186 lines
28 KiB
XML
186 lines
28 KiB
XML
<document id="9867C9A0B755113BAC9F37711126E611" ID-CLB-Dataset="110554" ID-DOI="10.1649/0010-065X-69.mo4.167" ID-GBIF-Dataset="459316b8-ecf9-4bcb-a14b-100658d73012" ID-ISSN="1938-4394" ID-Zenodo-Dep="4911739" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623095475750" checkinUser="carolina" docAuthor="Flores, Gustavo E. & Aballay, Fernando H." docDate="2015" docId="182B9F50FFEDFFDDFF2AFB4CFB58FB19" docLanguage="en" docName="TheColeopBull.69.mo14.167-179.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 14) 69" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Vaniosus Kulzer 1956" docType="treatment" docVersion="9" lastPageNumber="172" masterDocId="E412E728FFE8FFD8FFDCFFAFFFCCFFEA" masterDocTitle="Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson" masterLastPageNumber="179" masterPageNumber="167" pageNumber="172" updateTime="1699002214655" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="898A2C7D23FE75425AA2D763BF0FB8F8">
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<mods:title id="40C288EEA8CBF096907C1937C61EA9CB">Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson</mods:title>
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<mods:name id="C1D013E486EBCE4B2AA1D8C25EE63CBD" type="personal">
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<mods:roleTerm id="848D72006F8AD488DA81E473FCECAA6A">Author</mods:roleTerm>
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<mods:namePart id="979A5E17AB7B9373E9C7CF147D8565C5">Flores, Gustavo E.</mods:namePart>
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<mods:affiliation id="E08DE89B4D0EE78B12833DB9AF9602FE">CONICET, Laboratorio de Entomología Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT-CONICET Mendoza) Casilla de correo 507, 5500 Mendoza, ARGENTINA</mods:affiliation>
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<mods:name id="D0D7B600C092C7643BE4489863EE4BB2" type="personal">
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<mods:roleTerm id="CFAD4F45BFBDE5789DACE0C67C01AF97">Author</mods:roleTerm>
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<mods:namePart id="6131D50C3516F5B2031BEC6660A3FCC9">Aballay, Fernando H.</mods:namePart>
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<mods:affiliation id="97481BC59A097DFEE4CD06D8F58CA33F">CONICET, Laboratorio de Entomología Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT-CONICET Mendoza) Casilla de correo 507, 5500 Mendoza, ARGENTINA</mods:affiliation>
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<mods:typeOfResource id="28330FF5D7C46EB045557096579C7ED0">text</mods:typeOfResource>
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<mods:title id="39D142091344507113CAE047E9D9F540">The Coleopterists Bulletin</mods:title>
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<mods:part id="A60F59CC5C5276F280BA41EF5F0F28FC">
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<mods:date id="876B5087E1F2FDAD583FFD87FF3738C7">2015</mods:date>
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<mods:detail id="CFF5A50D8EC0F8FC58F37394EBE6CAD1" type="series">
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<mods:title id="8CF21947180FF6FBCD78EB25EF915C62">mo 14</mods:title>
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<mods:number id="815919E034C31260FB1171EC4C97174A">2015-12-18</mods:number>
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<mods:number id="0E289F514FE79C397CDC4D34529EEF81">69</mods:number>
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<mods:start id="CB1D644AFD171365B8162B993BCAC052">167</mods:start>
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<mods:identifier id="88FA6FD812DC00CA273822059B193F81" type="CLB-Dataset">110554</mods:identifier>
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<mods:identifier id="F394F1AAFF7781368B41749C074C6AB4" type="DOI">10.1649/0010-065X-69.mo4.167</mods:identifier>
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<treatment id="182B9F50FFEDFFDDFF2AFB4CFB58FB19" ID-DOI="http://doi.org/10.5281/zenodo.6950036" ID-GBIF-Taxon="198139692" ID-Zenodo-Dep="6950036" LSID="urn:lsid:plazi:treatment:182B9F50FFEDFFDDFF2AFB4CFB58FB19" httpUri="http://treatment.plazi.org/id/182B9F50FFEDFFDDFF2AFB4CFB58FB19" lastPageNumber="172" pageId="5" pageNumber="172">
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<subSubSection id="D8987DCDFFEDFFDDFF2AFB4CFDECFB12" box="[246,544,1251,1272]" pageId="5" pageNumber="172" type="nomenclature">
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<paragraph id="903D2E46FFEDFFDDFF2AFB4CFDECFB12" blockId="5.[246,544,1251,1299]" box="[246,544,1251,1272]" pageId="5" pageNumber="172">
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<heading id="CB75992AFFEDFFDDFF2AFB4CFDECFB12" box="[246,544,1251,1272]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="172" reason="2">
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<emphasis id="A2F6F254FFEDFFDDFF2AFB4CFDECFB12" bold="true" box="[246,544,1251,1272]" pageId="5" pageNumber="172">
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Genus
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<taxonomicName id="578255C5FFEDFFDDFEE2FB4CFDECFB12" authority="Kulzer, 1956" authorityName="Kulzer" authorityYear="1956" box="[318,544,1251,1272]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="genus">
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<emphasis id="A2F6F254FFEDFFDDFEE2FB4CFE5BFB12" bold="true" box="[318,407,1251,1272]" italics="true" pageId="5" pageNumber="172">Vaniosus</emphasis>
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<bibRefCitation id="F41353B7FFEDFFDDFE43FB4CFDECFB12" author="Kulzer" box="[415,544,1251,1272]" firstAuthor="Kulzer" pageId="5" pageNumber="172" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer, 1956</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D8987DCDFFEDFFDDFEDEFB51FDDFFAF9" box="[258,531,1277,1299]" pageId="5" pageNumber="172" type="description">
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<paragraph id="903D2E46FFEDFFDDFEDEFB51FDDFFAF9" blockId="5.[246,544,1251,1299]" box="[258,531,1277,1299]" pageId="5" pageNumber="172">
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(
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<figureCitation id="08B932C3FFEDFFDDFED5FB51FEA4FAF9" box="[265,360,1277,1299]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Figs. 2–4</figureCitation>
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,
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<figureCitation id="08B932C3FFEDFFDDFEA8FB51FE57FAF9" box="[372,411,1278,1299]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">6, 8</figureCitation>
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,
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<figureCitation id="08B932C3FFEDFFDDFE74FB51FE0EFAF9" box="[424,450,1278,1299]" captionStart="Figs" captionStartId="4.[197,237,1686,1705]" captionTargetBox="[173,659,923,1663]" captionTargetId="figure-306@4.[172,659,923,1663]" captionText="Figs. 9–10. Antennomeres 9–11.9) Thinobatis rufipes rufipes; 10) Vaniosus profana." figureDoi="http://doi.org/10.5281/zenodo.6941556" httpUri="https://zenodo.org/record/6941556/files/figure.png" pageId="5" pageNumber="172">10</figureCitation>
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, 12, 21)
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</paragraph>
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</subSubSection>
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<subSubSection id="D8987DCDFFEDFFDDFF4BFA86FE52FAD7" box="[151,414,1320,1342]" pageId="5" pageNumber="172" type="reference_group">
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<paragraph id="903D2E46FFEDFFDDFF4BFA86FE52FAD7" blockId="5.[151,638,1320,1368]" box="[151,414,1320,1342]" pageId="5" pageNumber="172">
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<treatmentCitationGroup id="B0920968FFEDFFDDFF4BFA86FE52FAD7" box="[151,414,1320,1342]" pageId="5" pageNumber="172">
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<treatmentCitation id="11230857FFEDFFDDFF4BFA86FE56FAD7" author="Kulzer" box="[151,410,1320,1342]" page="896" pageId="5" pageNumber="172" year="1956">
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<taxonomicName id="578255C5FFEDFFDDFF4BFA86FE56FAD7" authority="Kulzer 1956: 896" authorityName="Kulzer" authorityPageNumber="896" authorityYear="1956" box="[151,410,1320,1342]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="genus">
