1765,1791]" pageId="4" pageNumber="35">
<emphasis id="C5EAEA83FFB9A56AFF35B894569CF909" bold="true" box="[198,485,1765,1791]" pageId="4" pageNumber="35">Diagnostic description.</emphasis>
Adult male (measurements in Table).
</paragraph>
<paragraph id="F7213691FFB9A56AFF35B97D52ECF81C" blockId="4.[151,1437,1417,2030]" pageId="4" pageNumber="35">
Ground colour of thorax, scutellum, haltere, legs and abdomen green to brownish green; antennal pedicel, tentorium, scutal stripes, postnotum and sternum brown to dark brown or black (rarely light brown or orange in freshly emerged specimens); wing membrane pale, greenish, with C, M and radial veins somewhat darker, olive. Frontal tubercles usually absent or present as tiny swellings. Third palpomere shorter than fourth. Wing membrane below veins M and R4+5, including almost entire area under M3+4, Cu1 and An covered with macrotrichia; very short proximal section of R4+5 and 1/3 part of Cu and neighbouring false veins bare (
<figureCitation id="6FA52A14FFB9A56AFAB3B9A152FDF81C" box="[1344,1412,2000,2026]" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1259,1283]" captionTargetBox="[174,1412,196,1223]" captionTargetId="figure@3.[154,1439,158,1256]" captionTargetPageId="3" captionText="FIGURES 1 3. Male wing. 1: Tanytarsus brundini, 2: T. curticornis, 3: T. salmelai sp. n." httpUri="https://zenodo.org/record/189528/files/figure.png" pageId="4" pageNumber="35">Fig. 1</figureCitation>
).
</paragraph>
<caption id="A3E16619FFB8A56BFF64B9715463F8E1" box="[151,794,1792,1816]" httpUri="https://zenodo.org/record/189529/files/figure.png" pageId="5" pageNumber="36" targetBox="[223,1368,188,1743]" targetPageId="5">
<paragraph id="F7213691FFB8A56BFF64B9715463F8E1" blockId="5.[151,794,1792,1816]" box="[151,794,1792,1816]" pageId="5" pageNumber="36">
<emphasis id="C5EAEA83FFB8A56BFF64B971566BF8EE" bold="true" box="[151,274,1792,1816]" pageId="5" pageNumber="36">FIGURE 4</emphasis>
.
<taxonomicName id="309E4D12FFB8A56BFED3B970568CF8EE" box="[288,501,1793,1816]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="salmelai" status="sp. nov.">
<emphasis id="C5EAEA83FFB8A56BFED3B970568CF8EE" box="[288,501,1793,1816]" italics="true" pageId="5" pageNumber="36">Tanytarsus salmelai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DED957F8FFB8A56BFE0FB971554EF8EE" box="[508,567,1792,1816]" pageId="5" pageNumber="36" rank="species">
<emphasis id="C5EAEA83FFB8A56BFE0FB971554EF8EE" bold="true" box="[508,567,1792,1816]" pageId="5" pageNumber="36">sp. n.</emphasis>
</taxonomicNameLabel>
, male. Hypopygium.
</paragraph>
</caption>
<paragraph id="F7213691FFB8A568FF64B93853D4FF21" blockId="5.[151,1437,1865,2003]" lastBlockId="6.[151,1436,152,330]" lastPageId="6" lastPageNumber="37" pageId="5" pageNumber="36">
Gonostylus with slight narrowing distally. Anal tergite with pair of median setae. Basilateral setae absent. Anal point stout, armed with 47 (usually 4 or 5) spinulae, usually with lanceolate apical expansion (
<figureCitation id="6FA52A14FFB8A56BFAB1B91F57D1F85B" captionStart="FIGURES 5 18" captionStartId="6.[151,274,1982,2006]" captionTargetBox="[242,1340,373,1967]" captionTargetId="figure@6.[234,1353,373,1968]" captionTargetPageId="6" captionText="FIGURES 5 18. Variability in hypopygial anal point. 5 9: Tanytarsus brundini, 10 14: T. curticornis, 15 18: T. salmelai sp. n." httpUri="https://zenodo.org/record/189530/files/figure.png" pageId="5" pageNumber="36">Figs 5 9</figureCitation>
). Superior volsella with enlarged anterolateral part and posteromedian lobe, median margin concave. Pearshaped apical lobe of digitus relatively small in comparison with stout superior volsella, bearing narrow conical tip (
<figureCitation id="6FA52A14FFBBA568FED8BEE956FEFF44" box="[299,391,152,178]" captionStart="FIGURES 19 27" captionStartId="8.[151,269,1787,1811]" captionTargetBox="[177,1384,204,1746]" captionTargetId="figure@8.[162,1424,194,1793]" captionTargetPageId="8" captionText="FIGURES 19 27. Hypopygial volsellae. 19 21: superior volsella and digitus, 22 24: median volsella and inner margin of coxite, 25 27: inferior volsella. 19, 22, 25: Tanytarsus brundini; 20, 23, 26: T. curticornis; 21, 24, 27: T. salmelai sp. n." httpUri="https://zenodo.org/record/189531/files/figure.png" pageId="6" pageNumber="37">Fig. 19</figureCitation>
). Inner margin of coxite above median volsella distinctly concave (
<figureCitation id="6FA52A14FFBBA568FB39BEE95251FF44" box="[1226,1320,152,178]" captionStart="FIGURES 19 27" captionStartId="8.[151,269,1787,1811]" captionTargetBox="[177,1384,204,1746]" captionTargetId="figure@8.[162,1424,194,1793]" captionTargetPageId="8" captionText="FIGURES 19 27. Hypopygial volsellae. 19 21: superior volsella and digitus, 22 24: median volsella and inner margin of coxite, 25 27: inferior volsella. 19, 22, 25: Tanytarsus brundini; 20, 23, 26: T. curticornis; 21, 24, 27: T. salmelai sp. n." httpUri="https://zenodo.org/record/189531/files/figure.png" pageId="6" pageNumber="37">Fig. 22</figureCitation>
). Inferior volsella slender, apically rounded, slightly wrinkled on its dorsomedian surface (
<figureCitation id="6FA52A14FFBBA568FBBBBECC53E6FF21" box="[1096,1183,189,215]" captionStart="FIGURES 19 27" captionStartId="8.[151,269,1787,1811]" captionTargetBox="[177,1384,204,1746]" captionTargetId="figure@8.[162,1424,194,1793]" captionTargetPageId="8" captionText="FIGURES 19 27. Hypopygial volsellae. 19 21: superior volsella and digitus, 22 24: median volsella and inner margin of coxite, 25 27: inferior volsella. 19, 22, 25: Tanytarsus brundini; 20, 23, 26: T. curticornis; 21, 24, 27: T. salmelai sp. n." httpUri="https://zenodo.org/record/189531/files/figure.png" pageId="6" pageNumber="37">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BF84651AFFBBA569FF35BE93550AFC65" lastPageId="7" lastPageNumber="38" pageId="6" pageNumber="37" type="discussion">
<paragraph id="F7213691FFBBA568FF35BE935240FEBC" blockId="6.[151,1436,152,330]" pageId="6" pageNumber="37">
<emphasis id="C5EAEA83FFBBA568FF35BE935628FF0A" bold="true" box="[198,337,226,252]" pageId="6" pageNumber="37">Discussion.</emphasis>
<bibRefCitation id="930F4B60FFBBA568FEA8BE935552FF0A" author="Lindeberg" box="[347,555,226,252]" pageId="6" pageNumber="42" refString="Lindeberg, B. (1963) Taxonomy, biology and biometry of Tanytarsus curticornis Kieff. and T. brundini n. sp. (Dipt., Chironomidae). Annales Entomologici Fennici, 29, 118 - 130." type="journal article" year="1963">Lindeberg (1963)</bibRefCitation>
considered adult males of
<taxonomicName id="309E4D12FFBBA568FC82BE935326FF0D" box="[881,1119,226,251]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="brundini">
<emphasis id="C5EAEA83FFBBA568FC82BE935326FF0D" box="[881,1119,226,251]" italics="true" pageId="6" pageNumber="37">Tanytarsus brundini</emphasis>
</taxonomicName>
and
<taxonomicName id="309E4D12FFBBA568FB6FBE935244FF0D" box="[1180,1341,226,251]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="curticornis">
<emphasis id="C5EAEA83FFBBA568FB6FBE935244FF0D" box="[1180,1341,226,251]" italics="true" pageId="6" pageNumber="37">T. curticornis</emphasis>
</taxonomicName>
as very similar based on their hypopygial structure. The male of
<taxonomicName id="309E4D12FFBBA568FCCFBF785353FED4" box="[828,1066,265,290]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="brundini">
<emphasis id="C5EAEA83FFBBA568FCCFBF785353FED4" box="[828,1066,265,290]" italics="true" pageId="6" pageNumber="37">Tanytarsus brundini</emphasis>
</taxonomicName>
(1) can be easily distinguished from those of
<taxonomicName id="309E4D12FFBBA568FECFBF4156A3FEBF" box="[316,474,304,329]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="curticornis">
<emphasis id="C5EAEA83FFBBA568FECFBF4156A3FEBF" box="[316,474,304,329]" italics="true" pageId="6" pageNumber="37">T. curticornis</emphasis>
</taxonomicName>
(2) and
<taxonomicName id="309E4D12FFBBA568FDCFBF4155C5FEBF" box="[572,700,304,329]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="salmelai">
<emphasis id="C5EAEA83FFBBA568FDCFBF4155C5FEBF" box="[572,700,304,329]" italics="true" pageId="6" pageNumber="37">T. salmelai</emphasis>
</taxonomicName>
(3) by the combination of characters presented below.
</paragraph>
<caption id="A3E16619FFBBA568FF64B9CF564DF800" httpUri="https://zenodo.org/record/189530/files/figure.png" pageId="6" pageNumber="37" targetBox="[242,1340,373,1967]" targetPageId="6">
<paragraph id="F7213691FFBBA568FF64B9CF564DF800" blockId="6.[151,1435,1982,2038]" pageId="6" pageNumber="37">
<emphasis id="C5EAEA83FFBBA568FF64B9CF562BF820" bold="true" box="[151,338,1982,2006]" pageId="6" pageNumber="37">FIGURES 518</emphasis>
. Variability in hypopygial anal point. 59:
<taxonomicName id="309E4D12FFBBA568FCB3B9CF535DF823" box="[832,1060,1982,2005]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="brundini">
<emphasis id="C5EAEA83FFBBA568FCB3B9CF535DF823" box="[832,1060,1982,2005]" italics="true" pageId="6" pageNumber="37">Tanytarsus brundini</emphasis>
</taxonomicName>
, 1014:
<taxonomicName id="309E4D12FFBBA568FB7AB9CF5259F823" box="[1161,1312,1982,2005]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="curticornis">
<emphasis id="C5EAEA83FFBBA568FB7AB9CF5259F823" box="[1161,1312,1982,2005]" italics="true" pageId="6" pageNumber="37">T. curticornis</emphasis>
</taxonomicName>
, 1518:
<taxonomicName id="309E4D12FFBBA568FA7BB9CF5788F803" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="salmelai" status="sp. nov.">
<emphasis id="C5EAEA83FFBBA568FA7BB9CF5788F803" italics="true" pageId="6" pageNumber="37">T. salmelai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DED957F8FFBBA568FF0BB9AF564DF800" box="[248,308,2014,2038]" pageId="6" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="F7213691FFBAA569FF35BEE9578BFE11" blockId="7.[151,1437,152,915]" pageId="7" pageNumber="38">
At least one pair of median setae on the anal tergite present (1, 3) or median setae absent (2). Basilateral setae absent (1, 3) or single strong seta on each side of the anal tergite present (2). Anal point slightly lanceolate, usually with 4 or 5 spinulae (1), anal point broadly rounded or slightly cut apically, usually with 7 or 8 spinulae (2) or anal point distinctly narrowed and strongly elongated, usually with 4 spinulae (3) (
<figureCitation id="6FA52A14FFBAA569FAB0BF7D5622FEBA" captionStart="FIGURES 5 18" captionStartId="6.[151,274,1982,2006]" captionTargetBox="[242,1340,373,1967]" captionTargetId="figure@6.[234,1353,373,1968]" captionTargetPageId="6" captionText="FIGURES 5 18. Variability in hypopygial anal point. 5 9: Tanytarsus brundini, 10 14: T. curticornis, 15 18: T. salmelai sp. n." httpUri="https://zenodo.org/record/189530/files/figure.png" pageId="7" pageNumber="38">Figs 5 9, 1014, 1518</figureCitation>
respectively). Superior volsella concave (1) or transversally cut (2, 3) (
<figureCitation id="6FA52A14FFBAA569FB48BF435231FEBA" box="[1211,1352,306,332]" captionStart="FIGURES 19 27" captionStartId="8.[151,269,1787,1811]" captionTargetBox="[177,1384,204,1746]" captionTargetId="figure@8.[162,1424,194,1793]" captionTargetPageId="8" captionText="FIGURES 19 27. Hypopygial volsellae. 19 21: superior volsella and digitus, 22 24: median volsella and inner margin of coxite, 25 27: inferior volsella. 19, 22, 25: Tanytarsus brundini; 20, 23, 26: T. curticornis; 21, 24, 27: T. salmelai sp. n." httpUri="https://zenodo.org/record/189531/files/figure.png" pageId="7" pageNumber="38">Figs 1921</figureCitation>
). Pearshaped lobe of digitus with narrow conical tip, relatively small in comparison with superior volsella (1, 3) or pear-shaped lobe of digitus roundish, relatively stout in comparison with superior volsella (2) (
<figureCitation id="6FA52A14FFBAA569FAF7BFF152F6FE6C" box="[1284,1423,384,410]" captionStart="FIGURES 19 27" captionStartId="8.[151,269,1787,1811]" captionTargetBox="[177,1384,204,1746]" captionTargetId="figure@8.[162,1424,194,1793]" captionTargetPageId="8" captionText="FIGURES 19 27. Hypopygial volsellae. 19 21: superior volsella and digitus, 22 24: median volsella and inner margin of coxite, 25 27: inferior volsella. 19, 22, 25: Tanytarsus brundini; 20, 23, 26: T. curticornis; 21, 24, 27: T. salmelai sp. n." httpUri="https://zenodo.org/record/189531/files/figure.png" pageId="7" pageNumber="38">Figs 1921</figureCitation>
). Inner margin of coxite above median volsella distinctly concave (1), straight (2) or slightly concave (3) (
<figureCitation id="6FA52A14FFBAA569FA9BBFD7579CFE11" captionStart="FIGURES 19 27" captionStartId="8.[151,269,1787,1811]" captionTargetBox="[177,1384,204,1746]" captionTargetId="figure@8.[162,1424,194,1793]" captionTargetPageId="8" captionText="FIGURES 19 27. Hypopygial volsellae. 19 21: superior volsella and digitus, 22 24: median volsella and inner margin of coxite, 25 27: inferior volsella. 19, 22, 25: Tanytarsus brundini; 20, 23, 26: T. curticornis; 21, 24, 27: T. salmelai sp. n." httpUri="https://zenodo.org/record/189531/files/figure.png" pageId="7" pageNumber="38">Figs 2224</figureCitation>
).
</paragraph>
<paragraph id="F7213691FFBAA569FF36BF855607FD0F" blockId="7.[151,1437,152,915]" pageId="7" pageNumber="38">
The shape of inferior volsellae of the species compared is also distinct when the structures are examined in properly mounted specimens (
<figureCitation id="6FA52A14FFBAA569FDCFBC6B55AAFDC2" box="[572,723,538,564]" captionStart="FIGURES 19 27" captionStartId="8.[151,269,1787,1811]" captionTargetBox="[177,1384,204,1746]" captionTargetId="figure@8.[162,1424,194,1793]" captionTargetPageId="8" captionText="FIGURES 19 27. Hypopygial volsellae. 19 21: superior volsella and digitus, 22 24: median volsella and inner margin of coxite, 25 27: inferior volsella. 19, 22, 25: Tanytarsus brundini; 20, 23, 26: T. curticornis; 21, 24, 27: T. salmelai sp. n." httpUri="https://zenodo.org/record/189531/files/figure.png" pageId="7" pageNumber="38">Figs 2527</figureCitation>
). The wing of
<taxonomicName id="309E4D12FFBAA569FC63BC6B53E9FDC5" box="[912,1168,538,563]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="salmelai">
<emphasis id="C5EAEA83FFBAA569FC63BC6B53E9FDC5" box="[912,1168,538,563]" italics="true" pageId="7" pageNumber="38">Tanytarsus salmelai</emphasis>
</taxonomicName>
has a distinct shape, colouration and chaetotaxy (
<figureCitation id="6FA52A14FFBAA569FE17BC305552FDAD" box="[484,555,577,603]" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1259,1283]" captionTargetBox="[174,1412,196,1223]" captionTargetId="figure@3.[154,1439,158,1256]" captionTargetPageId="3" captionText="FIGURES 1 3. Male wing. 1: Tanytarsus brundini, 2: T. curticornis, 3: T. salmelai sp. n." httpUri="https://zenodo.org/record/189528/files/figure.png" pageId="7" pageNumber="38">Fig. 3</figureCitation>
), whereas the wings of
<taxonomicName id="309E4D12FFBAA569FCB2BC3054BAFDAC" box="[833,963,577,602]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="brundini">
<emphasis id="C5EAEA83FFBAA569FCB2BC3054BAFDAC" box="[833,963,577,602]" italics="true" pageId="7" pageNumber="38">T. brundini</emphasis>
</taxonomicName>
and
<taxonomicName id="309E4D12FFBAA569FC0FBC3053EEFDAC" box="[1020,1175,577,602]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="curticornis">
<emphasis id="C5EAEA83FFBAA569FC0FBC3053EEFDAC" box="[1020,1175,577,602]" italics="true" pageId="7" pageNumber="38">T. curticornis</emphasis>
</taxonomicName>
are very similar (
<figureCitation id="6FA52A14FFBAA569FA9BBC30578BFD74" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1259,1283]" captionTargetBox="[174,1412,196,1223]" captionTargetId="figure@3.[154,1439,158,1256]" captionTargetPageId="3" captionText="FIGURES 1 3. Male wing. 1: Tanytarsus brundini, 2: T. curticornis, 3: T. salmelai sp. n." httpUri="https://zenodo.org/record/189528/files/figure.png" pageId="7" pageNumber="38">Figs 1 and 2</figureCitation>
). Slight differences can be observed in the chaetotaxy, i.e. the number of macrotrichia in cells under M3+4, Cu1, An and on veins R4+5, Cu and neighbouring false veins (higher number in
<taxonomicName id="309E4D12FFBAA569FB3ABCFE522CFD5E" box="[1225,1365,655,680]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="brundini">
<emphasis id="C5EAEA83FFBAA569FB3ABCFE522CFD5E" box="[1225,1365,655,680]" italics="true" pageId="7" pageNumber="38">T. brundini</emphasis>
</taxonomicName>
). For differences in measurable characters of males
<taxonomicName id="309E4D12FFBAA569FD36BCC95344FD24" authority="Lindeberg" authorityName="Lindeberg" box="[709,1085,696,722]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="brundini">
<emphasis id="C5EAEA83FFBAA569FD36BCC954CEFD27" box="[709,951,696,721]" italics="true" pageId="7" pageNumber="38">Tanytarsus brundini</emphasis>
Lindeberg
</taxonomicName>
,
<taxonomicName id="309E4D12FFBAA569FBB9BCC95233FD24" authority="Kieffer" authorityName="Kieffer" box="[1098,1354,696,722]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="curticornis">
<emphasis id="C5EAEA83FFBAA569FBB9BCC95395FD27" box="[1098,1260,696,721]" italics="true" pageId="7" pageNumber="38">T. curticornis</emphasis>
Kieffer
</taxonomicName>
and
<taxonomicName id="309E4D12FFBAA569FA75BCC95783FD0E" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="salmelai">
<emphasis id="C5EAEA83FFBAA569FA75BCC95783FD0E" italics="true" pageId="7" pageNumber="38">T. salmelai</emphasis>
</taxonomicName>
, see Table.
</paragraph>
<paragraph id="F7213691FFBAA569FF36BD74550AFC65" blockId="7.[151,1437,152,915]" pageId="7" pageNumber="38">
In addition, Lindeberg (l.c.) based his separation of
<taxonomicName id="309E4D12FFBAA569FC8BBD775306FCE9" box="[888,1151,774,799]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="brundini">
<emphasis id="C5EAEA83FFBAA569FC8BBD775306FCE9" box="[888,1151,774,799]" italics="true" pageId="7" pageNumber="38">Tanytarsus brundini</emphasis>
</taxonomicName>
and
<taxonomicName id="309E4D12FFBAA569FB32BD77520AFCE9" box="[1217,1395,774,799]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="38" phylum="Arthropoda" rank="species" species="curticornis">
<emphasis id="C5EAEA83FFBAA569FB32BD77520AFCE9" box="[1217,1395,774,799]" italics="true" pageId="7" pageNumber="38">T. curticornis</emphasis>
</taxonomicName>
on observations of their swarming behaviour. We confirm that swarms containing both species appear rarely. Only one sample collected at Vesijärvi contained specimens of both species flying together in a large swarm consisting of several tanytarsine species.
</paragraph>
</subSubSection>
</treatment>
</document>