treatments-xml/data/E7/6C/C5/E76CC555D018FFE6FF615C7BF1A9FA3F.xml

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<mods:title id="55E779C5F3610D1570BD55F94EF94F07">Taxonomic revision of Mesamphiagrion Kennedy, 1920 from Colombia (Odonata: Coenagrionidae), with the description of four new species</mods:title>
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<mods:namePart id="C62F4ECB60BAA9851C938396195F7ABC">Bota-Sierra, Cornelio Andrés</mods:namePart>
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<mods:namePart id="CCADE608B541ABEF3CB47CCF2DA35976">Echeverri, Martha Isabel Wolff</mods:namePart>
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<mods:date id="38646C1F1BDE3B041D8581906AD7941B">2013</mods:date>
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<subSubSection id="27DF27C8D018FFEBFF615C7BF112FB12" pageId="5" pageNumber="406" type="nomenclature">
<paragraph id="6F7A7443D018FFEBFF615C7BF065FBD4" blockId="5.[151,1436,1086,1182]" box="[151,827,1086,1112]" pageId="5" pageNumber="406">
<heading id="3432C32FD018FFEBFF615C7BF065FBD4" bold="true" box="[151,827,1086,1112]" fontSize="11" level="1" pageId="5" pageNumber="406" reason="1">
<emphasis id="5DB1A851D018FFEBFF615C7BF065FBD4" bold="true" box="[151,827,1086,1112]" pageId="5" pageNumber="406">
<taxonomicName id="A8C50FC0D018FFEBFF615C7BF188FBD4" ID-CoL="9X3JK" authority="Bota-Sierra" authorityName="Bota-Sierra" box="[151,726,1086,1112]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="gaudiimontanum" status="sp. nov.">
<emphasis id="5DB1A851D018FFEBFF615C7BF163FBD4" bold="true" box="[151,573,1086,1112]" italics="true" pageId="5" pageNumber="406">Mesamphiagrion gaudiimontanum</emphasis>
Bota-Sierra
</taxonomicName>
<taxonomicNameLabel id="4682152AD018FFEBFD285C7BF065FBD4" box="[734,827,1086,1112]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6F7A7443D018FFEBFF615C24F112FB12" blockId="5.[151,1436,1086,1182]" pageId="5" pageNumber="406">
<figureCitation id="F7FE68C6D018FFEBFF615C24F3BDFBF6" box="[151,227,1121,1146]" captionStart="FIGURE 3" captionStartId="17.[151,250,1955,1977]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@17.[204,1382,193,1921]" captionTargetPageId="17" captionText="FIGURE 3. Mesamphiagrion santainense (Antioquia, Belmira): (a) Male holotype; (b) Female paratype. M. gaudiimontanum (Antioquia, Belmira) Mature males: (c) Dorsal view; (d) Lateral view." httpUri="https://zenodo.org/record/246831/files/figure.png" pageId="5" pageNumber="406">Figs. 3</figureCitation>
c–d (habitus ♂), 5a–c (ontogenic changes ♂), 6a–d (ontogenic changes ♀), 7a–f (diagnostic traits), 13a–b (S7– 
<specimenCount id="79C3BFCAD018FFEBFF3A5CC3F3B6FB12" box="[204,232,1158,1182]" pageId="5" pageNumber="406" type="generic">10</specimenCount>
♂), 14k–l (S7–10 ♀), 16 (map)
</paragraph>
</subSubSection>
<subSubSection id="27DF27C8D018FFEBFF615C88F775FAA2" pageId="5" pageNumber="406" type="etymology">
<paragraph id="6F7A7443D018FFEBFF615C88F1B7FA86" blockId="5.[151,1437,1229,2010]" pageId="5" pageNumber="406">
<emphasis id="5DB1A851D018FFEBFF615C88F240FB6A" bold="true" box="[151,286,1229,1254]" pageId="5" pageNumber="406">Etymology.</emphasis>
From Latin 
<emphasis id="5DB1A851D018FFEBFE5B5C88F151FB6A" box="[429,527,1229,1254]" italics="true" pageId="5" pageNumber="406">gaudium</emphasis>
, happiness, and 
<emphasis id="5DB1A851D018FFEBFD3A5C8BF01CFB69" box="[716,834,1230,1253]" italics="true" pageId="5" pageNumber="406">montanum</emphasis>
, of the mountains. The name refers to its beauty and conspicuous presence in the highlands where it lives.
</paragraph>
<paragraph id="6F7A7443D018FFEBFF315D50F775FAA2" blockId="5.[151,1437,1229,2010]" box="[199,1067,1301,1326]" pageId="5" pageNumber="406">
<emphasis id="5DB1A851D018FFEBFF315D50F2E1FAA2" bold="true" box="[199,447,1301,1326]" pageId="5" pageNumber="406">Specimens examined</emphasis>
(
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: male 
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, allotype, 48 
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).
</paragraph>
</subSubSection>
<subSubSection id="27DF27C8D018FFE9FF315D7FF2CFFD40" lastPageId="7" lastPageNumber="408" pageId="5" pageNumber="406" type="description">
<paragraph id="6F7A7443D018FFEBFF315D7FF270F951" blockId="5.[151,1437,1229,2010]" pageId="5" pageNumber="406">
CEUA: Antioquia: 
<specimenCount id="79C3BFCAD018FFEBFE575D7FF293FADD" box="[417,461,1338,1362]" pageId="5" pageNumber="406" type="generic">1 ♂</specimenCount>
, Municipality Bello: Corregimiento San Félix: Township Sabanalarga: Subpáramo Las Baldías: 
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<geoCoordinate id="0AF11284D018FFEBFE725D1BF145FAFA" box="[388,539,1373,1398]" direction="west" orientation="longitude" pageId="5" pageNumber="406" precision="15" value="-75.649445">W 75°38'58&quot;</geoCoordinate>
<quantity id="A83DD9A6D018FFEBFDDF5D1BF124FAFA" box="[553,634,1374,1398]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="406" unit="m" value="3100.0">3100m</quantity>
, 
<date id="1B7B5283D018FFEBFD7D5D18F053FAFA" box="[651,781,1373,1398]" pageId="5" pageNumber="406" value="2013-07-16">16.vii.2013</date>
, Leg: J. R. Albertino &amp; A. L. Montoya. Municipality Belmira: Township Río Arriba: 
<taxonomicName id="A8C50FC0D018FFEBFE0D5DC7F10EFA16" box="[507,592,1410,1434]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="409" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
El Morro, 
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<geoCoordinate id="0AF11284D018FFEBFCA85DC7F0A8FA16" box="[862,1014,1409,1434]" direction="west" orientation="longitude" pageId="5" pageNumber="406" precision="1" value="-75.669106">W 75°40'8.8&quot;</geoCoordinate>
<quantity id="A83DD9A6D018FFEBFC0B5DC7F708FA15" box="[1021,1110,1410,1434]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="5" pageNumber="406" unit="m" value="3270.0">3270 m</quantity>
: 
<specimenCount id="79C3BFCAD018FFEBFB965DC7F7DAFA16" box="[1120,1156,1410,1434]" pageId="5" pageNumber="406" type="generic">1♂</specimenCount>
, 
<date id="1B7B5283D018FFEBFB665DC7F65FFA16" box="[1168,1281,1410,1434]" pageId="5" pageNumber="406" value="2008-05-04">04.v.2008</date>
, Leg: C. Bota &amp; A.L. Montoya. 
<specimenCount id="79C3BFCAD018FFEBFE9A5DE3F2C7FA31" box="[364,409,1446,1469]" pageId="5" pageNumber="406" type="generic">7 ♂</specimenCount>
and 3 ♀, 
<date id="1B7B5283D018FFEBFDE35DE0F1C2FA32" box="[533,668,1445,1470]" pageId="5" pageNumber="406" value="2010-04-02" valueMax="2010-04-04" valueMin="2010-04-02">2–4.iv.2010</date>
, Leg: C. Bota. 
<specimenCount id="79C3BFCAD018FFEBFCA55DE3F0DDFA31" box="[851,899,1446,1470]" pageId="5" pageNumber="406" type="generic">2 ♂</specimenCount>
, 
<date id="1B7B5283D018FFEBFC645DE0F0A3FA32" box="[914,1021,1445,1470]" pageId="5" pageNumber="406" value="2011-01-15">15.i.2011</date>
, Leg: L. Ríos, A. L. Montoya &amp; J. Cardona. 
<typeStatus id="B07ECAE1D018FFEBFEF15D8CF22CFA6E" box="[263,370,1481,1506]" pageId="5" pageNumber="406" type="holotype">Holotype</typeStatus>
, Allotype and 
<specimenCount id="79C3BFCAD018FFEBFDD75D8FF10FFA6D" box="[545,593,1482,1506]" pageId="5" pageNumber="406" type="generic">3 ♂</specimenCount>
, 
<date id="1B7B5283D018FFEBFD965D8FF18EFA6E" box="[608,720,1482,1506]" pageId="5" pageNumber="406" value="2011-05-10">10.v.2011</date>
, Leg: C. Bota &amp; J. D. Castaño. Alto del Indio, 
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<geoCoordinate id="0AF11284D018FFEBFF615DA8F3A4F98A" box="[151,250,1517,1542]" direction="west" orientation="longitude" pageId="5" pageNumber="406" precision="925" value="-75.7">W75°42'</geoCoordinate>
22.0&quot; 
<quantity id="A83DD9A6D018FFEBFEB35DA8F2C1F989" box="[325,415,1517,1542]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="5" pageNumber="406" unit="m" value="2950.0">2950 m</quantity>
: 6 ♀ and 
<specimenCount id="79C3BFCAD018FFEBFDFB5DABF164F989" box="[525,570,1518,1542]" pageId="5" pageNumber="406" type="generic">1 ♂</specimenCount>
, 
<date id="1B7B5283D018FFEBFDB15DA8F1EBF98A" box="[583,693,1517,1542]" pageId="5" pageNumber="406" value="2012-03-03">3.iii.2012</date>
; 
<specimenCount id="79C3BFCAD018FFEBFD345DABF1B2F989" box="[706,748,1518,1542]" pageId="5" pageNumber="406" type="generic">2 ♂</specimenCount>
and 4 ♀, 
<date id="1B7B5283D018FFEBFCA85DA8F0B1F98A" box="[862,1007,1517,1542]" pageId="5" pageNumber="406" value="2012-07-07" valueMax="2012-07-08" valueMin="2012-07-07">7–8.vii.2012</date>
, Leg: A. Clavijo. 
<taxonomicName id="A8C50FC0D018FFEBFB4C5DABF651F98A" box="[1210,1295,1518,1542]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="409" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
de Sabanas, 
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<geoCoordinate id="0AF11284D018FFEBFEDD5E54F294F9A6" box="[299,458,1553,1578]" direction="west" orientation="longitude" pageId="5" pageNumber="406" precision="1" value="-75.64569">W75°38'44.5&quot;</geoCoordinate>
<quantity id="A83DD9A6D018FFEBFE255E57F170F9A5" box="[467,558,1554,1578]" metricMagnitude="3" metricUnit="m" metricValue="3.13" pageId="5" pageNumber="406" unit="m" value="3130.0">3130 m</quantity>
: 
<specimenCount id="79C3BFCAD018FFEBFDCD5E54F139F9A5" box="[571,615,1553,1578]" pageId="5" pageNumber="406" type="generic">5 ♂</specimenCount>
and 1 ♀, 
<date id="1B7B5283D018FFEBFD285E57F000F9A6" box="[734,862,1554,1578]" pageId="5" pageNumber="406" value="2011-10-01" valueMax="2011-10-03" valueMin="2011-10-01">1–3.x.2011</date>
, Leg: C. Bota &amp; J.D. Castaño. 
<specimenCount id="79C3BFCAD018FFEBFB3E5E57F7ABF9A5" box="[1224,1269,1554,1578]" pageId="5" pageNumber="406" type="generic">2 ♂</specimenCount>
and 1 ♀, path from the town toward El Morro, 
<geoCoordinate id="0AF11284D018FFEBFDF55E70F10CF9C2" box="[515,594,1589,1614]" direction="north" orientation="latitude" pageId="5" pageNumber="406" precision="925" value="6.616667">6°37'N</geoCoordinate>
<geoCoordinate id="0AF11284D018FFEBFDAE5E70F1E5F9C2" box="[600,699,1589,1614]" direction="west" orientation="longitude" pageId="5" pageNumber="406" precision="925" value="-75.666664">75°40'W</geoCoordinate>
<quantity id="A83DD9A6D018FFEBFD375E70F048F9C1" box="[705,790,1589,1614]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="5" pageNumber="406" unit="m" value="2950.0">2950 m</quantity>
approx., 
<date id="1B7B5283D018FFEBFC885E70F75FF9C2" box="[894,1025,1589,1614]" pageId="5" pageNumber="406" value="2012-07-16">16.vii.2012</date>
. Leg: C. Bota, C. Moreno, C. Flórez, J. Caly, J. del Río, J. Sierra, K. Mejía &amp; M. Moreno. 
<specimenCount id="79C3BFCAD018FFEBFCF45E1FF06EF9FD" box="[770,816,1626,1650]" pageId="5" pageNumber="406" type="generic">2 ♂</specimenCount>
, Corregimiento Labores, Finca El Paraíso, 
<geoCoordinate id="0AF11284D018FFEBFAD75E1FF624F9FE" box="[1313,1402,1625,1650]" direction="north" orientation="latitude" pageId="5" pageNumber="406" precision="925" value="6.6666665">N 6°40'</geoCoordinate>
<geoCoordinate id="0AF11284D018FFEBFA745E1FF3BCF91A" direction="west" orientation="longitude" pageId="5" pageNumber="406" precision="925" value="-75.63333">W 75°38'</geoCoordinate>
, 
<date id="1B7B5283D018FFEBFF1B5E38F22AF91A" box="[237,372,1661,1686]" pageId="5" pageNumber="406" value="2012-04-05" valueMax="2012-04-07" valueMin="2012-04-05">5–7.iv.2012</date>
, Leg: C. Moreno, S. Bota, T. Bota &amp; C. Bota. Municipality San José de la Montaña, 
<taxonomicName id="A8C50FC0D018FFEBFAD15E3BF623F91A" box="[1319,1405,1662,1686]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="409" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
<collectingCountry id="17D234D3D018FFEBFA755E38F3BAF936" name="Republic of the Congo" pageId="5" pageNumber="406">El Congo</collectingCountry>
, 
<geoCoordinate id="0AF11284D018FFEBFF195EE4F224F936" box="[239,378,1697,1722]" direction="north" orientation="latitude" pageId="5" pageNumber="406" precision="1" value="6.7604446">N6°45'37.6&quot;</geoCoordinate>
<geoCoordinate id="0AF11284D018FFEBFE765EE4F17EF936" box="[384,544,1697,1722]" direction="west" orientation="longitude" pageId="5" pageNumber="406" precision="1" value="-75.590004">W75°35'24.0&quot;</geoCoordinate>
<quantity id="A83DD9A6D018FFEBFDD05EE7F126F936" box="[550,632,1698,1722]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="406" unit="m" value="3200.0">3200m</quantity>
: 5 ♀ and 
<specimenCount id="79C3BFCAD018FFEBFD175EE7F053F935" box="[737,781,1698,1722]" pageId="5" pageNumber="406" type="generic">3 ♂</specimenCount>
, 
<date id="1B7B5283D018FFEBFCEE5EE4F0E5F936" box="[792,955,1697,1722]" pageId="5" pageNumber="406" value="2011-09-10" valueMax="2011-09-14" valueMin="2011-09-10">10–14.ix.2011</date>
, Leg: L. Ríos &amp; J. Zapata. 1 ♀, 
<date id="1B7B5283D018FFEBFAE85EE4F6C6F936" box="[1310,1432,1697,1722]" pageId="5" pageNumber="406" value="2011-11-05">05.xi.2011</date>
, Leg: C. Bota.
</paragraph>
<paragraph id="6F7A7443D018FFEBFF315EACF78EF81E" blockId="5.[151,1437,1229,2010]" pageId="5" pageNumber="406">
<emphasis id="5DB1A851D018FFEBFF315EACF207F88E" bold="true" box="[199,345,1769,1794]" pageId="5" pageNumber="406">Description.</emphasis>
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. 
<emphasis id="5DB1A851D018FFEBFE2E5EACF141F88E" bold="true" box="[472,543,1769,1794]" pageId="5" pageNumber="406">Head.</emphasis>
Labium cream along edges becoming light blue toward center. Mandible base blue. Labrum blue with a black small medio-basal spot and dorsolateral black margins. Gena blue. Anteclypeus blue. Postclypeus black. Frons black, with blue postocular spots touching eye margin, with pruinescence, in anterior view area bordering gena and clypeus on each side light blue, divided by a black medial stripe over clypeus. Black antennae. Rear of head dorsally cream and ventrally pale blue (
<figureCitation id="F7FE68C6D018FFEBFBF05F3CF70CF81E" box="[1030,1106,1913,1938]" captionStart="FIGURE 3" captionStartId="17.[151,250,1955,1977]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@17.[204,1382,193,1921]" captionTargetPageId="17" captionText="FIGURE 3. Mesamphiagrion santainense (Antioquia, Belmira): (a) Male holotype; (b) Female paratype. M. gaudiimontanum (Antioquia, Belmira) Mature males: (c) Dorsal view; (d) Lateral view." httpUri="https://zenodo.org/record/246831/files/figure.png" pageId="5" pageNumber="406">Figs. 3</figureCitation>
c–d, 5b–c).
</paragraph>
<paragraph id="6F7A7443D018FFE8FF315FD8F16AFE79" blockId="5.[151,1437,1229,2010]" lastBlockId="6.[151,1436,151,1977]" lastPageId="6" lastPageNumber="407" pageId="5" pageNumber="406">
<emphasis id="5DB1A851D018FFEBFF315FD8F278F83A" bold="true" box="[199,294,1949,1974]" pageId="5" pageNumber="406">Thorax.</emphasis>
Prothorax black, with anterior lobe of pronotum light blue, propleura light blue. Medial lobe of posterior prothoracic lobe developed into a caudally projected squarish plate with smoothly rounded margins and posterior edge slightly concave, each lateral edge approximately half length of distal edge. Pterothorax black with blue antehumeral stripe, wide blue stripe covering mesepimeron and metepisternum, metepimeron light blue, and venter cream. Coxa light blue. External side of remainder of leg black and medial side light blue with blue rounded spot near base of femur. Tarsi, spurs, and claws black (
<figureCitation id="F7FE68C6D01BFFE8FD015941F01DFE91" box="[759,835,260,285]" captionStart="FIGURE 3" captionStartId="17.[151,250,1955,1977]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@17.[204,1382,193,1921]" captionTargetPageId="17" captionText="FIGURE 3. Mesamphiagrion santainense (Antioquia, Belmira): (a) Male holotype; (b) Female paratype. M. gaudiimontanum (Antioquia, Belmira) Mature males: (c) Dorsal view; (d) Lateral view." httpUri="https://zenodo.org/record/246831/files/figure.png" pageId="6" pageNumber="407">Figs. 3</figureCitation>
c–d, 5b–c). Six external spurs on right metafemur and seven on left metafemur, as long as space between them or shorter, gradually increasing in size toward apex. Six external spurs on right metatibia and seven on left metatibia, as long as space between them or shorter, gradually decreasing in size toward apex. Tarsal claws with well developed supplementary tooth. Wings hyaline. Pt blue, ratio between distal and proximal length about 1:1. CuP reaching CuPAA slightly distal to confluence of CuPAA with hind margin of wing. Px 
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FW, Px 
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right HW, 
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left HW. 
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2 branching between Px 4 and 
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FW, between Px 3 and 
<quantity id="A83DD9A6D01BFFE8FE925999F2D1FE79" box="[356,399,476,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="407" unit="in" value="4.0">4 in</quantity>
HW (
<figureCitation id="F7FE68C6D01BFFE8FE225999F149FE79" box="[468,535,476,501]" captionStart="FIGURE 5" captionStartId="20.[151,250,1906,1928]" captionTargetBox="[181,1405,429,1883]" captionTargetId="figure@20.[181,1405,427,1885]" captionTargetPageId="20" captionText="FIGURE 5. Ontogenic changes in Mesamphiagrion gaudiimontanum males (Antioquia, Belmira): (a) Teneral; (b) Mature; (c) Middle age parasitized by mites (Photograph by Camilo Flórez-V.)." httpUri="https://zenodo.org/record/246833/files/figure.png" pageId="6" pageNumber="407">Fig. 5</figureCitation>
b).
</paragraph>
<paragraph id="6F7A7443D01BFFE8FF3159BAF098FCF3" blockId="6.[151,1436,151,1977]" pageId="6" pageNumber="407">
<emphasis id="5DB1A851D01BFFE8FF3159BAF21DFD94" bold="true" box="[199,323,511,536]" pageId="6" pageNumber="407">Abdomen.</emphasis>
Dorsum black and lateral terga cream except: distal half of S1 lateral terga light blue, S2 lateral terga bluish, S7 dorsum with a blue spot occupying distal 5/6, S3–7 with a proximal incomplete blue ring, S8–9 dorsum blue (
<figureCitation id="F7FE68C6D01BFFE8FEC65A02F22FFDEC" box="[304,369,583,608]" captionStart="FIGURE 3" captionStartId="17.[151,250,1955,1977]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@17.[204,1382,193,1921]" captionTargetPageId="17" captionText="FIGURE 3. Mesamphiagrion santainense (Antioquia, Belmira): (a) Male holotype; (b) Female paratype. M. gaudiimontanum (Antioquia, Belmira) Mature males: (c) Dorsal view; (d) Lateral view." httpUri="https://zenodo.org/record/246831/files/figure.png" pageId="6" pageNumber="407">Fig. 3</figureCitation>
c–d, 5b–c, 12b). Genital ligula (as in figures 7d–e) with apex concave with a pair of lateroapical lobes with blunt distal portion and proximal portion terminating in a sharp hook, lateromedial lobes rounded in ectal view, and inner medial process as long as lateromedial lobes in lateral view. Cercus (
<figureCitation id="F7FE68C6D01BFFE8FB7F5ACAF78BFD24" box="[1161,1237,655,680]" captionStart="FIGURE 7" captionStartId="22.[151,250,1817,1839]" captionTargetBox="[328,1261,193,1765]" captionTargetId="figure@22.[320,1262,193,1771]" captionTargetPageId="22" captionText="FIGURE 7. Mesamphiagrion gaianii (Paratype, Venezuela, Trujillo, Páramo La Cristalina) vs. M. gaudiimontanum (Antioquia, Belmira) (traits of M. gaianii on left): (a) Lateral view of caudal appendages; (b) Posterior view of left cercus; (c) Medio-dorsal view of cercus; (d) Ectal view of genital ligula; (e) M. gaudiimontanum lateral view of genital ligula; Female paratype of M. gaudiimontanum: (f) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Inner process. Illustrations drawn by: Rosser Garrison (a to d), Gloria Mora (e), and Natalia Uribe (f)." httpUri="https://zenodo.org/record/246835/files/figure.png" pageId="6" pageNumber="407">Figs. 7</figureCitation>
a–c) external side black, medial side blue, shorter than S10 length, with elongated dorsal process finishing in hook directed ventrally, this being most distal point of animal, apex of external ventroapical process blunt, inner ventro-apical process rising as concave carina in laterodorsal view, ventrobasal process terminating in blunt point, ventroapical processes located midway between dorsal and ventrobasal processes in posteromedial view. Paraproct light blue with black tip and distally directed cylindrical apical process, which is about half cercus length or slightly longer. Total length 
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. Abdomen length 
<quantity id="A83DD9A6D01BFFE8FE405B2DF156FCF3" box="[438,520,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="407" unit="mm" value="23.0">23 mm</quantity>
. FW length 
<quantity id="A83DD9A6D01BFFE8FD665B2DF1BCFCF3" box="[656,738,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="407" unit="mm" value="18.0">18 mm</quantity>
. HW length 
<quantity id="A83DD9A6D01BFFE8FC995B2DF09CFCF3" box="[879,962,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="407" unit="mm" value="17.0">17 mm</quantity>
.
</paragraph>
<paragraph id="6F7A7443D01BFFE8FF315BC9F23AFB13" blockId="6.[151,1436,151,1977]" pageId="6" pageNumber="407">
<emphasis id="5DB1A851D01BFFE8FF315BC9F1F7FC29" bold="true" box="[199,681,908,933]" pageId="6" pageNumber="407">
Variations in male 
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. Mature.
</emphasis>
Pruinescence on head present or absent. Rear of head cream. Metapleural black stripe continuous or interrupted once or twice. Six to nine external metafemoral spurs. Four to seven external metatibial spurs. 11–13 Px in FW, 
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HW. 11–13 Px in FW, 10–11 Px in HW. 
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2 branching between Px 4 and 
<quantity id="A83DD9A6D01BFFE8FE905BB2F2CCFB9C" box="[358,402,1015,1040]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="407" unit="in" value="6.0">6 in</quantity>
FW, between Px 3 and 
<quantity id="A83DD9A6D01BFFE8FD685BB2F194FB9C" box="[670,714,1015,1040]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="407" unit="in" value="5.0">5 in</quantity>
HW. In one specimen, S7 pale blue dorsal spot extending along distal 2/3 without reaching distal margin (
<figureCitation id="F7FE68C6D01BFFE8FD9D5C59F1E3FBB8" box="[619,701,1052,1077]" captionStart="FIGURE 12" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[201,1388,193,1937]" captionTargetId="figure@31.[201,1388,193,1938]" captionTargetPageId="31" captionText="FIGURE 12. Mesamphiagrion tamaense (Santander, Suratá): (a) Mature female; (b) Juvenile female; (c) Mature male; (d) Juvenile male." httpUri="https://zenodo.org/record/246840/files/figure.png" pageId="6" pageNumber="407">Fig. 12</figureCitation>
a). S10 sometimes with small proximal light blue spot and medial distal tip blue. Paraproct cream, sometimes with process bluish. Apex of ligula more or less markedly concave, in some cases almost straight. Total length 
<quantity id="A83DD9A6D01BFFE8FD9B5C21F1B2FBF0" box="[621,748,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="6" pageNumber="407" unit="mm" value="30.0" valueMax="31.0" valueMin="29.0">29–31 mm</quantity>
. Abdominal length 
<quantity id="A83DD9A6D01BFFE8FC275C21F70EFBF0" box="[977,1104,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="6" pageNumber="407" unit="mm" value="23.0" valueMax="24.0" valueMin="22.0">22–24 mm</quantity>
. FW length 
<quantity id="A83DD9A6D01BFFE8FB175C21F63EFBF0" box="[1249,1376,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="6" pageNumber="407" unit="mm" value="18.5" valueMax="19.0" valueMin="18.0">18–19 mm</quantity>
. HW length 
<quantity id="A83DD9A6D01BFFE8FF125CC2F23EFB13" box="[228,352,1159,1184]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="6" pageNumber="407" unit="mm" value="17.0" valueMax="18.0" valueMin="16.0">16–18 mm</quantity>
.
</paragraph>
<paragraph id="6F7A7443D01BFFE8FF315CE9F063FABC" blockId="6.[151,1436,151,1977]" pageId="6" pageNumber="407">
<emphasis id="5DB1A851D01BFFE8FF315CE9F26FFB49" bold="true" box="[199,305,1196,1221]" pageId="6" pageNumber="407">Juvenile.</emphasis>
Labium cream. Base of mandible cream. Labrum blue. Gena blue. Anteclypeus light blue and postclypeus reddish brown. Frons as in mature males but paler. Prothorax brown. Pterothorax as in mature males but black zones brown. Coxa light brown. Pt red. S1–4 as in mature males but red instead of black. S4 with a distal black spot. S5–6 black dorsally and red laterally (
<figureCitation id="F7FE68C6D01BFFE8FD345D52F05DFABC" box="[706,771,1303,1328]" captionStart="FIGURE 5" captionStartId="20.[151,250,1906,1928]" captionTargetBox="[181,1405,429,1883]" captionTargetId="figure@20.[181,1405,427,1885]" captionTargetPageId="20" captionText="FIGURE 5. Ontogenic changes in Mesamphiagrion gaudiimontanum males (Antioquia, Belmira): (a) Teneral; (b) Mature; (c) Middle age parasitized by mites (Photograph by Camilo Flórez-V.)." httpUri="https://zenodo.org/record/246833/files/figure.png" pageId="6" pageNumber="407">Fig. 5</figureCitation>
a–b).
</paragraph>
<paragraph id="6F7A7443D01BFFE8FF315D79F1CEFAF4" blockId="6.[151,1436,151,1977]" pageId="6" pageNumber="407">
<emphasis id="5DB1A851D01BFFE8FF315D79F1CAFAD9" bold="true" box="[199,660,1340,1365]" pageId="6" pageNumber="407">Allotype (androchrome female). Head.</emphasis>
Color pattern as in 
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but paler, postocular spots not reaching eye margin and occipital bar cream (
<figureCitation id="F7FE68C6D01BFFE8FDC65D1AF12DFAF4" box="[560,627,1375,1400]" captionStart="FIGURE 6" captionStartId="21.[151,250,1942,1964]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@21.[204,1382,193,1921]" captionTargetPageId="21" captionText="FIGURE 6. Ontogenic changes in both morphs of Mesamphiagrion gaudiimontanum females (Antioquia, Belmira): Androchrome: (a) Juvenile; (b) Mature. Gynochrome: (c) Juvenile parasitized by mites (Photograph by Cintia Moreno); (d) Mature." httpUri="https://zenodo.org/record/246834/files/figure.png" pageId="6" pageNumber="407">Fig. 6</figureCitation>
b).
</paragraph>
<paragraph id="6F7A7443D01BFFE8FF315DC1F134F9F9" blockId="6.[151,1436,151,1977]" pageId="6" pageNumber="407">
<emphasis id="5DB1A851D01BFFE8FF315DC1F278FA11" bold="true" box="[199,294,1412,1437]" pageId="6" pageNumber="407">Thorax.</emphasis>
Coloration as in mature males but dark brown instead of black (
<figureCitation id="F7FE68C6D01BFFE8FC075DC1F76AFA11" box="[1009,1076,1412,1437]" captionStart="FIGURE 6" captionStartId="21.[151,250,1942,1964]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@21.[204,1382,193,1921]" captionTargetPageId="21" captionText="FIGURE 6. Ontogenic changes in both morphs of Mesamphiagrion gaudiimontanum females (Antioquia, Belmira): Androchrome: (a) Juvenile; (b) Mature. Gynochrome: (c) Juvenile parasitized by mites (Photograph by Cintia Moreno); (d) Mature." httpUri="https://zenodo.org/record/246834/files/figure.png" pageId="6" pageNumber="407">Fig. 6</figureCitation>
b). Medial lobe of posterior lobe of pronotum well developed, with lateral margin as long as about 1/3 of posterior margin, its distal edge strongly concave medially. Mesepisternal plate approximately flat and triangular, black with blue apex (
<figureCitation id="F7FE68C6D01BFFE8FB035D89F663FA69" box="[1269,1341,1484,1509]" captionStart="FIGURE 7" captionStartId="22.[151,250,1817,1839]" captionTargetBox="[328,1261,193,1765]" captionTargetId="figure@22.[320,1262,193,1771]" captionTargetPageId="22" captionText="FIGURE 7. Mesamphiagrion gaianii (Paratype, Venezuela, Trujillo, Páramo La Cristalina) vs. M. gaudiimontanum (Antioquia, Belmira) (traits of M. gaianii on left): (a) Lateral view of caudal appendages; (b) Posterior view of left cercus; (c) Medio-dorsal view of cercus; (d) Ectal view of genital ligula; (e) M. gaudiimontanum lateral view of genital ligula; Female paratype of M. gaudiimontanum: (f) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Inner process. Illustrations drawn by: Rosser Garrison (a to d), Gloria Mora (e), and Natalia Uribe (f)." httpUri="https://zenodo.org/record/246835/files/figure.png" pageId="6" pageNumber="407">Fig. 7</figureCitation>
f). Coxa bluish cream. Remainder of leg as in 
<typeStatus id="B07ECAE1D01BFFE8FDB75DAAF1FDF984" box="[577,675,1519,1544]" pageId="6" pageNumber="407" type="holotype">holotype</typeStatus>
but brown instead of black and cream instead of blue. Six external spurs on right metafemur and seven on left metafemur. Five external spurs on right metatibia and six on left metatibia. Pt reddish brown. 12 Px in right FW, 
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left FW, 10 Px in HW. 
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2 branching between Px 5 and 
<quantity id="A83DD9A6D01BFFE8FA875E72F6C2F9DC" box="[1393,1436,1591,1616]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="407" unit="in" value="6.0">6 in</quantity>
FW, between Px 4 and 
<quantity id="A83DD9A6D01BFFE8FE6C5E19F29AF9F9" box="[410,452,1628,1653]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="407" unit="in" value="5.0">5 in</quantity>
HW (
<figureCitation id="F7FE68C6D01BFFE8FDFF5E19F112F9F9" box="[521,588,1628,1653]" captionStart="FIGURE 6" captionStartId="21.[151,250,1942,1964]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@21.[204,1382,193,1921]" captionTargetPageId="21" captionText="FIGURE 6. Ontogenic changes in both morphs of Mesamphiagrion gaudiimontanum females (Antioquia, Belmira): Androchrome: (a) Juvenile; (b) Mature. Gynochrome: (c) Juvenile parasitized by mites (Photograph by Cintia Moreno); (d) Mature." httpUri="https://zenodo.org/record/246834/files/figure.png" pageId="6" pageNumber="407">Fig. 6</figureCitation>
b).
</paragraph>
<paragraph id="6F7A7443D01BFFE8FF315E3AF2F4F834" blockId="6.[151,1436,151,1977]" pageId="6" pageNumber="407">
<emphasis id="5DB1A851D01BFFE8FF315E3AF21DF914" bold="true" box="[199,323,1663,1688]" pageId="6" pageNumber="407">Abdomen.</emphasis>
Dorsum black and venter cream, except for blue lateral spots on S1–2, distal margin of S1 light blue, S2–3 dorsally brown, S4–5 same but with black distal 1/4, S3–7 with incomplete cream ring along proximal margin, S7 with blue spot located on middle of segment dorsum covering approximately half, S8 with light blue spot covering basal 3/4, S9 with longitudinal media blue line running from center to distal margin. S10 with median blue stripe widening distally (
<figureCitation id="F7FE68C6D01BFFE8FDB35F4AF1CBF8A4" box="[581,661,1807,1832]" captionStart="FIGURE 6" captionStartId="21.[151,250,1942,1964]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@21.[204,1382,193,1921]" captionTargetPageId="21" captionText="FIGURE 6. Ontogenic changes in both morphs of Mesamphiagrion gaudiimontanum females (Antioquia, Belmira): Androchrome: (a) Juvenile; (b) Mature. Gynochrome: (c) Juvenile parasitized by mites (Photograph by Cintia Moreno); (d) Mature." httpUri="https://zenodo.org/record/246834/files/figure.png" pageId="6" pageNumber="407">Figs. 6</figureCitation>
b, 13k). Black line from S2 proximal margin to vulvar spine, vulvar spine black. Ovipositor, cercus, and paraproct cream. Subbasal plate of ovipositor triangular. Ovipositor apex reaching level of apex of paraproct, stylus reaching apex of cercus. Cercus brown, conical, slightly longer than half of S10. Paraproct creamy blue, truncate posteriorly. Total length 
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. Abdomen length 
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. FW length 
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. HW length 
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.
</paragraph>
<paragraph id="6F7A7443D01AFFE9FF3158D2F6C9FF7B" blockId="7.[151,1437,151,1616]" pageId="7" pageNumber="408">
<emphasis id="5DB1A851D01AFFE9FF3158D2F06BFF3C" bold="true" box="[199,821,151,176]" pageId="7" pageNumber="408">
Variation in mature female androchrome 
<typeStatus id="B07ECAE1D01AFFE9FD4E58D2F06EFF3C" box="[696,816,151,176]" pageId="7" pageNumber="408" type="paratype">paratypes</typeStatus>
.
</emphasis>
Five to eight metafemoral external spurs. Five to eight metatibial external spurs. 10–14 Px in FW, 9–12 Px in HW. 
<collectingCountry id="17D234D3D01AFFE9FCC758F9F00AFF58" box="[817,852,188,212]" name="Poland" pageId="7" pageNumber="408">RP</collectingCountry>
2 branching between the Px 4 and 
<quantity id="A83DD9A6D01AFFE9FB2558F9F7A0FF59" box="[1235,1278,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="408" unit="in" value="6.0">6 in</quantity>
FWs, in HWs between Px 3 and 5. Total length 
<quantity id="A83DD9A6D01AFFE9FDFB58A5F1D7FF7B" box="[525,649,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="7" pageNumber="408" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">28–32 mm</quantity>
. Abdomen length 
<quantity id="A83DD9A6D01AFFE9FCA0589AF08CFF7B" box="[854,978,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="7" pageNumber="408" unit="mm" value="23.0" valueMax="25.0" valueMin="21.0">21–25 mm</quantity>
. FW 
<quantity id="A83DD9A6D01AFFE9FBF858A5F7D4FF7B" box="[1038,1162,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="7" pageNumber="408" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 mm</quantity>
. HW length 
<quantity id="A83DD9A6D01AFFE9FAE1589AF6CDFF7B" box="[1303,1427,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="7" pageNumber="408" unit="mm" value="17.5" valueMax="19.0" valueMin="16.0">16–19 mm</quantity>
.
</paragraph>
<paragraph id="6F7A7443D01AFFE9FF315941F2E3FE5C" blockId="7.[151,1437,151,1616]" pageId="7" pageNumber="408">
<emphasis id="5DB1A851D01AFFE9FF315941F06DFE91" bold="true" box="[199,819,260,285]" pageId="7" pageNumber="408">
Variation in female gynochrome 
<typeStatus id="B07ECAE1D01AFFE9FD955941F185FE91" box="[611,731,260,285]" pageId="7" pageNumber="408" type="paratype">paratypes</typeStatus>
. Head.
</emphasis>
As in androchrome females (
<figureCitation id="F7FE68C6D01AFFE9FB6E5941F7BEFE91" box="[1176,1248,260,285]" captionStart="FIGURE 6" captionStartId="21.[151,250,1942,1964]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@21.[204,1382,193,1921]" captionTargetPageId="21" captionText="FIGURE 6. Ontogenic changes in both morphs of Mesamphiagrion gaudiimontanum females (Antioquia, Belmira): Androchrome: (a) Juvenile; (b) Mature. Gynochrome: (c) Juvenile parasitized by mites (Photograph by Cintia Moreno); (d) Mature." httpUri="https://zenodo.org/record/246834/files/figure.png" pageId="7" pageNumber="408">Fig. 6</figureCitation>
c–d) but labium cream, sometimes blue toward center. Base of mandibles light yellow, becoming yellow toward apex. Labrum brown, with small brown to black medio-basal spot. Gena yellow. Anteclypeus bluish gray to gray. Postclypeus dark brown with or without pruinescence. Frons with or without pruinescence, in anterior view with area above clypeus black, in dorsal view brown, occipital edge cream, ocelli surrounded by black rings, postocular spots iridescent green or absent.
</paragraph>
<paragraph id="6F7A7443D01AFFE9FF315999F7B3FDEC" blockId="7.[151,1437,151,1616]" pageId="7" pageNumber="408">
<emphasis id="5DB1A851D01AFFE9FF315999F278FE79" bold="true" box="[199,294,476,501]" pageId="7" pageNumber="408">Thorax.</emphasis>
Prothorax dark brown with gray propleura, with or without pruinescence. Pterothorax as in 
<typeStatus id="B07ECAE1D01AFFE9FACC5999F6C2FE79" box="[1338,1436,476,501]" pageId="7" pageNumber="408" type="holotype">holotype</typeStatus>
but brown instead of black and green instead of blue, becoming darker with age and looking completely brown when covered with a thick layer of pruinescence (
<figureCitation id="F7FE68C6D01AFFE9FD495A61F05FFDB1" box="[703,769,548,573]" captionStart="FIGURE 6" captionStartId="21.[151,250,1942,1964]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@21.[204,1382,193,1921]" captionTargetPageId="21" captionText="FIGURE 6. Ontogenic changes in both morphs of Mesamphiagrion gaudiimontanum females (Antioquia, Belmira): Androchrome: (a) Juvenile; (b) Mature. Gynochrome: (c) Juvenile parasitized by mites (Photograph by Cintia Moreno); (d) Mature." httpUri="https://zenodo.org/record/246834/files/figure.png" pageId="7" pageNumber="408">Fig. 6</figureCitation>
c–d). Coxa brown with or without pruinescence. Remainder of leg brown with black spines or brown with femur medial surface gray. Pt red or brown to bluish.
</paragraph>
<paragraph id="6F7A7443D01AFFE9FF315A29F2CFFD40" blockId="7.[151,1437,151,1616]" pageId="7" pageNumber="408">
<emphasis id="5DB1A851D01AFFE9FF315A29F21DFD09" bold="true" box="[199,323,620,645]" pageId="7" pageNumber="408">Abdomen.</emphasis>
Dorsum black and sides cream, except proximal 3/4 of S3–4 and S1–2 which are brown on mature females and red on juvenile females (
<figureCitation id="F7FE68C6D01AFFE9FDA35ACAF1F9FD24" box="[597,679,655,680]" captionStart="FIGURE 6" captionStartId="21.[151,250,1942,1964]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@21.[204,1382,193,1921]" captionTargetPageId="21" captionText="FIGURE 6. Ontogenic changes in both morphs of Mesamphiagrion gaudiimontanum females (Antioquia, Belmira): Androchrome: (a) Juvenile; (b) Mature. Gynochrome: (c) Juvenile parasitized by mites (Photograph by Cintia Moreno); (d) Mature." httpUri="https://zenodo.org/record/246834/files/figure.png" pageId="7" pageNumber="408">Figs. 6</figureCitation>
c–d, 13l). Juveniles with small blue spot on dorsum of S10 and sometimes also on S9.
</paragraph>
</subSubSection>
<subSubSection id="27DF27C8D01AFFE9FF315A92F0E0FB64" pageId="7" pageNumber="408" type="diagnosis">
<paragraph id="6F7A7443D01AFFE9FF315A92F0E0FB64" blockId="7.[151,1437,151,1616]" pageId="7" pageNumber="408">
<emphasis id="5DB1A851D01AFFE9FF315A92F21FFD7C" bold="true" box="[199,321,727,752]" pageId="7" pageNumber="408">Diagnosis.</emphasis>
Although in juvenile specimens the pt is red, in mature males it is blue and distinguishes this species from all other males of the genus. The cercus dorsal process is longer than ventrobasal external process, a character state that (
<figureCitation id="F7FE68C6D01AFFE9FE775B5AF292FCBB" box="[385,460,799,824]" captionStart="FIGURE 7" captionStartId="22.[151,250,1817,1839]" captionTargetBox="[328,1261,193,1765]" captionTargetId="figure@22.[320,1262,193,1771]" captionTargetPageId="22" captionText="FIGURE 7. Mesamphiagrion gaianii (Paratype, Venezuela, Trujillo, Páramo La Cristalina) vs. M. gaudiimontanum (Antioquia, Belmira) (traits of M. gaianii on left): (a) Lateral view of caudal appendages; (b) Posterior view of left cercus; (c) Medio-dorsal view of cercus; (d) Ectal view of genital ligula; (e) M. gaudiimontanum lateral view of genital ligula; Female paratype of M. gaudiimontanum: (f) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Inner process. Illustrations drawn by: Rosser Garrison (a to d), Gloria Mora (e), and Natalia Uribe (f)." httpUri="https://zenodo.org/record/246835/files/figure.png" pageId="7" pageNumber="408">Figs 7</figureCitation>
a, c) groups this species with 
<emphasis id="5DB1A851D01AFFE9FCDC5B64F27EFCD1" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="A8C50FC0D01AFFE9FCDC5B64F0E1FCB4" box="[810,959,801,824]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tamaense">M. tamaense</taxonomicName>
, 
<taxonomicName id="A8C50FC0D01AFFE9FC3B5B64F70BFCB4" box="[973,1109,799,824]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="nataliae">M. nataliae</taxonomicName>
, 
<taxonomicName id="A8C50FC0D01AFFE9FB955B64F780FCB4" box="[1123,1246,801,824]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaianii">M. gaianii</taxonomicName>
, 
<taxonomicName id="A8C50FC0D01AFFE9FB1B5B64F631FCB4" box="[1261,1391,799,824]" class="Insecta" family="Coenagrionidae" genus="Acanthagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="laterale">M. laterale</taxonomicName>
, 
<taxonomicName id="A8C50FC0D01AFFE9FA885B64F242FCD1" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="demarmelsi">M. demarmelsi</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="A8C50FC0D01AFFE9FEA05B00F290FCD1" box="[342,462,836,861]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="dunklei">
<emphasis id="5DB1A851D01AFFE9FEA05B00F290FCD1" box="[342,462,836,861]" italics="true" pageId="7" pageNumber="408">M. dunklei</emphasis>
</taxonomicName>
. The elongate ventrobasal process ending in a blunt apex (
<figureCitation id="F7FE68C6D01AFFE9FBAB5B01F7F7FCD1" box="[1117,1193,836,861]" captionStart="FIGURE 7" captionStartId="22.[151,250,1817,1839]" captionTargetBox="[328,1261,193,1765]" captionTargetId="figure@22.[320,1262,193,1771]" captionTargetPageId="22" captionText="FIGURE 7. Mesamphiagrion gaianii (Paratype, Venezuela, Trujillo, Páramo La Cristalina) vs. M. gaudiimontanum (Antioquia, Belmira) (traits of M. gaianii on left): (a) Lateral view of caudal appendages; (b) Posterior view of left cercus; (c) Medio-dorsal view of cercus; (d) Ectal view of genital ligula; (e) M. gaudiimontanum lateral view of genital ligula; Female paratype of M. gaudiimontanum: (f) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Inner process. Illustrations drawn by: Rosser Garrison (a to d), Gloria Mora (e), and Natalia Uribe (f)." httpUri="https://zenodo.org/record/246835/files/figure.png" pageId="7" pageNumber="408">Figs. 7</figureCitation>
b–c) separates it from 
<taxonomicName id="A8C50FC0D01AFFE9FF615B2CF248FC0C" box="[151,278,871,896]" class="Insecta" family="Coenagrionidae" genus="Acanthagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="laterale">
<emphasis id="5DB1A851D01AFFE9FF615B2CF248FC0C" box="[151,278,871,896]" italics="true" pageId="7" pageNumber="408">M. laterale</emphasis>
</taxonomicName>
, 
<emphasis id="5DB1A851D01AFFE9FED05B2CF10DFC0C" box="[294,595,871,896]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="A8C50FC0D01AFFE9FED05B2CF2E4FC0C" box="[294,442,873,896]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tamaense">M. tamaense</taxonomicName>
, 
<taxonomicName id="A8C50FC0D01AFFE9FE3E5B2CF111FC0C" box="[456,591,871,896]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="nataliae">M. nataliae</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="A8C50FC0D01AFFE9FD785B2CF05BFC0C" box="[654,773,873,896]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaianii">
<emphasis id="5DB1A851D01AFFE9FD785B2CF05BFC0C" box="[654,773,873,896]" italics="true" pageId="7" pageNumber="408">M. gaianii</emphasis>
</taxonomicName>
, being similar to 
<taxonomicName id="A8C50FC0D01AFFE9FC225B2CF710FC0C" box="[980,1102,871,896]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="dunklei">
<emphasis id="5DB1A851D01AFFE9FC225B2CF710FC0C" box="[980,1102,871,896]" italics="true" pageId="7" pageNumber="408">M. dunklei</emphasis>
</taxonomicName>
. The cercus of 
<taxonomicName id="A8C50FC0D01AFFE9FAF35B2CF622FC0C" box="[1285,1404,873,896]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaianii">
<emphasis id="5DB1A851D01AFFE9FAF35B2CF622FC0C" box="[1285,1404,873,896]" italics="true" pageId="7" pageNumber="408">M. gaianii</emphasis>
</taxonomicName>
in posterior view is also narrower than the cercus of 
<taxonomicName id="A8C50FC0D01AFFE9FD025BC8F083FC29" box="[756,989,908,933]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaudiimontanum">
<emphasis id="5DB1A851D01AFFE9FD025BC8F083FC29" box="[756,989,908,933]" italics="true" pageId="7" pageNumber="408">M. gaudiimontanum</emphasis>
</taxonomicName>
, in which the internal carina is more developed (
<figureCitation id="F7FE68C6D01AFFE9FEED5BEAF23EFC44" box="[283,352,943,968]" captionStart="FIGURE 7" captionStartId="22.[151,250,1817,1839]" captionTargetBox="[328,1261,193,1765]" captionTargetId="figure@22.[320,1262,193,1771]" captionTargetPageId="22" captionText="FIGURE 7. Mesamphiagrion gaianii (Paratype, Venezuela, Trujillo, Páramo La Cristalina) vs. M. gaudiimontanum (Antioquia, Belmira) (traits of M. gaianii on left): (a) Lateral view of caudal appendages; (b) Posterior view of left cercus; (c) Medio-dorsal view of cercus; (d) Ectal view of genital ligula; (e) M. gaudiimontanum lateral view of genital ligula; Female paratype of M. gaudiimontanum: (f) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Inner process. Illustrations drawn by: Rosser Garrison (a to d), Gloria Mora (e), and Natalia Uribe (f)." httpUri="https://zenodo.org/record/246835/files/figure.png" pageId="7" pageNumber="408">Fig. 7</figureCitation>
b). The wider medial lobe of posterior prothoracic lobe separates 
<taxonomicName id="A8C50FC0D01AFFE9FBA65BF4F66BFC44" box="[1104,1333,943,968]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaudiimontanum">
<emphasis id="5DB1A851D01AFFE9FBA65BF4F66BFC44" box="[1104,1333,943,968]" italics="true" pageId="7" pageNumber="408">M. gaudiimontanum</emphasis>
</taxonomicName>
from 
<emphasis id="5DB1A851D01AFFE9FA8B5BF4F27FFC61" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="A8C50FC0D01AFFE9FA8B5BF4F243FC61" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="demarmelsi">M. demarmelsi</taxonomicName>
.
</emphasis>
Shape of genital ligula (
<figureCitation id="F7FE68C6D01AFFE9FDCB5B91F1D5FC61" box="[573,651,980,1005]" captionStart="FIGURE 7" captionStartId="22.[151,250,1817,1839]" captionTargetBox="[328,1261,193,1765]" captionTargetId="figure@22.[320,1262,193,1771]" captionTargetPageId="22" captionText="FIGURE 7. Mesamphiagrion gaianii (Paratype, Venezuela, Trujillo, Páramo La Cristalina) vs. M. gaudiimontanum (Antioquia, Belmira) (traits of M. gaianii on left): (a) Lateral view of caudal appendages; (b) Posterior view of left cercus; (c) Medio-dorsal view of cercus; (d) Ectal view of genital ligula; (e) M. gaudiimontanum lateral view of genital ligula; Female paratype of M. gaudiimontanum: (f) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Inner process. Illustrations drawn by: Rosser Garrison (a to d), Gloria Mora (e), and Natalia Uribe (f)." httpUri="https://zenodo.org/record/246835/files/figure.png" pageId="7" pageNumber="408">Figs. 7</figureCitation>
d–e) clearly separates it from 
<emphasis id="5DB1A851D01AFFE9FC155B90F6CBFC61" box="[995,1429,980,1005]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="A8C50FC0D01AFFE9FC155B90F736FC61" box="[995,1128,980,1005]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="nataliae">M. nataliae</taxonomicName>
, 
<taxonomicName id="A8C50FC0D01AFFE9FB805B90F657FC60" box="[1142,1289,981,1004]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tamaense">M. tamaense</taxonomicName>
, 
<taxonomicName id="A8C50FC0D01AFFE9FAE15B90F6CBFC61" box="[1303,1429,980,1005]" class="Insecta" family="Coenagrionidae" genus="Acanthagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="laterale">M. laterale</taxonomicName>
</emphasis>
, and 
<taxonomicName id="A8C50FC0D01AFFE9FF3A5BBCF217FB9C" box="[204,329,1015,1040]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="dunklei">
<emphasis id="5DB1A851D01AFFE9FF3A5BBCF217FB9C" box="[204,329,1015,1040]" italics="true" pageId="7" pageNumber="408">M. dunklei</emphasis>
</taxonomicName>
but resembles 
<emphasis id="5DB1A851D01AFFE9FDF75BBCF1DDFB9C" box="[513,643,1017,1040]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="A8C50FC0D01AFFE9FDF75BBCF121FB9C" box="[513,639,1017,1040]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaianii">M. gaianii</taxonomicName>
.
</emphasis>
Females are distinguished by the medial lobe of posterior lobe of pronotum well developed, with lateral margin as long as about 1/3 of posterior margin and distal edge strongly concave medially (
<figureCitation id="F7FE68C6D01AFFE9FE8F5C7AF29DFBD4" box="[377,451,1087,1112]" captionStart="FIGURE 7" captionStartId="22.[151,250,1817,1839]" captionTargetBox="[328,1261,193,1765]" captionTargetId="figure@22.[320,1262,193,1771]" captionTargetPageId="22" captionText="FIGURE 7. Mesamphiagrion gaianii (Paratype, Venezuela, Trujillo, Páramo La Cristalina) vs. M. gaudiimontanum (Antioquia, Belmira) (traits of M. gaianii on left): (a) Lateral view of caudal appendages; (b) Posterior view of left cercus; (c) Medio-dorsal view of cercus; (d) Ectal view of genital ligula; (e) M. gaudiimontanum lateral view of genital ligula; Female paratype of M. gaudiimontanum: (f) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Inner process. Illustrations drawn by: Rosser Garrison (a to d), Gloria Mora (e), and Natalia Uribe (f)." httpUri="https://zenodo.org/record/246835/files/figure.png" pageId="7" pageNumber="408">Fig. 7</figureCitation>
f). Androchrome females differ further from females of 
<taxonomicName id="A8C50FC0D01AFFE9FB9B5C04F7B6FBD4" box="[1133,1256,1089,1112]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaianii">
<emphasis id="5DB1A851D01AFFE9FB9B5C04F7B6FBD4" box="[1133,1256,1089,1112]" italics="true" pageId="7" pageNumber="408">M. gaianii</emphasis>
</taxonomicName>
(
<figureCitation id="F7FE68C6D01AFFE9FB095C7AF60BFBD4" box="[1279,1365,1087,1112]" captionStart="FIGURE 13. S 7 – 10" captionStartId="32.[151,250,1386,1408]" captionTargetBox="[378,1208,193,1364]" captionTargetId="figure@32.[378,1209,193,1364]" captionTargetPageId="32" captionText="FIGURE 13. S 7 – 10 color patterns of males: (a) and (b) Mesamphiagrion gaudiimontanum; (c) and (d) M. risi; (e) M. rosseri; (f) and (g) M. santainense; (h) M. tamaense; (i) and (j) M. occultum; (k) M. demarmelsi; (l) M. laterale; (m) M. ovigerum; (n) M. nataliae." httpUri="https://zenodo.org/record/246841/files/figure.png" pageId="7" pageNumber="408">Fig. 13</figureCitation>
r), 
<taxonomicName id="A8C50FC0D01AFFE9FA8B5C04F244FBF1" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="demarmelsi">
<emphasis id="5DB1A851D01AFFE9FA8B5C04F244FBF1" italics="true" pageId="7" pageNumber="408">M. demarmelsi</emphasis>
</taxonomicName>
(
<figureCitation id="F7FE68C6D01AFFE9FEDE5C21F2DBFBF0" box="[296,389,1124,1149]" captionStart="FIGURE 13. S 7 – 10" captionStartId="32.[151,250,1386,1408]" captionTargetBox="[378,1208,193,1364]" captionTargetId="figure@32.[378,1209,193,1364]" captionTargetPageId="32" captionText="FIGURE 13. S 7 – 10 color patterns of males: (a) and (b) Mesamphiagrion gaudiimontanum; (c) and (d) M. risi; (e) M. rosseri; (f) and (g) M. santainense; (h) M. tamaense; (i) and (j) M. occultum; (k) M. demarmelsi; (l) M. laterale; (m) M. ovigerum; (n) M. nataliae." httpUri="https://zenodo.org/record/246841/files/figure.png" pageId="7" pageNumber="408">Fig. 13s</figureCitation>
), and some females of 
<taxonomicName id="A8C50FC0D01AFFE9FD705C20F05CFBF1" box="[646,770,1124,1149]" class="Insecta" family="Coenagrionidae" genus="Acanthagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="laterale">
<emphasis id="5DB1A851D01AFFE9FD705C20F05CFBF1" box="[646,770,1124,1149]" italics="true" pageId="7" pageNumber="408">M. laterale</emphasis>
</taxonomicName>
(
<figureCitation id="F7FE68C6D01AFFE9FCE75C21F001FBF0" box="[785,863,1124,1149]" captionStart="FIGURE 13. S 7 – 10" captionStartId="32.[151,250,1386,1408]" captionTargetBox="[378,1208,193,1364]" captionTargetId="figure@32.[378,1209,193,1364]" captionTargetPageId="32" captionText="FIGURE 13. S 7 – 10 color patterns of males: (a) and (b) Mesamphiagrion gaudiimontanum; (c) and (d) M. risi; (e) M. rosseri; (f) and (g) M. santainense; (h) M. tamaense; (i) and (j) M. occultum; (k) M. demarmelsi; (l) M. laterale; (m) M. ovigerum; (n) M. nataliae." httpUri="https://zenodo.org/record/246841/files/figure.png" pageId="7" pageNumber="408">Fig. 13</figureCitation>
o–q) by having a blue spot on S7 (
<figureCitation id="F7FE68C6D01AFFE9FB295C21F675FBF1" box="[1247,1323,1124,1149]" captionStart="FIGURE 6" captionStartId="21.[151,250,1942,1964]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@21.[204,1382,193,1921]" captionTargetPageId="21" captionText="FIGURE 6. Ontogenic changes in both morphs of Mesamphiagrion gaudiimontanum females (Antioquia, Belmira): Androchrome: (a) Juvenile; (b) Mature. Gynochrome: (c) Juvenile parasitized by mites (Photograph by Cintia Moreno); (d) Mature." httpUri="https://zenodo.org/record/246834/files/figure.png" pageId="7" pageNumber="408">Figs. 6</figureCitation>
a–b, 13k), and from 
<taxonomicName id="A8C50FC0D01AFFE9FEF25CCCF2DBFB2C" box="[260,389,1159,1184]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="nataliae">
<emphasis id="5DB1A851D01AFFE9FEF25CCCF2DBFB2C" box="[260,389,1159,1184]" italics="true" pageId="7" pageNumber="408">M. nataliae</emphasis>
</taxonomicName>
(
<figureCitation id="F7FE68C6D01AFFE9FE625CC2F2BCFB2C" box="[404,482,1159,1184]" captionStart="FIGURE 13. S 7 – 10" captionStartId="32.[151,250,1386,1408]" captionTargetBox="[378,1208,193,1364]" captionTargetId="figure@32.[378,1209,193,1364]" captionTargetPageId="32" captionText="FIGURE 13. S 7 – 10 color patterns of males: (a) and (b) Mesamphiagrion gaudiimontanum; (c) and (d) M. risi; (e) M. rosseri; (f) and (g) M. santainense; (h) M. tamaense; (i) and (j) M. occultum; (k) M. demarmelsi; (l) M. laterale; (m) M. ovigerum; (n) M. nataliae." httpUri="https://zenodo.org/record/246841/files/figure.png" pageId="7" pageNumber="408">Fig. 13</figureCitation>
a–c) and 
<taxonomicName id="A8C50FC0D01AFFE9FDBD5CCCF19CFB2C" box="[587,706,1159,1184]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="dunklei">
<emphasis id="5DB1A851D01AFFE9FDBD5CCCF19CFB2C" box="[587,706,1159,1184]" italics="true" pageId="7" pageNumber="408">M. dunklei</emphasis>
</taxonomicName>
by presenting different shaped spots on S7–10 (
<figureCitation id="F7FE68C6D01AFFE9FB285CC2F675FB2C" box="[1246,1323,1159,1184]" captionStart="FIGURE 6" captionStartId="21.[151,250,1942,1964]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@21.[204,1382,193,1921]" captionTargetPageId="21" captionText="FIGURE 6. Ontogenic changes in both morphs of Mesamphiagrion gaudiimontanum females (Antioquia, Belmira): Androchrome: (a) Juvenile; (b) Mature. Gynochrome: (c) Juvenile parasitized by mites (Photograph by Cintia Moreno); (d) Mature." httpUri="https://zenodo.org/record/246834/files/figure.png" pageId="7" pageNumber="408">Figs. 6</figureCitation>
a–b, 13k). Juveniles of 
<emphasis id="5DB1A851D01AFFE9FED55CE8F18EFB48" box="[291,720,1196,1221]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="A8C50FC0D01AFFE9FED55CE8F2FCFB49" box="[291,418,1196,1221]" class="Insecta" family="Coenagrionidae" genus="Acanthagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="laterale">M. laterale</taxonomicName>
, 
<taxonomicName id="A8C50FC0D01AFFE9FE5B5CE8F16EFB49" box="[429,560,1196,1221]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="nataliae">M. nataliae</taxonomicName>
, 
<taxonomicName id="A8C50FC0D01AFFE9FDCD5CE8F195FB48" box="[571,715,1197,1220]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tamaense">M. tamaense</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="A8C50FC0D01AFFE9FCF35CE8F027FB48" box="[773,889,1197,1220]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaianii">
<emphasis id="5DB1A851D01AFFE9FCF35CE8F027FB48" box="[773,889,1197,1220]" italics="true" pageId="7" pageNumber="408">M. gaianii</emphasis>
</taxonomicName>
have red coloration only on S1–3, whereas in 
<taxonomicName id="A8C50FC0D01AFFE9FA8B5CE8F20BFB64" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaudiimontanum">
<emphasis id="5DB1A851D01AFFE9FA8B5CE8F20BFB64" italics="true" pageId="7" pageNumber="408">M. gaudiimontanum</emphasis>
</taxonomicName>
the red coloration extends through S1–6 (
<figureCitation id="F7FE68C6D01AFFE9FCDA5C8AF027FB64" box="[812,889,1231,1256]" captionStart="FIGURE 5" captionStartId="20.[151,250,1906,1928]" captionTargetBox="[181,1405,429,1883]" captionTargetId="figure@20.[181,1405,427,1885]" captionTargetPageId="20" captionText="FIGURE 5. Ontogenic changes in Mesamphiagrion gaudiimontanum males (Antioquia, Belmira): (a) Teneral; (b) Mature; (c) Middle age parasitized by mites (Photograph by Camilo Flórez-V.)." httpUri="https://zenodo.org/record/246833/files/figure.png" pageId="7" pageNumber="408">Figs. 5</figureCitation>
a, 6a).
</paragraph>
</subSubSection>
<subSubSection id="27DF27C8D01AFFE6FF315CB1F6C3FCC1" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408" type="discussion">
<paragraph id="6F7A7443D01AFFE9FF315CB1F67EF984" blockId="7.[151,1437,151,1616]" pageId="7" pageNumber="408">
<emphasis id="5DB1A851D01AFFE9FF315CB1F265FA81" bold="true" box="[199,315,1268,1293]" pageId="7" pageNumber="408">Remarks.</emphasis>
This species is difficult to diagnose because of its similarity to 
<emphasis id="5DB1A851D01AFFE9FBE85CB0F7C2FA80" box="[1054,1180,1269,1292]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="A8C50FC0D01AFFE9FBE85CB0F7C6FA80" box="[1054,1176,1269,1292]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaianii">M. gaianii</taxonomicName>
.
</emphasis>
R. W. Garrison (
<emphasis id="5DB1A851D01AFFE9FA925CB0F3BFFABC" italics="true" pageId="7" pageNumber="408">pers. comm.</emphasis>
) suggested, mainly based on diagnostic traits of the males (
<figureCitation id="F7FE68C6D01AFFE9FC565D52F0AFFABC" box="[928,1009,1303,1328]" captionStart="FIGURE 7" captionStartId="22.[151,250,1817,1839]" captionTargetBox="[328,1261,193,1765]" captionTargetId="figure@22.[320,1262,193,1771]" captionTargetPageId="22" captionText="FIGURE 7. Mesamphiagrion gaianii (Paratype, Venezuela, Trujillo, Páramo La Cristalina) vs. M. gaudiimontanum (Antioquia, Belmira) (traits of M. gaianii on left): (a) Lateral view of caudal appendages; (b) Posterior view of left cercus; (c) Medio-dorsal view of cercus; (d) Ectal view of genital ligula; (e) M. gaudiimontanum lateral view of genital ligula; Female paratype of M. gaudiimontanum: (f) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Inner process. Illustrations drawn by: Rosser Garrison (a to d), Gloria Mora (e), and Natalia Uribe (f)." httpUri="https://zenodo.org/record/246835/files/figure.png" pageId="7" pageNumber="408">Figs. 7</figureCitation>
a–d), that populations of this species found in the northern Cordillera Central could correspond to a population of 
<taxonomicName id="A8C50FC0D01AFFE9FBE25D78F7D5FAD8" box="[1044,1163,1341,1364]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaianii">
<emphasis id="5DB1A851D01AFFE9FBE25D78F7D5FAD8" box="[1044,1163,1341,1364]" italics="true" pageId="7" pageNumber="408">M. gaianii</emphasis>
</taxonomicName>
with certain variations. However, we found subtle differences in these characters that are preserved in the Santa Inés populations, as well as strong differences in the females, which lead us to propose the hypothesis that populations present in this part of 
<collectingCountry id="17D234D3D01AFFE9FF615DE2F25BFA4C" box="[151,261,1447,1472]" name="Colombia" pageId="7" pageNumber="408">Colombia</collectingCountry>
are a lineage whose evolutionary history is sufficiently different from 
<taxonomicName id="A8C50FC0D01AFFE9FB875DECF7B3FA4C" box="[1137,1261,1449,1472]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaianii">
<emphasis id="5DB1A851D01AFFE9FB875DECF7B3FA4C" box="[1137,1261,1449,1472]" italics="true" pageId="7" pageNumber="408">M. gaianii</emphasis>
</taxonomicName>
as to provide reproductive isolation, not only due to geographical and ecological separation but also to variation of several morphological traits directly involved in mating, and therefore they can be considered different species.
</paragraph>
<paragraph id="6F7A7443D01AFFE9FF315E51F074F9DC" blockId="7.[151,1437,151,1616]" pageId="7" pageNumber="408">
Color polymorphism was observed in females of this species (
<figureCitation id="F7FE68C6D01AFFE9FC725E51F095F9A1" box="[900,971,1556,1581]" captionStart="FIGURE 6" captionStartId="21.[151,250,1942,1964]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@21.[204,1382,193,1921]" captionTargetPageId="21" captionText="FIGURE 6. Ontogenic changes in both morphs of Mesamphiagrion gaudiimontanum females (Antioquia, Belmira): Androchrome: (a) Juvenile; (b) Mature. Gynochrome: (c) Juvenile parasitized by mites (Photograph by Cintia Moreno); (d) Mature." httpUri="https://zenodo.org/record/246834/files/figure.png" pageId="7" pageNumber="408">Figs 6</figureCitation>
a–d), with gynochrome and androchrome individuals found at a ratio of approximately 3:1 (
<tableCitation id="224741F8D01AFFE9FD335E72F046F9DC" box="[709,792,1591,1616]" captionStart="TABLE 1" captionStartId="7.[151,239,1660,1683]" captionText="TABLE 1. Number of examined females of M. gaudiimontanum." pageId="7" pageNumber="408">Table 1</tableCitation>
).
</paragraph>
<caption id="3BBA24CBD01AFFE9FF615E39F011F91F" box="[151,847,1659,1683]" pageId="7" pageNumber="408">
<paragraph id="6F7A7443D01AFFE9FF615E39F011F91F" blockId="7.[151,847,1659,1683]" box="[151,847,1659,1683]" pageId="7" pageNumber="408">
<emphasis id="5DB1A851D01AFFE9FF615E39F254F91F" bold="true" box="[151,266,1660,1683]" pageId="7" pageNumber="408">TABLE 1.</emphasis>
Number of examined females of 
<taxonomicName id="A8C50FC0D01AFFE9FD865E39F017F91F" box="[624,841,1659,1683]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="gaudiimontanum">
<emphasis id="5DB1A851D01AFFE9FD865E39F017F91F" box="[624,841,1659,1683]" italics="true" pageId="7" pageNumber="408">M. gaudiimontanum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="6F7A7443D01AFFE9FF695EEBF7DFF962" blockId="7.[159,1421,1710,1774]" pageId="7" pageNumber="408">
Androchrome teneral females Gynochrome teneral females Androchrome mature females Gynochrome mature females 3 
<date id="1B7B5283D01AFFE9FE145E9CF7DFF962" box="[482,1153,1752,1774]" pageId="7" pageNumber="408" value="2010-04-07">7 4 10</date>
</paragraph>
<paragraph id="6F7A7443D01AFFE9FF315F6BF27FF85B" blockId="7.[151,1436,1838,2007]" pageId="7" pageNumber="408">
<emphasis id="5DB1A851D01AFFE9FF315F6BF293F8CB" bold="true" box="[199,461,1838,1863]" pageId="7" pageNumber="408">Habitat and biology.</emphasis>
Adults inhabit páramos (high mountain wet tropical ecosystems unique to the Neotropics) and montane forest edges dominated by the oak 
<taxonomicName id="A8C50FC0D01AFFE9FCA25F16F772F8E7" box="[852,1068,1874,1899]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="408" phylum="Tracheophyta" rank="species" species="humboldti">
<emphasis id="5DB1A851D01AFFE9FCA25F16F772F8E7" box="[852,1068,1874,1899]" italics="true" pageId="7" pageNumber="408">Quercus humboldti</emphasis>
</taxonomicName>
. Mature males were found near peatlands or lakes, immatures and females from the edge of the oak forest to open areas. Larvae were collected in ponds, peatlands, and small pools with 
<emphasis id="5DB1A851D01AFFE9FD8C5FDFF1A9F83F" box="[634,759,1946,1971]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="A8C50FC0D01AFFE9FD8C5FDFF1AAF83F" box="[634,756,1946,1971]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="7" pageNumber="408" phylum="Bryophyta" rank="genus">Sphagnum</taxonomicName>
.
</emphasis>
Larvae in captivity were observed using 
<taxonomicName id="A8C50FC0D01AFFE9FB015FDFF633F83F" box="[1271,1389,1946,1971]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="7" pageNumber="408" phylum="Bryophyta" rank="genus">
<emphasis id="5DB1A851D01AFFE9FB015FDFF633F83F" box="[1271,1389,1946,1971]" italics="true" pageId="7" pageNumber="408">Sphagnum</emphasis>
</taxonomicName>
for camouflage.
</paragraph>
<paragraph id="6F7A7443D015FFE6FF3158DDF1D9FE2A" blockId="8.[151,1437,151,916]" pageId="8" pageNumber="409">
Mature males were found close to peatlands perched on vegetation and 
<taxonomicName id="A8C50FC0D015FFE6FC0158D2F733FF3C" box="[1015,1133,151,176]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="8" pageNumber="409" phylum="Bryophyta" rank="genus">
<emphasis id="5DB1A851D015FFE6FC0158D2F733FF3C" box="[1015,1133,151,176]" italics="true" pageId="8" pageNumber="409">Sphagnum</emphasis>
</taxonomicName>
, usually in great numbers. Fights between males were common at sunny times, sometimes, after which males perched with their wings open at about 45 degrees (
<figureCitation id="F7FE68C6D015FFE6FE70589BF294FF7B" box="[390,458,222,247]" captionStart="FIGURE 5" captionStartId="20.[151,250,1906,1928]" captionTargetBox="[181,1405,429,1883]" captionTargetId="figure@20.[181,1405,427,1885]" captionTargetPageId="20" captionText="FIGURE 5. Ontogenic changes in Mesamphiagrion gaudiimontanum males (Antioquia, Belmira): (a) Teneral; (b) Mature; (c) Middle age parasitized by mites (Photograph by Camilo Flórez-V.)." httpUri="https://zenodo.org/record/246833/files/figure.png" pageId="8" pageNumber="409">Fig. 5</figureCitation>
b), while sometimes bobbing their abdomen up to the level of thorax and down again repeatedly, a behavior also observed in 
<emphasis id="5DB1A851D015FFE6FDAF5947F1AFFE95" box="[601,753,258,281]" italics="true" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FDAF5947F1B2FE95" box="[601,748,258,281]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="tamaense">M. tamaense</taxonomicName>
.
</emphasis>
Machado (2012) suggested blue pterostigmata could have a territorial defense function, nevertheless field studies will need to be done in order to determine if this hypothesis has merit (Corbet 1999). Some fights between males were observed, in which they chased each other until one or both left the area. It was also common to find two males on the same or a nearby perch without fighting. Females were observed both far from and near water.
</paragraph>
<paragraph id="6F7A7443D015FFE6FF3159F4F61CFDDB" blockId="8.[151,1437,151,916]" pageId="8" pageNumber="409">
Alejandra Clavijo (
<emphasis id="5DB1A851D015FFE6FE5459F7F172FE45" box="[418,556,434,457]" italics="true" pageId="8" pageNumber="409">pers. comm.</emphasis>
) observed some pairs in copula, around noon, in sunny weather in 
<date id="1B7B5283D015FFE6FADB59F4F6C2FE46" box="[1325,1436,433,458]" pageId="8" pageNumber="409" value="2012-07">July 2012</date>
near a small stream with 
<taxonomicName id="A8C50FC0D015FFE6FE475991F179FE61" box="[433,551,468,493]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="8" pageNumber="409" phylum="Bryophyta" rank="genus">
<emphasis id="5DB1A851D015FFE6FE475991F179FE61" box="[433,551,468,493]" italics="true" pageId="8" pageNumber="409">Sphagnum</emphasis>
</taxonomicName>
; all females were gynochrome. On one occasion, CAB observed a male trying to capture a gynochrome female on a sunny morning, both perched on 
<taxonomicName id="A8C50FC0D015FFE6FC4A59B2F76CFD9C" box="[956,1074,503,528]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="8" pageNumber="409" phylum="Bryophyta" rank="genus">
<emphasis id="5DB1A851D015FFE6FC4A59B2F76CFD9C" box="[956,1074,503,528]" italics="true" pageId="8" pageNumber="409">Sphagnum</emphasis>
</taxonomicName>
; the male was performing short flights toward the female, and she was avoiding him until finally the female flew away. An androchrome female with wings quite deteriorated was collected with S7–10 clearly muddy, which indicated it was laying eggs.
</paragraph>
<paragraph id="6F7A7443D015FFE6FF315A24F6C2FD4C" blockId="8.[151,1437,151,916]" pageId="8" pageNumber="409">
Specimens perched on vegetation with rosette-like leaves, including frailejon (
<taxonomicName id="A8C50FC0D015FFE6FBC15A24F7C1FDF6" box="[1079,1183,609,634]" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="409" phylum="Tracheophyta" rank="genus">
<emphasis id="5DB1A851D015FFE6FBC15A24F7C1FDF6" box="[1079,1183,609,634]" italics="true" pageId="8" pageNumber="409">Espeletia</emphasis>
</taxonomicName>
sp.), or hedgehog-like leaves as some grasses. When disturbed, individuals often dropped into the center of the plant or tried to hind behind a leaf (also observed on 
<taxonomicName id="A8C50FC0D015FFE6FE0E5AECF12AFD4C" box="[504,628,679,704]" class="Insecta" family="Coenagrionidae" genus="Acanthagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="laterale">
<emphasis id="5DB1A851D015FFE6FE0E5AECF12AFD4C" box="[504,628,679,704]" italics="true" pageId="8" pageNumber="409">M. laterale</emphasis>
</taxonomicName>
and 
<taxonomicName id="A8C50FC0D015FFE6FD5C5AECF067FD4C" box="[682,825,681,704]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="tamaense">
<emphasis id="5DB1A851D015FFE6FD5C5AECF067FD4C" box="[682,825,681,704]" italics="true" pageId="8" pageNumber="409">M. tamaense</emphasis>
</taxonomicName>
). Some adults were parasitized by mites (
<figureCitation id="F7FE68C6D015FFE6FAFC5AE2F606FD4C" box="[1290,1368,679,704]" captionStart="FIGURE 5" captionStartId="20.[151,250,1906,1928]" captionTargetBox="[181,1405,429,1883]" captionTargetId="figure@20.[181,1405,427,1885]" captionTargetPageId="20" captionText="FIGURE 5. Ontogenic changes in Mesamphiagrion gaudiimontanum males (Antioquia, Belmira): (a) Teneral; (b) Mature; (c) Middle age parasitized by mites (Photograph by Camilo Flórez-V.)." httpUri="https://zenodo.org/record/246833/files/figure.png" pageId="8" pageNumber="409">Figs. 5</figureCitation>
c, 6c).
</paragraph>
<paragraph id="6F7A7443D015FFE6FF315A8EF6C3FCC1" blockId="8.[151,1437,151,916]" pageId="8" pageNumber="409">
This species co-occurs with the odonate species listed in 
<tableCitation id="224741F8D015FFE6FCBE5A8EF0C5FD6F" box="[840,923,715,740]" captionStart="TABLE 2" captionStartId="8.[151,239,959,982]" captionText="TABLE 2. Odonata collected with M. gaudiimontanum and M. santainense." pageId="8" pageNumber="409">Table 2</tableCitation>
, plus 
<emphasis id="5DB1A851D015FFE6FC165A89F61AFD6F" box="[992,1348,716,739]" italics="true" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FC165A89F7D5FD6F" box="[992,1163,716,739]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="santainense">M. santainense</taxonomicName>
; 
<taxonomicName id="A8C50FC0D015FFE6FB6C5A89F61AFD6F" box="[1178,1348,716,739]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="santainense">M. santainense</taxonomicName>
</emphasis>
has not been recorded at 
<taxonomicName id="A8C50FC0D015FFE6FEAB5AABF2ECFC8A" box="[349,434,750,774]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
ecosystems. Other odonates collected here were new species of 
<emphasis id="5DB1A851D015FFE6FB6E5AABF6C2FC8B" box="[1176,1436,750,775]" italics="true" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FB6E5AABF7A0FC8B" box="[1176,1278,750,775]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">Ischnura</taxonomicName>
, 
<taxonomicName id="A8C50FC0D015FFE6FAFA5AABF6C6FC8B" box="[1292,1432,750,775]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">Oxyallagma</taxonomicName>
,
</emphasis>
and 
<emphasis id="5DB1A851D015FFE6FF3C5B54F22EFCA6" box="[202,368,785,810]" italics="true" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FF3C5B54F232FCA6" box="[202,364,785,810]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">Rhionaeschna</taxonomicName>
,
</emphasis>
the descriptions of which are in preparation by the authors. 
<taxonomicName id="A8C50FC0D015FFE6FBC95B54F796FCA6" box="[1087,1224,785,810]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="5DB1A851D015FFE6FBC95B54F796FCA6" box="[1087,1224,785,810]" italics="true" pageId="8" pageNumber="409">Oxyallagma</emphasis>
</taxonomicName>
has thus far been recorded only from 
<collectingCountry id="17D234D3D015FFE6FE805B70F2F4FCC0" box="[374,426,821,844]" name="Peru" pageId="8" pageNumber="409">Peru</collectingCountry>
and 
<collectingCountry id="17D234D3D015FFE6FE165B71F163FCC1" box="[480,573,820,845]" name="Ecuador" pageId="8" pageNumber="409">Ecuador</collectingCountry>
(Garrison 
<emphasis id="5DB1A851D015FFE6FD405B73F1B0FCC1" box="[694,750,820,845]" italics="true" pageId="8" pageNumber="409">et al.</emphasis>
2010) and 
<taxonomicName id="A8C50FC0D015FFE6FC9B5B70F717FCC1" box="[877,1097,821,845]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="paramo">
<emphasis id="5DB1A851D015FFE6FC9B5B70F717FCC1" box="[877,1097,821,845]" italics="true" pageId="8" pageNumber="409">Sympetrum paramo</emphasis>
</taxonomicName>
thus far only from 
<collectingCountry id="17D234D3D015FFE6FAD45B71F6C7FCC1" box="[1314,1433,820,845]" name="Venezuela" pageId="8" pageNumber="409">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="27DF27C8D015FFE6FF315B12F0DCFBBE" pageId="8" pageNumber="409" type="distribution">
<paragraph id="6F7A7443D015FFE6FF315B12F726FC18" blockId="8.[151,1437,151,916]" pageId="8" pageNumber="409">
<emphasis id="5DB1A851D015FFE6FF315B12F23EFCFC" bold="true" box="[199,352,855,880]" pageId="8" pageNumber="409">Distribution.</emphasis>
Known only from the mountain range of Santa Inés, and Las Badías in the northwestern Cordillera Central, Department of Antioquia, between 2,700 and 
<quantity id="A83DD9A6D015FFE6FC875B3EF08AFC18" box="[881,980,891,916]" metricMagnitude="0" metricUnit="m" metricValue="3.27" pageId="8" pageNumber="409" unit="m" value="3.27">3,270 m</quantity>
.a.s.l. (
<figureCitation id="F7FE68C6D015FFE6FBE15B3EF734FC18" box="[1047,1130,891,916]" captionStart="FIGURE 16" captionStartId="36.[151,250,1359,1381]" captionTargetBox="[151,1436,193,1337]" captionTargetId="figure@36.[151,1436,193,1338]" captionTargetPageId="36" captionText="FIGURE 16. New distributional records of Mesamphiagrion demarmelsi, M. gaudiimontanum, M. ovigerum and M. risi in Colombia." httpUri="https://zenodo.org/record/246844/files/figure.png" pageId="8" pageNumber="409">Fig. 16</figureCitation>
).
</paragraph>
<caption id="3BBA24CBD015FFE6FF615BFAF09DFC5B" box="[151,963,958,983]" pageId="8" pageNumber="409">
<paragraph id="6F7A7443D015FFE6FF615BFAF09DFC5B" blockId="8.[151,963,958,983]" box="[151,963,958,983]" pageId="8" pageNumber="409">
<emphasis id="5DB1A851D015FFE6FF615BFAF256FC5A" bold="true" box="[151,264,959,982]" pageId="8" pageNumber="409">TABLE 2.</emphasis>
<taxonomicName id="A8C50FC0D015FFE6FEE75BFAF235FC5A" box="[273,363,959,982]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
collected with 
<taxonomicName id="A8C50FC0D015FFE6FDF85B85F1B9FC5A" box="[526,743,958,983]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="gaudiimontanum">
<emphasis id="5DB1A851D015FFE6FDF85B85F1B9FC5A" box="[526,743,958,983]" italics="true" pageId="8" pageNumber="409">M. gaudiimontanum</emphasis>
</taxonomicName>
and 
<taxonomicName id="A8C50FC0D015FFE6FCEC5B85F09EFC5B" box="[794,960,959,983]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="santainense">
<emphasis id="5DB1A851D015FFE6FCEC5B85F06EFC5B" box="[794,816,960,983]" italics="true" pageId="8" pageNumber="409">M</emphasis>
. santainense
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="6F7A7443D015FFE6FF695BB7F009FB84" blockId="8.[159,1203,1010,1404]" box="[159,855,1010,1032]" pageId="8" pageNumber="409">Family Genus Species</paragraph>
<paragraph id="6F7A7443D015FFE6FF695C59F0DCFBBE" blockId="8.[159,1203,1010,1404]" box="[159,898,1052,1074]" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FF695C59F247FBBE" box="[159,281,1052,1074]" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
<emphasis id="5DB1A851D015FFE6FE195C59F07EFBBE" box="[495,800,1052,1074]" italics="true" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FE195C59F122FBBE" box="[495,636,1052,1074]" class="Insecta" family="Libellulidae" genus="Erythrodiplax" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">Erythrodiplax</taxonomicName>
<taxonomicName id="A8C50FC0D015FFE6FCFC5C58F07EFBBE" box="[778,800,1053,1074]" class="Insecta" family="Libellulidae" genus="Erythrodiplax" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="cauca">E.</taxonomicName>
</emphasis>
cf. 
<emphasis id="5DB1A851D015FFE6FCB05C58F0DCFBBE" box="[838,898,1053,1074]" italics="true" pageId="8" pageNumber="409">cauca</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="27DF27C8D015FFE6FE195C03F1A9FA3F" pageId="8" pageNumber="409" type="description">
<paragraph id="6F7A7443D015FFE6FE195C03F0EBFBD7" blockId="8.[159,1203,1010,1404]" box="[495,949,1093,1115]" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FE195C03F13FFBD7" box="[495,609,1094,1115]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="5DB1A851D015FFE6FE195C03F13FFBD7" box="[495,609,1094,1115]" italics="true" pageId="8" pageNumber="409">Sympetrum</emphasis>
</taxonomicName>
<taxonomicName id="A8C50FC0D015FFE6FCFC5C03F0EBFBD7" authority="(Selys)" authorityName="Selys" baseAuthorityName="Selys" box="[778,949,1093,1115]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="gilvum">
<emphasis id="5DB1A851D015FFE6FCFC5C03F036FBD7" box="[778,872,1093,1115]" italics="true" pageId="8" pageNumber="409">S. gilvum</emphasis>
(Selys)
</taxonomicName>
</paragraph>
<paragraph id="6F7A7443D015FFE6FCFC5C2AF0A2FB08" blockId="8.[159,1203,1010,1404]" box="[778,1020,1134,1156]" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FCFC5C2AF0ACFB08" authority="DeMarmels" authorityName="DeMarmels" box="[778,1010,1134,1156]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="paramo">
<emphasis id="5DB1A851D015FFE6FCFC5C2AF02CFB08" box="[778,882,1135,1156]" italics="true" pageId="8" pageNumber="409">S. paramo</emphasis>
DeMarmels
</taxonomicName>
*
</paragraph>
<paragraph id="6F7A7443D015FFE6FF695CD2F094FB21" blockId="8.[159,1203,1010,1404]" box="[159,970,1175,1198]" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FF695CD2F234FB21" box="[159,362,1175,1197]" class="Insecta" family="Megapodagrionidae" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="family">Megapodagrionidae</taxonomicName>
<taxonomicName id="A8C50FC0D015FFE6FE195CD2F1CEFB21" box="[495,656,1175,1197]" class="Insecta" family="Megapodagrionidae" genus="Teinopodagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="5DB1A851D015FFE6FE195CD2F1CEFB21" box="[495,656,1175,1197]" italics="true" pageId="8" pageNumber="409">Teinopodagrion</emphasis>
</taxonomicName>
<taxonomicName id="A8C50FC0D015FFE6FCFC5CDCF094FB21" authority="(Selys)" authorityName="Selys" baseAuthorityName="Selys" box="[778,970,1175,1198]" class="Insecta" family="Megapodagrionidae" genus="Teinopodagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="oscillans">
<emphasis id="5DB1A851D015FFE6FCFC5CDCF022FB21" box="[778,892,1175,1198]" italics="true" pageId="8" pageNumber="409">T. oscillans</emphasis>
(Selys)
</taxonomicName>
</paragraph>
<paragraph id="6F7A7443D015FFE6FF695C84F0E4FA8C" blockId="8.[159,1203,1010,1404]" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FF695C84F261FB5B" box="[159,319,1217,1239]" class="Insecta" family="Coenagrionidae" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="family">Coenagrionidae</taxonomicName>
<emphasis id="5DB1A851D015FFE6FE195C84F0D8FB5B" box="[495,902,1217,1239]" italics="true" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FE195C84F135FB5B" box="[495,619,1217,1239]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">Oxyallagma</taxonomicName>
<taxonomicName id="A8C50FC0D015FFE6FCFC5C84F0D8FB5B" box="[778,902,1217,1239]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus" status="sp. nov.">Oxyallagma</taxonomicName>
</emphasis>
<emphasis id="5DB1A851D015FFE6FC7A5C84F082FB5B" bold="true" box="[908,988,1217,1239]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="4682152AD015FFE6FC7A5C84F082FB5B" box="[908,988,1217,1239]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="5DB1A851D015FFE6FE195CAFF03AFA8C" box="[495,868,1258,1280]" italics="true" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FE195CAFF117FA8C" box="[495,585,1258,1280]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">Ischnura</taxonomicName>
<taxonomicName id="A8C50FC0D015FFE6FCFC5CAFF03AFA8C" box="[778,868,1258,1280]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus" status="sp. nov.">Ischnura</taxonomicName>
</emphasis>
<emphasis id="5DB1A851D015FFE6FC9C5CAFF0E4FA8C" bold="true" box="[874,954,1258,1280]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="4682152AD015FFE6FC9C5CAFF0E4FA8C" box="[874,954,1258,1280]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="6F7A7443D015FFE6FF695D51F7EDFAA6" blockId="8.[159,1203,1010,1404]" box="[159,1203,1300,1322]" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FF695D51F254FAA6" box="[159,266,1300,1322]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="family">Aeshnidae</taxonomicName>
<emphasis id="5DB1A851D015FFE6FE195D51F0C9FAA6" box="[495,919,1300,1322]" italics="true" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FE195D51F121FAA6" box="[495,639,1300,1322]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">Rhionaeschna</taxonomicName>
<taxonomicName id="A8C50FC0D015FFE6FCFC5D50F0C9FAA6" box="[778,919,1300,1322]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="brevicercia">R. brevicercia</taxonomicName>
</emphasis>
(Muzón &amp; von Ellenrieder)
</paragraph>
<paragraph id="6F7A7443D015FFE6FCFC5D7BF0DEFADF" blockId="8.[159,1203,1010,1404]" box="[778,896,1341,1363]" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FCFC5D7BF0DEFADF" box="[778,896,1341,1363]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="marchali">R. marchali</taxonomicName>
</paragraph>
<paragraph id="6F7A7443D015FFE6FCFC5D23F0A2FAF0" blockId="8.[159,1203,1010,1404]" box="[778,1020,1382,1404]" pageId="8" pageNumber="409">
<taxonomicName id="A8C50FC0D015FFE6FCFC5D23F0C7FAF0" box="[778,921,1382,1404]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="5DB1A851D015FFE6FCFC5D23F0C7FAF0" box="[778,921,1382,1404]" italics="true" pageId="8" pageNumber="409">Rhionaeschna</emphasis>
</taxonomicName>
<emphasis id="5DB1A851D015FFE6FC695D23F0AEFAF0" bold="true" box="[927,1008,1382,1404]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="4682152AD015FFE6FC695D23F0AEFAF0" box="[927,1008,1382,1404]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
*
</paragraph>
<paragraph id="6F7A7443D015FFE6FF615DD9F1A9FA3F" blockId="8.[151,759,1436,1460]" box="[151,759,1436,1460]" pageId="8" pageNumber="409">
*Not collected with 
<taxonomicName id="A8C50FC0D015FFE6FE865DD8F14FFA38" box="[368,529,1437,1460]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="santainense">
<emphasis id="5DB1A851D015FFE6FE865DD8F14FFA38" box="[368,529,1437,1460]" italics="true" pageId="8" pageNumber="409">M. santainense</emphasis>
</taxonomicName>
(
<taxonomicName id="A8C50FC0D015FFE6FDE95DD9F12CFA3F" box="[543,626,1436,1459]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
inhabitans).
</paragraph>
</subSubSection>
</treatment>
</document>