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<emphasis id="A2F6F254FFEDFFDDFF4BFA86FF20FAD4" box="[151,236,1321,1342]" italics="true" pageId="5" pageNumber="172">Vaniosus</emphasis>
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<bibRefCitation id="F41353B7FFEDFFDDFF2FFA87FE56FAD7" author="Kulzer" box="[243,410,1320,1341]" firstAuthor="Kulzer" page="896" pageId="5" pageNumber="172" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer 1956: 896</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="D8987DCDFFEDFFDDFE78FA87FDE4FAB2" pageId="5" pageNumber="172" type="type_taxon">
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<paragraph id="903D2E46FFEDFFDDFE78FA87FDE4FAB2" blockId="5.[151,638,1320,1368]" pageId="5" pageNumber="172">
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<typeStatus id="4F3990E4FFEDFFDDFE78FA87FE1FFAD7" box="[420,467,1320,1341]" pageId="5" pageNumber="172">Type</typeStatus>
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species:
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<taxonomicName id="578255C5FFEDFFDDFDF4FA86FE54FAB2" authority="Kulzer, 1956" authorityName="Kulzer" authorityYear="1956" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="paradoxus">
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<emphasis id="A2F6F254FFEDFFDDFDF4FA86FED8FAB2" italics="true" pageId="5" pageNumber="172">Vaniosus paradoxus</emphasis>
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<bibRefCitation id="F41353B7FFEDFFDDFEC0FAECFE54FAB2" author="Kulzer" box="[284,408,1347,1368]" firstAuthor="Kulzer" pageId="5" pageNumber="172" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer, 1956</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="D8987DCDFFEDFFDDFF73FAC1FBD2FD00" pageId="5" pageNumber="172" type="description">
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||
<paragraph id="903D2E46FFEDFFDDFF73FAC1FC77FDDA" blockId="5.[151,638,1390,1729]" lastBlockId="5.[685,1173,194,800]" pageId="5" pageNumber="172">
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<emphasis id="A2F6F254FFEDFFDDFF73FAC1FEE7FA69" bold="true" box="[175,299,1390,1411]" pageId="5" pageNumber="172">Description.</emphasis>
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In addition to the characters stated above: Mandibles finer, acute, base 1.5X wider than the apex, both mandibles with a dorsal cusp on basal half longer on the right than on the left mandible (
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<figureCitation id="08B932C3FFEDFFDDFEDDFA77FEF3FA07" box="[257,319,1496,1517]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 6</figureCitation>
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), both mandibles with a ventral fossa (
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<figureCitation id="08B932C3FFEDFFDDFF07FA5DFED5F9ED" box="[219,281,1522,1543]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 8</figureCitation>
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); labrum exposed, partially sclerotized; clypeus short, not covering the labrum, anterior margin triangular nearly at same level with antennal insertions, armed with a row of teeth directed downwards, in the apex with a large tooth prolonged anterad (
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<figureCitation id="08B932C3FFEDFFDDFEBCF9D8FE6CF966" box="[352,416,1655,1676]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 6</figureCitation>
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); prementum entirely membranous, nearly concealed beneath mentum; anterior margin of mentum straight or slightly concave (
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<figureCitation id="08B932C3FFEDFFDDFD37FF6DFCE5FF3D" box="[747,809,194,215]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 8</figureCitation>
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); eyes small, protruding out, upper margin lacking carina; length of epicanthus 2.5x the length of eye (
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<figureCitation id="08B932C3FFEDFFDDFC86FF58FC58FEE6" box="[858,916,247,268]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 6</figureCitation>
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); antennae surpassing posterior margin of pronotum, antennomere 1 sticks out distal half of antennal insertion (
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<figureCitation id="08B932C3FFEDFFDDFC33FE83FBE7FEAB" box="[1007,1067,300,321]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 8</figureCitation>
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), antennomeres 9 and 10 cylindrical, not expanded outwards, longer than wide, antennomere 11 equal in length to 10 (
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<figureCitation id="08B932C3FFEDFFDDFC8CFED3FC56FE7B" box="[848,922,380,401]" captionStart="Figs" captionStartId="4.[197,237,1686,1705]" captionTargetBox="[173,659,923,1663]" captionTargetId="figure-306@4.[172,659,923,1663]" captionText="Figs. 9–10. Antennomeres 9–11.9) Thinobatis rufipes rufipes; 10) Vaniosus profana." figureDoi="http://doi.org/10.5281/zenodo.6941556" httpUri="https://zenodo.org/record/6941556/files/figure.png" pageId="5" pageNumber="172">Fig. 10</figureCitation>
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). Pronotum with anterior angles obtuse, slopeing towards venter (
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<figureCitation id="08B932C3FFEDFFDDFBF0FE38FB44FE46" box="[1068,1160,406,428]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Figs. 2–4</figureCitation>
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). Metasternum twice the length of mesosternum. Metacoxae separated by less than half metacoxal width; metacoxal width 3X metacoxal length; distance between meso– and metacoxae twice the metacoxal length (
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||
<figureCitation id="08B932C3FFEDFFDDFCB9FDB4FC63FDDA" box="[869,943,539,560]" captionStart="Figs" captionStartId="4.[731,771,973,992]" captionTargetBox="[707,1193,194,950]" captionTargetId="figure-325@4.[707,1194,193,950]" captionTargetPageId="4" captionText="Figs. 11–12. Meso– and metasterna. 11) Thinobatis rufipes rufipes; 12) Vaniosus profana." figureDoi="http://doi.org/10.5281/zenodo.4911745" httpUri="https://zenodo.org/record/4911745/files/figure.png" pageId="5" pageNumber="172">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="903D2E46FFEDFFDDFD19FD99FBD2FD00" blockId="5.[685,1173,194,800]" pageId="5" pageNumber="172">
|
||
Female genitalia (
|
||
<figureCitation id="08B932C3FFEDFFDDFCA1FD99FC06FDA1" box="[893,970,566,587]" captionStart="Figs" captionStartId="9.[175,215,1662,1681]" captionTargetBox="[151,1171,1074,1639]" captionTargetId="figure-36@9.[151,1173,1074,1640]" captionTargetPageId="9" captionText="Figs. 21–22. Ovipositor in ventral view, spiculum and internal female reproductive tract. 21) Vaniosus profana; 22) Achanius (Ambigatus) rufonitens. Abbreviations: c = coxite; o = oviduct; p = paraproct; s = spermatheca; sag = spermathecal accessory gland; sp = spiculum; v = vagina. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911753" httpUri="https://zenodo.org/record/4911753/files/figure.png" pageId="5" pageNumber="172">Fig. 21</figureCitation>
|
||
). Spiculum without arms. Paraprocts long (2.0 <P/C ≤ 3.0), basal lobe of coxite not extended over paraproct. Vagina saccate. Spermathecal accesory gland longer than vagina, with annulate duct. Spermatheca single, compact, locular, at the end of vagina, near the duct of spermathecal accessory gland.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D8987DCDFFEDFFDDFD19FD5FFB4FFC8B" pageId="5" pageNumber="172" type="discussion">
|
||
<paragraph id="903D2E46FFEDFFDDFD19FD5FFC13FCF5" blockId="5.[685,1173,194,800]" pageId="5" pageNumber="172">
|
||
<emphasis id="A2F6F254FFEDFFDDFD19FD5FFCB9FCEF" bold="true" box="[709,885,752,773]" pageId="5" pageNumber="172">Species included.</emphasis>
|
||
<taxonomicName id="578255C5FFEDFFDDFC5EFD5FFB58FCEF" authority="Kulzer" authorityName="Kulzer" authorityYear="1956" box="[898,1172,752,773]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="paradoxus">
|
||
<emphasis id="A2F6F254FFEDFFDDFC5EFD5FFB86FCEF" box="[898,1098,752,773]" italics="true" pageId="5" pageNumber="172">Vaniosus paradoxus</emphasis>
|
||
Kulzer
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="578255C5FFEDFFDDFD04FCA4FC17FCF5" authority="(Kulzer)" baseAuthorityName="Kulzer" box="[728,987,778,800]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="profana">
|
||
<emphasis id="A2F6F254FFEDFFDDFD04FCA4FC4DFCCA" box="[728,897,779,800]" italics="true" pageId="5" pageNumber="172">Vaniosus profana</emphasis>
|
||
(Kulzer)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="903D2E46FFEDFFDDFD62FCE4FB4FFC8B" blockId="5.[702,1155,843,867]" box="[702,1155,843,867]" pageId="5" pageNumber="172">
|
||
<emphasis id="A2F6F254FFEDFFDDFD62FCE4FC71FC8B" bold="true" box="[702,957,843,866]" pageId="5" pageNumber="172">KEY TO THE SPECIES OF</emphasis>
|
||
<taxonomicName id="578255C5FFEDFFDDFC19FCE4FB4FFC8B" authority="KULZER" authorityName="Kulzer, Evaniosomini. Doyen" authorityYear="1994" box="[965,1155,843,867]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A2F6F254FFEDFFDDFC19FCE4FBE9FC8A" bold="true" box="[965,1061,843,867]" italics="true" pageId="5" pageNumber="172">VANIOSUS</emphasis>
|
||
<emphasis id="A2F6F254FFEDFFDDFBF3FCE4FB4FFC8B" bold="true" box="[1071,1155,843,866]" pageId="5" pageNumber="172">KULZER</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D8987DCDFFEDFFDDFD71FCD2FB58FB19" pageId="5" pageNumber="172" type="key">
|
||
<keyStep id="2B7636E3FFEDFFDDFD71FCD2FB58FB19" pageId="5" pageNumber="172">
|
||
<key id="E4137977FFEDFFDDFD71FCD2FB58FB19" pageId="5" pageNumber="172">
|
||
<paragraph id="903D2E46FFEDFFDDFD71FCD2FB5FFBDB" blockId="5.[685,1172,893,1267]" pageId="5" pageNumber="172">
|
||
<keyLead id="2B738D73FFEDFFDDFD71FCD2FB5FFBDB" pageId="5" pageNumber="172">
|
||
1. Head widest at epicanthus; antennomere 1 longer than 3, antennomere 2 longer than 3; posterior angles of pronotum very protruding, pointed outwards (
|
||
<figureCitation id="08B932C3FFEDFFDDFC7DFC63FC2EFC0B" box="[929,994,972,993]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Fig. 4</figureCitation>
|
||
); humeri not elevated; hind wings absent; first tarsomere of hind legs longer than 2–4 combined (
|
||
<figureCitation id="08B932C3FFEDFFDDFB8DFBADFB47FBFD" box="[1105,1163,1026,1047]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Fig. 3</figureCitation>
|
||
) .........................
|
||
<taxonomicName id="578255C5FFEDFFDDFCA2FBB3FB5FFBDB" authority="Kulzer" authorityName="Kulzer" authorityYear="1956" box="[894,1171,1052,1073]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="paradoxus">
|
||
<emphasis id="A2F6F254FFEDFFDDFCA2FBB3FB5FFBDB" bold="true" box="[894,1171,1052,1073]" pageId="5" pageNumber="172">
|
||
<emphasis id="A2F6F254FFEDFFDDFCA2FBB3FB8AFBDB" bold="true" box="[894,1094,1052,1073]" italics="true" pageId="5" pageNumber="172">Vaniosus paradoxus</emphasis>
|
||
Kulzer
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="903D2E46FFEDFFDDFD71FB90FB58FB19" blockId="5.[685,1172,893,1267]" pageId="5" pageNumber="172">
|
||
<keyLead id="2B738D73FFEDFFDDFD71FB90FB58FB19" pageId="5" pageNumber="172">
|
||
1′. Head widest at eyes; antennomere 1 as long as 3, antennomere 2 shorter than 3 (
|
||
<figureCitation id="08B932C3FFEDFFDDFB96FBF6FB4AFB84" box="[1098,1158,1113,1134]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 8</figureCitation>
|
||
); posterior angles of pronotum obtuse, not protruding; humeri elevated; hind wings present, reduced, as long as elytra, not folded; first tarsomere of hind legs as long as 2–3 combined (
|
||
<figureCitation id="08B932C3FFEDFFDDFCF8FB71FC97FB19" box="[804,859,1246,1267]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Fig. 2</figureCitation>
|
||
).......
|
||
<taxonomicName id="578255C5FFEDFFDDFC52FB71FB58FB19" authority="(Kulzer)" baseAuthorityName="Kulzer" box="[910,1172,1246,1267]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="profana">
|
||
<emphasis id="A2F6F254FFEDFFDDFC52FB71FB58FB19" bold="true" box="[910,1172,1246,1267]" pageId="5" pageNumber="172">
|
||
<emphasis id="A2F6F254FFEDFFDDFC52FB71FBFAFB19" bold="true" box="[910,1078,1246,1267]" italics="true" pageId="5" pageNumber="172">Vaniosus profana</emphasis>
|
||
(Kulzer)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |