422 lines
50 KiB
XML
422 lines
50 KiB
XML
<document id="57021FE21B5358C87C5D820D2A1E0B36" ID-CLB-Dataset="5605" ID-DOI="10.11646/zootaxa.3795.5.2" ID-GBIF-Dataset="87d17125-337d-4679-bdcd-102f6b022ad2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915083" ID-ZooBank="9394634C-9836-4973-868B-BDEE414E4EA8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623198152603" checkinUser="felipe" docAuthor="Petzold, Alice, Vargas-Ramírez, Mario, Kehlmaier, Christian, Vamberger, Melita, Branch, William R., Preez, Louis Du, Hofmeyr, Margaretha D., Meyer, Leon, Schleicher, Alfred, Široký, Pavel & Fritz, Uwe" docDate="2014" docId="CB02879FF939FFD0FF74FE47FDFDFE4A" docLanguage="en" docName="zt03795p548.pdf" docOrigin="Zootaxa 3795 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pelomedusa neumanni Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz 2014, sp. nov." docType="treatment" docVersion="6" lastPageNumber="539" masterDocId="373BFFE7F937FFC0FFE3FFB8FFC9FFBF" masterDocTitle="A revision of African helmeted terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species" masterLastPageNumber="548" masterPageNumber="523" pageNumber="537" updateTime="1699006367747" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="999A38015D942AF9B90D1F32AD4E2BA0">A revision of African helmeted terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species</mods:title>
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<mods:namePart id="001116E9400A899C7343EBD9F1E9CECA">Petzold, Alice</mods:namePart>
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<mods:affiliation id="CA6DB562B5B2E4BD74D0E00AD50C9B77">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="C78D12A2D108C44669267086EEA1C444">Vargas-Ramírez, Mario</mods:namePart>
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<mods:affiliation id="68F851B5282E71F04FB95AD995E21937">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="9C0FB09B69E12172FC5304A19B1FA6BC">Kehlmaier, Christian</mods:namePart>
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<mods:affiliation id="2127E246B670CC541445782126EB7F99">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="526800F10AD993C964AC6A22B33C8F1C">Vamberger, Melita</mods:namePart>
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<mods:affiliation id="0AF9D394D4535784DA3A391659C26D46">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="1587DE4D69B2AB3FCCD1FA55C485692D">Branch, William R.</mods:namePart>
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<mods:affiliation id="6EC9FBAD7E5DAB8E6A41931E4AB64878">Department of Herpetology, Port Elizabeth Museum, P. O. Box 13147, Humewood 6013, South Africa & Department of Zoology, P. O. Box 77000, Nelson Mandela Metropolitan University, Port Elizabeth 6031, South Africa</mods:affiliation>
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<mods:namePart id="D9FFE4B11875D37D3837B1C13DE3DA17">Preez, Louis Du</mods:namePart>
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<mods:affiliation id="CB2D7223F67FA60366FE376EC26D1DF2">Unit for Environmental Science & Management, North-West University, Potchefstroom Campus, Private Bag X 6001, Potchefstroom 2520, South Africa</mods:affiliation>
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<mods:namePart id="6EFFFFA714EE73B361270E77ED6045B1">Hofmeyr, Margaretha D.</mods:namePart>
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<mods:affiliation id="4E678475BE34A4CD4BC1B1373AF5CB4C">Chelonian Biodiversity and Conservation, Department of Biodiversity and Conservation Biology, University of the Western Cape, Private Bag X 17, Bellville 7535, South Africa</mods:affiliation>
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<mods:namePart id="87D51BAF28AAD04D57B4F033BEFF2FBB">Meyer, Leon</mods:namePart>
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<mods:affiliation id="BCB034009E11AE894C2EBF23077888E6">Unit for Environmental Science & Management, North-West University, Potchefstroom Campus, Private Bag X 6001, Potchefstroom 2520, South Africa</mods:affiliation>
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<mods:namePart id="8020FCFF5A948F204621441FBCD6FA44">Schleicher, Alfred</mods:namePart>
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<mods:affiliation id="CBDAA8DF4C16143FE38C6F1FB2F2F40C">P. O. Box 30566, Windhoek, Namibia</mods:affiliation>
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<mods:namePart id="4C525BEB1479A7132242081885E62EC3">Široký, Pavel</mods:namePart>
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<mods:affiliation id="5410AB468B50C8AF6D8E4D9E47B42E0E">Department of Biology and Wildlife Diseases, Faculty of Veterinary Hygiene and Ecology, University of Veterinary and Pharmaceutical Sciences, Palackého tř. 1 / 3, 612 42 Brno, Czech Republic</mods:affiliation>
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<mods:namePart id="FF2A2DA98801544F3ED11BCCAE5413B1">Fritz, Uwe</mods:namePart>
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<mods:affiliation id="54B1833A70A8419675BEEBDCD695815D">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:title id="8C3B6E2050036280A2C3680FC1004CB5">Zootaxa</mods:title>
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<mods:part id="B38C94BDD1E0379885F41ADD420641F4">
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<mods:date id="9C61720109024C82CEE587B0F9DD2B12">2014</mods:date>
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<mods:number id="95527090EE14B011375BB9A35DDC38C9">2014-05-15</mods:number>
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<mods:number id="CBFAACCAC52401B31F677B0F067D4A9C">3795</mods:number>
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<mods:number id="3B636D1D3DC0BC238D90D2462D491DE8">5</mods:number>
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<mods:identifier id="52F509E0C59601B86515153C78A2C6DF" type="DOI">10.11646/zootaxa.3795.5.2</mods:identifier>
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<mods:identifier id="D8D31A665A515D233A94EA3AE1F87D13" type="GBIF-Dataset">87d17125-337d-4679-bdcd-102f6b022ad2</mods:identifier>
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<treatment id="CB02879FF939FFD0FF74FE47FDFDFE4A" ID-DOI="http://doi.org/10.5281/zenodo.4915097" ID-GBIF-Taxon="183403857" ID-Zenodo-Dep="4915097" LSID="urn:lsid:plazi:treatment:CB02879FF939FFD0FF74FE47FDFDFE4A" httpUri="http://treatment.plazi.org/id/CB02879FF939FFD0FF74FE47FDFDFE4A" lastPageId="16" lastPageNumber="539" pageId="14" pageNumber="537">
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<subSubSection id="0BB16502F939FFCEFF74FE47FDD8FDA5" box="[151,529,511,538]" pageId="14" pageNumber="537" type="nomenclature">
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<paragraph id="43143689F939FFCEFF74FE47FDD8FDA5" blockId="14.[151,529,511,538]" box="[151,529,511,538]" pageId="14" pageNumber="537">
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<heading id="185C81E5F939FFCEFF74FE47FDD8FDA5" bold="true" box="[151,529,511,538]" fontSize="11" level="1" pageId="14" pageNumber="537" reason="1">
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<emphasis id="71DFEA9BF939FFCEFF74FE47FDD8FDA5" bold="true" box="[151,529,511,538]" pageId="14" pageNumber="537">
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<taxonomicName id="84AB4D0AF939FFCEFF74FE47FE64FDA6" authority="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz, 2014" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[151,429,511,537]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="neumanni" status="sp. nov.">
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<emphasis id="71DFEA9BF939FFCEFF74FE47FE64FDA6" bold="true" box="[151,429,511,537]" italics="true" pageId="14" pageNumber="537">Pelomedusa neumanni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6AEC57E0F939FFCEFE57FDB8FDD8FDA5" box="[436,529,512,538]" pageId="14" pageNumber="537" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0BB16502F939FFCEFF74FDFEFC1FFAB4" pageId="14" pageNumber="537" type="diagnosis">
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<paragraph id="43143689F939FFCEFF74FDFEFC1FFAB4" blockId="14.[151,1437,582,1873]" pageId="14" pageNumber="537">
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<emphasis id="71DFEA9BF939FFCEFF74FDFEFEDAFDE0" bold="true" box="[151,275,582,607]" pageId="14" pageNumber="537">Diagnosis:</emphasis>
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Medium-sized helmeted terrapins with a known maximum straight carapacial length of
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<quantity id="84539B6CF939FFCEFAD6FDFEFA50FDE1" box="[1333,1433,582,607]" metricMagnitude="-1" metricUnit="m" metricValue="1.94" pageId="14" pageNumber="537" unit="cm" value="19.4">19.4 cm</quantity>
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. Pectoral scutes rectangular with wide midseam contact. One large undivided temporal head scale. Two small barbels under chin. Colouration variable; specimens with light horn-coloured carapace and completely yellow plastron and individuals with brownish plastron known to occur; soft parts ventrally lighter than dorsally.
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<taxonomicName id="84AB4D0AF939FFCEFF74FD6EFE5CFD51" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[151,405,726,751]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="neumanni">
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<emphasis id="71DFEA9BF939FFCEFF74FD6EFE5CFD51" box="[151,405,726,751]" italics="true" pageId="14" pageNumber="537">Pelomedusa neumanni</emphasis>
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</taxonomicName>
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differs from all other
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<taxonomicName id="84AB4D0AF939FFCEFD6CFD6EFCB9FD50" box="[655,880,726,751]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="71DFEA9BF939FFCEFD6CFD6EFCD1FD50" box="[655,792,726,751]" italics="true" pageId="14" pageNumber="537">Pelomedusa</emphasis>
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species
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</taxonomicName>
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except possibly
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<taxonomicName id="84AB4D0AF939FFCEFBCFFD6FFB53FD50" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[1068,1178,726,751]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="gehafie">
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<emphasis id="71DFEA9BF939FFCEFBCFFD6FFB53FD50" box="[1068,1178,726,751]" italics="true" pageId="14" pageNumber="537">P. gehafie</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="84AB4D0AF939FFCEFB4AFD6FFB30FD50" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1193,1273,726,751]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="kobe">
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<emphasis id="71DFEA9BF939FFCEFB4AFD6FFB30FD50" box="[1193,1273,726,751]" italics="true" pageId="14" pageNumber="537">P. kobe</emphasis>
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</taxonomicName>
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and candidate species B (in which the respective character states are unknown) by the presence of thymine (T) instead of cytosine (C) at position 57, by the presence of thymine (T) instead of cytosine (C) or guanine (G) at position 349, and by the presence of adenine (A) instead of cytosine (C) or guanine (G) at position 353 of the 360-bp-long reference alignment of the 12S rRNA gene (Supporting Information).
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<taxonomicName id="84AB4D0AF939FFCEFCD8FCDEFBF3FCC1" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[827,1082,870,895]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="neumanni">
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<emphasis id="71DFEA9BF939FFCEFCD8FCDEFBF3FCC1" box="[827,1082,870,895]" italics="true" pageId="14" pageNumber="537">Pelomedusa neumanni</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="84AB4D0AF939FFCEFB2CFCDFFAF4FCC0" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[1231,1341,870,895]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="gehafie">
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<emphasis id="71DFEA9BF939FFCEFB2CFCDFFAF4FCC0" box="[1231,1341,870,895]" italics="true" pageId="14" pageNumber="537">P. gehafie</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="84AB4D0AF939FFCEFAA8FCDFFA55FCC0" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1355,1436,870,895]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="kobe">
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<emphasis id="71DFEA9BF939FFCEFAA8FCDFFA55FCC0" box="[1355,1436,870,895]" italics="true" pageId="14" pageNumber="537">P. kobe</emphasis>
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</taxonomicName>
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and candidate species B by the presence of thymine (T) instead of cytosine (C) at position 116, by the presence of adenine (A) or thymine (T) instead of cytosine (C) or guanine (G) at position 147, and by the presence of adenine (A) instead of guanine (G) at position 223. In addition,
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<taxonomicName id="84AB4D0AF939FFCEFCC2FC6BFC67FC55" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[801,942,979,1002]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="neumanni">
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||
<emphasis id="71DFEA9BF939FFCEFCC2FC6BFC67FC55" box="[801,942,979,1002]" italics="true" pageId="14" pageNumber="537">P. neumanni</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="84AB4D0AF939FFCEFBAFFC6BFB09FC54" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[1100,1216,978,1003]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF939FFCEFBAFFC6BFB09FC54" box="[1100,1216,978,1003]" italics="true" pageId="14" pageNumber="537">P. gehafie</emphasis>
|
||
</taxonomicName>
|
||
by the presence of adenine (A) instead of guanine (G) at position 256, by the presence of cytosine (C) instead of adenine (A) at position 268, and by the presence of cytosine (C) instead of thymine (T) at position 345.
|
||
<taxonomicName id="84AB4D0AF939FFCEFB7FFBA2FA52FB8D" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1180,1435,1050,1075]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF939FFCEFB7FFBA2FA52FB8D" box="[1180,1435,1050,1075]" italics="true" pageId="14" pageNumber="537">Pelomedusa neumanni</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="84AB4D0AF939FFCEFEC7FB87FEB1FBE8" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[292,376,1086,1111]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="kobe">
|
||
<emphasis id="71DFEA9BF939FFCEFEC7FB87FEB1FBE8" box="[292,376,1086,1111]" italics="true" pageId="14" pageNumber="537">P. kobe</emphasis>
|
||
</taxonomicName>
|
||
by the presence of cytosine (C) instead of thymine (T) at position 298.
|
||
<taxonomicName id="84AB4D0AF939FFCEFB7EFB86FA52FBE9" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1181,1435,1086,1111]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF939FFCEFB7EFB86FA52FBE9" box="[1181,1435,1086,1111]" italics="true" pageId="14" pageNumber="537">Pelomedusa neumanni</emphasis>
|
||
</taxonomicName>
|
||
differs from candidate species B by the presence of cytosine (C) instead of thymine (T) at positions 103, 268 and 274, by the presence of thymine (T) instead of cytosine (C) at positions 109, 125, 152, 287 and 326, by the presence of cytosine (C) instead of adenine (A) at positions 122, 166 and 303, by a gap instead of thymine (T) at positions 126 and 168, by the presence of adenine (A) instead of guanine (G) at positions 148, 154, 180 and 223, and by the presence of thymine (T) instead of adenine (A) at position 302.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F939FFCEFF24FAAFFD56FAEC" pageId="14" pageNumber="537" type="materials_examined">
|
||
<paragraph id="43143689F939FFCEFF24FAAFFD56FAEC" blockId="14.[151,1437,582,1873]" pageId="14" pageNumber="537">
|
||
<materialsCitation id="F3C33CD4F939FFCEFF24FAAFFD56FAEC" ID-GBIF-Occurrence="3319794314" accessionNumber="V 74974" collectingDate="2003-09-11" collectionCode="V" collectorName="P. Siroky" country="Kenya" latitude="0.284" location="National Museum Prague" longLatPrecision="13" longitude="34.742" pageId="14" pageNumber="537" specimenCode="NMP6" specimenCount="1" stateProvince="Kakamega" typeStatus="holotype">
|
||
<emphasis id="71DFEA9BF939FFCEFF24FAAFFEF4FA90" bold="true" box="[199,317,1303,1327]" pageId="14" pageNumber="537">
|
||
<typeStatus id="9C10882BF939FFCEFF24FAAFFEF1FA90" box="[199,312,1303,1327]" pageId="14" pageNumber="537" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<location id="46746052F939FFCEFEA5FAAEFDA3FA90" LSID="urn:lsid:plazi:treatment:CB02879FF939FFD0FF74FE47FDFDFE4A:46746052F939FFCEFEA5FAAEFDA3FA90" box="[326,618,1302,1327]" country="Kenya" latitude="0.284" longLatPrecision="13" longitude="34.742" name="National Museum Prague" pageId="14" pageNumber="537" stateProvince="Kakamega">National Museum Prague</location>
|
||
(
|
||
<specimenCode id="130D9EF2F939FFCEFD98FAAEFD0EFA90" box="[635,711,1302,1327]" collectionCode="NMP" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="14" pageNumber="537">NMP6</specimenCode>
|
||
<accessionNumber id="5CF8AB6AF939FFCEFD24FAAEFCE7FA90" box="[711,814,1302,1327]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V74974" pageId="14" pageNumber="537" type="EnaNcbi">
|
||
<collectionCode id="25BAAE4CF939FFCEFD24FAAEFD12FA90" box="[711,731,1302,1327]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="537" type="Museum">V</collectionCode>
|
||
74974
|
||
</accessionNumber>
|
||
, adult male,
|
||
<collectingRegion id="816FF86BF939FFCEFC26FAAEFB89FA90" box="[965,1088,1302,1327]" country="Kenya" name="Kakamega" pageId="14" pageNumber="537">Kakamega</collectingRegion>
|
||
,
|
||
<collectingCountry id="3BBC7619F939FFCEFBAEFAAFFB50FA90" box="[1101,1177,1303,1327]" name="Kenya" pageId="14" pageNumber="537">Kenya</collectingCountry>
|
||
,
|
||
<geoCoordinate id="269F504EF939FFCEFB44FAAEFADDFA90" box="[1191,1300,1302,1327]" degrees="0" direction="north" minutes="17.04" orientation="latitude" pageId="14" pageNumber="537" precision="9" value="0.284">N0°17.04</geoCoordinate>
|
||
<geoCoordinate id="269F504EF939FFCEFAFEFAAEFA51FA90" box="[1309,1432,1302,1327]" degrees="34" direction="east" minutes="44.52" orientation="longitude" pageId="14" pageNumber="537" precision="9" value="34.742">E34°44.52</geoCoordinate>
|
||
; leg.
|
||
<collectorName id="EE5E535FF939FFCEFF24FA83FEE6FAEC" box="[199,303,1338,1363]" pageId="14" pageNumber="537">P. Široký</collectorName>
|
||
,
|
||
<date id="37151049F939FFCEFED9FA82FDDFFAEC" box="[314,534,1338,1363]" pageId="14" pageNumber="537" value="2003-09-11">
|
||
<collectingDate id="2751E9A1F939FFCEFED9FA82FDDFFAEC" box="[314,534,1338,1363]" pageId="14" pageNumber="537" value="2003-09-11">11 September 2003</collectingDate>
|
||
</date>
|
||
;
|
||
<figureCitation id="DB902A0CF939FFCEFDC1FA82FDAAFAEC" box="[546,611,1338,1363]" captionStart="FIGURE 4" captionStartId="15.[151,250,1913,1935]" captionTargetBox="[187,1380,204,1862]" captionTargetId="figure-68@15.[151,1436,143,1882]" captionTargetPageId="15" captionText="FIGURE 4. Dorsal and ventral aspects of the holotype of Pelomedusa neumanni sp. nov. (NMP6V 74974, adult male, Kakamega, Kenya; top) and of Pelomedusa schweinfurthi sp. nov. (SMF 56161, female, Liria, Central Equatoria, South Sudan; bottom). Scale bars, 3 cm. Photos: P. Široký and E. Morawa." figureDoi="http://doi.org/10.5281/zenodo.4915093" httpUri="https://zenodo.org/record/4915093/files/figure.png" pageId="14" pageNumber="537">Fig. 4</figureCitation>
|
||
top).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F939FFCEFF24FAE6FBC5F9F0" pageId="14" pageNumber="537" type="description">
|
||
<paragraph id="43143689F939FFCEFF24FAE6FB98FA5C" blockId="14.[151,1437,582,1873]" pageId="14" pageNumber="537">
|
||
<emphasis id="71DFEA9BF939FFCEFF24FAE6FDD8FAC8" bold="true" box="[199,529,1374,1399]" pageId="14" pageNumber="537">
|
||
Description of the
|
||
<typeStatus id="9C10882BF939FFCEFE41FAE7FDC4FAC8" box="[418,525,1375,1399]" pageId="14" pageNumber="537" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Dry shell plus head in ethanol. Straight carapacial length
|
||
<quantity id="84539B6CF939FFCEFB75FAE6FB3BFAC9" box="[1174,1266,1374,1399]" metricMagnitude="-1" metricUnit="m" metricValue="1.66" pageId="14" pageNumber="537" unit="cm" value="16.6">16.6 cm</quantity>
|
||
, plastral length
|
||
<quantity id="84539B6CF939FFCEFF74FA3AFF31FA25" box="[151,248,1410,1435]" metricMagnitude="-1" metricUnit="m" metricValue="1.49" pageId="14" pageNumber="537" unit="cm" value="14.9">14.9 cm</quantity>
|
||
. Carapace wide, dark brown, in dorsal view with pronounced waist. Epidermal scutes of vertebral 2–4 missing, anterior carapacial margin slightly mutilated. Plastron dark coloured; pectoral scutes with midseam contact. Head dorsally dark coloured; throat yellowish, two small barbels under chin.
|
||
</paragraph>
|
||
<paragraph id="43143689F939FFCEFF24FA57FBC5F9F0" blockId="14.[151,1437,582,1873]" pageId="14" pageNumber="537">
|
||
<materialsCitation id="F3C33CD4F939FFCEFF24FA57FBC1F9F0" ID-GBIF-Occurrence="3319794319" accessionNumber="ZMB 28356, NMW 24452" collectionCode="ZMB, NMW" country="Tanzania" county="Museum" location="Berlin" municipality="Naturkunde" pageId="14" pageNumber="537" specimenCode="ZFMK 81951" specimenCount="1" stateProvince="Kakamega" typeStatus="paratype">
|
||
<emphasis id="71DFEA9BF939FFCEFF24FA57FE81F9B8" bold="true" box="[199,328,1519,1543]" pageId="14" pageNumber="537">
|
||
<typeStatus id="9C10882BF939FFCEFF24FA57FE8AF9B8" box="[199,323,1519,1543]" pageId="14" pageNumber="537" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectingCounty id="AA754E05F939FFCEFEB0FA57FE7CF9B9" box="[339,437,1519,1542]" pageId="14" pageNumber="537">Museum</collectingCounty>
|
||
für
|
||
<collectingMunicipality id="A370ACF3F939FFCEFE08FA56FDBBF9B8" box="[491,626,1518,1543]" pageId="14" pageNumber="537">Naturkunde</collectingMunicipality>
|
||
,
|
||
<location id="46746052F939FFCEFD62FA56FD0EF9B8" LSID="urn:lsid:plazi:treatment:CB02879FF939FFD0FF74FE47FDFDFE4A:46746052F939FFCEFD62FA56FD0EF9B8" box="[641,711,1518,1543]" country="Tanzania" county="Museum" municipality="Naturkunde" name="Berlin" pageId="14" pageNumber="537" stateProvince="Kakamega">Berlin</location>
|
||
(
|
||
<collectionCode id="25BAAE4CF939FFCEFD3AFA56FCDEF9B8" box="[729,791,1518,1543]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="14" pageNumber="537" type="Museum">ZMB</collectionCode>
|
||
28356, hatchling,
|
||
<location id="46746052F939FFCEFC17FA57FB6BF9B8" LSID="urn:lsid:plazi:treatment:CB02879FF939FFD0FF74FE47FDFDFE4A:46746052F939FFCEFC17FA57FB6BF9B8" box="[1012,1186,1518,1543]" country="Tanzania" county="Museum" municipality="Naturkunde" name="Mangara River" pageId="14" pageNumber="537" stateProvince="Kakamega">Mangara River</location>
|
||
, Manyara,
|
||
<collectingCountry id="3BBC7619F939FFCEFAC5FA56FA59F9B8" box="[1318,1424,1518,1543]" name="Tanzania" pageId="14" pageNumber="537">Tanzania</collectingCountry>
|
||
);
|
||
<location id="46746052F939FFCEFF74F9AAFDE9F994" LSID="urn:lsid:plazi:treatment:CB02879FF939FFD0FF74FE47FDFDFE4A:46746052F939FFCEFF74F9AAFDE9F994" box="[151,544,1554,1579]" country="Tanzania" county="Museum" municipality="Naturkunde" name="Naturhistorisches Museum Wien" pageId="14" pageNumber="537" stateProvince="Kakamega">Naturhistorisches Museum Wien</location>
|
||
(
|
||
<collectionCode id="25BAAE4CF939FFCEFDD9F9AAFD4DF994" box="[570,644,1554,1579]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="537">NMW</collectionCode>
|
||
24452, male,
|
||
<location id="46746052F939FFCEFCAAF9AAFC3DF994" LSID="urn:lsid:plazi:treatment:CB02879FF939FFD0FF74FE47FDFDFE4A:46746052F939FFCEFCAAF9AAFC3DF994" box="[841,1012,1554,1579]" country="Tanzania" county="Museum" municipality="Naturkunde" name="Lake Victoria" pageId="14" pageNumber="537" stateProvince="Kakamega">Lake Victoria</location>
|
||
);
|
||
<location id="46746052F939FFCEFBF0F9AAFEAEF9F0" LSID="urn:lsid:plazi:treatment:CB02879FF939FFD0FF74FE47FDFDFE4A:46746052F939FFCEFBF0F9AAFEAEF9F0" country="Tanzania" county="Museum" municipality="Naturkunde" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="14" pageNumber="537" stateProvince="Kakamega">Zoologisches Forschungsmuseum Alexander Koenig</location>
|
||
,
|
||
<location id="46746052F939FFCEFE90F98FFE66F9F1" LSID="urn:lsid:plazi:treatment:CB02879FF939FFD0FF74FE47FDFDFE4A:46746052F939FFCEFE90F98FFE66F9F1" box="[371,431,1591,1614]" country="Tanzania" county="Museum" municipality="Naturkunde" name="Bonn" pageId="14" pageNumber="537" stateProvince="Kakamega">Bonn</location>
|
||
(
|
||
<specimenCode id="130D9EF2F939FFCEFE5DF98EFD97F9F0" box="[446,606,1590,1615]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="14" pageNumber="537" type="Museum">ZFMK 81951</specimenCode>
|
||
, hatchling,
|
||
<collectingRegion id="816FF86BF939FFCEFD3CF98EFC9EF9F0" box="[735,855,1590,1615]" country="Kenya" name="Kakamega" pageId="14" pageNumber="537">Kakamega</collectingRegion>
|
||
<location id="46746052F939FFCEFCBDF98FFC6EF9F1" LSID="urn:lsid:plazi:treatment:CB02879FF939FFD0FF74FE47FDFDFE4A:46746052F939FFCEFCBDF98FFC6EF9F1" box="[862,935,1591,1614]" country="Tanzania" county="Museum" municipality="Naturkunde" name="Forest" pageId="14" pageNumber="537" stateProvince="Kakamega">Forest</location>
|
||
,
|
||
<collectingCountry id="3BBC7619F939FFCEFC51F98EFBC9F9F0" box="[946,1024,1590,1615]" name="Kenya" pageId="14" pageNumber="537">Kenya</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F939FFCEFF24F9E2FD7AF95D" pageId="14" pageNumber="537" type="etymology">
|
||
<paragraph id="43143689F939FFCEFF24F9E2FD7AF95D" blockId="14.[151,1437,582,1873]" pageId="14" pageNumber="537">
|
||
<emphasis id="71DFEA9BF939FFCEFF24F9E2FE6EF9CC" bold="true" box="[199,423,1626,1651]" pageId="14" pageNumber="537">Derivatio nominis:</emphasis>
|
||
We dedicate this new species to Oscar Neumann (
|
||
<date id="37151049F939FFCEFC04F9E2FB7FF9CC" box="[999,1206,1626,1651]" pageId="14" pageNumber="537" value="1867-09-03">3 September 1867</date>
|
||
–
|
||
<date id="37151049F939FFCEFB26F9E2FA95F9CC" box="[1221,1372,1626,1651]" pageId="14" pageNumber="537" value="1946-05-17">17 May 1946</date>
|
||
) who undertook in the late 19
|
||
<superScript id="B4DE9BC1F939FFCEFE4DF9C6FE73F933" attach="left" box="[430,442,1662,1676]" fontSize="6" pageId="14" pageNumber="537">th</superScript>
|
||
and early 20
|
||
<superScript id="B4DE9BC1F939FFCEFDB6F9C6FDA8F933" attach="left" box="[597,609,1662,1676]" fontSize="6" pageId="14" pageNumber="537">th</superScript>
|
||
centuries influential expeditions to East Africa,
|
||
<collectingCountry id="3BBC7619F939FFCEFB75F939FB3DF925" box="[1174,1268,1665,1690]" name="Somalia" pageId="14" pageNumber="537">Somalia</collectingCountry>
|
||
,
|
||
<collectingCountry id="3BBC7619F939FFCEFAE0F939FAACF925" box="[1283,1381,1665,1690]" name="Ethiopia" pageId="14" pageNumber="537">Ethiopia</collectingCountry>
|
||
, and
|
||
<collectingCountry id="3BBC7619F939FFCEFF74F91DFF28F901" box="[151,225,1701,1726]" name="Sudan" pageId="14" pageNumber="537">Sudan</collectingCountry>
|
||
. Neumann worked voluntarily for over 40 years in the Museum für Naturkunde,
|
||
<collectingRegion id="816FF86BF939FFCEFB98F91DFB0DF901" box="[1147,1220,1701,1726]" country="Germany" name="Berlin" pageId="14" pageNumber="537">Berlin</collectingRegion>
|
||
, but was forced by the Nazi government to leave
|
||
<collectingCountry id="3BBC7619F939FFCEFE04F971FD86F95D" box="[487,591,1737,1762]" name="Germany" pageId="14" pageNumber="537">Germany</collectingCountry>
|
||
in 1941.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F939FFCEFF24F955FAFEF8EF" pageId="14" pageNumber="537" type="distribution">
|
||
<paragraph id="43143689F939FFCEFF24F955FAFEF8EF" blockId="14.[151,1437,582,1873]" pageId="14" pageNumber="537">
|
||
<emphasis id="71DFEA9BF939FFCEFF24F955FEABF8B9" bold="true" box="[199,354,1773,1798]" pageId="14" pageNumber="537">Distribution:</emphasis>
|
||
<collectingCountry id="3BBC7619F939FFCEFE8AF956FE7BF8B9" box="[361,434,1774,1798]" name="Kenya" pageId="14" pageNumber="537">Kenya</collectingCountry>
|
||
(
|
||
<bibRefCitation id="273A4B78F939FFCEFE23F955FD3CF8B9" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[448,757,1773,1798]" pageId="14" pageNumber="537" pagination="428 - 440" refId="ref18768" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="71DFEA9BF939FFCEFD9EF956FD7CF8B9" box="[637,693,1773,1798]" italics="true" pageId="14" pageNumber="537">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
) and
|
||
<collectingCountry id="3BBC7619F939FFCEFCD1F955FC5EF8B9" box="[818,919,1773,1798]" name="Tanzania" pageId="14" pageNumber="537">Tanzania</collectingCountry>
|
||
(this study). The genetically verified
|
||
<typeStatus id="9C10882BF939FFCEFADFF956FA55F8B9" box="[1340,1436,1774,1798]" pageId="14" pageNumber="537" type="paratype">paratype</typeStatus>
|
||
ZMB 28356, collected by Oscar Neumann in the late 19
|
||
<superScript id="B4DE9BC1F939FFCEFCFBF8A9FCEDF8A0" attach="left" box="[792,804,1809,1823]" fontSize="6" pageId="14" pageNumber="537">th</superScript>
|
||
century in present-day
|
||
<collectingCountry id="3BBC7619F939FFCEFBD1F8ACFB53F892" box="[1074,1178,1812,1837]" name="Tanzania" pageId="14" pageNumber="537">Tanzania</collectingCountry>
|
||
, provides evidence for the occurrence of
|
||
<taxonomicName id="84AB4D0AF939FFCEFE83F881FE23F8EF" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[352,490,1849,1872]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF939FFCEFE83F881FE23F8EF" box="[352,490,1849,1872]" italics="true" pageId="14" pageNumber="537">P. neumanni</emphasis>
|
||
</taxonomicName>
|
||
in close proximity to genetically verified records of
|
||
<taxonomicName id="84AB4D0AF939FFCEFBDBF881FB40F8EE" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1080,1161,1848,1873]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="kobe">
|
||
<emphasis id="71DFEA9BF939FFCEFBDBF881FB40F8EE" box="[1080,1161,1848,1873]" italics="true" pageId="14" pageNumber="537">P. kobe</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="84AB4D0AF939FFCEFB5CF881FAF9F8EE" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[1215,1328,1848,1873]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="537" phylum="Chordata" rank="species" species="subrufa">
|
||
<emphasis id="71DFEA9BF939FFCEFB5CF881FAF9F8EE" box="[1215,1328,1848,1873]" italics="true" pageId="14" pageNumber="537">P. subrufa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="17D46601F938FFCFFF74F8C1FD34F86C" ID-DOI="http://doi.org/10.5281/zenodo.4915093" ID-Zenodo-Dep="4915093" httpUri="https://zenodo.org/record/4915093/files/figure.png" pageId="15" pageNumber="538" startId="15.[151,250,1913,1935]" targetBox="[187,1380,204,1862]" targetPageId="15">
|
||
<paragraph id="43143689F938FFCFFF74F8C1FD34F86C" blockId="15.[151,1436,1913,2003]" pageId="15" pageNumber="538">
|
||
<emphasis id="71DFEA9BF938FFCFFF74F8C1FEDEF830" bold="true" box="[151,279,1913,1935]" pageId="15" pageNumber="538">FIGURE 4.</emphasis>
|
||
Dorsal and ventral aspects of the holotype of
|
||
<taxonomicName id="84AB4D0AF938FFCFFCF6F8C1FBC9F830" authority="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz, 2014" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[789,1024,1913,1935]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="538" phylum="Chordata" rank="species" species="neumanni" status="sp. nov.">
|
||
<emphasis id="71DFEA9BF938FFCFFCF6F8C1FBC9F830" box="[789,1024,1913,1935]" italics="true" pageId="15" pageNumber="538">Pelomedusa neumanni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="71DFEA9BF938FFCFFBE8F8C1FBA8F830" bold="true" box="[1035,1121,1913,1935]" pageId="15" pageNumber="538">
|
||
<taxonomicNameLabel id="6AEC57E0F938FFCFFBE8F8C1FBA8F830" box="[1035,1121,1913,1935]" pageId="15" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NMP6
|
||
<accessionNumber id="5CF8AB6AF938FFCFFB5BF8C1FAD1F830" box="[1208,1304,1913,1935]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V74974" pageId="15" pageNumber="538" type="EnaNcbi">V 74974</accessionNumber>
|
||
, adult male, Kakamega, Kenya; top) and of
|
||
<taxonomicName id="84AB4D0AF938FFCFFE30F822FD17F80F" authority="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz, 2014" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[467,734,1946,1968]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="538" phylum="Chordata" rank="species" species="schweinfurthi" status="sp. nov.">
|
||
<emphasis id="71DFEA9BF938FFCFFE30F822FD17F80F" box="[467,734,1946,1968]" italics="true" pageId="15" pageNumber="538">Pelomedusa schweinfurthi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="71DFEA9BF938FFCFFD07F822FCFDF80F" bold="true" box="[740,820,1946,1968]" pageId="15" pageNumber="538">
|
||
<taxonomicNameLabel id="6AEC57E0F938FFCFFD07F822FCFDF80F" box="[740,820,1946,1968]" pageId="15" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(SMF 56161, female, Liria, Central Equatoria, South Sudan; bottom). Scale bars, 3 cm. Photos: P. Široký and E. Morawa.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0BB16502F927FFD0FF24FF20FDFDFE4A" pageId="16" pageNumber="539" type="discussion">
|
||
<paragraph id="43143689F927FFD0FF24FF20FA8BFEA2" blockId="16.[151,1437,151,501]" pageId="16" pageNumber="539">
|
||
<emphasis id="71DFEA9BF927FFD0FF24FF20FEF4FF0F" bold="true" box="[199,317,152,176]" pageId="16" pageNumber="539">Remarks:</emphasis>
|
||
<taxonomicName id="84AB4D0AF927FFD0FEAEFF2FFD9AFF10" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[333,595,151,176]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF927FFD0FEAEFF2FFD9AFF10" box="[333,595,151,176]" italics="true" pageId="16" pageNumber="539">Pelomedusa neumanni</emphasis>
|
||
</taxonomicName>
|
||
corresponds to mtDNA lineage V of
|
||
<bibRefCitation id="273A4B78F927FFD0FBD5FF2FFA51FF0F" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[1078,1432,151,176]" pageId="16" pageNumber="539" pagination="428 - 440" refId="ref18768" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="71DFEA9BF927FFD0FB19FF21FAF2FF0F" box="[1274,1339,151,176]" italics="true" pageId="16" pageNumber="539">et al.</emphasis>
|
||
(2010)
|
||
</bibRefCitation>
|
||
. According to our phylogenetic analyses of mtDNA sequences,
|
||
<taxonomicName id="84AB4D0AF927FFD0FC82FF05FC25FF6B" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[865,1004,189,212]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF927FFD0FC82FF05FC25FF6B" box="[865,1004,189,212]" italics="true" pageId="16" pageNumber="539">P. neumanni</emphasis>
|
||
</taxonomicName>
|
||
belongs to the northern species group of
|
||
<taxonomicName id="84AB4D0AF927FFD0FF56FF67FEF7FF47" authorityName="Wagler" authorityYear="1830" box="[181,318,223,248]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="genus">
|
||
<emphasis id="71DFEA9BF927FFD0FF56FF67FEF7FF47" box="[181,318,223,248]" italics="true" pageId="16" pageNumber="539">Pelomedusa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="84AB4D0AF927FFD0FEAFFF67FD80FF48" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[332,585,223,248]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF927FFD0FEAFFF67FD80FF48" box="[332,585,223,248]" italics="true" pageId="16" pageNumber="539">Pelomedusa neumanni</emphasis>
|
||
</taxonomicName>
|
||
is phylogenetically most closely related to
|
||
<taxonomicName id="84AB4D0AF927FFD0FBCDFF59FB52FF47" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[1070,1179,223,248]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF927FFD0FBCDFF59FB52FF47" box="[1070,1179,223,248]" italics="true" pageId="16" pageNumber="539">P. gehafie</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="84AB4D0AF927FFD0FB31FF59FAEBFF47" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1234,1314,223,248]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="kobe">
|
||
<emphasis id="71DFEA9BF927FFD0FB31FF59FAEBFF47" box="[1234,1314,223,248]" italics="true" pageId="16" pageNumber="539">P. kobe</emphasis>
|
||
</taxonomicName>
|
||
. However, the sister group relationships within the clade comprising these three species are not well resolved (
|
||
<figureCitation id="DB902A0CF927FFD0FB13FEBCFAF8FEA2" box="[1264,1329,260,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,999,1021]" captionTargetBox="[167,1426,219,969]" captionTargetId="figure-480@3.[151,1436,199,975]" captionTargetPageId="3" captionText="FIGURE 1. Maximum Likelihood tree for helmeted terrapins (Pelomedusa spp.) using 1848 bp of mitochondrial DNA (12S, cyt b, ND4+tRNAs), rooted with Pelusios sinuatus. Terminal clades collapsed to cartoons. The topology of the Bayesian 50% majority rule consensus tree was identical. Numbers along branches indicate bootstrap support and clade support under BI (posterior probabilities) greater than 50 or 0.95, respectively. Asterisks represent maximum support under both methods. Colours and symbol correspond to the map (Fig. 2). On the right, proposed species names indicated; in brackets, mtDNA lineages. For clade membership of individual samples, see Table S1. The questionable sample MTD T 5484 from Swellendam District (Western Cape), South Africa, is highlighted in red (see text). The shown terrapins are Pelomedusa neumanni (top; Kakamega, Kenya) and P. galeata (bottom; Port Elizabeth, South Africa—photos: H. Prokop and W.R. Branch)." figureDoi="http://doi.org/10.5281/zenodo.4915085" httpUri="https://zenodo.org/record/4915085/files/figure.png" pageId="16" pageNumber="539">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="43143689F927FFD0FF24FE9FFF26FE14" blockId="16.[151,1437,151,501]" pageId="16" pageNumber="539">
|
||
The ranges of
|
||
<taxonomicName id="84AB4D0AF927FFD0FE92FE91FE37FE80" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[369,510,296,320]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF927FFD0FE92FE91FE37FE80" box="[369,510,296,320]" italics="true" pageId="16" pageNumber="539">P. neumanni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="84AB4D0AF927FFD0FDECFE91FDAAFEFF" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[527,611,295,320]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="kobe">
|
||
<emphasis id="71DFEA9BF927FFD0FDECFE91FDAAFEFF" box="[527,611,295,320]" italics="true" pageId="16" pageNumber="539">P. kobe</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="84AB4D0AF927FFD0FD7CFE91FCDAFEFF" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[671,787,295,320]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="subrufa">
|
||
<emphasis id="71DFEA9BF927FFD0FD7CFE91FCDAFEFF" box="[671,787,295,320]" italics="true" pageId="16" pageNumber="539">P. subrufa</emphasis>
|
||
</taxonomicName>
|
||
abut or overlap in the East African savannah. To avoid taxonomic misidentifications, we included only genetically verified museum specimens in our
|
||
<typeStatus id="9C10882BF927FFD0FAF5FEF4FA8FFEDB" box="[1302,1350,332,356]" pageId="16" pageNumber="539">type</typeStatus>
|
||
series. However, it should be noted that all
|
||
<specimenCount id="55ADFD00F927FFD0FDDDFED7FD11FE37" box="[574,728,367,392]" pageId="16" pageNumber="539" type="generic">18 specimens</specimenCount>
|
||
from
|
||
<collectingCountry id="3BBC7619F927FFD0FCC2FEC8FCA3FE37" box="[801,874,368,392]" name="Kenya" pageId="16" pageNumber="539">Kenya</collectingCountry>
|
||
we studied have pectoral scutes in wide midline contact.
|
||
</paragraph>
|
||
<paragraph id="43143689F927FFD0FF24FE00FDFDFE4A" blockId="16.[151,1437,151,501]" pageId="16" pageNumber="539">
|
||
For the monogenean flatworm
|
||
<taxonomicName id="84AB4D0AF927FFD0FDDFFE0FFC9CFE6F" authorityName="Euzet & Combes" authorityYear="1965" box="[572,853,439,464]" class="Monogenea" family="Polystomatidae" genus="Polystomoides" kingdom="Animalia" order="Polystomatidea" pageId="16" pageNumber="539" phylum="Platyhelminthes" rank="species" species="chabaudi">
|
||
<emphasis id="71DFEA9BF927FFD0FDDFFE0FFC9CFE6F" box="[572,853,439,464]" italics="true" pageId="16" pageNumber="539">Polystomoides chabaudi</emphasis>
|
||
</taxonomicName>
|
||
possibly parasitizing
|
||
<taxonomicName id="84AB4D0AF927FFD0FBBDFE0FFAAAFE70" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1118,1379,439,464]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF927FFD0FBBDFE0FFAAAFE70" box="[1118,1379,439,464]" italics="true" pageId="16" pageNumber="539">Pelomedusa neumanni</emphasis>
|
||
</taxonomicName>
|
||
, see below under
|
||
<taxonomicName id="84AB4D0AF927FFD0FECAFE65FE53FE4A" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[297,410,476,501]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" sensu="stricto" species="subrufa">
|
||
<emphasis id="71DFEA9BF927FFD0FECAFE65FE53FE4A" box="[297,410,476,501]" italics="true" pageId="16" pageNumber="539">P. subrufa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AEC57E0F927FFD0FE42FE64FDF9FE4A" box="[417,560,476,501]" pageId="16" pageNumber="539" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |