315 lines
36 KiB
XML
315 lines
36 KiB
XML
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<mods:title id="7BA553F4F065BDDBB80E9BE06A61901E">A revision of African helmeted terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species</mods:title>
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<mods:affiliation id="1279E8EDEF09C8A08FEDF12A91496F5D">Department of Herpetology, Port Elizabeth Museum, P. O. Box 13147, Humewood 6013, South Africa & Department of Zoology, P. O. Box 77000, Nelson Mandela Metropolitan University, Port Elizabeth 6031, South Africa</mods:affiliation>
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<mods:affiliation id="088E3CAEAF8B90B5D3A824922E476BC3">Unit for Environmental Science & Management, North-West University, Potchefstroom Campus, Private Bag X 6001, Potchefstroom 2520, South Africa</mods:affiliation>
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<mods:affiliation id="7D9B55D1E0E39E33477B81A37FA7302B">Chelonian Biodiversity and Conservation, Department of Biodiversity and Conservation Biology, University of the Western Cape, Private Bag X 17, Bellville 7535, South Africa</mods:affiliation>
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<mods:namePart id="B4F43605B99754B44C7698D3A960AACC">Schleicher, Alfred</mods:namePart>
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<mods:affiliation id="907C3768D04C7322C686F11BAC0CB2E5">P. O. Box 30566, Windhoek, Namibia</mods:affiliation>
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<mods:namePart id="38F0174F755A198DAE38C14B8D629329">Široký, Pavel</mods:namePart>
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<mods:affiliation id="CB2574C0DC0D18D85D2C8E1BC07B2BD7">Department of Biology and Wildlife Diseases, Faculty of Veterinary Hygiene and Ecology, University of Veterinary and Pharmaceutical Sciences, Palackého tř. 1 / 3, 612 42 Brno, Czech Republic</mods:affiliation>
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<mods:namePart id="EDB5F5495B68013F30B9DE46F8017B35">Fritz, Uwe</mods:namePart>
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<mods:affiliation id="A56759B9B46131783DA3428E25486E94">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:date id="6F68447856571F98DB60637BBAE91A81">2014</mods:date>
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<treatment id="CB02879FF93CFFCCFF74F97FFF3DFAEA" ID-DOI="http://doi.org/10.5281/zenodo.5082955" ID-GBIF-Taxon="183403855" ID-Zenodo-Dep="5082955" LSID="urn:lsid:plazi:treatment:CB02879FF93CFFCCFF74F97FFF3DFAEA" httpUri="http://treatment.plazi.org/id/CB02879FF93CFFCCFF74F97FFF3DFAEA" lastPageId="12" lastPageNumber="535" pageId="11" pageNumber="534">
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<subSubSection id="0BB16502F93CFFCBFF74F97FFD98F95D" box="[151,593,1735,1762]" pageId="11" pageNumber="534" type="nomenclature">
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<paragraph id="43143689F93CFFCBFF74F97FFD98F95D" blockId="11.[151,593,1735,1762]" box="[151,593,1735,1762]" pageId="11" pageNumber="534">
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<taxonomicName id="84AB4D0AF93CFFCBFF74F97FFD98F95D" authority="(Ruppell, 1835)" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[151,593,1735,1762]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="534" phylum="Chordata" rank="species" species="gehafie">
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<emphasis id="71DFEA9BF93CFFCBFF74F97FFD98F95D" bold="true" box="[151,593,1735,1762]" pageId="11" pageNumber="534">
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<emphasis id="71DFEA9BF93CFFCBFF74F97FFE43F95E" bold="true" box="[151,394,1735,1761]" italics="true" pageId="11" pageNumber="534">Pelomedusa gehafie</emphasis>
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(
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<bibRefCitation id="273A4B78F93CFFCBFE7AF970FD80F95D" author="Ruppell, E." box="[409,585,1736,1762]" pageId="11" pageNumber="534" refId="ref17558" refString="Ruppell, E. (1835) Neue Wirbelthiere zu der Fauna von Abyssinien gehorig. Amphibien. Siegmund Schmerber, Frankfurt am Main, 18 pp., 6 plates. http: // dx. doi. org / 10.5962 / bhl. title. 53778" type="book" year="1835">Rüppell, 1835</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="0BB16502F93CFFCBFF74F8B6FD47F81A" pageId="11" pageNumber="534" type="reference_group">
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<paragraph id="43143689F93CFFCBFF74F8B6FBBBF8FA" blockId="11.[151,1423,1806,1957]" pageId="11" pageNumber="534">
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1835
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<taxonomicName id="84AB4D0AF93CFFCBFF02F8B6FE3AF89A" authority="Ruppell" authorityName="Ruppell" box="[225,499,1806,1829]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurodira" pageId="11" pageNumber="534" phylum="Chordata" rank="species" species="gehafie">
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<emphasis id="71DFEA9BF93CFFCBFF02F8B6FE50F89A" box="[225,409,1806,1829]" italics="true" pageId="11" pageNumber="534">Pentonyx gehafie</emphasis>
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Rüppell
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</taxonomicName>
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—Type locality: eastern slope of coastal mountains,
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<collectingCountry id="3BBC7619F93CFFCBFBC0F8B6FBA6F89A" box="[1059,1135,1806,1829]" name="Eritrea" pageId="11" pageNumber="534">Eritrea</collectingCountry>
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;
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<typeStatus id="9C10882BF93CFFCBFB99F8B6FB15F89A" box="[1146,1244,1806,1829]" pageId="11" pageNumber="534" type="lectotype">lectotype</typeStatus>
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(
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<bibRefCitation id="273A4B78F93CFFCBFB0AF8B6FA4AF89A" author="Mertens, R." box="[1257,1411,1806,1829]" pageId="11" pageNumber="534" pagination="139 - 142" refId="ref17133" refString="Mertens, R. (1937) Bemerkungen uber die Rassen von Pelomedusa subrufa (La Cepede). Zoologischer Anzeiger, 117, 139 - 142." type="journal article" year="1937">Mertens 1937</bibRefCitation>
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): Senckenberg-Museum, Frankfurt am Main, SMF 7947 (
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<figureCitation id="DB902A0CF93CFFCBFCD6F896FCBDF8FA" box="[821,884,1838,1861]" captionStart="FIGURE 5" captionStartId="18.[151,250,1918,1940]" captionTargetBox="[226,1382,244,1853]" captionTargetId="figure-68@18.[151,1436,145,1884]" captionTargetPageId="18" captionText="FIGURE 5. Dorsal and ventral aspects of the holotype of Pelomedusa somalica sp. nov. (BMNH 1970.1481, adult, Borama district, Awdal, Somaliland/Somalia; top) and of Pelomedusa variabilis sp. nov. (SMF 58075, subadult male, Gold Coast, Ghana; bottom). Scale bars, 3 cm. Photos: A. Petzold and E. Morawa." figureDoi="http://doi.org/10.5281/zenodo.4915095" httpUri="https://zenodo.org/record/4915095/files/figure.png" pageId="11" pageNumber="534">Fig. 5</figureCitation>
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top in
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<bibRefCitation id="273A4B78F93CFFCBFC5CF896FBA2F8FA" author="Fritz, U. & Petzold, A. & Kehlmaier, C. & Kindler, C. & Campbell, P. & Hofmeyr, M. D. & Branch, W. R." box="[959,1131,1838,1861]" pageId="11" pageNumber="534" pagination="501 - 522" refId="ref16261" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. & Branch, W. R. (2014) Disentangling the Pelomedusa complex using type specimens and historical DNA. Zootaxa, 3795 (5), 501 - 522 http: // dx. doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
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Fritz
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||
<emphasis id="71DFEA9BF93CFFCBFC1BF896FBE7F8FA" box="[1016,1070,1838,1861]" italics="true" pageId="11" pageNumber="534">et al.</emphasis>
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2014
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</bibRefCitation>
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)
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</paragraph>
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<paragraph id="43143689F93CFFCBFF74F8F6FD47F81A" blockId="11.[151,1423,1806,1957]" pageId="11" pageNumber="534">
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1910
|
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<taxonomicName id="84AB4D0AF93CFFCBFF02F8F6FCF7F8DA" authority="Vaillant & Grandidier" authorityName="Vaillant & Grandidier" authorityYear="1910" box="[225,830,1870,1893]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="534" phylum="Chordata" rank="variety" species="galeata" variety="disjuncta">
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||
<emphasis id="71DFEA9BF93CFFCBFF02F8F6FE73F8DA" box="[225,442,1870,1893]" italics="true" pageId="11" pageNumber="534">Pelomedusa galeata</emphasis>
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var.
|
||
<emphasis id="71DFEA9BF93CFFCBFE0DF8F6FD99F8DA" box="[494,592,1870,1893]" italics="true" pageId="11" pageNumber="534">disjuncta</emphasis>
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||
Vaillant & Grandidier
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||
</taxonomicName>
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||
—Restricted type locality (by
|
||
<typeStatus id="9C10882BF93CFFCBFB61F8F6FB2DF8DA" box="[1154,1252,1870,1893]" pageId="11" pageNumber="534" type="lectotype">lectotype</typeStatus>
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designation,
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||
<bibRefCitation id="273A4B78F93CFFCBFF02F8D6FE58F83A" author="Fritz, U. & Petzold, A. & Kehlmaier, C. & Kindler, C. & Campbell, P. & Hofmeyr, M. D. & Branch, W. R." box="[225,401,1902,1925]" pageId="11" pageNumber="534" pagination="501 - 522" refId="ref16261" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. & Branch, W. R. (2014) Disentangling the Pelomedusa complex using type specimens and historical DNA. Zootaxa, 3795 (5), 501 - 522 http: // dx. doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
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Fritz
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||
<emphasis id="71DFEA9BF93CFFCBFEF9F8D6FE86F83A" box="[282,335,1902,1925]" italics="true" pageId="11" pageNumber="534">et al.</emphasis>
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2014
|
||
</bibRefCitation>
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): eastern slope of coastal mountains,
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<collectingCountry id="3BBC7619F93CFFCBFCFEF8D6FCA0F83A" box="[797,873,1902,1925]" name="Eritrea" pageId="11" pageNumber="534">Eritrea</collectingCountry>
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;
|
||
<materialsCitation id="F3C33CD4F93CFFCBFC90F8D6FD47F81A" ID-GBIF-Occurrence="3319794313" collectionCode="MNHN" pageId="11" pageNumber="534" specimenCode="MNHN 7870" specimenCount="1" typeStatus="lectotype">
|
||
<typeStatus id="9C10882BF93CFFCBFC90F8D6FC1CF83A" box="[883,981,1902,1925]" pageId="11" pageNumber="534" type="lectotype">lectotype</typeStatus>
|
||
(
|
||
<bibRefCitation id="273A4B78F93CFFCBFC00F8D6FB5DF83A" author="Fritz, U. & Petzold, A. & Kehlmaier, C. & Kindler, C. & Campbell, P. & Hofmeyr, M. D. & Branch, W. R." box="[995,1172,1902,1925]" pageId="11" pageNumber="534" pagination="501 - 522" refId="ref16261" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. & Branch, W. R. (2014) Disentangling the Pelomedusa complex using type specimens and historical DNA. Zootaxa, 3795 (5), 501 - 522 http: // dx. doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
|
||
Fritz
|
||
<emphasis id="71DFEA9BF93CFFCBFBFDF8D6FB9DF83A" box="[1054,1108,1902,1925]" italics="true" pageId="11" pageNumber="534">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
): Muséum National d’Histoire naturelle, Paris,
|
||
<specimenCode id="130D9EF2F93CFFCBFDE3F836FD47F81A" box="[512,654,1934,1957]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="534">MNHN 7870</specimenCode>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F93BFFCCFF74FF2FFC6FFCE2" pageId="12" pageNumber="535" type="diagnosis">
|
||
<paragraph id="43143689F93BFFCCFF74FF2FFC6FFCE2" blockId="12.[151,1437,151,1365]" pageId="12" pageNumber="535">
|
||
<emphasis id="71DFEA9BF93BFFCCFF74FF2FFEDAFF0F" bold="true" box="[151,275,151,176]" pageId="12" pageNumber="535">Diagnosis:</emphasis>
|
||
Medium-sized, light-coloured helmeted terrapins with a known maximum straight carapacial length of
|
||
<quantity id="84539B6CF93BFFCCFF74FF04FF3CFF6C" box="[151,245,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.78" pageId="12" pageNumber="535" unit="cm" value="17.8">17.8 cm</quantity>
|
||
. Pectoral scutes triangular and in adults widely separated from plastral midseam (in hatchlings the tips of the triangular pectorals may meet at the plastral midseam). One large undivided temporal scale on each side of head. Two small barbels under chin. Carapace unpatterned light coloured. Plastron in adults completely yellow.
|
||
<taxonomicName id="84AB4D0AF93BFFCCFF74FE9FFEB7FEFF" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[151,382,295,320]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF93BFFCCFF74FE9FFEB7FEFF" box="[151,382,295,320]" italics="true" pageId="12" pageNumber="535">Pelomedusa gehafie</emphasis>
|
||
</taxonomicName>
|
||
differs from all other
|
||
<taxonomicName id="84AB4D0AF93BFFCCFD6AFE9FFCA4FEFF" box="[649,877,295,320]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="71DFEA9BF93BFFCCFD6AFE9FFCDBFEFF" box="[649,786,295,320]" italics="true" pageId="12" pageNumber="535">Pelomedusa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName id="84AB4D0AF93BFFCCFC2FFE91FB4DFEFF" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[972,1156,295,320]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="schweinfurthi">
|
||
<emphasis id="71DFEA9BF93BFFCCFC2FFE91FB4DFEFF" box="[972,1156,295,320]" italics="true" pageId="12" pageNumber="535">P. schweinfurthi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="84AB4D0AF93BFFCCFB20FE91FA81FEFF" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1219,1352,295,320]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="somalica">
|
||
<emphasis id="71DFEA9BF93BFFCCFB20FE91FA81FEFF" box="[1219,1352,295,320]" italics="true" pageId="12" pageNumber="535">P. somalica</emphasis>
|
||
</taxonomicName>
|
||
by the presence of guanine (G) instead of adenine (A) at position 256 of the 360-bp-long reference alignment of the 12S rRNA gene (Supporting Information).
|
||
<taxonomicName id="84AB4D0AF93BFFCCFDB0FED7FCF1FE37" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[595,824,367,392]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF93BFFCCFDB0FED7FCF1FE37" box="[595,824,367,392]" italics="true" pageId="12" pageNumber="535">Pelomedusa gehafie</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="84AB4D0AF93BFFCCFC37FEC9FB42FE37" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[980,1163,367,392]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="schweinfurthi">
|
||
<emphasis id="71DFEA9BF93BFFCCFC37FEC9FB42FE37" box="[980,1163,367,392]" italics="true" pageId="12" pageNumber="535">P. schweinfurthi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="84AB4D0AF93BFFCCFB24FEC9FA83FE37" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1223,1354,367,392]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="somalica">
|
||
<emphasis id="71DFEA9BF93BFFCCFB24FEC9FA83FE37" box="[1223,1354,367,392]" italics="true" pageId="12" pageNumber="535">P. somalica</emphasis>
|
||
</taxonomicName>
|
||
by the presence of thymine (T) instead of cytosine (C) at positions 125, 267, 287 and 345, by the presence of adenine (A) instead of guanine (G) at position 180, by the presence of guanine (G) instead of adenine (A) at position 223, by the presence of cytosine (C) instead of thymine (T) at position 236, and by the presence of adenine (A) instead of cytosine (C) or thymine (T) at position 268. Furthermore,
|
||
<taxonomicName id="84AB4D0AF93BFFCCFCC9FDB9FC55FDA7" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[810,924,511,536]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF93BFFCCFCC9FDB9FC55FDA7" box="[810,924,511,536]" italics="true" pageId="12" pageNumber="535">P. gehafie</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="84AB4D0AF93BFFCCFBCCFDB9FB2AFDA7" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1071,1251,511,536]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="schweinfurthi">
|
||
<emphasis id="71DFEA9BF93BFFCCFBCCFDB9FB2AFDA7" box="[1071,1251,511,536]" italics="true" pageId="12" pageNumber="535">P. schweinfurthi</emphasis>
|
||
</taxonomicName>
|
||
by the presence of cytosine (C) or guanine (G) instead of adenine (A) at position 147, by the presence of adenine (A) instead of guanine (G) at positions 148 and 297, by the presence of cytosine (C) instead of adenine (A) at positions 166, 191 and 303, by the presence of thymine (T) instead of cytosine (C) at positions 266 and 326, by the presence of cytosine (C) instead of thymine (T) at position 279, by the presence of cytosine (C) instead of a gap at position 298, and by the presence of guanine (G) instead of adenine (A) at position 305. In addition to the above mentioned characters,
|
||
<taxonomicName id="84AB4D0AF93BFFCCFEF5FD61FE4AFD4F" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[278,387,727,752]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF93BFFCCFEF5FD61FE4AFD4F" box="[278,387,727,752]" italics="true" pageId="12" pageNumber="535">P. gehafie</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="84AB4D0AF93BFFCCFDF4FD61FD5EFD4F" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[535,663,727,752]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="somalica">
|
||
<emphasis id="71DFEA9BF93BFFCCFDF4FD61FD5EFD4F" box="[535,663,727,752]" italics="true" pageId="12" pageNumber="535">P. somalica</emphasis>
|
||
</taxonomicName>
|
||
by the presence of adenine (A) instead of thymine (T) at position 94, by the presence of cytosine (C) instead of thymine (T) at positions 95, 122 and 131, by the presence of adenine (A) instead of cytosine (C) at position 123, by the presence of thymine (T) instead of cytosine (C) at position 124, and by the presence of adenine (A) instead of guanine (G) at position 332.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F93BFFCCFF24FCDFFCB6FBE7" pageId="12" pageNumber="535" type="distribution">
|
||
<paragraph id="43143689F93BFFCCFF24FCDFFCB6FBE7" blockId="12.[151,1437,151,1365]" pageId="12" pageNumber="535">
|
||
<emphasis id="71DFEA9BF93BFFCCFF24FCDFFEABFC3F" bold="true" box="[199,354,871,896]" pageId="12" pageNumber="535">Distribution:</emphasis>
|
||
Genetically verified specimens of
|
||
<taxonomicName id="84AB4D0AF93BFFCCFD04FCD1FC9CFC3F" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[743,853,871,896]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF93BFFCCFD04FCD1FC9CFC3F" box="[743,853,871,896]" italics="true" pageId="12" pageNumber="535">P. gehafie</emphasis>
|
||
</taxonomicName>
|
||
are only known from the
|
||
<typeStatus id="9C10882BF93BFFCCFB90FCD0FB6AFC3F" box="[1139,1187,872,896]" pageId="12" pageNumber="535">type</typeStatus>
|
||
locality in
|
||
<collectingCountry id="3BBC7619F93BFFCCFAC2FCDFFAA4FC3F" box="[1313,1389,871,896]" name="Eritrea" pageId="12" pageNumber="535">Eritrea</collectingCountry>
|
||
and from a second questionable site (Gebel Arary, Naturhistorisches Museum Wien, NMW 24448). The latter locality could to be identified with Jabal Karari, Omdurman near
|
||
<collectingRegion id="816FF86BF93BFFCCFCFCFC17FC5AFC77" box="[799,915,943,968]" country="Sudan" name="Al Khartoum" pageId="12" pageNumber="535">Khartoum</collectingRegion>
|
||
,
|
||
<collectingCountry id="3BBC7619F93BFFCCFC7CFC17FC21FC77" box="[927,1000,943,968]" name="Sudan" pageId="12" pageNumber="535">Sudan</collectingCountry>
|
||
, because the specimen originates from Joseph Russegger (
|
||
<date id="37151049F93BFFCCFE99FC6CFD94FC52" box="[378,605,980,1005]" pageId="12" pageNumber="535" value="1802-11-18">18 November 1802</date>
|
||
–
|
||
<date id="37151049F93BFFCCFD8FFC6CFCD9FC52" box="[620,784,980,1005]" pageId="12" pageNumber="535" value="1863-06-20">20 June 1863</date>
|
||
), who is known to have collected near
|
||
<collectingRegion id="816FF86BF93BFFCCFB11FC6CFAAEFC52" box="[1266,1383,980,1005]" country="Sudan" name="Al Khartoum" pageId="12" pageNumber="535">Khartoum</collectingRegion>
|
||
. We tentatively identify therefore Gebel Arary with Jabal Karari and show this site in our map (
|
||
<figureCitation id="DB902A0CF93BFFCCFB78FC4FFB29FBAF" box="[1179,1248,1015,1040]" captionStart="FIGURE 2" captionStartId="4.[151,250,1074,1096]" captionTargetBox="[158,1430,183,1051]" captionTargetId="figure-447@4.[151,1436,173,1057]" captionTargetPageId="4" captionText="FIGURE 2. Genetically verified records of Pelomedusa species. Question marks denote doubtful or uncertain localities." figureDoi="http://doi.org/10.5281/zenodo.4915087" httpUri="https://zenodo.org/record/4915087/files/figure.png" pageId="12" pageNumber="535">Fig. 2</figureCitation>
|
||
) with a question mark. In the Museum für Naturkunde, Berlin, there is a further genetically verified and morphologically typical specimen originating in “
|
||
<collectingCountry id="3BBC7619F93BFFCCFE53FB87FDEDFBE7" box="[432,548,1087,1112]" name="Ethiopia" pageId="12" pageNumber="535">Abyssinia</collectingCountry>
|
||
” (ZMB 15693, coll. W. Jesse).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F93BFFCCFF24FBDCFF3DFAEA" pageId="12" pageNumber="535" type="discussion">
|
||
<paragraph id="43143689F93BFFCCFF24FBDCFF3DFAEA" blockId="12.[151,1437,151,1365]" pageId="12" pageNumber="535">
|
||
<emphasis id="71DFEA9BF93BFFCCFF24FBDCFEF4FBC3" bold="true" box="[199,317,1124,1148]" pageId="12" pageNumber="535">Remarks:</emphasis>
|
||
This is a morphologically distinctive species (
|
||
<emphasis id="71DFEA9BF93BFFCCFCBFFBDCFCB1FBC2" box="[860,888,1124,1149]" italics="true" pageId="12" pageNumber="535">cf.</emphasis>
|
||
the figures in
|
||
<bibRefCitation id="273A4B78F93BFFCCFBC7FBDCFB29FBC2" author="Fritz, U. & Petzold, A. & Kehlmaier, C. & Kindler, C. & Campbell, P. & Hofmeyr, M. D. & Branch, W. R." box="[1060,1248,1124,1149]" pageId="12" pageNumber="535" pagination="501 - 522" refId="ref16261" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. & Branch, W. R. (2014) Disentangling the Pelomedusa complex using type specimens and historical DNA. Zootaxa, 3795 (5), 501 - 522 http: // dx. doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
|
||
Fritz
|
||
<emphasis id="71DFEA9BF93BFFCCFB80FBDDFB57FBC2" box="[1123,1182,1124,1149]" italics="true" pageId="12" pageNumber="535">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
and this study).
|
||
<taxonomicName id="84AB4D0AF93BFFCCFF74FB3FFEB4FB1F" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[151,381,1159,1184]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF93BFFCCFF74FB3FFEB4FB1F" box="[151,381,1159,1184]" italics="true" pageId="12" pageNumber="535">Pelomedusa gehafie</emphasis>
|
||
</taxonomicName>
|
||
corresponds to mtDNA lineage X of
|
||
<bibRefCitation id="273A4B78F93BFFCCFCDBFB3FFBC2FB1F" author="Fritz, U. & Petzold, A. & Kehlmaier, C. & Kindler, C. & Campbell, P. & Hofmeyr, M. D. & Branch, W. R." box="[824,1035,1159,1184]" pageId="12" pageNumber="535" pagination="501 - 522" refId="ref16261" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. & Branch, W. R. (2014) Disentangling the Pelomedusa complex using type specimens and historical DNA. Zootaxa, 3795 (5), 501 - 522 http: // dx. doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
|
||
Fritz
|
||
<emphasis id="71DFEA9BF93BFFCCFC94FB31FC7AFB1F" box="[887,947,1159,1184]" italics="true" pageId="12" pageNumber="535">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
. It belongs to the northern species group of
|
||
<taxonomicName id="84AB4D0AF93BFFCCFEE7FB14FE44FB7A" authorityName="Wagler" authorityYear="1830" box="[260,397,1196,1221]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="genus">
|
||
<emphasis id="71DFEA9BF93BFFCCFEE7FB14FE44FB7A" box="[260,397,1196,1221]" italics="true" pageId="12" pageNumber="535">Pelomedusa</emphasis>
|
||
</taxonomicName>
|
||
, which is well supported in phylogenetic analyses of mtDNA. These reveal that the East African species
|
||
<taxonomicName id="84AB4D0AF93BFFCCFEBAFB69FE23FB58" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[345,490,1232,1256]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF93BFFCCFEBAFB69FE23FB58" box="[345,490,1232,1256]" italics="true" pageId="12" pageNumber="535">P. neumanni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="84AB4D0AF93BFFCCFDCDFB69FD4FFB57" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[558,646,1231,1256]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="kobe">
|
||
<emphasis id="71DFEA9BF93BFFCCFDCDFB69FD4FFB57" box="[558,646,1231,1256]" italics="true" pageId="12" pageNumber="535">P. kobe</emphasis>
|
||
</taxonomicName>
|
||
are closely related to
|
||
<taxonomicName id="84AB4D0AF93BFFCCFC7CFB69FBDCFB57" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[927,1045,1231,1256]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF93BFFCCFC7CFB69FBDCFB57" box="[927,1045,1231,1256]" italics="true" pageId="12" pageNumber="535">P. gehafie</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DB902A0CF93BFFCCFBC9FB77FBBAFB57" box="[1066,1139,1231,1256]" captionStart="FIGURE 1" captionStartId="3.[151,250,999,1021]" captionTargetBox="[167,1426,219,969]" captionTargetId="figure-480@3.[151,1436,199,975]" captionTargetPageId="3" captionText="FIGURE 1. Maximum Likelihood tree for helmeted terrapins (Pelomedusa spp.) using 1848 bp of mitochondrial DNA (12S, cyt b, ND4+tRNAs), rooted with Pelusios sinuatus. Terminal clades collapsed to cartoons. The topology of the Bayesian 50% majority rule consensus tree was identical. Numbers along branches indicate bootstrap support and clade support under BI (posterior probabilities) greater than 50 or 0.95, respectively. Asterisks represent maximum support under both methods. Colours and symbol correspond to the map (Fig. 2). On the right, proposed species names indicated; in brackets, mtDNA lineages. For clade membership of individual samples, see Table S1. The questionable sample MTD T 5484 from Swellendam District (Western Cape), South Africa, is highlighted in red (see text). The shown terrapins are Pelomedusa neumanni (top; Kakamega, Kenya) and P. galeata (bottom; Port Elizabeth, South Africa—photos: H. Prokop and W.R. Branch)." figureDoi="http://doi.org/10.5281/zenodo.4915085" httpUri="https://zenodo.org/record/4915085/files/figure.png" pageId="12" pageNumber="535">Fig. 1</figureCitation>
|
||
). These two species are morphologically easy to tell apart from
|
||
<taxonomicName id="84AB4D0AF93BFFCCFDB6FB4DFD0DFAB2" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[597,708,1268,1293]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF93BFFCCFDB6FB4DFD0DFAB2" box="[597,708,1268,1293]" italics="true" pageId="12" pageNumber="535">P. gehafie</emphasis>
|
||
</taxonomicName>
|
||
. Adults of
|
||
<taxonomicName id="84AB4D0AF93BFFCCFCA0FB4DFC7BFAB2" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[835,946,1268,1293]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="535" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF93BFFCCFCA0FB4DFC7BFAB2" box="[835,946,1268,1293]" italics="true" pageId="12" pageNumber="535">P. gehafie</emphasis>
|
||
</taxonomicName>
|
||
consistently have triangular pectoral scutes not reaching the plastral midseam, whereas the pectorals in the two other species are always in contact in the midline.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |