466 lines
51 KiB
XML
466 lines
51 KiB
XML
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<mods:title id="87BC05DCBB7D95CB058F9AC151AAB047">A revision of African helmeted terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species</mods:title>
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<mods:namePart id="62D352BEEEB11B2952F7846AE04E4B4C">Fritz, Uwe</mods:namePart>
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<treatment id="CB02879FF924FFD4FF74FF2FFAD1FE14" ID-DOI="http://doi.org/10.5281/zenodo.5082935" ID-GBIF-Taxon="183403862" ID-Zenodo-Dep="5082935" LSID="urn:lsid:plazi:treatment:CB02879FF924FFD4FF74FF2FFAD1FE14" httpUri="http://treatment.plazi.org/id/CB02879FF924FFD4FF74FF2FFAD1FE14" lastPageId="20" lastPageNumber="543" pageId="19" pageNumber="542">
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<emphasis id="71DFEA9BF924FFD3FF74FF2FFE46FF0E" bold="true" box="[151,399,151,177]" italics="true" pageId="19" pageNumber="542">Pelomedusa subrufa</emphasis>
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(
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<bibRefCitation id="273A4B78F924FFD3FE7DFF20FDB0FF0D" author="Bonnaterre, P. J." box="[414,633,152,178]" pageId="19" pageNumber="542" refId="ref14973" refString="Bonnaterre, P. J. (1789) Tableau encyclopedique et methodique des trois regnes de la nature. Erpetologie. Panckoucke, Hotel de Thou, Paris, xxviii + 70 + (1) pp., 35 plates. http: // dx. doi. org / 10.5962 / bhl. title. 59326" type="book" year="1789">Bonnaterre, 1789</bibRefCitation>
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)
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</emphasis>
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1789
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<emphasis id="71DFEA9BF924FFD3FF02FF66FE42FF4A" box="[225,395,222,245]" italics="true" pageId="19" pageNumber="542">Testudo subrufa</emphasis>
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Bonnaterre
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—Restricted type locality (Bour 1982): Taolañaro,
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<collectingCountry id="3BBC7619F924FFD3FBCAFF66FB64FF4A" box="[1065,1197,222,245]" name="Madagascar" pageId="19" pageNumber="542">Madagascar</collectingCountry>
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;
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: Muséum
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National d’Histoire naturelle, Paris, MNHN 7970 (
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Fig.
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in
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<bibRefCitation id="273A4B78F924FFD3FC83FF46FBC5FEAA" author="Fritz, U. & Petzold, A. & Kehlmaier, C. & Kindler, C. & Campbell, P. & Hofmeyr, M. D. & Branch, W. R." box="[864,1036,254,277]" pageId="19" pageNumber="542" pagination="501 - 522" refId="ref16261" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. & Branch, W. R. (2014) Disentangling the Pelomedusa complex using type specimens and historical DNA. Zootaxa, 3795 (5), 501 - 522 http: // dx. doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
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Fritz
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<emphasis id="71DFEA9BF924FFD3FC7AFF46FC06FEAA" box="[921,975,254,277]" italics="true" pageId="19" pageNumber="542">et al.</emphasis>
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2014
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) 1798
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<taxonomicName id="84AB4D0AF924FFD3FF02FEA6FE24FE8A" authority="Donndorff" authorityName="Donndorff" authorityYear="1798" box="[225,493,286,309]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="badia">
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<emphasis id="71DFEA9BF924FFD3FF02FEA6FEBFFE8A" box="[225,374,286,309]" italics="true" pageId="19" pageNumber="542">Testudo badia</emphasis>
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Donndorff
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(nomen novum) 1798
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<taxonomicName id="84AB4D0AF924FFD3FF02FE86FDCBFEEA" authority="Suckow" authorityName="Suckow" authorityYear="1798" box="[225,514,318,341]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="rubicunda">
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<emphasis id="71DFEA9BF924FFD3FF02FE86FE6EFEEA" box="[225,423,318,341]" italics="true" pageId="19" pageNumber="542">Testudo rubicunda</emphasis>
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Suckow
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</taxonomicName>
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(nomen novum) 1935
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<taxonomicName id="84AB4D0AF924FFD3FF02FEE6FD40FECA" authority="Hewitt" authorityName="Hewitt" authorityYear="1935" box="[225,649,350,373]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="subSpecies" species="galeata" subSpecies="damarensis">
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<emphasis id="71DFEA9BF924FFD3FF02FEE6FDF3FECA" box="[225,570,350,373]" italics="true" pageId="19" pageNumber="542">Pelomedusa galeata damarensis</emphasis>
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Hewitt
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</taxonomicName>
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—Type locality: Quickborn near Okahandja,
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;
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(
|
||
<bibRefCitation id="273A4B78F924FFD3FAA6FEE6FA5DFECA" author="Fritz, U. & Petzold, A. & Kehlmaier, C. & Kindler, C. & Campbell, P. & Hofmeyr, M. D. & Branch, W. R." box="[1349,1428,350,373]" pageId="19" pageNumber="542" pagination="501 - 522" refId="ref16261" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. & Branch, W. R. (2014) Disentangling the Pelomedusa complex using type specimens and historical DNA. Zootaxa, 3795 (5), 501 - 522 http: // dx. doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
|
||
Fritz
|
||
<emphasis id="71DFEA9BF924FFD3FA63FEE6FA5DFECA" box="[1408,1428,350,373]" italics="true" pageId="19" pageNumber="542">et</emphasis>
|
||
</bibRefCitation>
|
||
</paragraph>
|
||
<paragraph id="43143689F924FFD3FF02FEC6FA4AFE0A" blockId="19.[151,1428,222,469]" pageId="19" pageNumber="542">
|
||
<emphasis id="71DFEA9BF924FFD3FF02FEC6FF34FE2A" box="[225,253,382,405]" italics="true" pageId="19" pageNumber="542">al.</emphasis>
|
||
2014): Port Elizabeth Museum, PEM R14953 (Fig. 7 top in
|
||
<bibRefCitation id="273A4B78F924FFD3FC61FEC6FBE6FE2A" author="Fritz, U. & Petzold, A. & Kehlmaier, C. & Kindler, C. & Campbell, P. & Hofmeyr, M. D. & Branch, W. R." box="[898,1071,382,405]" pageId="19" pageNumber="542" pagination="501 - 522" refId="ref16261" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. & Branch, W. R. (2014) Disentangling the Pelomedusa complex using type specimens and historical DNA. Zootaxa, 3795 (5), 501 - 522 http: // dx. doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
|
||
Fritz
|
||
<emphasis id="71DFEA9BF924FFD3FC5FFEC6FC38FE2A" box="[956,1009,382,405]" italics="true" pageId="19" pageNumber="542">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
) 1937
|
||
<taxonomicName id="84AB4D0AF924FFD3FF02FE26FD4CFE0A" authority="Mertens" authorityName="Mertens" authorityYear="1937" box="[225,645,414,437]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="subSpecies" species="subrufa" subSpecies="wettsteini">
|
||
<emphasis id="71DFEA9BF924FFD3FF02FE26FDE1FE0A" box="[225,552,414,437]" italics="true" pageId="19" pageNumber="542">Pelomedusa subrufa wettsteini</emphasis>
|
||
Mertens
|
||
</taxonomicName>
|
||
—Type locality: Majunga (
|
||
<collectingRegion id="816FF86BF924FFD3FC44FE26FBEAFE0A" box="[935,1059,414,437]" country="Madagascar" name="Mahajanga" pageId="19" pageNumber="542">Mahajanga</collectingRegion>
|
||
), western
|
||
<collectingCountry id="3BBC7619F924FFD3FB6DFE26FADBFE0A" box="[1166,1298,414,437]" name="Madagascar" pageId="19" pageNumber="542">Madagascar</collectingCountry>
|
||
;
|
||
<typeStatus id="9C10882BF924FFD3FAFEFE26FAB6FE0A" box="[1309,1407,414,437]" pageId="19" pageNumber="542" type="holotype">holotype</typeStatus>
|
||
:
|
||
</paragraph>
|
||
<paragraph id="43143689F924FFD3FF02FE06FB83FE6A" blockId="19.[151,1428,222,469]" box="[225,1098,446,469]" pageId="19" pageNumber="542">
|
||
Senckenberg-Museum, Frankfurt am Main, SMF 7958 (Fig.
|
||
<quantity id="84539B6CF924FFD3FC84FE06FC59FE6A" box="[871,912,446,469]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="19" pageNumber="542" unit="in" value="8.0">8 in</quantity>
|
||
<bibRefCitation id="273A4B78F924FFD3FC74FE06FB8AFE6A" author="Fritz, U. & Petzold, A. & Kehlmaier, C. & Kindler, C. & Campbell, P. & Hofmeyr, M. D. & Branch, W. R." box="[919,1091,446,469]" pageId="19" pageNumber="542" pagination="501 - 522" refId="ref16261" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. & Branch, W. R. (2014) Disentangling the Pelomedusa complex using type specimens and historical DNA. Zootaxa, 3795 (5), 501 - 522 http: // dx. doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
|
||
Fritz
|
||
<emphasis id="71DFEA9BF924FFD3FC33FE06FBCFFE6A" box="[976,1030,446,469]" italics="true" pageId="19" pageNumber="542">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F924FFD3FF74FDBAFB99FBC0" pageId="19" pageNumber="542" type="diagnosis">
|
||
<paragraph id="43143689F924FFD3FF74FDBAFB99FBC0" blockId="19.[151,1437,514,1979]" pageId="19" pageNumber="542">
|
||
<emphasis id="71DFEA9BF924FFD3FF74FDBAFEDAFDA4" bold="true" box="[151,275,514,539]" pageId="19" pageNumber="542">Diagnosis:</emphasis>
|
||
Small to medium-sized helmeted terrapins with a known maximum straight carapacial length of
|
||
<quantity id="84539B6CF924FFD3FA88FDBAFF74FD81" metricMagnitude="-1" metricUnit="m" metricValue="1.97" pageId="19" pageNumber="542" unit="cm" value="19.7">19.7 cm</quantity>
|
||
. Yet, the shell length of most adults is below
|
||
<quantity id="84539B6CF924FFD3FD2BFD9EFCDAFD81" box="[712,787,550,575]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="19" pageNumber="542" unit="cm" value="14.0">14 cm</quantity>
|
||
. Pectoral scutes in most specimens with broad contact at plastral midseam; however, terrapins with narrow contact or, exceptionally, with triangular pectorals without midseam contact occur. One large undivided temporal scale on each side of head. Two small barbels under chin. Carapace uniform light to dark brown. Plastron in larger individuals mostly light coloured, in younger specimens often with dark markings.
|
||
<taxonomicName id="84AB4D0AF924FFD3FE33FD0EFD72FD70" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[464,699,694,719]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" sensu="stricto" species="subrufa">
|
||
<emphasis id="71DFEA9BF924FFD3FE33FD0EFD72FD70" box="[464,699,694,719]" italics="true" pageId="19" pageNumber="542">Pelomedusa subrufa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AEC57E0F924FFD3FD26FD0FFC9CFD70" box="[709,853,694,719]" pageId="19" pageNumber="542" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
differs from all other
|
||
<taxonomicName id="84AB4D0AF924FFD3FB81FD0EFA8EFD70" box="[1122,1351,694,719]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="71DFEA9BF924FFD3FB81FD0EFB22FD70" box="[1122,1259,694,719]" italics="true" pageId="19" pageNumber="542">Pelomedusa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by the presence of cytosine (C) instead of thymine (T) at position 117, by the presence of thymine (T) instead of adenine (A) at position 169, by the presence of thymine (T) instead of adenine (A) or cytosine (C) at position 191, by the presence of guanine (G) instead of adenine (A) at positions 226 and 296, by the presence of adenine (A) instead of cytosine (C) at position 280, and by the presence of cytosine (C) or thymine (T) instead of adenine (A) or guanine (G) at position 289 of the 360-bp-long reference alignment of the 12S rRNA gene (Supporting Information). In addition,
|
||
<taxonomicName id="84AB4D0AF924FFD3FEE0FC37FEBFFC18" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[259,374,910,935]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" sensu="stricto" species="subrufa">
|
||
<emphasis id="71DFEA9BF924FFD3FEE0FC37FEBFFC18" box="[259,374,910,935]" italics="true" pageId="19" pageNumber="542">P. subrufa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AEC57E0F924FFD3FE9CFC37FDC4FC18" box="[383,525,910,935]" pageId="19" pageNumber="542" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
differs from all other
|
||
<taxonomicName id="84AB4D0AF924FFD3FCECFC36FC3BFC18" box="[783,1010,910,935]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="71DFEA9BF924FFD3FCECFC36FC51FC18" box="[783,920,910,935]" italics="true" pageId="19" pageNumber="542">Pelomedusa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName id="84AB4D0AF924FFD3FBADFC37FB09FC18" baseAuthorityName="Schoepff" baseAuthorityYear="1792" box="[1102,1216,910,935]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="71DFEA9BF924FFD3FBADFC37FB09FC18" box="[1102,1216,910,935]" italics="true" pageId="19" pageNumber="542">P. galeata</emphasis>
|
||
</taxonomicName>
|
||
by the presence of cytosine (C) instead of adenine (A) or guanine (G) at position 148, by the presence of guanine (G) instead of adenine (A) at position 159, by the presence of cytosine (C) instead of thymine (T) at position 167, and by the presence of guanine (G) instead of adenine (A) or thymine (T) at position 343. Moreover,
|
||
<taxonomicName id="84AB4D0AF924FFD3FB75FC43FAC1FBAC" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[1174,1288,1018,1043]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" sensu="stricto" species="subrufa">
|
||
<emphasis id="71DFEA9BF924FFD3FB75FC43FAC1FBAC" box="[1174,1288,1018,1043]" italics="true" pageId="19" pageNumber="542">P. subrufa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AEC57E0F924FFD3FAF3FC43FA55FBAC" box="[1296,1436,1018,1043]" pageId="19" pageNumber="542" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
differs from all other
|
||
<taxonomicName id="84AB4D0AF924FFD3FE69FBA6FDA5FB88" box="[394,620,1054,1079]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="71DFEA9BF924FFD3FE69FBA6FDDAFB88" box="[394,531,1054,1079]" italics="true" pageId="19" pageNumber="542">Pelomedusa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName id="84AB4D0AF924FFD3FD27FBA7FCFCFB88" baseAuthorityName="Schoepff" baseAuthorityYear="1792" box="[708,821,1054,1079]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="71DFEA9BF924FFD3FD27FBA7FCFCFB88" box="[708,821,1054,1079]" italics="true" pageId="19" pageNumber="542">P. galeata</emphasis>
|
||
</taxonomicName>
|
||
and possibly
|
||
<taxonomicName id="84AB4D0AF924FFD3FC31FBA7FB88FB88" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[978,1089,1054,1079]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="71DFEA9BF924FFD3FC31FBA7FB88FB88" box="[978,1089,1054,1079]" italics="true" pageId="19" pageNumber="542">P. gehafie</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="84AB4D0AF924FFD3FBACFBA7FB69FB88" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1103,1184,1054,1079]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="kobe">
|
||
<emphasis id="71DFEA9BF924FFD3FBACFBA7FB69FB88" box="[1103,1184,1054,1079]" italics="true" pageId="19" pageNumber="542">P. kobe</emphasis>
|
||
</taxonomicName>
|
||
and candidate species B (in which the respective character states are unknown) by the presence of thymine (T) instead of cytosine (C) at position 26 and by the presence of cytosine (C) instead of thymine (T) at position 38.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F924FFD3FF24FB32FECAFAB0" pageId="19" pageNumber="542" type="distribution">
|
||
<paragraph id="43143689F924FFD3FF24FB32FECAFAB0" blockId="19.[151,1437,514,1979]" pageId="19" pageNumber="542">
|
||
<emphasis id="71DFEA9BF924FFD3FF24FB32FEABFB1C" bold="true" box="[199,354,1162,1187]" pageId="19" pageNumber="542">Distribution:</emphasis>
|
||
Southern
|
||
<collectingCountry id="3BBC7619F924FFD3FE02FB32FDFDFB1C" box="[481,564,1162,1187]" name="Angola" pageId="19" pageNumber="542">Angola</collectingCountry>
|
||
(this study),
|
||
<collectingCountry id="3BBC7619F924FFD3FD39FB33FC82FB1D" box="[730,843,1163,1186]" name="Botswana" pageId="19" pageNumber="542">Botswana</collectingCountry>
|
||
, southeastern
|
||
<collectingCountry id="3BBC7619F924FFD3FC16FB32FA5EFB1C" box="[1013,1431,1162,1187]" name="Democratic Republic of the Congo" pageId="19" pageNumber="542">Democratic Republic of the Congo</collectingCountry>
|
||
,
|
||
<collectingCountry id="3BBC7619F924FFD3FF74FB16FF25FB78" box="[151,236,1198,1223]" name="Malawi" pageId="19" pageNumber="542">Malawi</collectingCountry>
|
||
(
|
||
<bibRefCitation id="273A4B78F924FFD3FF18FB16FDFAFB78" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[251,563,1198,1223]" pageId="19" pageNumber="542" pagination="428 - 440" refId="ref18768" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="71DFEA9BF924FFD3FE5BFB17FE39FB78" box="[440,496,1198,1223]" italics="true" pageId="19" pageNumber="542">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
),
|
||
<collectingCountry id="3BBC7619F924FFD3FDA5FB16FD6FFB78" box="[582,678,1198,1223]" name="Namibia" pageId="19" pageNumber="542">Namibia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="273A4B78F924FFD3FD56FB16FCB3FB78" author="Wong, R. A. & Fong, J. J. & Papenfuss, T. J." box="[693,890,1198,1223]" pageId="19" pageNumber="542" pagination="575 - 585" refId="ref19006" refString="Wong, R. A., Fong, J. J. & Papenfuss, T. J. (2010) Phylogeography of the African helmeted terrapin, Pelomedusa subrufa: genetic structure, dispersal, and human introduction. Proceedings of the California Academy of Sciences, 61, 575 - 585." type="journal article" year="2010">
|
||
Wong
|
||
<emphasis id="71DFEA9BF924FFD3FCE3FB17FCF0FB78" box="[768,825,1198,1223]" italics="true" pageId="19" pageNumber="542">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
; this study),
|
||
<collectingCountry id="3BBC7619F924FFD3FBE9FB16FB53FB78" box="[1034,1178,1198,1223]" name="South Africa" pageId="19" pageNumber="542">South Africa</collectingCountry>
|
||
(
|
||
<collectingRegion id="816FF86BF924FFD3FB4BFB16FADDFB78" box="[1192,1300,1198,1223]" country="South Africa" name="Limpopo" pageId="19" pageNumber="542">Limpopo</collectingRegion>
|
||
; this study), and
|
||
<collectingRegion id="816FF86BF924FFD3FF24FB6AFE57FB54" box="[199,414,1234,1259]" country="Tanzania" name="Kilimanjaro" pageId="19" pageNumber="542">Kilimanjaro region</collectingRegion>
|
||
of
|
||
<collectingCountry id="3BBC7619F924FFD3FE26FB6AFDE3FB54" box="[453,554,1234,1259]" name="Tanzania" pageId="19" pageNumber="542">Tanzania</collectingCountry>
|
||
(this study). Introduced to
|
||
<collectingCountry id="3BBC7619F924FFD3FC83FB6AFC2EFB54" box="[864,999,1234,1259]" name="Madagascar" pageId="19" pageNumber="542">Madagascar</collectingCountry>
|
||
(
|
||
<bibRefCitation id="273A4B78F924FFD3FC14FB6AFAF9FB54" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[1015,1328,1234,1259]" pageId="19" pageNumber="542" pagination="428 - 440" refId="ref18768" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="71DFEA9BF924FFD3FB57FB6BFB27FB54" box="[1204,1262,1234,1259]" italics="true" pageId="19" pageNumber="542">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="273A4B78F924FFD3FADEFB6BFF31FAB0" author="Wong, R. A. & Fong, J. J. & Papenfuss, T. J." pageId="19" pageNumber="542" pagination="575 - 585" refId="ref19006" refString="Wong, R. A., Fong, J. J. & Papenfuss, T. J. (2010) Phylogeography of the African helmeted terrapin, Pelomedusa subrufa: genetic structure, dispersal, and human introduction. Proceedings of the California Academy of Sciences, 61, 575 - 585." type="journal article" year="2010">
|
||
Wong
|
||
<emphasis id="71DFEA9BF924FFD3FA6BFB6BFF7DFAB0" italics="true" pageId="19" pageNumber="542">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F924FFD4FF24FAA3FAD1FE14" lastPageId="20" lastPageNumber="543" pageId="19" pageNumber="542" type="discussion">
|
||
<paragraph id="43143689F924FFD3FF24FAA3FBD8FAE9" blockId="19.[151,1437,514,1979]" pageId="19" pageNumber="542">
|
||
<emphasis id="71DFEA9BF924FFD3FF24FAA3FEF4FA8C" bold="true" box="[199,317,1307,1331]" pageId="19" pageNumber="542">Remarks:</emphasis>
|
||
<taxonomicName id="84AB4D0AF924FFD3FEA4FAA2FDF8FA8C" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[327,561,1306,1331]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" sensu="stricto" species="subrufa">
|
||
<emphasis id="71DFEA9BF924FFD3FEA4FAA2FDF8FA8C" box="[327,561,1306,1331]" italics="true" pageId="19" pageNumber="542">Pelomedusa subrufa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AEC57E0F924FFD3FDD8FAA3FD03FA8C" box="[571,714,1306,1331]" pageId="19" pageNumber="542" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
corresponds to mtDNA lineage VIII of
|
||
<bibRefCitation id="273A4B78F924FFD3FB41FAA2FF2CFAE8" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." pageId="19" pageNumber="542" pagination="428 - 440" refId="ref18768" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="71DFEA9BF924FFD3FA83FAA3FA55FA8C" box="[1376,1436,1306,1331]" italics="true" pageId="19" pageNumber="542">et al.</emphasis>
|
||
(2010)
|
||
</bibRefCitation>
|
||
. Besides
|
||
<taxonomicName id="84AB4D0AF924FFD3FEAEFA87FE74FAE8" baseAuthorityName="Schoepff" baseAuthorityYear="1792" box="[333,445,1342,1367]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="71DFEA9BF924FFD3FEAEFA87FE74FAE8" box="[333,445,1342,1367]" italics="true" pageId="19" pageNumber="542">P. galeata</emphasis>
|
||
</taxonomicName>
|
||
, it is one of the two southern species of
|
||
<taxonomicName id="84AB4D0AF924FFD3FC62FA86FBC3FAE8" authorityName="Wagler" authorityYear="1830" box="[897,1034,1342,1367]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="genus">
|
||
<emphasis id="71DFEA9BF924FFD3FC62FA86FBC3FAE8" box="[897,1034,1342,1367]" italics="true" pageId="19" pageNumber="542">Pelomedusa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="43143689F924FFD3FF24FADBFBA2FA58" blockId="19.[151,1437,514,1979]" pageId="19" pageNumber="542">
|
||
In the course of the present study, we recorded
|
||
<taxonomicName id="84AB4D0AF924FFD3FD30FADBFC8DFAC4" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[723,836,1378,1403]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" sensu="stricto" species="subrufa">
|
||
<emphasis id="71DFEA9BF924FFD3FD30FADBFC8DFAC4" box="[723,836,1378,1403]" italics="true" pageId="19" pageNumber="542">P. subrufa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AEC57E0F924FFD3FCA8FADBFC1CFAC4" box="[843,981,1378,1403]" pageId="19" pageNumber="542" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
for the first time for
|
||
<collectingCountry id="3BBC7619F924FFD3FB5CFADAFA98FAC4" box="[1215,1361,1378,1403]" name="South Africa" pageId="19" pageNumber="542">South Africa</collectingCountry>
|
||
, based on one sample from the western border region of the Kruger Park (Hoedspruit,
|
||
<collectingRegion id="816FF86BF924FFD3FBD3FA3EFB50FA20" box="[1072,1177,1414,1439]" country="South Africa" name="Limpopo" pageId="19" pageNumber="542">Limpopo</collectingRegion>
|
||
,
|
||
<collectingCountry id="3BBC7619F924FFD3FB44FA3EFAF6FA20" box="[1191,1343,1414,1439]" name="South Africa" pageId="19" pageNumber="542">South Africa</collectingCountry>
|
||
). To the west and south, the next genetically verified records are approximately
|
||
<quantity id="84539B6CF924FFD3FC59FA12FBC6FA7C" box="[954,1039,1450,1475]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="542" unit="km" value="300.0">300 km</quantity>
|
||
distant and are
|
||
<taxonomicName id="84AB4D0AF924FFD3FB22FA13FAF8FA7C" baseAuthorityName="Schoepff" baseAuthorityYear="1792" box="[1217,1329,1450,1475]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="71DFEA9BF924FFD3FB22FA13FAF8FA7C" box="[1217,1329,1450,1475]" italics="true" pageId="19" pageNumber="542">P. galeata</emphasis>
|
||
</taxonomicName>
|
||
. It seems possible that the ranges of the two species abut or overlap in northeastern
|
||
<collectingCountry id="3BBC7619F924FFD3FC30FA76FBAFFA58" box="[979,1126,1486,1511]" name="South Africa" pageId="19" pageNumber="542">South Africa</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="43143689F924FFD3FF24FA4BFED6F8B8" blockId="19.[151,1437,514,1979]" pageId="19" pageNumber="542">
|
||
In the Zoologische Staatssammlung München there are genetically verified specimens of
|
||
<taxonomicName id="84AB4D0AF924FFD3FB3EFA4BFA9BF9B4" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[1245,1362,1522,1547]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" sensu="stricto" species="subrufa">
|
||
<emphasis id="71DFEA9BF924FFD3FB3EFA4BFA9BF9B4" box="[1245,1362,1522,1547]" italics="true" pageId="19" pageNumber="542">P. subrufa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AEC57E0F924FFD3FABDFA4BFF14F990" pageId="19" pageNumber="542" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
collected on
|
||
<date id="37151049F924FFD3FE96F9AEFE37F990" box="[373,510,1558,1583]" pageId="19" pageNumber="542" value="1937-05-03">3 May 1937</date>
|
||
by Wolfgang Uthmöller (
|
||
<date id="37151049F924FFD3FCC1F9AEFC28F990" box="[802,993,1558,1583]" pageId="19" pageNumber="542" value="1904-10-25">25 October 1904</date>
|
||
–
|
||
<date id="37151049F924FFD3FC13F9AEFB6DF990" box="[1008,1188,1558,1583]" pageId="19" pageNumber="542" value="1951-08-25">25 August 1951</date>
|
||
) at the northern slope of the
|
||
<collectingRegion id="816FF86BF924FFD3FF01F982FEA5F9EC" box="[226,364,1594,1619]" country="Tanzania" name="Kilimanjaro" pageId="19" pageNumber="542">Kilimanjaro</collectingRegion>
|
||
,
|
||
<collectingCountry id="3BBC7619F924FFD3FE9AF982FE17F9EC" box="[377,478,1594,1619]" name="Tanzania" pageId="19" pageNumber="542">Tanzania</collectingCountry>
|
||
(ZSM 320/1937:1–4), and additional genetically verified specimens with the date
|
||
<date id="37151049F924FFD3FA6DF982FED9F9C8" pageId="19" pageNumber="542" value="1937-05-05">5 May 1937</date>
|
||
originating from the same collector, but bearing only “Tanganyika” as locality data (ZSM 285/ 1937:1–3). The latter specimens represent another species (
|
||
<taxonomicName id="84AB4D0AF924FFD3FCD4F93BFC40F924" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[823,905,1666,1691]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="kobe">
|
||
<emphasis id="71DFEA9BF924FFD3FCD4F93BFC40F924" box="[823,905,1666,1691]" italics="true" pageId="19" pageNumber="542">P. kobe</emphasis>
|
||
</taxonomicName>
|
||
, see above), suggesting that both taxa occur in close geographical proximity or even sympatry in
|
||
<collectingCountry id="3BBC7619F924FFD3FD37F91EFCF5F900" box="[724,828,1702,1727]" name="Tanzania" pageId="19" pageNumber="542">Tanzania</collectingCountry>
|
||
. This is also supported by further genetically verified specimens of
|
||
<taxonomicName id="84AB4D0AF924FFD3FED1F973FE4AF95C" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[306,387,1738,1763]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="kobe">
|
||
<emphasis id="71DFEA9BF924FFD3FED1F973FE4AF95C" box="[306,387,1738,1763]" italics="true" pageId="19" pageNumber="542">P. kobe</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<collectingRegion id="816FF86BF924FFD3FE10F972FD5EF95C" box="[499,663,1738,1763]" country="Tanzania" name="Arusha" pageId="19" pageNumber="542">Arusha region</collectingRegion>
|
||
,
|
||
<collectingCountry id="3BBC7619F924FFD3FD41F972FCC0F95C" box="[674,777,1738,1763]" name="Tanzania" pageId="19" pageNumber="542">Tanzania</collectingCountry>
|
||
, from where also the new species
|
||
<taxonomicName id="84AB4D0AF924FFD3FB6EF973FADEF95D" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1165,1303,1739,1762]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF924FFD3FB6EF973FADEF95D" box="[1165,1303,1739,1762]" italics="true" pageId="19" pageNumber="542">P. neumanni</emphasis>
|
||
</taxonomicName>
|
||
is recorded (see above).
|
||
</paragraph>
|
||
<paragraph id="43143689F924FFD3FF24F8AAFBBBF8F0" blockId="19.[151,1437,514,1979]" pageId="19" pageNumber="542">
|
||
The identity of mtDNA sequences of
|
||
<taxonomicName id="84AB4D0AF924FFD3FD71F8ABFCC0F894" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[658,777,1810,1835]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" sensu="stricto" species="subrufa">
|
||
<emphasis id="71DFEA9BF924FFD3FD71F8ABFCC0F894" box="[658,777,1810,1835]" italics="true" pageId="19" pageNumber="542">P. subrufa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AEC57E0F924FFD3FCF5F8ABFC61F894" box="[790,936,1810,1835]" pageId="19" pageNumber="542" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
from
|
||
<collectingCountry id="3BBC7619F924FFD3FC1BF8AAFB94F894" box="[1016,1117,1810,1835]" name="Tanzania" pageId="19" pageNumber="542">Tanzania</collectingCountry>
|
||
and
|
||
<collectingCountry id="3BBC7619F924FFD3FB43F8AAFAEEF894" box="[1184,1319,1810,1835]" name="Madagascar" pageId="19" pageNumber="542">Madagascar</collectingCountry>
|
||
points to
|
||
<collectingCountry id="3BBC7619F924FFD3FF74F88EFF35F8F0" box="[151,252,1846,1871]" name="Tanzania" pageId="19" pageNumber="542">Tanzania</collectingCountry>
|
||
as geographical source region for the introduced Malagasy helmeted terrapins.
|
||
</paragraph>
|
||
<paragraph id="43143689F924FFD3FF24F8E2FD05F804" blockId="19.[151,1437,514,1979]" pageId="19" pageNumber="542">
|
||
Among 94 museum specimens of
|
||
<taxonomicName id="84AB4D0AF924FFD3FDA6F8E3FD7EF8CC" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[581,695,1882,1907]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="542" phylum="Chordata" rank="species" species="subrufa">
|
||
<emphasis id="71DFEA9BF924FFD3FDA6F8E3FD7EF8CC" box="[581,695,1882,1907]" italics="true" pageId="19" pageNumber="542">P. subrufa</emphasis>
|
||
</taxonomicName>
|
||
, two terrapins have triangular pectorals without midseam contact, and 19 terrapins have pectoral scutes with narrow plastral midseam contact, whilst the remaining specimens have pectoral scutes with broad contact at the midseam.
|
||
</paragraph>
|
||
<paragraph id="43143689F923FFD4FF24FF2FFE4BFEFF" blockId="20.[151,1437,151,429]" pageId="20" pageNumber="543">
|
||
<taxonomicName id="84AB4D0AF923FFD4FF24FF2FFE64FF0F" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[199,429,151,176]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="subrufa">
|
||
<emphasis id="71DFEA9BF923FFD4FF24FF2FFE64FF0F" box="[199,429,151,176]" italics="true" pageId="20" pageNumber="543">Pelomedusa subrufa</emphasis>
|
||
</taxonomicName>
|
||
can cope with extremely arid conditions. For
|
||
<collectingCountry id="3BBC7619F923FFD4FC51FF2FFBDCFF0F" box="[946,1045,151,176]" name="Namibia" pageId="20" pageNumber="543">Namibia</collectingCountry>
|
||
, there are records from the mouths of temporary streams (so-called riviers) in the Namib Desert (
|
||
<bibRefCitation id="273A4B78F923FFD4FCABFF04FC15FF6A" author="Branch, B." box="[840,988,188,213]" pageId="20" pageNumber="543" refId="ref15176" refString="Branch, B. (2008) Tortoises, Terrapins & Turtles of Africa. Struik Publishers, Cape Town, 128 pp." type="book" year="2008">Branch 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="273A4B78F923FFD4FC04FF04FAC6FF6A" author="Boycott, R. C. & Bourquin, O." box="[999,1295,188,213]" pageId="20" pageNumber="543" pagination="1788" refId="ref15032" refString="Boycott, R. C. & Bourquin, O. (2008) Pelomedusa subrufa (Lacepede 1788), helmeted turtle, helmeted terrapin. In: Rhodin, A. G. J., Pritchard, P. C. H., van Dijk, P. P., Saumure, R. A., Buhlmann, K. A. & Iverson, J. B. (Eds.), Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN / SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs, No. 5. Chelonian Research Foundation, Lunenburg, MA, pp. 007.1 - 007.6. http: // dx. doi. org / 10.3854 / crm. 5.007. subrufa. v 1.2008" type="book chapter" year="2008">Boycott & Bourquin 2008</bibRefCitation>
|
||
; vouchers in the Museum für Naturkunde, Berlin). It is well known that Namibian terrapins burrow underground during arid periods, and if there is no rain, they may evidently survive up to six years burrowed (A. Schleicher, pers. observ.,
|
||
<collectingRegion id="816FF86BF923FFD4FF74FE9FFECCFEFF" box="[151,261,295,320]" country="Namibia" name="Omaheke" pageId="20" pageNumber="543">Omaheke</collectingRegion>
|
||
,
|
||
<collectingCountry id="3BBC7619F923FFD4FEF3FE9FFEBFFEFF" box="[272,374,295,320]" name="Namibia" pageId="20" pageNumber="543">Namibia</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="43143689F923FFD4FF24FEF4FAD1FE14" blockId="20.[151,1437,151,429]" pageId="20" pageNumber="543">
|
||
The monogenean flatworm
|
||
<taxonomicName id="84AB4D0AF923FFD4FDE3FEF4FBECFEDA" authority="Euzet & Combes, 1965" authorityName="Euzet & Combes" authorityYear="1965" box="[512,1061,332,357]" class="Monogenea" family="Polystomatidae" genus="Polystomoides" kingdom="Animalia" order="Polystomatidea" pageId="20" pageNumber="543" phylum="Platyhelminthes" rank="species" species="chabaudi">
|
||
<emphasis id="71DFEA9BF923FFD4FDE3FEF4FCDDFEDA" box="[512,788,332,357]" italics="true" pageId="20" pageNumber="543">Polystomoides chabaudi</emphasis>
|
||
<bibRefCitation id="273A4B78F923FFD4FCFFFEF4FBECFEDA" author="Euzet, L. & Combes, C." box="[796,1061,332,357]" pageId="20" pageNumber="543" pagination="40" refId="ref15622" refString="Euzet, L. & Combes, C. (1965) Parasites des cheloniens malgaches. Polystomoides chabaudi n. sp. (Monogenea) chez la tortue d'eau douce Pelomedusa subrufa Lacepede 1788. Annales de Parasitologie Humaine et Comparee, 40, 455 - 450." type="journal article" year="1965">Euzet & Combes, 1965</bibRefCitation>
|
||
</taxonomicName>
|
||
was described from
|
||
<taxonomicName id="84AB4D0AF923FFD4FAF0FEF4FF24FE37" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" sensu="stricto" species="subrufa">
|
||
<emphasis id="71DFEA9BF923FFD4FAF0FEF4FF24FE37" italics="true" pageId="20" pageNumber="543">Pelomedusa subrufa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6AEC57E0F923FFD4FF16FEC8FE4AFE37" box="[245,387,367,392]" pageId="20" pageNumber="543" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
(collected at Betioky,
|
||
<collectingCountry id="3BBC7619F923FFD4FD64FED7FCDDFE37" box="[647,788,367,392]" name="Madagascar" pageId="20" pageNumber="543">Madagascar</collectingCountry>
|
||
). The same polystomatid species was also reported from helmeted terrapins from the
|
||
<collectingRegion id="816FF86BF923FFD4FE37FE2CFDF0FE12" box="[468,569,404,429]" country="Uganda" name="Kampala" pageId="20" pageNumber="543">Kampala</collectingRegion>
|
||
area,
|
||
<collectingCountry id="3BBC7619F923FFD4FD9EFE2CFD1DFE12" box="[637,724,404,429]" name="Uganda" pageId="20" pageNumber="543">Uganda</collectingCountry>
|
||
(
|
||
<bibRefCitation id="273A4B78F923FFD4FD00FE2CFCB0FE12" author="Tinsley, R. C." box="[739,889,404,429]" pageId="20" pageNumber="543" pagination="251 - 263" refId="ref18436" refString="Tinsley, R. C. (1973) Observations on Polystomatidae (Monogenoidea) from East Africa with a description of Polystoma makereri n. sp. Zeitschrift fur Parasitenkunde, 42, 251 - 263. http: // dx. doi. org / 10.1007 / bf 00328887" type="journal article" year="1973">Tinsley 1973</bibRefCitation>
|
||
), that is, probably from
|
||
<taxonomicName id="84AB4D0AF923FFD4FB64FE2DFAD8FE13" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1159,1297,405,428]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="543" phylum="Chordata" rank="species" species="neumanni">
|
||
<emphasis id="71DFEA9BF923FFD4FB64FE2DFAD8FE13" box="[1159,1297,405,428]" italics="true" pageId="20" pageNumber="543">P. neumanni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |