1140 lines
136 KiB
XML
1140 lines
136 KiB
XML
<document id="3F033EEADD81719FF3B01EF3018EC2DB" ID-DOI="10.15298/rusentj.32.1.03" ID-ISSN="0132-8069" ID-Zenodo-Dep="10949096" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712198792598" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2023" docId="2F4187C3FFC5167F79DBFAC12301704E" docLanguage="en" docName="RussEntomolJ.32.1.16-39.pdf" docOrigin="Russian Entomological Journal 32 (1)" docSource="http://dx.doi.org/10.15298/rusentj.32.1.03" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.4:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="4" docTitle="Argutor (Argutor) Dejean 1821" docType="treatment" docVersion="4" lastPageNumber="34" masterDocId="D378FFBBFFCB166D7966FF9A2163720F" masterDocTitle="New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam" masterLastPageNumber="39" masterPageNumber="16" pageNumber="30" updateTime="1731592677940" updateUser="julia">
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<mods:titleInfo id="B54D9DC58158286D60F4990F57B77FFF">
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<mods:title id="E271BAF1FB40824A14190ADAC067A9BC">New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam</mods:title>
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</mods:titleInfo>
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<mods:name id="2535A0FBE4C15762E59F4BC4397FF01A" type="personal">
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<mods:namePart id="EFD878EE571D185C65BB5CDCB19AD3FB">Fedorenko, D. N.</mods:namePart>
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<mods:typeOfResource id="D563C594022FEA327B32241A8F22101A">text</mods:typeOfResource>
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<mods:title id="08642E34A376DBA5286A510D0EE8AC63">Russian Entomological Journal</mods:title>
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<mods:part id="114C1D071DAE7D406B1B2918385FF17C">
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<mods:date id="65CBD431455343E6F963162E266E15E7">2023</mods:date>
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<mods:number id="CDF9785C562B2D8B4948E0ECD6E697EB">2023-03-31</mods:number>
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<mods:number id="B7CCBEC83B48C0F322C83F171B5D9760">32</mods:number>
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<mods:classification id="34BAA4D727A3AFCD24C442581A56FDAE">journal article</mods:classification>
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<mods:identifier id="575153F2C39E967FCC31729A98686D6F" type="DOI">10.15298/rusentj.32.1.03</mods:identifier>
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<mods:identifier id="3D4C321F4313DD8422E02A2D81C7A168" type="ISSN">0132-8069</mods:identifier>
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<treatment id="2F4187C3FFC5167F79DBFAC12301704E" LSID="urn:lsid:plazi:treatment:2F4187C3FFC5167F79DBFAC12301704E" httpUri="http://treatment.plazi.org/id/2F4187C3FFC5167F79DBFAC12301704E" lastPageId="18" lastPageNumber="34" pageId="14" pageNumber="30">
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<paragraph id="A75736D5FFC5166379DBFAC123EB777D" blockId="14.[189,776,1371,2069]" box="[189,648,1371,1394]" pageId="14" pageNumber="30">
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<heading id="FC1F81B9FFC5166379DBFAC123EB777D" box="[189,648,1371,1394]" fontSize="8" level="2" pageId="14" pageNumber="30" reason="6">
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<smallCapsWord id="A1B1A009FFC516637989FAFA206B777E" baselines="1389" box="[239,264,1376,1393]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="to" pageId="14" pageNumber="30">TO</smallCapsWord>
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<smallCapsWord id="A1B1A009FFC516637869FAFA201C777E" baselines="1389" box="[271,383,1376,1393]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="subgenera" pageId="14" pageNumber="30">SUBGENERA</smallCapsWord>
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<smallCapsWord id="A1B1A009FFC5166378E0FAFA20FD777E" baselines="1389" box="[390,414,1376,1393]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="of" pageId="14" pageNumber="30">OF</smallCapsWord>
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<smallCapsWord id="A1B1A009FFC5166378C2FAFA20AB777F" baselines="1388" box="[420,456,1376,1392]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="the" pageId="14" pageNumber="30">THE</smallCapsWord>
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||
<taxonomicName id="60E84D56FFC5166378A9FAC1234A777F" ID-CoL="8LTWJ" authority="Dejean, 1821" authorityName="Dejean" authorityPageNumber="11" authorityYear="1821" box="[463,553,1371,1394]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<smallCapsWord id="A1B1A009FFC5166378A9FAC1234A777F" baselines="1388,1388" box="[463,553,1371,1394]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Argutor" pageId="14" pageNumber="30">
|
||
<emphasis id="959CEAC7FFC5166378A9FAC1234A777F" box="[463,553,1371,1394]" italics="true" pageId="14" pageNumber="30">ARGUTOR</emphasis>
|
||
</smallCapsWord>
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</taxonomicName>
|
||
<smallCapsWord id="A1B1A009FFC516637B56FAFA23E2777E" baselines="1389" box="[560,641,1376,1393]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="lineage" pageId="14" pageNumber="30">LINEAGE</smallCapsWord>
|
||
:
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFF2655EFFC5166379DBFAE52417771F" pageId="14" pageNumber="30" type="key">
|
||
<key id="D37961E4FFC5166379DBFAE52417771F" pageId="14" pageNumber="30">
|
||
<paragraph id="A75736D5FFC5166379DBFAE5231577EA" blockId="14.[189,776,1371,2069]" pageId="14" pageNumber="30">
|
||
<keyLead id="1C1995E0FFC5166379DBFAE5231577EA" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC5166379DBFAE5231577EA" pageId="14" pageNumber="30">1(10) Elytra with parascutellar seta each, mostly elongated and more or less parallel-sided. Body mostly macropterous; metepisternum long, often much longer than wide. Tarsomere 5 ventrally setose to glabrous.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC5166379DBFA70226774E0" blockId="14.[189,776,1371,2069]" pageId="14" pageNumber="30">
|
||
<keyLead id="1C1995E0FFC5166379DBFA70226774E0" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC5166379DBFA70226774E0" pageId="14" pageNumber="30">
|
||
2(3) Prosternal process apically beaded. Tarsi without or with a median longitudinal sulcus in addition to dorsolateral sulci in
|
||
<emphasis id="959CEAC7FFC51663785AF9BA202C743A" box="[316,335,1568,1589]" italics="true" pageId="14" pageNumber="30">ta</emphasis>
|
||
2 and
|
||
<emphasis id="959CEAC7FFC5166378F1F9BA20C9743A" box="[407,426,1568,1589]" italics="true" pageId="14" pageNumber="30">ta</emphasis>
|
||
3, conspicuous anterior and more shallow posterior;
|
||
<emphasis id="959CEAC7FFC5166378FBF9A120C9745F" box="[413,426,1595,1616]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
3 with 1–3 lateral setae. Pronotum with single, inner, basolateral sulcus on each side. Elytra with parascutellar striole missing and three (discal) setae, those d2 and d3 adjoining or proximate to stria 2. Spermatheca undifferentiated to nearly differentiated, medium-sized, annulated toward apex;
|
||
<emphasis id="959CEAC7FFC516637B4CF92523AA74DB" box="[554,713,1727,1748]" italics="true" pageId="14" pageNumber="30">bursa copulatrix</emphasis>
|
||
medium-sized to long. — Holarctic. .......................
|
||
<emphasis id="959CEAC7FFC516637BD7F940226774E0" box="[689,772,1754,1775]" italics="true" pageId="14" pageNumber="30">
|
||
<taxonomicName id="60E84D56FFC516637BD7F940226274E0" authorityName="Dejean" authorityYear="1821" box="[689,769,1754,1775]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">Argutor</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC5166379DBF96F2387754F" blockId="14.[189,776,1371,2069]" pageId="14" pageNumber="30">
|
||
<keyLead id="1C1995E0FFC5166379DBF96F2387754F" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC5166379DBF96F2387754F" pageId="14" pageNumber="30">
|
||
3(2) Prosternal process not apically beaded;
|
||
<emphasis id="959CEAC7FFC516637B06F96F23117505" box="[608,626,1781,1802]" italics="true" pageId="14" pageNumber="30">ta</emphasis>
|
||
2 and
|
||
<emphasis id="959CEAC7FFC516637BCBF96F23DC7505" box="[685,703,1781,1802]" italics="true" pageId="14" pageNumber="30">ta</emphasis>
|
||
3 bisulcate dorsolaterally, posterior sulcus ranging from shallow to indistinct. Pronotum mostly bisulcate on each side.
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC5166379DBF8DE226775D0" blockId="14.[189,776,1371,2069]" pageId="14" pageNumber="30">
|
||
<keyLead id="1C1995E0FFC5166379DBF8DE226775D0" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC5166379DBF8DE226775D0" pageId="14" pageNumber="30">
|
||
4(5) Elytron with 1–3 discal setae, d2 and/or d3 adjoining stria 2; parascutellar striole long;
|
||
<emphasis id="959CEAC7FFC516637B64F8FA2376757A" box="[514,533,1888,1909]" italics="true" pageId="14" pageNumber="30">ta</emphasis>
|
||
2 and
|
||
<emphasis id="959CEAC7FFC516637B30F8FA230A757A" box="[598,617,1888,1909]" italics="true" pageId="14" pageNumber="30">ta</emphasis>
|
||
3 mostly with a distinct posterior dorsolateral sulcus;
|
||
<emphasis id="959CEAC7FFC516637B2EF8E12336759F" box="[584,597,1915,1936]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
3 asetose laterally. Female gonosubcoxite IX with 3–4 latero-apical setae. Spermatheca and
|
||
<emphasis id="959CEAC7FFC5166378FAF835232675CB" box="[412,581,1967,1988]" italics="true" pageId="14" pageNumber="30">bursa copulatrix</emphasis>
|
||
medium-sized. — Holarctic. ..........................................................
|
||
<emphasis id="959CEAC7FFC516637BC8F850226775D0" box="[686,772,1994,2015]" italics="true" pageId="14" pageNumber="30">
|
||
<taxonomicName id="60E84D56FFC516637BC8F850226375D0" authorityName="Gozis" authorityYear="1886" box="[686,768,1994,2015]" pageId="14" pageNumber="38" rank="subGenus" subGenus="Phonias">Phonias</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC5166379DBF87E25837364" blockId="14.[189,776,1371,2069]" lastBlockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">
|
||
<keyLead id="1C1995E0FFC5166379DBF87E22647A1A" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC5166379DBF87E22647A1A" pageId="14" pageNumber="30">
|
||
5(4) Elytron with 1–3 discal setae adjoining stria 3; parascutellar striole long to missing;
|
||
<emphasis id="959CEAC7FFC516637B73F79A234B7A1A" box="[533,552,2048,2069]" italics="true" pageId="14" pageNumber="30">ta</emphasis>
|
||
2 and
|
||
<emphasis id="959CEAC7FFC516637B39F79A23117A1A" box="[607,626,2048,2069]" italics="true" pageId="14" pageNumber="30">ta</emphasis>
|
||
3 with posterior
|
||
</keyStep>
|
||
</keyLead>
|
||
<keyLead id="1C1995E0FFC516637A34FEA125837364" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC516637A34FEA125837364" pageId="14" pageNumber="30">dorsolateral sulcus vestigial or indistinct. Female gonosubcoxite IX without latero-apical setae.</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC516637A4AFEEA2417704F" blockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">
|
||
<keyLead id="1C1995E0FFC516637A4AFEEA2417704F" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC516637A4AFEEA2417704F" pageId="14" pageNumber="30">
|
||
6(7) Pronotum much wider at base than at apex, PB/PA 1.4– 1.6, with lateral margin slightly explanate between fine lateral bead and a distinct sublateral line inside. Elytron with 2–3 discal setae;
|
||
<emphasis id="959CEAC7FFC516637D42FE5B255273D9" box="[1060,1073,449,470]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
3 with 1–3 lateral setae (occasionally absent from one side).
|
||
<emphasis id="959CEAC7FFC516637D31FE46259A73FF" box="[1111,1273,475,497]" italics="true" pageId="14" pageNumber="30">Bursa copulatrix</emphasis>
|
||
and, accordingly, internal sac of aedeagus increased much in length (
|
||
<figureCitation id="3FD32A50FFC516637A3FFD8B22CC7028" box="[857,943,529,551]" captionStart="Figs 100–104" captionStartId="20.[226,264,1897,1916]" captionTargetBox="[197,1386,626,1869]" captionTargetId="figure-320@20.[189,1399,625,1869]" captionTargetPageId="20" captionText="Figs 100–104. Reproductive tract in female: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — bursa copulatrix; dv — diverticulum; ov — common oviduct; rp — receptacle; sc — seminal canal; sg — spermathecal gland; 63, 65–67 — ventral aspect; 64, 68 — dorsal aspect; sgd — spermathecal gland duct; sp — spermatheca. Scale bars 1 mm. Рис. 100–104. Репродуктивный тракт самки: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — копулЯтивнаЯ сумка; dv — дивертикул; ov — непарный Яйцевод; rp — семеприемник; sc — семепровод; sg — ЖелеЗа сперматеки; sgd — проток ЖелеЗы сперматеки; sp — сперматека. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949120" httpUri="https://zenodo.org/record/10949120/files/figure.png" pageId="14" pageNumber="30">Fig. 100</figureCitation>
|
||
), spermatheca short. — Eastern Palearctic. ... ....................................................................
|
||
<emphasis id="959CEAC7FFC516637D98FDB12417704F" box="[1278,1396,555,576]" italics="true" pageId="14" pageNumber="30">
|
||
<taxonomicName id="60E84D56FFC516637D98FDB12413704F" authorityName="Motschulsky" authorityYear="1866" box="[1278,1392,555,576]" class="Insecta" family="Carabidae" genus="Badistrinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">Badistrinus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC516637A4AFDDC247970A3" blockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">
|
||
<keyLead id="1C1995E0FFC516637A4AFDDC247970A3" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC516637A4AFDDC247970A3" pageId="14" pageNumber="30">
|
||
7(6) Pronotum convex up to fine lateral bead, rounded on sides, with base less wide relative to apex, PB/PA ~1.3. Elytron with 1–2 discal setae;
|
||
<emphasis id="959CEAC7FFC516637D1BFDE625E9709E" box="[1149,1162,636,657]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
3 without or with single lateral seta.
|
||
<emphasis id="959CEAC7FFC516637AAFFD0D250F70A4" box="[969,1132,662,684]" italics="true" pageId="14" pageNumber="30">Bursa copulatrix</emphasis>
|
||
moderately long.
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC516637A4AFD2B241770F3" blockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">
|
||
<keyLead id="1C1995E0FFC516637A4AFD2B241770F3" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC516637A4AFD2B241770F3" pageId="14" pageNumber="30">
|
||
8(9) Elytron with two discal setae, d2 and d3. Body small, BL 5–5.5 mm; elytra elongated. Spermatheca short. — Eastern Palearctic. ..................................?
|
||
<taxonomicName id="60E84D56FFC516637DB5FD7C244970F4" authorityName="Motschulsky" authorityYear="1866" box="[1235,1322,742,763]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC516637DB5FD7C244970F4" box="[1235,1322,742,763]" italics="true" pageId="14" pageNumber="30">Pledarus</emphasis>
|
||
</taxonomicName>
|
||
(part.).
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC516637A4AFC9B24177169" blockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">
|
||
<keyLead id="1C1995E0FFC516637A4AFC9B24177169" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC516637A4AFC9B24177169" pageId="14" pageNumber="30">
|
||
9(8) Elytron with single discal seta, d3. Body larger, BL
|
||
<quantity id="60109B30FFC516637C34FC9B2219713E" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="14" pageNumber="30" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
|
||
; elytra shorter, sides rounded. Spermatheca long, capitate and corrugated (
|
||
<figureCitation id="3FD32A50FFC516637D26FCAC25F77143" box="[1088,1172,822,844]" captionStart="Figs 100–104" captionStartId="20.[226,264,1897,1916]" captionTargetBox="[197,1386,626,1869]" captionTargetId="figure-320@20.[189,1399,625,1869]" captionTargetPageId="20" captionText="Figs 100–104. Reproductive tract in female: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — bursa copulatrix; dv — diverticulum; ov — common oviduct; rp — receptacle; sc — seminal canal; sg — spermathecal gland; 63, 65–67 — ventral aspect; 64, 68 — dorsal aspect; sgd — spermathecal gland duct; sp — spermatheca. Scale bars 1 mm. Рис. 100–104. Репродуктивный тракт самки: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — копулЯтивнаЯ сумка; dv — дивертикул; ov — непарный Яйцевод; rp — семеприемник; sc — семепровод; sg — ЖелеЗа сперматеки; sgd — проток ЖелеЗы сперматеки; sp — сперматека. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949120" httpUri="https://zenodo.org/record/10949120/files/figure.png" pageId="14" pageNumber="30">Fig. 103</figureCitation>
|
||
). — Eastern Palearctic. ........................................................................
|
||
<emphasis id="959CEAC7FFC516637C70FCCB24177169" box="[1302,1396,849,870]" italics="true" pageId="14" pageNumber="30">
|
||
<taxonomicName id="60E84D56FFC516637C70FCCB24137169" authorityName="Motschulsky" authorityYear="1866" box="[1302,1392,849,870]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">Pledarus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC516637A4AFCF1256671E3" blockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">
|
||
<keyLead id="1C1995E0FFC516637A4AFCF1256671E3" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC516637A4AFCF1256671E3" pageId="14" pageNumber="30">
|
||
10(1) Elytra without parascutellar seta, mostly shorter and more rounded on sides. Tarsomere 5 setose or setulose ventrally. Metepisternum shorter;
|
||
<emphasis id="959CEAC7FFC516637DFCFC3B25CE71B9" box="[1178,1197,929,950]" italics="true" pageId="14" pageNumber="30">ta</emphasis>
|
||
2 and
|
||
<emphasis id="959CEAC7FFC516637D81FC3B259971B9" box="[1255,1274,929,950]" italics="true" pageId="14" pageNumber="30">ta</emphasis>
|
||
3 with posterior dorsolateral sulcus missing or imperceptible on tarsomeres 1–2 only.
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyLead id="1C1995E0FFC516637A4AFC6B2417771F" pageId="14" pageNumber="30">
|
||
<keyStep id="1C1C2E70FFC516637A4AFC6B2417771F" pageId="14" pageNumber="30">
|
||
<paragraph id="A75736D5FFC516637A4AFC6B24177658" blockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">
|
||
11(12) Elytron with single discal seta, d3 adjoining stria 3; parascutellar striole short to missing. Pronotum with a deep fovea between inner and outer basolateral sulcus. Body apterous. ....................................
|
||
<taxonomicName id="60E84D56FFC516637DB4FBDB244A7659" authorityName="Motschulsky" authorityYear="1866" box="[1234,1321,1089,1110]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC516637DB4FBDB244A7659" box="[1234,1321,1089,1110]" italics="true" pageId="14" pageNumber="30">Pledarus</emphasis>
|
||
</taxonomicName>
|
||
(part.).
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC516637A4AFBC12417771F" blockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">
|
||
12(11) Elytron with three discal setae, d1 adjoining stria 3, d2 and d3 adjoining stria 2; parascutellar striole long. Pronotum more or less cordate, with single, inner, basolateral sulcus on each side; basal angles slightly acute to subrectangular, not toothed. Tarsomere 5 ventrally setose. Body mostly brachypterous to apterous. — Holarctic. ........... .....................................................................
|
||
<emphasis id="959CEAC7FFC516637C64FB612417771F" box="[1282,1396,1275,1296]" italics="true" pageId="14" pageNumber="30">
|
||
<taxonomicName id="60E84D56FFC516637C64FB612413771F" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[1282,1392,1275,1296]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">Omaseulus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</keyStep>
|
||
</keyLead>
|
||
</key>
|
||
</subSubSection>
|
||
<subSubSection id="EFF2655EFFC5167F7A34FA8C2301704E" lastPageId="18" lastPageNumber="34" pageId="14" pageNumber="30" type="discussion">
|
||
<paragraph id="A75736D5FFC516637A34FA8C25DF776E" blockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">Some significant characters vary between species of this lineage considerably, as well as between individuals in some of them. These characters are as follows.</paragraph>
|
||
<paragraph id="A75736D5FFC516637A34FAFC256B74FE" blockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">
|
||
1) Lateral sulci of
|
||
<emphasis id="959CEAC7FFC516637D60FAFD25707773" box="[1030,1043,1383,1404]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
2 and
|
||
<emphasis id="959CEAC7FFC516637D36FAFD253E7773" box="[1104,1117,1383,1404]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
3. — This sulcus is deep and well traceable in
|
||
<emphasis id="959CEAC7FFC516637AA8FA1B22B87799" box="[974,987,1409,1430]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
2, while being more shallow in
|
||
<taxonomicName id="60E84D56FFC516637C63FA1B24137799" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[1285,1392,1409,1430]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC516637C63FA1B24137799" box="[1285,1392,1409,1430]" italics="true" pageId="14" pageNumber="30">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
. Fine and deep sulcus that runs on at least apical 3/4–4/5
|
||
<emphasis id="959CEAC7FFC516637C21FA06243777BE" box="[1351,1364,1436,1457]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
3 is characteristic of
|
||
<taxonomicName id="60E84D56FFC516637AABFA2C257A77C4" authorityName="Dejean" authorityYear="1821" box="[973,1049,1462,1483]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC516637AABFA2C257A77C4" box="[973,1049,1462,1483]" italics="true" pageId="14" pageNumber="30">Argutor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFC516637D2FFA2C25FB77C4" authorityName="Gozis" authorityYear="1886" box="[1097,1176,1462,1483]" pageId="14" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFC516637D2FFA2C25FB77C4" box="[1097,1176,1462,1483]" italics="true" pageId="14" pageNumber="30">Phonias</emphasis>
|
||
</taxonomicName>
|
||
, but
|
||
<taxonomicName id="60E84D56FFC516637DACFA2D242E77C4" ID-CoL="4Q3FS" authorityName="" baseAuthorityName="Motschulsky" baseAuthorityYear="1866" box="[1226,1357,1462,1484]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="species" species="chameleon">
|
||
<emphasis id="959CEAC7FFC516637DACFA2D242E77C4" box="[1226,1357,1462,1484]" italics="true" pageId="14" pageNumber="30">P. chameleon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFC516637A4AFA4B257077E9" authority="Csiki, 1930" authorityName="Csiki" authorityYear="1930" box="[812,1043,1489,1511]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="species" species="taxonyis">
|
||
<emphasis id="959CEAC7FFC516637A4AFA4B22F977E9" box="[812,922,1489,1510]" italics="true" pageId="14" pageNumber="30">P. taxonyis</emphasis>
|
||
Csiki, 1930
|
||
</taxonomicName>
|
||
have this sulcus either shallow and shorter or vague, respectively. It is traceable in apical 2/3 tibia in
|
||
<taxonomicName id="60E84D56FFC516637A10F99C22867414" authorityName="Motschulsky" authorityYear="1866" box="[886,997,1542,1563]" class="Insecta" family="Carabidae" genus="Badistrinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC516637A10F99C22867414" box="[886,997,1542,1563]" italics="true" pageId="14" pageNumber="30">Badistrinus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFC516637D73F99C250F7414" authorityName="Motschulsky" authorityYear="1866" box="[1045,1132,1542,1563]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC516637D73F99C250F7414" box="[1045,1132,1542,1563]" italics="true" pageId="14" pageNumber="30">Pledarus</emphasis>
|
||
</taxonomicName>
|
||
, while ranging from nearly entire in some specimens of
|
||
<taxonomicName id="60E84D56FFC516637D06F9BB246F7439" ID-CoL="8LFDQ" authorityName="Tschitscherine" authorityYear="1889" box="[1120,1292,1569,1590]" class="Insecta" family="Carabidae" genus="Eurythorax" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="species" species="haptoderoides">
|
||
<emphasis id="959CEAC7FFC516637D06F9BB246F7439" box="[1120,1292,1569,1590]" italics="true" pageId="14" pageNumber="30">P. haptoderoides</emphasis>
|
||
</taxonomicName>
|
||
to hardly traceable in apical 1/3 tibia in
|
||
<taxonomicName id="60E84D56FFC516637D3FF9A625B3745F" authorityName="A. Morawitz" authorityYear="1862" box="[1113,1232,1595,1617]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="species" species="neglectus">
|
||
<emphasis id="959CEAC7FFC516637D3FF9A625B3745F" box="[1113,1232,1595,1617]" italics="true" pageId="14" pageNumber="30">P. neglectus</emphasis>
|
||
</taxonomicName>
|
||
and some specimens of
|
||
<taxonomicName id="60E84D56FFC516637AEEF9CD22857464" authorityName="Jedlicka" authorityYear="1930" box="[904,998,1622,1644]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="species" species="goschi">
|
||
<emphasis id="959CEAC7FFC516637AEEF9CD22857464" box="[904,998,1622,1644]" italics="true" pageId="14" pageNumber="30">P. goschi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="60E84D56FFC516637A91F9CC25017464" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[1015,1122,1622,1643]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC516637A91F9CC25017464" box="[1015,1122,1622,1643]" italics="true" pageId="14" pageNumber="30">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
have the most reduced
|
||
<emphasis id="959CEAC7FFC516637C3BF9CD24097463" box="[1373,1386,1623,1644]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
3 sulcus that ranges from shallow in apical 2/3–1/2 (
|
||
<taxonomicName id="60E84D56FFC516637C76F9EB22DC74AE" authority="(Sturm, 1824))" baseAuthorityName="Sturm" baseAuthorityYear="1824" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="species" species="diligens">
|
||
<emphasis id="959CEAC7FFC516637C76F9EB24157489" box="[1296,1398,1649,1670]" italics="true" pageId="14" pageNumber="30">P. diligens</emphasis>
|
||
(Sturm, 1824))
|
||
</taxonomicName>
|
||
through vague in apical 3/5–1/2 (the other species examined) to nearly indistinct (some specimens of many species) or missing (some specimens of
|
||
<taxonomicName id="60E84D56FFC516637C70F95B229E74FE" authority="Tschitscherine, 1899" authorityName="Tschitscherine" authorityYear="1899" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="species" species="eobius">
|
||
<emphasis id="959CEAC7FFC516637C70F95B241574D9" box="[1302,1398,1729,1750]" italics="true" pageId="14" pageNumber="30">P. eobius</emphasis>
|
||
<bibRefCitation id="C3794B24FFC516637A4AF941229E74FE" author="Tschitscherine T." box="[812,1021,1755,1777]" pageId="14" pageNumber="30" pagination="269 - 283" refId="ref20745" refString="Tschitscherine T. 1899. Tschitscherine T. 1899. Carabiques nouveaux ou peu connus // L'Abeille. Vol. 29. P. 269 - 283." type="journal article" year="1899">Tschitschérine, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC516637A34F96C24317554" blockId="14.[812,1399,315,2071]" pageId="14" pageNumber="30">
|
||
These sulci are most likely to be plesiomorphic character state in Pterostichitae, which follows from the fact that pronouncedly sulcate tibiae are peculiar to higher taxa such as,
|
||
<emphasis id="959CEAC7FFC516637A4AF8DD22337553" box="[812,848,1863,1884]" italics="true" pageId="14" pageNumber="30">e.g.</emphasis>
|
||
, Abacetina, Cratocerini, Morionini, Panagaeini, etc.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC516627A34F8FB23E673D9" blockId="14.[812,1399,315,2071]" lastBlockId="15.[189,775,315,791]" lastPageId="15" lastPageNumber="31" pageId="14" pageNumber="30">
|
||
2) Protibia anterior face with a median longitudinal sulcus. — This sulcus is not characteristic of
|
||
<taxonomicName id="60E84D56FFC516637DB7F8E12429759F" ID-CoL="8KVYP" authorityName="Bonelli" authorityYear="1810" box="[1233,1354,1915,1936]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC516637DB7F8E12429759F" box="[1233,1354,1915,1936]" italics="true" pageId="14" pageNumber="30">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
, but some species of the
|
||
<taxonomicName id="60E84D56FFC516637A90F80C252175A4" authorityName="Dejean" authorityYear="1821" box="[1014,1090,1942,1963]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC516637A90F80C252175A4" box="[1014,1090,1942,1963]" italics="true" pageId="14" pageNumber="30">Argutor</emphasis>
|
||
</taxonomicName>
|
||
lineage have a vestigial sulcus that runs on basal 1/3–2/5
|
||
<emphasis id="959CEAC7FFC516637D2DF82B253B75C9" box="[1099,1112,1969,1990]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
1. Out of the other species examined, only
|
||
<taxonomicName id="60E84D56FFC516637AAEF856241575EF" ID-CoL="4Q3PX" authority="Reitter, 1887" authorityName="Reitter" authorityYear="1887" box="[968,1398,1995,2017]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="species" species="pseudopedius" subGenus="Falsargutor">
|
||
<emphasis id="959CEAC7FFC516637AAEF85622BE75EE" box="[968,989,1996,2017]" italics="true" pageId="14" pageNumber="30">P.</emphasis>
|
||
(
|
||
<emphasis id="959CEAC7FFC516637A8DF851250375EF" box="[1003,1120,1995,2016]" italics="true" pageId="14" pageNumber="30">Falsargutor</emphasis>
|
||
)
|
||
<emphasis id="959CEAC7FFC516637D08F851259275EF" box="[1134,1265,1995,2016]" italics="true" pageId="14" pageNumber="30">pseudopedius</emphasis>
|
||
Reitter, 1887
|
||
</taxonomicName>
|
||
was found to have the anterior face of
|
||
<emphasis id="959CEAC7FFC516637DDBF87D25A975F3" box="[1213,1226,2023,2044]" italics="true" pageId="14" pageNumber="30">ti</emphasis>
|
||
1 with a fine yet complete sulcus, combined with another special feature, a row of a few short setae running along the inner margin below the antennal cleaner fissure. This species also has the abdominal sternite VIII and tergite VIII in female rather similar to those seen within the
|
||
<taxonomicName id="60E84D56FFC4166278F8FE16208973AE" authorityName="Dejean" authorityYear="1821" box="[414,490,396,417]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC4166278F8FE16208973AE" box="[414,490,396,417]" italics="true" pageId="15" pageNumber="31">Argutor</emphasis>
|
||
</taxonomicName>
|
||
lineage, but spermatheca is differentiated, which is characteristic of the
|
||
<taxonomicName id="60E84D56FFC416627BEBFE3C226573B4" ID-CoL="8KVYP" authorityName="Bonelli" authorityYear="1810" box="[653,774,422,443]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC416627BEBFE3C226573B4" box="[653,774,422,443]" italics="true" pageId="15" pageNumber="31">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
lineage D sensu Sasakawa and Kubota [2007].
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC416627984FE4125D273FE" blockId="15.[189,775,315,791]" lastBlockId="15.[812,1399,315,791]" pageId="15" pageNumber="31">
|
||
3) Dorsolateral sulci in
|
||
<emphasis id="959CEAC7FFC4166278BCFE46208E73FE" box="[474,493,476,497]" italics="true" pageId="15" pageNumber="31">ta</emphasis>
|
||
2 and
|
||
<emphasis id="959CEAC7FFC416627B57FE46232773FE" box="[561,580,476,497]" italics="true" pageId="15" pageNumber="31">ta</emphasis>
|
||
3. — The tarsi are generally bisulcate within the lineage, the anterior (outer) sulcus being deeper than the posterior (inner) one. This difference is by comparison slight in
|
||
<taxonomicName id="60E84D56FFC416627B57FDB6231E704E" authorityName="Dejean" authorityYear="1821" box="[561,637,556,577]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC416627B57FDB6231E704E" box="[561,637,556,577]" italics="true" pageId="15" pageNumber="31">Argutor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFC416627BD7FDB12263704F" authorityName="Gozis" authorityYear="1886" box="[689,768,555,576]" pageId="15" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFC416627BD7FDB12263704F" box="[689,768,555,576]" italics="true" pageId="15" pageNumber="31">Phonias</emphasis>
|
||
</taxonomicName>
|
||
, which have the sulci best developed. These are wide, deep and thence conspicuous on tarsomeres 1–4 (
|
||
<taxonomicName id="60E84D56FFC416627BE3FDFB23BC7079" ID-CoL="4Q3FW" baseAuthorityName="Bates" baseAuthorityYear="1883" box="[645,735,609,630]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="dulcis">
|
||
<emphasis id="959CEAC7FFC416627BE3FDFB23BC7079" box="[645,735,609,630]" italics="true" pageId="15" pageNumber="31">P. dulcis</emphasis>
|
||
</taxonomicName>
|
||
) or slightly finer in
|
||
<emphasis id="959CEAC7FFC416627831FDE62009709E" box="[343,362,636,657]" italics="true" pageId="15" pageNumber="31">ta</emphasis>
|
||
2 and still more so in
|
||
<emphasis id="959CEAC7FFC416627B5DFDE6232D709E" box="[571,590,636,657]" italics="true" pageId="15" pageNumber="31">ta</emphasis>
|
||
3 (
|
||
<taxonomicName id="60E84D56FFC416627B01FDE62386709F" ID-CoL="4Q3FZ" authorityName="A. Morawitz" authorityYear="1862" box="[615,741,635,657]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="sulcitarsis">
|
||
<emphasis id="959CEAC7FFC416627B01FDE62386709F" box="[615,741,635,657]" italics="true" pageId="15" pageNumber="31">P. sulcitarsis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFC416627B94FDE6206870A4" ID-CoL="QZHZ" authorityName="Panzer" authorityYear="1796" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="959CEAC7FFC416627B94FDE6206870A4" italics="true" pageId="15" pageNumber="31">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
), or absent from mesotarsomere 4 (
|
||
<taxonomicName id="60E84D56FFC416627B03FD0D204970C9" ID-CoL="4Q3FY" authority="Apfelbeck, 1904" authorityName="Apfelbeck" authorityYear="1904" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="leonisi">
|
||
<emphasis id="959CEAC7FFC416627B03FD0D23A370A4" box="[613,704,662,684]" italics="true" pageId="15" pageNumber="31">P. leonisi</emphasis>
|
||
Apfelbeck, 1904
|
||
</taxonomicName>
|
||
; some specimens of
|
||
<taxonomicName id="60E84D56FFC416627890FD2B230370C9" ID-CoL="QZHZ" authorityName="Panzer" authorityYear="1796" box="[502,608,689,710]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="959CEAC7FFC416627890FD2B230370C9" box="[502,608,689,710]" italics="true" pageId="15" pageNumber="31">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
), combined with the posterior sulcus hardly traceable on mesotarsomeres 1 and 2 only (
|
||
<taxonomicName id="60E84D56FFC416627849FD7D20D170F4" ID-CoL="4Q3FS" authorityName="" baseAuthorityName="Motschulsky" baseAuthorityYear="1866" box="[303,434,742,764]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="chameleon">
|
||
<emphasis id="959CEAC7FFC416627849FD7D20D170F4" box="[303,434,742,764]" italics="true" pageId="15" pageNumber="31">P. chameleon</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="60E84D56FFC4166278A3FD7C237770F4" authorityName="Gozis" authorityYear="1886" box="[453,532,742,763]" pageId="15" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFC4166278A3FD7C237770F4" box="[453,532,742,763]" italics="true" pageId="15" pageNumber="31">Phonias</emphasis>
|
||
</taxonomicName>
|
||
have similar sulci except that the apical two (most species) or 2–3 (
|
||
<taxonomicName id="60E84D56FFC416627B15FC9B239C7119" authorityName="Solsky" authorityYear="1874" box="[627,767,769,790]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="stricticollis">
|
||
<emphasis id="959CEAC7FFC416627B15FC9B239C7119" box="[627,767,769,790]" italics="true" pageId="15" pageNumber="31">P. stricticollis</emphasis>
|
||
</taxonomicName>
|
||
) mesotarsomeres are not sulcate, or the mesotarsomeres 1 and 2 and the metatarsomeres 1–3 are only anteriorly sulcate (
|
||
<taxonomicName id="60E84D56FFC416627C07FECD221F7389" authorityName="Csiki" authorityYear="1930" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="taxonyis">
|
||
<emphasis id="959CEAC7FFC416627C07FECD221F7389" italics="true" pageId="15" pageNumber="31">P. taxonyis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="60E84D56FFC416627AF7FEEB252E7388" box="[913,1101,369,391]" pageId="15" pageNumber="31">
|
||
<emphasis id="959CEAC7FFC416627AF7FEEB25637389" box="[913,1024,369,390]" italics="true" pageId="15" pageNumber="31">Badistrinus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are slightly distinctive in having basal three or four tarsomeres anteriorly sulcate, in couple with the posterior sulci being very fine and traceable on basal 2–3 mesotarsomeres and basal four (
|
||
<taxonomicName id="60E84D56FFC416627DC0FE5B242C73D9" ID-CoL="8LFDQ" authorityName="Tschitscherine" authorityYear="1889" box="[1190,1359,449,470]" class="Insecta" family="Carabidae" genus="Eurythorax" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="haptoderoides">
|
||
<emphasis id="959CEAC7FFC416627DC0FE5B242C73D9" box="[1190,1359,449,470]" italics="true" pageId="15" pageNumber="31">P. haptoderoides</emphasis>
|
||
</taxonomicName>
|
||
) or only 1–2 (
|
||
<taxonomicName id="60E84D56FFC416627AF4FE46256073FF" ID-CoL="6SK65" authorityName="Motschulsky" authorityYear="1844" box="[914,1027,475,497]" class="Insecta" family="Carabidae" genus="Omaseus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="laticollis">
|
||
<emphasis id="959CEAC7FFC416627AF4FE46256073FF" box="[914,1027,475,497]" italics="true" pageId="15" pageNumber="31">P. laticollis</emphasis>
|
||
</taxonomicName>
|
||
) metatarsomeres.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC416627A34FE6C241470C8" blockId="15.[812,1399,315,791]" pageId="15" pageNumber="31">
|
||
<taxonomicName id="60E84D56FFC416627A34FE6C22CA7004" authorityName="Motschulsky" authorityYear="1866" box="[850,937,502,523]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC416627A34FE6C22CA7004" box="[850,937,502,523]" italics="true" pageId="15" pageNumber="31">Pledarus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFC416627ABEFE6C25207004" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[984,1091,502,523]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC416627ABEFE6C25207004" box="[984,1091,502,523]" italics="true" pageId="15" pageNumber="31">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
have the most reduced dorsolateral sulci, especially the posterior ones. Mesotarsomeres 1– 2 and metatarsomeres 1–3 mostly have the anterior sulci while the posterior sulci are either missing (many
|
||
<taxonomicName id="60E84D56FFC416627C79FDDC222B7079" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC416627C79FDDC222B7079" italics="true" pageId="15" pageNumber="31">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
) or hardly traceable on basal one or two tarsomeres, mostly metatarsomeres, in
|
||
<taxonomicName id="60E84D56FFC416627D55FDE125E9709F" authorityName="Motschulsky" authorityYear="1866" box="[1075,1162,635,656]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC416627D55FDE125E9709F" box="[1075,1162,635,656]" italics="true" pageId="15" pageNumber="31">Pledarus</emphasis>
|
||
</taxonomicName>
|
||
as well as in
|
||
<taxonomicName id="60E84D56FFC416627C75FDE6222B70A3" authorityName="A. Morawitz" authorityYear="1862" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="neglectus">
|
||
<emphasis id="959CEAC7FFC416627C75FDE6222B70A3" italics="true" pageId="15" pageNumber="31">P. neglectus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFC416627A31FD0D22D070A4" authorityName="Jedlicka" authorityYear="1930" box="[855,947,662,684]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="goschi">
|
||
<emphasis id="959CEAC7FFC416627A31FD0D22D070A4" box="[855,947,662,684]" italics="true" pageId="15" pageNumber="31">P. goschi</emphasis>
|
||
</taxonomicName>
|
||
, and some
|
||
<taxonomicName id="60E84D56FFC416627D4FFD0C25F770A4" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[1065,1172,662,683]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC416627D4FFD0C25F770A4" box="[1065,1172,662,683]" italics="true" pageId="15" pageNumber="31">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
such as
|
||
<taxonomicName id="60E84D56FFC416627D88FD0D256670C8" authority="(Tschitscherine, 1897)" baseAuthorityName="Tschitscherine" baseAuthorityYear="1897" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="jankowskyi">
|
||
<emphasis id="959CEAC7FFC416627D88FD0D241570A4" box="[1262,1398,662,684]" italics="true" pageId="15" pageNumber="31">P. jankowskyi</emphasis>
|
||
(
|
||
<bibRefCitation id="C3794B24FFC416627A55FD2B229E70C8" author="Tschitscherine T." box="[819,1021,689,711]" pageId="15" pageNumber="31" pagination="260 - 351" refId="ref20656" refString="Tschitscherine T. 1897. Materiaux pour servir a l'etude des feroniens. III // Horae Soc. Ent. Ross. T. 30. P. 260 - 351." type="journal article" year="1897">Tschitschérine, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="60E84D56FFC416627D44FD2B241070C8" authority="(Lutshnik, 1922)" baseAuthorityName="Lutshnik" baseAuthorityYear="1922" box="[1058,1395,689,711]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="morawitzianus">
|
||
<emphasis id="959CEAC7FFC416627D44FD2B25A970C9" box="[1058,1226,689,710]" italics="true" pageId="15" pageNumber="31">P. morawitzianus</emphasis>
|
||
(Lutshnik, 1922)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC4167D7A34FD51209373FE" blockId="15.[812,1399,315,791]" lastBlockId="16.[189,776,315,2071]" lastPageId="16" lastPageNumber="32" pageId="15" pageNumber="31">
|
||
4) Ventral setae of tarsomere 5. — The tarsomere is ventrally setose in some species of
|
||
<taxonomicName id="60E84D56FFC416627DE5FD7C258D70F4" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[1155,1262,742,763]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC416627DE5FD7C258D70F4" box="[1155,1262,742,763]" italics="true" pageId="15" pageNumber="31">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
or setulose in some others such as
|
||
<taxonomicName id="60E84D56FFC416627A8AFC9B25277119" authorityName="Tschitscherine" authorityYear="1899" box="[1004,1092,769,790]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="eobius">
|
||
<emphasis id="959CEAC7FFC416627A8AFC9B25277119" box="[1004,1092,769,790]" italics="true" pageId="15" pageNumber="31">P. eobius</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="60E84D56FFC4167D7D07FC9B207B735E" authority="A. Morawitz, 1862" authorityName="A. Morawitz" authorityYear="1862" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" lastPageId="16" lastPageNumber="32" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="morawitzianus">
|
||
<emphasis id="959CEAC7FFC416627D07FC9B246B7119" box="[1121,1288,769,790]" italics="true" pageId="15" pageNumber="31">P. morawitzianus</emphasis>
|
||
A. Morawitz, 1862
|
||
</taxonomicName>
|
||
, while it varies from setose to glabrous between species of the other three subgenera. The ventral setae are reduced considerably in size yet still distinct in
|
||
<taxonomicName id="60E84D56FFDB167D7BE5FEEB22637389" ID-CoL="4Q3FZ" authorityName="A. Morawitz" authorityYear="1862" box="[643,768,369,390]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="sulcitarsis">
|
||
<emphasis id="959CEAC7FFDB167D7BE5FEEB22637389" box="[643,768,369,390]" italics="true" pageId="16" pageNumber="32">P. sulcitarsis</emphasis>
|
||
</taxonomicName>
|
||
, but almost imperceptible in
|
||
<taxonomicName id="60E84D56FFDB167D78BCFE16238773AE" authority="(Lafer, 1983)" baseAuthorityName="Lafer" baseAuthorityYear="1983" box="[474,740,395,417]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="kerzhneri">
|
||
<emphasis id="959CEAC7FFDB167D78BCFE16233073AF" box="[474,595,395,417]" italics="true" pageId="16" pageNumber="32">P. kerzhneri</emphasis>
|
||
(Lafer, 1983)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D7B94FE16237173B3" ID-CoL="4Q4B4" authority="Jedlicka, 1962" authorityName="Jedlicka" authorityYear="1962" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="lutschniki" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFDB167D7B94FE16226473AE" box="[754,775,396,417]" italics="true" pageId="16" pageNumber="32">P.</emphasis>
|
||
(
|
||
<emphasis id="959CEAC7FFDB167D79A3FE3C207773B4" box="[197,276,422,443]" italics="true" pageId="16" pageNumber="32">Phonias</emphasis>
|
||
)
|
||
<emphasis id="959CEAC7FFDB167D7844FE3C20E373B4" box="[290,384,422,443]" italics="true" pageId="16" pageNumber="32">lutschniki</emphasis>
|
||
<bibRefCitation id="C3794B24FFDB167D78E0FE3C237173B3" author="Jedlicka A." box="[390,530,422,444]" pageId="16" pageNumber="32" pagination="177 - 346" refId="ref19769" refString="Jedlicka A. 1962. Monographie des Tribus Pterostichini aus Ostasien (Pterostichi, Trigonotomi, Myadi) (Coleoptera, Carabidae) // Entom. Abhandl. Mus. Tierk. Dresden. Bd. 26. Nr. 21. S. 177 - 346." type="journal article" year="1962">Jedlička, 1962</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="959CEAC7FFDB167D7B24FE3D233B73B3" box="[578,600,423,444]" italics="true" pageId="16" pageNumber="32">P.</emphasis>
|
||
<taxonomicName id="60E84D56FFDB167D7B38FE3C226373B4" ID-CoL="4Q49V" authorityName="Bates" authorityYear="1873" box="[606,768,422,443]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="longinquus" subGenus="Phonias">
|
||
(
|
||
<emphasis id="959CEAC7FFDB167D7B00FE3C23E273B4" box="[614,641,422,443]" italics="true" pageId="16" pageNumber="32">Ph</emphasis>
|
||
.)
|
||
<emphasis id="959CEAC7FFDB167D7BF3FE3C226373B4" box="[661,768,422,443]" italics="true" pageId="16" pageNumber="32">longinquus</emphasis>
|
||
</taxonomicName>
|
||
, from which it follows that the erection of
|
||
<taxonomicName id="60E84D56FFDB167D7B29FE5B23D273D9" ID-CoL="8LR59" authorityName="Jeanne" authorityYear="1988" box="[591,689,449,470]" class="Insecta" family="Carabidae" genus="Biphonias" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7B29FE5B23D273D9" box="[591,689,449,470]" italics="true" pageId="16" pageNumber="32">Biphonias</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<bibRefCitation id="C3794B24FFDB167D7BDAFE5B218C73FE" author="Jeanne C." pageId="16" pageNumber="32" pagination="69 - 87" refId="ref19639" refString="Jeanne C. 1988. Carabiques nouveaux ou remarquables (9 e note) // Bull. Soc. Linn. Bordeaux. Vol. 16. P. 69 - 87." type="journal article" year="1988">Jeanne, 1988</bibRefCitation>
|
||
] has been mere formality.
|
||
</paragraph>
|
||
<caption id="F397665DFFC416627984F8CB22597A1E" ID-DOI="http://doi.org/10.5281/zenodo.10949114" ID-Zenodo-Dep="10949114" httpUri="https://zenodo.org/record/10949114/files/figure.png" pageId="15" pageNumber="31" startId="15.[226,263,1873,1892]" targetBox="[193,1392,817,1839]" targetPageId="15" targetType="figure">
|
||
<paragraph id="A75736D5FFC416627984F8CB238675A2" blockId="15.[188,1399,1873,2065]" pageId="15" pageNumber="31">
|
||
Figs 63–68. Reproductive tract in female: 63 —
|
||
<taxonomicName id="60E84D56FFC416627BE8F8CB2233756B" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[654,848,1873,1892]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="austrinus" status="sp. nov.">
|
||
<emphasis id="959CEAC7FFC416627BE8F8CB2233756B" box="[654,848,1873,1892]" italics="true" pageId="15" pageNumber="31">Pterostichus austrinus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFC416627A31F8CB22E1756B" bold="true" box="[855,898,1873,1892]" pageId="15" pageNumber="31">
|
||
<taxonomicNameLabel id="8EAF57BCFFC416627A31F8CB22E1756B" box="[855,898,1873,1892]" pageId="15" pageNumber="31" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 64–65 —
|
||
<taxonomicName id="60E84D56FFC416627A81F8C8252C756B" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[999,1103,1873,1892]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="glabellus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFC416627A81F8C8252C756B" box="[999,1103,1873,1892]" italics="true" pageId="15" pageNumber="31">P. glabellus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFC416627D30F8CB25E2756B" bold="true" box="[1110,1153,1873,1892]" pageId="15" pageNumber="31">
|
||
<taxonomicNameLabel id="8EAF57BCFFC416627D30F8CB25E2756B" box="[1110,1153,1873,1892]" pageId="15" pageNumber="31" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 66 —
|
||
<taxonomicName id="60E84D56FFC416627DA0F8C821927573" authority="(Panzer, 1796)" baseAuthorityName="Panzer" baseAuthorityYear="1796" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="959CEAC7FFC416627DA0F8C8244B756B" box="[1222,1320,1874,1892]" italics="true" pageId="15" pageNumber="31">P. strenuus</emphasis>
|
||
(Panzer, 1796)
|
||
</taxonomicName>
|
||
; 67 —
|
||
<taxonomicName id="60E84D56FFC416627855F8F020CA7573" authorityName="" baseAuthorityName="Motschulsky" baseAuthorityYear="1866" box="[307,425,1897,1916]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="chameleon">
|
||
<emphasis id="959CEAC7FFC416627855F8F020CA7573" box="[307,425,1897,1916]" italics="true" pageId="15" pageNumber="31">P. chameleon</emphasis>
|
||
</taxonomicName>
|
||
; 68 —
|
||
<taxonomicName id="60E84D56FFC41662788BF8F023037573" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[493,608,1898,1916]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="unisetosus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFC41662788BF8F023037573" box="[493,608,1898,1916]" italics="true" pageId="15" pageNumber="31">P. unisetosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFC416627B01F8F323F17573" bold="true" box="[615,658,1897,1916]" pageId="15" pageNumber="31">
|
||
<taxonomicNameLabel id="8EAF57BCFFC416627B01F8F323F17573" box="[615,658,1897,1916]" pageId="15" pageNumber="31" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 63, 65–67 — ventral aspect; 64, 68 — dorsal aspect;
|
||
<emphasis id="959CEAC7FFC416627D12F8F324597573" box="[1140,1338,1897,1917]" italics="true" pageId="15" pageNumber="31">
|
||
<emphasis id="959CEAC7FFC416627D12F8F325EB7573" bold="true" box="[1140,1160,1897,1916]" italics="true" pageId="15" pageNumber="31">bc</emphasis>
|
||
— bursa copulatrix
|
||
</emphasis>
|
||
;
|
||
<emphasis id="959CEAC7FFC416627C21F8F324397573" bold="true" box="[1351,1370,1897,1916]" italics="true" pageId="15" pageNumber="31">bs</emphasis>
|
||
— bulbous structure;
|
||
<emphasis id="959CEAC7FFC416627804F81B2015759B" bold="true" box="[354,374,1921,1940]" italics="true" pageId="15" pageNumber="31">dv</emphasis>
|
||
— diverticulum;
|
||
<emphasis id="959CEAC7FFC416627B71F8182348759A" bold="true" box="[535,555,1922,1941]" italics="true" pageId="15" pageNumber="31">ov</emphasis>
|
||
— common oviduct;
|
||
<emphasis id="959CEAC7FFC416627B97F8182267759A" bold="true" box="[753,772,1922,1941]" italics="true" pageId="15" pageNumber="31">rp</emphasis>
|
||
— receptacle;
|
||
<emphasis id="959CEAC7FFC416627AE9F81822C2759A" bold="true" box="[911,929,1922,1941]" italics="true" pageId="15" pageNumber="31">sc</emphasis>
|
||
— seminal canal;
|
||
<emphasis id="959CEAC7FFC416627D2AF818253C759A" bold="true" box="[1100,1119,1922,1941]" italics="true" pageId="15" pageNumber="31">sg</emphasis>
|
||
— spermathecal gland;
|
||
<emphasis id="959CEAC7FFC416627C5DF81B243A759B" bold="true" box="[1339,1369,1921,1940]" italics="true" pageId="15" pageNumber="31">sgd</emphasis>
|
||
— spermathecal gland duct;
|
||
<emphasis id="959CEAC7FFC4166278F8F80020D275A2" bold="true" box="[414,433,1946,1965]" italics="true" pageId="15" pageNumber="31">sp</emphasis>
|
||
— spermatheca. Scale bars 1 mm.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFC416627984F82C22597A1E" blockId="15.[188,1399,1873,2065]" pageId="15" pageNumber="31">
|
||
Рис. 63–68. Репродуктивный тракт самки: 63 —
|
||
<taxonomicName id="60E84D56FFC416627BFBF82F223C75C7" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[669,863,1973,1992]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="austrinus" status="sp. nov.">
|
||
<emphasis id="959CEAC7FFC416627BFBF82F223C75C7" box="[669,863,1973,1992]" italics="true" pageId="15" pageNumber="31">Pterostichus austrinus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFC416627A02F82F22EC75C7" bold="true" box="[868,911,1973,1992]" pageId="15" pageNumber="31">
|
||
<taxonomicNameLabel id="8EAF57BCFFC416627A02F82F22EC75C7" box="[868,911,1973,1992]" pageId="15" pageNumber="31" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 64–65 —
|
||
<taxonomicName id="60E84D56FFC416627A96F82C253475C7" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[1008,1111,1973,1992]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="glabellus" status="sp.n.">
|
||
<emphasis id="959CEAC7FFC416627A96F82C253475C7" box="[1008,1111,1973,1992]" italics="true" pageId="15" pageNumber="31">P. glabellus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFC416627D3BF82F25EB75C7" bold="true" box="[1117,1160,1973,1992]" pageId="15" pageNumber="31">
|
||
<taxonomicNameLabel id="8EAF57BCFFC416627D3BF82F25EB75C7" box="[1117,1160,1973,1992]" pageId="15" pageNumber="31" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 66 —
|
||
<taxonomicName id="60E84D56FFC416627DAFF82C219275EF" authority="(Panzer, 1796)" baseAuthorityName="Panzer" baseAuthorityYear="1796" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="959CEAC7FFC416627DAFF82C244A75C7" box="[1225,1321,1974,1992]" italics="true" pageId="15" pageNumber="31">P. strenuus</emphasis>
|
||
(Panzer, 1796)
|
||
</taxonomicName>
|
||
; 67 —
|
||
<taxonomicName id="60E84D56FFC416627855F85420CA75EF" authorityName="" baseAuthorityName="Motschulsky" baseAuthorityYear="1866" box="[307,425,1997,2016]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="chameleon">
|
||
<emphasis id="959CEAC7FFC416627855F85420CA75EF" box="[307,425,1997,2016]" italics="true" pageId="15" pageNumber="31">P. chameleon</emphasis>
|
||
</taxonomicName>
|
||
; 68 —
|
||
<taxonomicName id="60E84D56FFC41662788BF854230375EF" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[493,608,1998,2016]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="31" phylum="Arthropoda" rank="species" species="unisetosus" status="sp.n">
|
||
<emphasis id="959CEAC7FFC41662788BF854230375EF" box="[493,608,1998,2016]" italics="true" pageId="15" pageNumber="31">P. unisetosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFC416627B01F85723EF75EF" bold="true" box="[615,652,1997,2016]" pageId="15" pageNumber="31">
|
||
<taxonomicNameLabel id="8EAF57BCFFC416627B01F85723EF75EF" box="[615,652,1997,2016]" pageId="15" pageNumber="31" rank="species">sp.n</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 63, 65–67 — вентрально; 64, 68 — дорЗально;
|
||
<emphasis id="959CEAC7FFC416627D2FF857253E75EF" bold="true" box="[1097,1117,1997,2016]" italics="true" pageId="15" pageNumber="31">bc</emphasis>
|
||
— копулЯтивнаЯ сумка;
|
||
<emphasis id="959CEAC7FFC416627C21F857243975EF" bold="true" box="[1351,1370,1997,2016]" italics="true" pageId="15" pageNumber="31">bs</emphasis>
|
||
— бульбарное обраЗование;
|
||
<emphasis id="959CEAC7FFC4166278C3F87F20DA75F7" bold="true" box="[421,441,2021,2040]" italics="true" pageId="15" pageNumber="31">dv</emphasis>
|
||
— дивертикул;
|
||
<emphasis id="959CEAC7FFC416627B2DF87F233C75F7" bold="true" box="[587,607,2021,2040]" italics="true" pageId="15" pageNumber="31">ov</emphasis>
|
||
— непарный Яйцевод;
|
||
<emphasis id="959CEAC7FFC416627A56F87F222075F7" bold="true" box="[816,835,2021,2040]" italics="true" pageId="15" pageNumber="31">rp</emphasis>
|
||
— семеприемник;
|
||
<emphasis id="959CEAC7FFC416627A89F87F256275F7" bold="true" box="[1007,1025,2021,2040]" italics="true" pageId="15" pageNumber="31">sc</emphasis>
|
||
— семепровод;
|
||
<emphasis id="959CEAC7FFC416627DF5F87F25C575F7" bold="true" box="[1171,1190,2021,2040]" italics="true" pageId="15" pageNumber="31">sg</emphasis>
|
||
— ЖелеЗа сперматеки;
|
||
<emphasis id="959CEAC7FFC4166279DAF86721B97A1F" bold="true" box="[188,218,2045,2064]" italics="true" pageId="15" pageNumber="31">sgd</emphasis>
|
||
— проток ЖелеЗы сперматеки;
|
||
<emphasis id="959CEAC7FFC416627B62F86723747A1F" bold="true" box="[516,535,2045,2064]" italics="true" pageId="15" pageNumber="31">sp</emphasis>
|
||
— сперматека. Масштаб 1 мм.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A75736D5FFDB167D7984FE6C238F70C9" blockId="16.[189,776,315,2071]" pageId="16" pageNumber="32">
|
||
5) Pronotal basolateral sulci. — Two, well-developed, sulci on each side are certain to be plesiomorphous character state. It is observed in
|
||
<taxonomicName id="60E84D56FFDB167D78F3FDB12361704F" authorityName="Motschulsky" authorityYear="1866" box="[405,514,555,576]" class="Insecta" family="Carabidae" genus="Badistrinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D78F3FDB12361704F" box="[405,514,555,576]" italics="true" pageId="16" pageNumber="32">Badistrinus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D7B68FDB12300704F" authorityName="Motschulsky" authorityYear="1866" box="[526,611,555,576]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7B68FDB12300704F" box="[526,611,555,576]" italics="true" pageId="16" pageNumber="32">Pledarus</emphasis>
|
||
</taxonomicName>
|
||
and many representatives of
|
||
<taxonomicName id="60E84D56FFDB167D7858FDDC20E87054" authorityName="Gozis" authorityYear="1886" box="[318,395,582,603]" pageId="16" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFDB167D7858FDDC20E87054" box="[318,395,582,603]" italics="true" pageId="16" pageNumber="32">Phonias</emphasis>
|
||
</taxonomicName>
|
||
, whereas the outer, shorter, sulcus has been reduced to a punctiform vestige or totally in
|
||
<taxonomicName id="60E84D56FFDB167D7BD0FDFB22627079" authorityName="Dejean" authorityYear="1821" box="[694,769,609,630]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7BD0FDFB22627079" box="[694,769,609,630]" italics="true" pageId="16" pageNumber="32">Argutor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D79DBFDE12045709F" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[189,294,635,656]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D79DBFDE12045709F" box="[189,294,635,656]" italics="true" pageId="16" pageNumber="32">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
and some species of
|
||
<taxonomicName id="60E84D56FFDB167D7892FDE12322709F" authorityName="Gozis" authorityYear="1886" box="[500,577,635,656]" pageId="16" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFDB167D7892FDE12322709F" box="[500,577,635,656]" italics="true" pageId="16" pageNumber="32">Phonias</emphasis>
|
||
</taxonomicName>
|
||
such as
|
||
<taxonomicName id="60E84D56FFDB167D7BF0FDE62263709E" authorityName="Csiki" authorityYear="1930" box="[662,768,636,657]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="taxonyis">
|
||
<emphasis id="959CEAC7FFDB167D7BF0FDE62263709E" box="[662,768,636,657]" italics="true" pageId="16" pageNumber="32">P. taxonyis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="60E84D56FFDB167D79DBFD0D20D170A3" authority="Jedlicka, 1936" authorityName="Jedlicka" authorityYear="1936" box="[189,434,662,684]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="arrowi">
|
||
<emphasis id="959CEAC7FFDB167D79DBFD0D207970A3" box="[189,282,663,684]" italics="true" pageId="16" pageNumber="32">P. arrowi</emphasis>
|
||
<bibRefCitation id="C3794B24FFDB167D7842FD0C20D170A3" author="Jedlicka A." box="[292,434,662,684]" pageId="16" pageNumber="32" pagination="133" refId="ref19718" refString="Jedlicka A. 1936. Novy cinsky Pterostichus (Car.) // Cas. Cesk. Spol. Ent. Vol. 33. P. 133." type="journal article" year="1936">Jedlička, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="60E84D56FFDB167D78D9FD0D232670A4" ID-CoL="4Q49V" authorityName="Bates" authorityYear="1873" box="[447,581,662,684]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="longinquus">
|
||
<emphasis id="959CEAC7FFDB167D78D9FD0D232670A4" box="[447,581,662,684]" italics="true" pageId="16" pageNumber="32">P. longinquus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="60E84D56FFDB167D7B33FD0D202F70C9" authority="(Motschulsky, 1866)" baseAuthorityName="Motschulsky" baseAuthorityYear="1866" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="ripensis">
|
||
<emphasis id="959CEAC7FFDB167D7B33FD0D23DE70A3" box="[597,701,663,684]" italics="true" pageId="16" pageNumber="32">P. ripensis</emphasis>
|
||
(
|
||
<bibRefCitation id="C3794B24FFDB167D7BABFD0C202570C9" author="Motschulsky V." pageId="16" pageNumber="32" pagination="227 - 313" refId="ref20070" refString="Motschulsky V. 1866. Enumeration des nouvelles especes de coleopteres rapportes de ses voyages. 4 - eme article (suite) // Bull. Soc. Nat. Mosc. T. 38 [1865]. P. 227 - 313." type="book chapter" year="1866">Motschulsky, 1866</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="60E84D56FFDB167D78E6FD2B236070C9" ID-CoL="6WLVF" baseAuthorityName="Tschitscherine" baseAuthorityYear="1897" box="[384,515,689,710]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="ussuriensis">
|
||
<emphasis id="959CEAC7FFDB167D78E6FD2B236070C9" box="[384,515,689,710]" italics="true" pageId="16" pageNumber="32">P. ussuriensis</emphasis>
|
||
</taxonomicName>
|
||
, as well as in
|
||
<taxonomicName id="60E84D56FFDB167D7BEBFD2B238570C9" authorityName="Jedlicka" authorityYear="1930" box="[653,742,689,710]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="goschi">
|
||
<emphasis id="959CEAC7FFDB167D7BEBFD2B238570C9" box="[653,742,689,710]" italics="true" pageId="16" pageNumber="32">P. goschi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDB167D7984FD5123337168" blockId="16.[189,776,315,2071]" pageId="16" pageNumber="32">
|
||
6) Elytral parascutellar striole and seta. — The striole is entire or long, posteriorly adjoining stria 1 or almost so in
|
||
<taxonomicName id="60E84D56FFDB167D79DBFC9B206F7119" authorityName="Gozis" authorityYear="1886" box="[189,268,769,790]" pageId="16" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFDB167D79DBFC9B206F7119" box="[189,268,769,790]" italics="true" pageId="16" pageNumber="32">Phonias</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFDB167D785AFC9B20C47119" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[316,423,769,790]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D785AFC9B20C47119" box="[316,423,769,790]" italics="true" pageId="16" pageNumber="32">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
while ranging from long to missing within
|
||
<taxonomicName id="60E84D56FFDB167D7867FC812013713F" authorityName="Motschulsky" authorityYear="1866" box="[257,368,795,816]" class="Insecta" family="Carabidae" genus="Badistrinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7867FC812013713F" box="[257,368,795,816]" italics="true" pageId="16" pageNumber="32">Badistrinus</emphasis>
|
||
</taxonomicName>
|
||
and from short to missing within
|
||
<taxonomicName id="60E84D56FFDB167D7BDBFC8121B17143" authorityName="Motschulsky" authorityYear="1866" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7BDBFC8121B17143" italics="true" pageId="16" pageNumber="32">Pledarus</emphasis>
|
||
</taxonomicName>
|
||
. The seta is absent from the elytra of
|
||
<taxonomicName id="60E84D56FFDB167D7B35FCAC23DD7144" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[595,702,822,843]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7B35FCAC23DD7144" box="[595,702,822,843]" italics="true" pageId="16" pageNumber="32">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFDB167D7B94FCAD23757169" authority="Sundukov, 2013" authorityName="Sundukov" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="larisae" subGenus="Pledarus">
|
||
<emphasis id="959CEAC7FFDB167D7B94FCAD22647143" box="[754,775,823,844]" italics="true" pageId="16" pageNumber="32">P.</emphasis>
|
||
(
|
||
<emphasis id="959CEAC7FFDB167D79A3FCCB207F7169" box="[197,284,849,870]" italics="true" pageId="16" pageNumber="32">Pledarus</emphasis>
|
||
)
|
||
<emphasis id="959CEAC7FFDB167D784AFCCB200C7169" box="[300,367,849,870]" italics="true" pageId="16" pageNumber="32">larisae</emphasis>
|
||
<bibRefCitation id="C3794B24FFDB167D7810FCCB23757169" author="Sundukov Yu. N." box="[374,534,849,871]" pageId="16" pageNumber="32" refId="ref20559" refString="Sundukov Yu. N. 2013. [An annotated catalogue of the ground beetles (Coleoptera: Caraboidea) of Sikhote-Alin]. Vladivostok: Dalnauka. 271 p. [In Russian]" type="book" year="2013">Sundukov, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
only.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDB167D7984FCF623D676D4" blockId="16.[189,776,315,2071]" pageId="16" pageNumber="32">
|
||
7) Elytral discal setae. — Primary, ground plan, pattern is peculiar to
|
||
<taxonomicName id="60E84D56FFDB167D7850FC1C20E17194" authorityName="Dejean" authorityYear="1821" box="[310,386,902,923]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7850FC1C20E17194" box="[310,386,902,923]" italics="true" pageId="16" pageNumber="32">Argutor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D78F2FC1C209C7194" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[404,511,902,923]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D78F2FC1C209C7194" box="[404,511,902,923]" italics="true" pageId="16" pageNumber="32">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
and some
|
||
<taxonomicName id="60E84D56FFDB167D7B1CFC1C23AA7194" authorityName="Gozis" authorityYear="1886" box="[634,713,902,923]" pageId="16" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFDB167D7B1CFC1C23AA7194" box="[634,713,902,923]" italics="true" pageId="16" pageNumber="32">Phonias</emphasis>
|
||
</taxonomicName>
|
||
. It is defined by the seta d1 adjoining stria 1, combined with the setae d2 and d3 approximate to or adjoining stria 2. Some of these setae vary individually in number to be not seldom either doubled on or absent from one or both sides. The remainder of
|
||
<taxonomicName id="60E84D56FFDB167D7859FB9120ED762F" authorityName="Gozis" authorityYear="1886" box="[319,398,1035,1056]" pageId="16" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFDB167D7859FB9120ED762F" box="[319,398,1035,1056]" italics="true" pageId="16" pageNumber="32">Phonias</emphasis>
|
||
</taxonomicName>
|
||
have only retained either the posterior two setae or but one. The setae d2 and d3 also range between stria 2 and middle of interval 3 individually, albeit occasionally, in a species, and variations of such a kind are certain to have given rise to the setation characteristic of
|
||
<taxonomicName id="60E84D56FFDB167D7B20FBEC23D07684" authorityName="Motschulsky" authorityYear="1866" box="[582,691,1142,1163]" class="Insecta" family="Carabidae" genus="Badistrinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7B20FBEC23D07684" box="[582,691,1142,1163]" italics="true" pageId="16" pageNumber="32">Badistrinus</emphasis>
|
||
</taxonomicName>
|
||
first and a reduced setation of
|
||
<taxonomicName id="60E84D56FFDB167D78EFFB0B20BD76A9" authorityName="Motschulsky" authorityYear="1866" box="[393,478,1169,1190]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D78EFFB0B20BD76A9" box="[393,478,1169,1190]" italics="true" pageId="16" pageNumber="32">Pledarus</emphasis>
|
||
</taxonomicName>
|
||
after. The setae adjoining stria 3 are characteristic of these two subgenera, which also is true of
|
||
<taxonomicName id="60E84D56FFDB167D79DBFB5D206D76D3" ID-CoL="78LVB" authorityName="Novoa" authorityYear="1979" box="[189,270,1223,1244]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="perisi">
|
||
<emphasis id="959CEAC7FFDB167D79DBFB5D206D76D3" box="[189,270,1223,1244]" italics="true" pageId="16" pageNumber="32">P. perisi</emphasis>
|
||
</taxonomicName>
|
||
placed by
|
||
<bibRefCitation id="C3794B24FFDB167D781EFB5C237B76D4" author="Bousquet Y." box="[376,536,1221,1243]" pageId="16" pageNumber="32" pagination="1 - 292" refId="ref19199" refString="Bousquet Y. 1999. Supraspecific classification of the Nearctic Pterostichini (Coleoptera: Carabidae) // Fabreries. Suppl. 9. Association des Entomologists Amateurs du Quebec, Quebec, Canada. P. 1 - 292." type="book chapter" year="1999">Bousquet [1999]</bibRefCitation>
|
||
within
|
||
<taxonomicName id="60E84D56FFDB167D7B04FB5C23CC76D4" authorityName="Gozis" authorityYear="1886" box="[610,687,1222,1243]" pageId="16" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFDB167D7B04FB5C23CC76D4" box="[610,687,1222,1243]" italics="true" pageId="16" pageNumber="32">Phonias</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDB167D7984FB7B209C76F9" blockId="16.[189,776,315,2071]" box="[226,511,1249,1271]" pageId="16" pageNumber="32">8) Female reproductive tract.</paragraph>
|
||
<paragraph id="A75736D5FFDB167D7984FB612354740E" blockId="16.[189,776,315,2071]" pageId="16" pageNumber="32">
|
||
(1)
|
||
<emphasis id="959CEAC7FFDB167D7860FB6620C9771F" box="[262,426,1275,1297]" italics="true" pageId="16" pageNumber="32">Bursa copulatrix</emphasis>
|
||
(
|
||
<emphasis id="959CEAC7FFDB167D78DFFB6120B3771F" box="[441,464,1275,1296]" italics="true" pageId="16" pageNumber="32">bc</emphasis>
|
||
). — When folded it is obtrapezoidal, broader before than behind, with apex invaginate and slightly sclerotized, which is characteristic of not only
|
||
<taxonomicName id="60E84D56FFDB167D7BB3FAAB206F776F" ID-CoL="8KVYP" authorityName="Bonelli" authorityYear="1810" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7BB3FAAB206F776F" italics="true" pageId="16" pageNumber="32">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
but also many other carabids. This primary
|
||
<emphasis id="959CEAC7FFDB167D7BB6FAD120437774" italics="true" pageId="16" pageNumber="32">bursa copulatrix</emphasis>
|
||
(
|
||
<emphasis id="959CEAC7FFDB167D7856FAFC20247774" box="[304,327,1382,1403]" italics="true" pageId="16" pageNumber="32">bc</emphasis>
|
||
1) tends to be increasingly elongated in some representatives. It has become telescoped, with intercalary section (
|
||
<emphasis id="959CEAC7FFDB167D7868FA01204677BF" box="[270,293,1435,1456]" italics="true" pageId="16" pageNumber="32">bc</emphasis>
|
||
2) having appeared to mediate between
|
||
<emphasis id="959CEAC7FFDB167D7BC6FA0123D477BF" box="[672,695,1435,1456]" italics="true" pageId="16" pageNumber="32">bc</emphasis>
|
||
1 and an elongated apical part (
|
||
<taxonomicName id="60E84D56FFDB167D78F5FA2D209877C4" ID-CoL="QZHZ" authorityName="Panzer" authorityYear="1796" box="[403,507,1462,1484]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="959CEAC7FFDB167D78F5FA2D209877C4" box="[403,507,1462,1484]" italics="true" pageId="16" pageNumber="32">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
), or has evolved into a very long tube which correlates to the internal sac of aedeagus in width (
|
||
<taxonomicName id="60E84D56FFDB167D7865FA762016740F" ID-CoL="6SK65" authorityName="Motschulsky" authorityYear="1844" box="[259,373,1515,1537]" class="Insecta" family="Carabidae" genus="Omaseus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="laticollis">
|
||
<emphasis id="959CEAC7FFDB167D7865FA762016740F" box="[259,373,1515,1537]" italics="true" pageId="16" pageNumber="32">P. laticollis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D78E4FA76234A740F" ID-CoL="8LFDQ" authorityName="Tschitscherine" authorityYear="1889" box="[386,553,1515,1537]" class="Insecta" family="Carabidae" genus="Eurythorax" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="haptoderoides">
|
||
<emphasis id="959CEAC7FFDB167D78E4FA76234A740F" box="[386,553,1515,1537]" italics="true" pageId="16" pageNumber="32">P. haptoderoides</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDB167D7984F99C20B17529" blockId="16.[189,776,315,2071]" pageId="16" pageNumber="32">
|
||
Basal sclerite of the seminal canal or bulbous structures as the bodies of probable sphincters at the oviduct-bursal junction have been found in no species of the lineage but
|
||
<taxonomicName id="60E84D56FFDB167D7B94F9A620467464" ID-CoL="4Q3FS" authorityName="" baseAuthorityName="Motschulsky" baseAuthorityYear="1866" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="chameleon">
|
||
<emphasis id="959CEAC7FFDB167D7B94F9A620467464" italics="true" pageId="16" pageNumber="32">P. chameleon</emphasis>
|
||
</taxonomicName>
|
||
. It has one bulb round the basal diverticulum and the other, larger, one at this junction (
|
||
<figureCitation id="3FD32A50FFDB167D7B45F9EB23097489" box="[547,618,1648,1671]" captionStart="Figs 63–68" captionStartId="15.[226,263,1873,1892]" captionTargetBox="[193,1392,817,1839]" captionTargetId="figure-183@15.[188,1398,813,1843]" captionTargetPageId="15" captionText="Figs 63–68. Reproductive tract in female: 63 — Pterostichus austrinus sp.n.; 64–65 — P. glabellus sp.n.; 66 — P. strenuus (Panzer, 1796); 67 — P. chameleon; 68 — P. unisetosus sp.n.; 63, 65–67 — ventral aspect; 64, 68 — dorsal aspect; bc — bursa copulatrix; bs — bulbous structure; dv — diverticulum; ov — common oviduct; rp — receptacle; sc — seminal canal; sg — spermathecal gland; sgd — spermathecal gland duct; sp — spermatheca. Scale bars 1 mm. Рис. 63–68. Репродуктивный тракт самки: 63 — Pterostichus austrinus sp.n.; 64–65 — P. glabellus sp.n.; 66 — P. strenuus (Panzer, 1796); 67 — P. chameleon; 68 — P. unisetosus sp.n; 63, 65–67 — вентрально; 64, 68 — дорЗально; bc — копулЯтивнаЯ сумка; bs — бульбарное обраЗование; dv — дивертикул; ov — непарный Яйцевод; rp — семеприемник; sc — семепровод; sg — ЖелеЗа сперматеки; sgd — проток ЖелеЗы сперматеки; sp — сперматека. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949114" httpUri="https://zenodo.org/record/10949114/files/figure.png" pageId="16" pageNumber="32">Fig. 67</figureCitation>
|
||
). These bulbous structures are characteristic of many
|
||
<taxonomicName id="60E84D56FFDB167D7B4EF91123C274AF" ID-CoL="8KVYP" authorityName="Bonelli" authorityYear="1810" box="[552,673,1675,1696]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7B4EF91123C274AF" box="[552,673,1675,1696]" italics="true" pageId="16" pageNumber="32">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
lineage D sensu Sasakawa and Kubota [2007] except
|
||
<taxonomicName id="60E84D56FFDB167D7B3CF93C219374D9" ID-CoL="6S4P" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" class="Insecta" family="Curculionidae" genus="Platysma" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7B3CF93C23D774B4" box="[602,692,1702,1723]" italics="true" pageId="16" pageNumber="32">Platysma</emphasis>
|
||
Bonelli, 1810
|
||
</taxonomicName>
|
||
, with its allies or derivatives such as,
|
||
<emphasis id="959CEAC7FFDB167D7B18F95B23C174D9" box="[638,674,1729,1750]" italics="true" pageId="16" pageNumber="32">e.g.</emphasis>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D7BD4F95B203074FE" authority="Stephens, 1835" authorityName="Stephens" authorityYear="1835" class="Insecta" family="Carabidae" genus="Adelosia" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7BD4F95B226574D9" box="[690,774,1729,1750]" italics="true" pageId="16" pageNumber="32">Adelosia</emphasis>
|
||
Stephens, 1835
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="60E84D56FFDB167D783BF94123F874FF" authority="Fischer-Waldheim, 1822" authorityName="Fischer-Waldheim" authorityYear="1822" box="[349,667,1755,1777]" class="Insecta" family="Carabidae" genus="Plectes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D783BF94120C074FF" box="[349,419,1755,1776]" italics="true" pageId="16" pageNumber="32">Plectes</emphasis>
|
||
Fischer-Waldheim, 1822
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="60E84D56FFDB167D7BC3F94120CD7504" authority="Fischer-Waldheim, 1823" authorityName="Fischer-Waldheim" authorityYear="1823" class="Insecta" family="Carabidae" genus="Myosodus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7BC3F941226574FF" box="[677,774,1755,1776]" italics="true" pageId="16" pageNumber="32">Myosodus</emphasis>
|
||
Fischer-Waldheim, 1823
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="60E84D56FFDB167D78D1F96C23B97504" authority="Chaudoir, 1838" authorityName="Chaudoir" authorityYear="1838" box="[439,730,1782,1804]" class="Insecta" family="Carabidae" genus="Metallophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D78D1F96C235E7504" box="[439,573,1782,1803]" italics="true" pageId="16" pageNumber="32">Metallophilus</emphasis>
|
||
Chaudoir, 1838
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="60E84D56FFDB167D79DBF88B20AD7529" authority="Sciaky, 1996" authorityName="Sciaky" authorityYear="1996" box="[189,462,1808,1831]" class="Insecta" family="Carabidae" genus="Sinoreophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D79DBF88B20207529" box="[189,323,1809,1830]" italics="true" pageId="16" pageNumber="32">Sinoreophilus</emphasis>
|
||
Sciaky, 1996
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDB167D798EF8B125CB7078" blockId="16.[189,776,315,2071]" lastBlockId="16.[812,1399,315,2070]" pageId="16" pageNumber="32">
|
||
(2) Spermatheca. — It enters
|
||
<emphasis id="959CEAC7FFDB167D7B60F8B1237E754F" box="[518,541,1835,1856]" italics="true" pageId="16" pageNumber="32">bc</emphasis>
|
||
just near oviduct-bursal junction, widely ranging between undifferentiated and nearly differentiated
|
||
<typeStatus id="78538877FFDB167D782FF8FB20107579" box="[329,371,1889,1910]" pageId="16" pageNumber="32">type</typeStatus>
|
||
as postulated by
|
||
<bibRefCitation id="C3794B24FFDB167D7B4FF8FB23B27579" author="Bousquet Y." box="[553,721,1888,1910]" pageId="16" pageNumber="32" pagination="1 - 292" refId="ref19199" refString="Bousquet Y. 1999. Supraspecific classification of the Nearctic Pterostichini (Coleoptera: Carabidae) // Fabreries. Suppl. 9. Association des Entomologists Amateurs du Quebec, Quebec, Canada. P. 1 - 292." type="book chapter" year="1999">Bousquet [1999]</bibRefCitation>
|
||
. It is rather short annulated, more or less fusiform, and mostly has 1–2 minute diverticula toward spermatheca-bursal junction. Both number and position of these diverticula are speciesspecific characters of slight or no individual variation [Sasakawa, 2004], which point of view I incline to share. For instance, there are two diverticula on either side of the gland duct-spermathecal junction, the proximal one being closer to this junction (
|
||
<taxonomicName id="60E84D56FFDB167D7ADEFECD25447364" authorityName="Bates" authorityYear="1883" box="[952,1063,342,364]" class="Insecta" family="Carabidae" genus="Plectes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="defossus">
|
||
<emphasis id="959CEAC7FFDB167D7ADEFECD25447364" box="[952,1063,342,364]" italics="true" pageId="16" pageNumber="32">P. defossus</emphasis>
|
||
</taxonomicName>
|
||
) or adjoining
|
||
<emphasis id="959CEAC7FFDB167D7DDEFECC25AC7364" box="[1208,1231,342,363]" italics="true" pageId="16" pageNumber="32">bc</emphasis>
|
||
(
|
||
<figureCitation id="3FD32A50FFDB167D7DB8FECC244A7363" box="[1246,1321,342,364]" captionStart="Figs 63–68" captionStartId="15.[226,263,1873,1892]" captionTargetBox="[193,1392,817,1839]" captionTargetId="figure-183@15.[188,1398,813,1843]" captionTargetPageId="15" captionText="Figs 63–68. Reproductive tract in female: 63 — Pterostichus austrinus sp.n.; 64–65 — P. glabellus sp.n.; 66 — P. strenuus (Panzer, 1796); 67 — P. chameleon; 68 — P. unisetosus sp.n.; 63, 65–67 — ventral aspect; 64, 68 — dorsal aspect; bc — bursa copulatrix; bs — bulbous structure; dv — diverticulum; ov — common oviduct; rp — receptacle; sc — seminal canal; sg — spermathecal gland; sgd — spermathecal gland duct; sp — spermatheca. Scale bars 1 mm. Рис. 63–68. Репродуктивный тракт самки: 63 — Pterostichus austrinus sp.n.; 64–65 — P. glabellus sp.n.; 66 — P. strenuus (Panzer, 1796); 67 — P. chameleon; 68 — P. unisetosus sp.n; 63, 65–67 — вентрально; 64, 68 — дорЗально; bc — копулЯтивнаЯ сумка; bs — бульбарное обраЗование; dv — дивертикул; ov — непарный Яйцевод; rp — семеприемник; sc — семепровод; sg — ЖелеЗа сперматеки; sgd — проток ЖелеЗы сперматеки; sp — сперматека. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949114" httpUri="https://zenodo.org/record/10949114/files/figure.png" pageId="16" pageNumber="32">Fig. 66</figureCitation>
|
||
). When single diverticulum is present it either adjoins
|
||
<emphasis id="959CEAC7FFDB167D7D80FEEB259E7389" box="[1254,1277,369,390]" italics="true" pageId="16" pageNumber="32">bc</emphasis>
|
||
(
|
||
<taxonomicName id="60E84D56FFDB167D7C6CFEEB222673AE" ID-CoL="4Q3FZ" authorityName="A. Morawitz" authorityYear="1862" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="sulcitarsis">
|
||
<emphasis id="959CEAC7FFDB167D7C6CFEEB222673AE" italics="true" pageId="16" pageNumber="32">P. sulcitarsis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D7A31FE1622BC73AF" ID-CoL="4Q3FS" authorityName="" baseAuthorityName="Motschulsky" baseAuthorityYear="1866" box="[855,991,395,417]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="chameleon">
|
||
<emphasis id="959CEAC7FFDB167D7A31FE1622BC73AF" box="[855,991,395,417]" italics="true" pageId="16" pageNumber="32">P. chameleon</emphasis>
|
||
</taxonomicName>
|
||
) or is situated between the gland duct-spermathecal junction and the spermatheca-bursal one (
|
||
<taxonomicName id="60E84D56FFDB167D7C04FE3D22E973D9" ID-CoL="4Q4B4" authorityName="Jedlicka" authorityYear="1962" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="lutschniki">
|
||
<emphasis id="959CEAC7FFDB167D7C04FE3D22E973D9" italics="true" pageId="16" pageNumber="32">P. lutschniki</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="60E84D56FFDB167D7AF8FE5B256C73D9" ID-CoL="6SK65" authorityName="Motschulsky" authorityYear="1844" box="[926,1039,449,470]" class="Insecta" family="Carabidae" genus="Omaseus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="laticollis">
|
||
<emphasis id="959CEAC7FFDB167D7AF8FE5B256C73D9" box="[926,1039,449,470]" italics="true" pageId="16" pageNumber="32">P. laticollis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="60E84D56FFDB167D7D7AFE5B25A173D9" ID-CoL="8LFDQ" authorityName="Tschitscherine" authorityYear="1889" box="[1052,1218,449,470]" class="Insecta" family="Carabidae" genus="Eurythorax" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="haptoderoides">
|
||
<emphasis id="959CEAC7FFDB167D7D7AFE5B25A173D9" box="[1052,1218,449,470]" italics="true" pageId="16" pageNumber="32">P. haptoderoides</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<figureCitation id="3FD32A50FFDB167D7D8EFE5B245F73D8" box="[1256,1340,449,471]" captionStart="Figs 100–104" captionStartId="20.[226,264,1897,1916]" captionTargetBox="[197,1386,626,1869]" captionTargetId="figure-320@20.[189,1399,625,1869]" captionTargetPageId="20" captionText="Figs 100–104. Reproductive tract in female: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — bursa copulatrix; dv — diverticulum; ov — common oviduct; rp — receptacle; sc — seminal canal; sg — spermathecal gland; 63, 65–67 — ventral aspect; 64, 68 — dorsal aspect; sgd — spermathecal gland duct; sp — spermatheca. Scale bars 1 mm. Рис. 100–104. Репродуктивный тракт самки: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — копулЯтивнаЯ сумка; dv — дивертикул; ov — непарный Яйцевод; rp — семеприемник; sc — семепровод; sg — ЖелеЗа сперматеки; sgd — проток ЖелеЗы сперматеки; sp — сперматека. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949120" httpUri="https://zenodo.org/record/10949120/files/figure.png" pageId="16" pageNumber="32">Fig. 100</figureCitation>
|
||
), or it has given rise to the spermathecal gland duct (
|
||
<taxonomicName id="60E84D56FFDB167D7D8DFE46240B73FE" box="[1259,1384,476,497]" class="Insecta" family="Carabidae" genus="Plectes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="usuriensis">
|
||
<emphasis id="959CEAC7FFDB167D7D8DFE46240B73FE" box="[1259,1384,476,497]" italics="true" pageId="16" pageNumber="32">P. usuriensis</emphasis>
|
||
</taxonomicName>
|
||
). Other species examined either have no distinct deverticula (
|
||
<taxonomicName id="60E84D56FFDB167D7A52FD8B22C37029" ID-CoL="QZHZ" authorityName="Panzer" authorityYear="1796" box="[820,928,529,550]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="vernalis">
|
||
<emphasis id="959CEAC7FFDB167D7A52FD8B22C37029" box="[820,928,529,550]" italics="true" pageId="16" pageNumber="32">P. vernalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D7AC9FD8B25647029" ID-CoL="4Q3FW" baseAuthorityName="Bates" baseAuthorityYear="1883" box="[943,1031,529,550]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="dulcis">
|
||
<emphasis id="959CEAC7FFDB167D7AC9FD8B25647029" box="[943,1031,529,550]" italics="true" pageId="16" pageNumber="32">P. dulcis</emphasis>
|
||
</taxonomicName>
|
||
) or, in contrast, have three ones. At least distal two of them are large and thence similar to underdeveloped spermathecae that begin on the gland duct-spermathecal junction (
|
||
<figureCitation id="3FD32A50FFDB167D7D74FDFB25FF7078" box="[1042,1180,609,631]" captionStart="Figs 100–104" captionStartId="20.[226,264,1897,1916]" captionTargetBox="[197,1386,626,1869]" captionTargetId="figure-320@20.[189,1399,625,1869]" captionTargetPageId="20" captionText="Figs 100–104. Reproductive tract in female: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — bursa copulatrix; dv — diverticulum; ov — common oviduct; rp — receptacle; sc — seminal canal; sg — spermathecal gland; 63, 65–67 — ventral aspect; 64, 68 — dorsal aspect; sgd — spermathecal gland duct; sp — spermatheca. Scale bars 1 mm. Рис. 100–104. Репродуктивный тракт самки: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — копулЯтивнаЯ сумка; dv — дивертикул; ov — непарный Яйцевод; rp — семеприемник; sc — семепровод; sg — ЖелеЗа сперматеки; sgd — проток ЖелеЗы сперматеки; sp — сперматека. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949120" httpUri="https://zenodo.org/record/10949120/files/figure.png" pageId="16" pageNumber="32">Figs 101–102</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDB167D7A34FDE125EC767E" blockId="16.[812,1399,315,2070]" pageId="16" pageNumber="32">
|
||
This junction also varies considerably in position from species to species. It mostly ranges within basal 1/3 spermatheca, while sometimes driving closer to its middle (
|
||
<taxonomicName id="60E84D56FFDB167D7C04FD2B22EC70EF" ID-CoL="4Q3FZ" authorityName="A. Morawitz" authorityYear="1862" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="sulcitarsis">
|
||
<emphasis id="959CEAC7FFDB167D7C04FD2B22EC70EF" italics="true" pageId="16" pageNumber="32">P. sulcitarsis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D7AF9FD56257970EF" ID-CoL="4Q4B4" authorityName="Jedlicka" authorityYear="1962" box="[927,1050,715,737]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="lutschniki">
|
||
<emphasis id="959CEAC7FFDB167D7AF9FD56257970EF" box="[927,1050,715,737]" italics="true" pageId="16" pageNumber="32">P. lutschniki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D7D4CFD5625E170EF" ID-CoL="4Q3FW" baseAuthorityName="Bates" baseAuthorityYear="1883" box="[1066,1154,715,737]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="dulcis">
|
||
<emphasis id="959CEAC7FFDB167D7D4CFD5625E170EF" box="[1066,1154,715,737]" italics="true" pageId="16" pageNumber="32">P. dulcis</emphasis>
|
||
</taxonomicName>
|
||
). Variation range of the spermatheca shape and length is much wider. The spermatheca is mostly rather short, subfusiform, more or less c- or sshaped, about as long as folded
|
||
<emphasis id="959CEAC7FFDB167D7D3DFC812511713F" box="[1115,1138,795,816]" italics="true" pageId="16" pageNumber="32">bc</emphasis>
|
||
and slightly exceeding this latter in width, broadest in about apical third, with no or not well differentiated receptacle and seminal canal. Rather slight modifications include spermathecae either clavate, more or less enlarged and broadened apicad (
|
||
<taxonomicName id="60E84D56FFDB167D7DCEFC1C24707194" ID-CoL="8LSFB" authorityName="Lutshnik" authorityYear="1929" box="[1192,1299,902,923]" class="Insecta" family="Carabidae" genus="Omaseulus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7DCEFC1C24707194" box="[1192,1299,902,923]" italics="true" pageId="16" pageNumber="32">Omaseulus</emphasis>
|
||
</taxonomicName>
|
||
) or long, slender and slightly wider in apical than in basal half (
|
||
<taxonomicName id="60E84D56FFDB167D7C04FC3B22F371DF" ID-CoL="4Q3FZ" authorityName="A. Morawitz" authorityYear="1862" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="sulcitarsis">
|
||
<emphasis id="959CEAC7FFDB167D7C04FC3B22F371DF" italics="true" pageId="16" pageNumber="32">P. sulcitarsis</emphasis>
|
||
</taxonomicName>
|
||
). Profound modification are two, spermatheca either very elongated, tubiform, somewhat corrugated, and capitate apically (
|
||
<figureCitation id="3FD32A50FFDB167D7A84FC6B25597608" box="[994,1082,1009,1031]" captionStart="Figs 100–104" captionStartId="20.[226,264,1897,1916]" captionTargetBox="[197,1386,626,1869]" captionTargetId="figure-320@20.[189,1399,625,1869]" captionTargetPageId="20" captionText="Figs 100–104. Reproductive tract in female: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — bursa copulatrix; dv — diverticulum; ov — common oviduct; rp — receptacle; sc — seminal canal; sg — spermathecal gland; 63, 65–67 — ventral aspect; 64, 68 — dorsal aspect; sgd — spermathecal gland duct; sp — spermatheca. Scale bars 1 mm. Рис. 100–104. Репродуктивный тракт самки: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — копулЯтивнаЯ сумка; dv — дивертикул; ov — непарный Яйцевод; rp — семеприемник; sc — семепровод; sg — ЖелеЗа сперматеки; sgd — проток ЖелеЗы сперматеки; sp — сперматека. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949120" httpUri="https://zenodo.org/record/10949120/files/figure.png" pageId="16" pageNumber="32">Fig. 103</figureCitation>
|
||
) or well-differentiated (
|
||
<taxonomicName id="60E84D56FFDB167D7C52FC6B2213762F" ID-CoL="4Q4B4" authorityName="Jedlicka" authorityYear="1962" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="lutschniki">
|
||
<emphasis id="959CEAC7FFDB167D7C52FC6B2213762F" italics="true" pageId="16" pageNumber="32">P. lutschniki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D7A19FB9622B5762F" ID-CoL="4Q3FW" baseAuthorityName="Bates" baseAuthorityYear="1883" box="[895,982,1035,1057]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="dulcis">
|
||
<emphasis id="959CEAC7FFDB167D7A19FB9622B5762F" box="[895,982,1035,1057]" italics="true" pageId="16" pageNumber="32">P. dulcis</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<figureCitation id="3FD32A50FFDB167D7A9BFB912530762E" box="[1021,1107,1035,1057]" captionStart="Figs 100–104" captionStartId="20.[226,264,1897,1916]" captionTargetBox="[197,1386,626,1869]" captionTargetId="figure-320@20.[189,1399,625,1869]" captionTargetPageId="20" captionText="Figs 100–104. Reproductive tract in female: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — bursa copulatrix; dv — diverticulum; ov — common oviduct; rp — receptacle; sc — seminal canal; sg — spermathecal gland; 63, 65–67 — ventral aspect; 64, 68 — dorsal aspect; sgd — spermathecal gland duct; sp — spermatheca. Scale bars 1 mm. Рис. 100–104. Репродуктивный тракт самки: 100 — Pterostichus haptoderoides; 101 — P. batdaiensis sp.n.; 102 — P. stricticollis; 103 — P. gibbicollis; 104 — P. dulcis; bc — копулЯтивнаЯ сумка; dv — дивертикул; ov — непарный Яйцевод; rp — семеприемник; sc — семепровод; sg — ЖелеЗа сперматеки; sgd — проток ЖелеЗы сперматеки; sp — сперматека. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949120" httpUri="https://zenodo.org/record/10949120/files/figure.png" pageId="16" pageNumber="32">Fig. 104</figureCitation>
|
||
), angled between narrow and elongated seminal canal and shortened and broadened receptacle, except only that the gland duct-spermathecal junction is proximal to (
|
||
<emphasis id="959CEAC7FFDB167D7AA5FBC622B4767E" box="[963,983,1116,1137]" italics="true" pageId="16" pageNumber="32">vs</emphasis>
|
||
. just in) the angle.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDB167D7A34FBEC22AD77C3" blockId="16.[812,1399,315,2070]" pageId="16" pageNumber="32">
|
||
It follows that the structures such as undifferentiated spermatheca, differentiated spermatheca, with well-developed receptacle and seminal canal, spermathecal diverticula and bulbous structures at the oviduct-bursal junction, occur in these or those members of the
|
||
<taxonomicName id="60E84D56FFDB167D7D0BFB7B25DA76F9" authorityName="Dejean" authorityYear="1821" box="[1133,1209,1249,1270]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7D0BFB7B25DA76F9" box="[1133,1209,1249,1270]" italics="true" pageId="16" pageNumber="32">Argutor</emphasis>
|
||
</taxonomicName>
|
||
lineage, and undifferentiated spermatheca with diverticulum or diverticula dominates over the other patterns. The fact that diverticulum also occurs in
|
||
<taxonomicName id="60E84D56FFDB167D7AECFAAB250A7749" ID-CoL="8KWT3" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" box="[906,1129,1329,1351]" class="Insecta" family="Carabidae" genus="Poecilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7AECFAAB22BE7749" box="[906,989,1329,1350]" italics="true" pageId="16" pageNumber="32">Poecilus</emphasis>
|
||
Bonelli, 1810
|
||
</taxonomicName>
|
||
, as a member of the lineage sister to
|
||
<taxonomicName id="60E84D56FFDB167D7AE0FAD1229C776F" ID-CoL="8KVYP" authorityName="Bonelli" authorityYear="1810" box="[902,1023,1355,1376]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7AE0FAD1229C776F" box="[902,1023,1355,1376]" italics="true" pageId="16" pageNumber="32">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
[Sasakawa, Kubota, 2007] while the most differentiated spermatheca has only been observed in
|
||
<taxonomicName id="60E84D56FFDB167D7C04FAFD22AA7799" ID-CoL="4Q3FW" baseAuthorityName="Bates" baseAuthorityYear="1883" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="dulcis" subGenus="Argutor">
|
||
<emphasis id="959CEAC7FFDB167D7C04FAFD24147773" box="[1378,1399,1383,1404]" italics="true" pageId="16" pageNumber="32">P.</emphasis>
|
||
(
|
||
<emphasis id="959CEAC7FFDB167D7A52FA1B22E37799" box="[820,896,1409,1430]" italics="true" pageId="16" pageNumber="32">Argutor</emphasis>
|
||
)
|
||
<emphasis id="959CEAC7FFDB167D7AF6FA1B22AA7799" box="[912,969,1409,1430]" italics="true" pageId="16" pageNumber="32">dulcis</emphasis>
|
||
</taxonomicName>
|
||
may suggest that this
|
||
<typeStatus id="78538877FFDB167D7DC0FA1B25B37799" box="[1190,1232,1409,1430]" pageId="16" pageNumber="32">type</typeStatus>
|
||
spermatheca has step by step evolved from undifferentiated one within the
|
||
<taxonomicName id="60E84D56FFDB167D7A4AFA2C221B77C4" authorityName="Dejean" authorityYear="1821" box="[812,888,1462,1483]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7A4AFA2C221B77C4" box="[812,888,1462,1483]" italics="true" pageId="16" pageNumber="32">Argutor</emphasis>
|
||
</taxonomicName>
|
||
lineage.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDB167D7A34FA4B252C759E" blockId="16.[812,1399,315,2070]" pageId="16" pageNumber="32">
|
||
<taxonomicName id="60E84D56FFDB167D7A34FA4B22A377E9" authorityName="Motschulsky" authorityYear="1866" box="[850,960,1489,1510]" class="Insecta" family="Carabidae" genus="Badistrinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7A34FA4B22A377E9" box="[850,960,1489,1510]" italics="true" pageId="16" pageNumber="32">Badistrinus</emphasis>
|
||
</taxonomicName>
|
||
is by comparison the most heterogeneous subgenus of this lineage, whereas it actually includes hardly more than four similar species (
|
||
<taxonomicName id="60E84D56FFDB167D7D31F99D25A07414" ID-CoL="6SK65" authorityName="Motschulsky" authorityYear="1844" box="[1111,1219,1542,1564]" class="Insecta" family="Carabidae" genus="Omaseus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="laticollis">
|
||
<emphasis id="959CEAC7FFDB167D7D31F99D25A07414" box="[1111,1219,1542,1564]" italics="true" pageId="16" pageNumber="32">P. laticollis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D7DA9F99D24137414" ID-CoL="8LFDQ" authorityName="Tschitscherine" authorityYear="1889" box="[1231,1392,1542,1564]" class="Insecta" family="Carabidae" genus="Eurythorax" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="haptoderoides">
|
||
<emphasis id="959CEAC7FFDB167D7DA9F99D24137414" box="[1231,1392,1542,1564]" italics="true" pageId="16" pageNumber="32">P. haptoderoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60E84D56FFDB167D7A4AF9BB22CF7439" ID-CoL="78M5S" baseAuthorityName="Tschitscherine" baseAuthorityYear="1897" box="[812,940,1569,1590]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="modicellus">
|
||
<emphasis id="959CEAC7FFDB167D7A4AF9BB22CF7439" box="[812,940,1569,1590]" italics="true" pageId="16" pageNumber="32">P. modicellus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="60E84D56FFDB167D7A86F9BB244D7439" ID-CoL="8MF58" authority="Habu et Tanaka, 1957" authorityName="Habu et Tanaka" authorityYear="1957" box="[992,1326,1568,1591]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="kajimurai">
|
||
<emphasis id="959CEAC7FFDB167D7A86F9BB25367439" box="[992,1109,1569,1590]" italics="true" pageId="16" pageNumber="32">P. kajimurai</emphasis>
|
||
Habu et Tanaka, 1957
|
||
</taxonomicName>
|
||
). These all share many characters specified in the key below, including extremely long
|
||
<emphasis id="959CEAC7FFDB167D7AA7F9CC253D7464" box="[961,1118,1622,1643]" italics="true" pageId="16" pageNumber="32">bursa copulatrix</emphasis>
|
||
and internal sac of aedeagus, certainly interdependent characters. The other species listed under the subgenus [
|
||
<bibRefCitation id="C3794B24FFDB167D7A98F91625E474AF" author="Lorenz W." box="[1022,1159,1675,1697]" pageId="16" pageNumber="32" refId="ref19861" refString="Lorenz W. 1998. Systematic list of extant ground beetles of the world (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 1 st ed. Tutzing. 502 pp." type="book" year="1998">Lorenz, 1998</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="C3794B24FFDB167D7DF3F91125A474AE" author="Lorenz W." box="[1173,1223,1675,1697]" pageId="16" pageNumber="32" refId="ref19903" refString="Lorenz W. 2005. Systematic list of extant ground beetles of the world (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd ed. Tutzing. 530 pp." type="book" year="2005">2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C3794B24FFDB167D7DB0F916241074AF" author="Bousquet Y." box="[1238,1395,1675,1697]" pageId="16" pageNumber="32" pagination="469 - 521" refId="ref19277" refString="Bousquet Y. 2003. Tribe Pterostichini Bonelli, 1810 // I. Lobl, A. Smetana (eds.). Catalogue of Palearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Stenstrup: Appolo Books. P. 469 - 521." type="book chapter" year="2003">Bousquet, 2003</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="C3794B24FFDB167D7A4AF93C223D74B4" author="Bousquet Y." box="[812,862,1702,1723]" pageId="16" pageNumber="32" pagination="675 - 755" refId="ref19326" refString="Bousquet Y. 2017. Tribe Pterostichini Bonelli, 1810 // I. Lobl, D. Lobl (eds.). Catalogue of Palearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Revised and updated edition. Leiden-Boston: Brill. P. 675 - 755." type="book chapter" year="2017">2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C3794B24FFDB167D7A0FF93C250A74B4" author="Makarov K. V. & Sundukov Yu. N." box="[873,1129,1701,1724]" pageId="16" pageNumber="32" pagination="362 - 372" refId="ref19973" refString="Makarov K. V., Sundukov Yu. N. 2022. On status and systematic position of Pterostichus goschi Jedlicka, 1930 (Coleoptera: Carabidae) // Russian Entomol. J. Vol. 31. No. 4. P. 362 - 372. doi: 10.15298 / rusentj. 31.4.05." type="journal article" year="2022">Makarov, Sundukov, 2022</bibRefCitation>
|
||
] are different. Smaller sized ones,
|
||
<taxonomicName id="60E84D56FFDB167D7A02F95B25F674D9" authority="A. Morawitz, 1862" authorityName="A. Morawitz" authorityYear="1862" box="[868,1173,1728,1751]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="neglectus">
|
||
<emphasis id="959CEAC7FFDB167D7A02F95B22BB74D9" box="[868,984,1729,1750]" italics="true" pageId="16" pageNumber="32">P. neglectus</emphasis>
|
||
A. Morawitz, 1862
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFDB167D7DA2F95B223D74FF" authority="Jedlicka, 1930" authorityName="Jedlicka" authorityYear="1930" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="goschi">
|
||
<emphasis id="959CEAC7FFDB167D7DA2F95B247F74D9" box="[1220,1308,1729,1750]" italics="true" pageId="16" pageNumber="32">P. goschi</emphasis>
|
||
<bibRefCitation id="C3794B24FFDB167D7C45F95B223D74FF" author="Jedlicka A." pageId="16" pageNumber="32" pagination="21 - 24" refId="ref19672" refString="Jedlicka A. 1930. Novi palaearkticti Carbicidi. (V. pokrac.). Neue palaeark. Carabiciden. (V. Folge.) // Cas. Cesk. Spol. Ent. Vol. 27. P. 21 - 24." type="journal article" year="1930">Jedlička, 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
, are more similar to
|
||
<taxonomicName id="60E84D56FFDB167D7D79F946246474FF" ID-CoL="GHQK" authorityName="Motschulsky" authorityYear="1844" box="[1055,1287,1755,1777]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="gibbicollis" subGenus="Pledarus">
|
||
<emphasis id="959CEAC7FFDB167D7D79F946255774FE" box="[1055,1076,1756,1777]" italics="true" pageId="16" pageNumber="32">P.</emphasis>
|
||
(
|
||
<emphasis id="959CEAC7FFDB167D7D27F94125F574FF" box="[1089,1174,1755,1776]" italics="true" pageId="16" pageNumber="32">Pledarus</emphasis>
|
||
)
|
||
<emphasis id="959CEAC7FFDB167D7DC2F941246474FF" box="[1188,1287,1755,1776]" italics="true" pageId="16" pageNumber="32">gibbicollis</emphasis>
|
||
</taxonomicName>
|
||
with which they share the left paramere quadrate, both
|
||
<emphasis id="959CEAC7FFDB167D7DB1F96C24157504" box="[1239,1398,1782,1803]" italics="true" pageId="16" pageNumber="32">bursa copulatrix</emphasis>
|
||
and endophallus shorter, and the pronotum similar in shape. However, the elytral discal setae are reduced to d
|
||
<quantity id="60109B30FFDB167D7D9FF8B1247F754E" box="[1273,1308,1835,1857]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="16" pageNumber="32" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName id="60E84D56FFDB167D7C47F8B12415754F" authorityName="Motschulsky" authorityYear="1866" box="[1313,1398,1835,1856]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7C47F8B12415754F" box="[1313,1398,1835,1856]" italics="true" pageId="16" pageNumber="32">Pledarus</emphasis>
|
||
</taxonomicName>
|
||
and spermatheca is unique in at least
|
||
<taxonomicName id="60E84D56FFDB167D7DE1F8DD24607554" ID-CoL="GHQK" authorityName="Motschulsky" authorityYear="1844" box="[1159,1283,1862,1884]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="gibbicollis">
|
||
<emphasis id="959CEAC7FFDB167D7DE1F8DD24607554" box="[1159,1283,1862,1884]" italics="true" pageId="16" pageNumber="32">P. gibbicollis</emphasis>
|
||
</taxonomicName>
|
||
, which may suggest that a separate species group or a subgenus is required for
|
||
<taxonomicName id="60E84D56FFDB167D7A28F8E622A1759F" authorityName="A. Morawitz" authorityYear="1862" box="[846,962,1915,1937]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="neglectus">
|
||
<emphasis id="959CEAC7FFDB167D7A28F8E622A1759F" box="[846,962,1915,1937]" italics="true" pageId="16" pageNumber="32">P. neglectus</emphasis>
|
||
</taxonomicName>
|
||
with its allies.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDB167C7A34F80C23D67004" blockId="16.[812,1399,315,2070]" lastBlockId="17.[189,775,315,684]" lastPageId="17" lastPageNumber="33" pageId="16" pageNumber="32">
|
||
In reassessing
|
||
<taxonomicName id="60E84D56FFDB167D7A8EF80C253D75A4" ID-CoL="8KVYP" authorityName="Bonelli" authorityYear="1810" box="[1000,1118,1942,1963]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7A8EF80C253D75A4" box="[1000,1118,1942,1963]" italics="true" pageId="16" pageNumber="32">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
from the Russian Far East
|
||
<bibRefCitation id="C3794B24FFDB167D7A4AF82A22B775C9" author="Sundukov Yu. N." box="[812,980,1968,1990]" pageId="16" pageNumber="32" refId="ref20559" refString="Sundukov Yu. N. 2013. [An annotated catalogue of the ground beetles (Coleoptera: Caraboidea) of Sikhote-Alin]. Vladivostok: Dalnauka. 271 p. [In Russian]" type="book" year="2013">Sundukov [2013]</bibRefCitation>
|
||
briefly reviewed and commented on seven subgenera that shared the
|
||
<emphasis id="959CEAC7FFDB167D7D43F856255675EE" box="[1061,1077,1996,2017]" italics="true" pageId="16" pageNumber="32">tr</emphasis>
|
||
3 with seta,
|
||
<emphasis id="959CEAC7FFDB167D7DCFF85625DD75EE" box="[1193,1214,1996,2017]" italics="true" pageId="16" pageNumber="32">cx</emphasis>
|
||
3 bisetose and long metepisternum, five of these subgenera belonging to the
|
||
<taxonomicName id="60E84D56FFDB167D7C59F87C222B7A19" authorityName="Dejean" authorityYear="1821" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDB167D7C59F87C222B7A19" italics="true" pageId="16" pageNumber="32">Argutor</emphasis>
|
||
</taxonomicName>
|
||
lineage. He transferred
|
||
<taxonomicName id="60E84D56FFDB167D7D5FF79B25DC7A1A" ID-CoL="4Q49V" authorityName="Bates" authorityYear="1873" box="[1081,1215,2048,2070]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="32" phylum="Arthropoda" rank="species" species="longinquus">
|
||
<emphasis id="959CEAC7FFDB167D7D5FF79B25DC7A1A" box="[1081,1215,2048,2070]" italics="true" pageId="16" pageNumber="32">P. longinquus</emphasis>
|
||
</taxonomicName>
|
||
and allied species from
|
||
<taxonomicName id="60E84D56FFDA167C7997FEA1205D735F" authorityName="Gozis" authorityYear="1886" box="[241,318,315,336]" pageId="17" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFDA167C7997FEA1205D735F" box="[241,318,315,336]" italics="true" pageId="17" pageNumber="33">Phonias</emphasis>
|
||
</taxonomicName>
|
||
to its formerly junior synonym,
|
||
<taxonomicName id="60E84D56FFDA167C7B11FEA123B4735F" ID-CoL="8LR59" authorityName="Jeanne" authorityYear="1988" box="[631,727,315,336]" class="Insecta" family="Carabidae" genus="Biphonias" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDA167C7B11FEA123B4735F" box="[631,727,315,336]" italics="true" pageId="17" pageNumber="33">Biphonias</emphasis>
|
||
</taxonomicName>
|
||
, and distinguished between the two subgenera chiefly by the elytral parascutellar seta either absent or present, respectively. Yet, this resurrection proved to be invalid since the
|
||
<typeStatus id="78538877FFDA167C7B1CFE1623C073AE" box="[634,675,396,417]" pageId="17" pageNumber="33">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="60E84D56FFDA167C79DBFE3C206973B4" authorityName="Gozis" authorityYear="1886" box="[189,266,422,443]" pageId="17" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFDA167C79DBFE3C206973B4" box="[189,266,422,443]" italics="true" pageId="17" pageNumber="33">Phonias</emphasis>
|
||
</taxonomicName>
|
||
had the parascutellar seta. Transfer of
|
||
<taxonomicName id="60E84D56FFDA167C7BF6FE3D226573B4" authorityName="A. Morawitz" authorityYear="1862" box="[656,774,422,444]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="species" species="neglectus">
|
||
<emphasis id="959CEAC7FFDA167C7BF6FE3D226573B4" box="[656,774,422,444]" italics="true" pageId="17" pageNumber="33">P. neglectus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFDA167C7980FE5B202373D9" authorityName="Jedlicka" authorityYear="1930" box="[230,320,449,470]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="species" species="goschi">
|
||
<emphasis id="959CEAC7FFDA167C7980FE5B202373D9" box="[230,320,449,470]" italics="true" pageId="17" pageNumber="33">P. goschi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="60E84D56FFDA167C781AFE5B208A73D9" authorityName="Motschulsky" authorityYear="1866" box="[380,489,449,470]" class="Insecta" family="Carabidae" genus="Badistrinus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDA167C781AFE5B208A73D9" box="[380,489,449,470]" italics="true" pageId="17" pageNumber="33">Badistrinus</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="60E84D56FFDA167C7B6FFE5B230A73D9" ID-CoL="8LR59" authorityName="Jeanne" authorityYear="1988" box="[521,617,449,470]" class="Insecta" family="Carabidae" genus="Biphonias" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDA167C7B6FFE5B230A73D9" box="[521,617,449,470]" italics="true" pageId="17" pageNumber="33">Biphonias</emphasis>
|
||
</taxonomicName>
|
||
was also wrong because the two species had the elytral discal setae adjoining stria 3, not stria 2 as was characteristic of
|
||
<taxonomicName id="60E84D56FFDA167C7B29FE6C23CC7004" ID-CoL="8LR59" authorityName="Jeanne" authorityYear="1988" box="[591,687,502,523]" class="Insecta" family="Carabidae" genus="Biphonias" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDA167C7B29FE6C23CC7004" box="[591,687,502,523]" italics="true" pageId="17" pageNumber="33">Biphonias</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDA167C7984FD8B254873FE" blockId="17.[189,775,315,684]" lastBlockId="17.[812,1399,315,684]" pageId="17" pageNumber="33">
|
||
The elytral discal setae d2 and d3 that adjoin stria 2 argue for
|
||
<taxonomicName id="60E84D56FFDA167C787AFDB62019704E" authorityName="Jedlicka" authorityYear="1936" box="[284,378,556,577]" class="Insecta" family="Carabidae" genus="Pledarus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="species" species="arrowi">
|
||
<emphasis id="959CEAC7FFDA167C787AFDB62019704E" box="[284,378,556,577]" italics="true" pageId="17" pageNumber="33">P. arrowi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFDA167C78CFFDB623AF704F" ID-CoL="4Q3GC" authority="Jedlicka, 1938" authorityName="Jedlicka" authorityYear="1938" box="[425,716,555,577]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="species" species="arrowianus">
|
||
<emphasis id="959CEAC7FFDA167C78CFFDB62356704E" box="[425,565,556,577]" italics="true" pageId="17" pageNumber="33">P. arrowianus</emphasis>
|
||
<bibRefCitation id="C3794B24FFDA167C7B5AFDB123AF704F" author="Jedlicka A." box="[572,716,555,577]" pageId="17" pageNumber="33" refId="ref19747" refString="Jedlicka A. 1938. Versuch einer Bestimmungstabelle der mir bekannten Pterostichus - Arten aus Ostasien. Praha. 12 S." type="book" year="1938">Jedlička, 1938</bibRefCitation>
|
||
</taxonomicName>
|
||
, to be transferred from
|
||
<taxonomicName id="60E84D56FFDA167C7803FDDC20B57054" authorityName="Motschulsky" authorityYear="1866" box="[357,470,582,603]" class="Insecta" family="Carabidae" genus="Badistrinus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDA167C7803FDDC20B57054" box="[357,470,582,603]" italics="true" pageId="17" pageNumber="33">Badistrinus</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="60E84D56FFDA167C789FFDDC232A7054" authorityName="Gozis" authorityYear="1886" box="[505,585,582,603]" pageId="17" pageNumber="38" rank="subGenus" subGenus="Phonias">
|
||
<emphasis id="959CEAC7FFDA167C789FFDDC232A7054" box="[505,585,582,603]" italics="true" pageId="17" pageNumber="33">Phonias</emphasis>
|
||
</taxonomicName>
|
||
. This latter subgenus seems to be primitivemost among the others as it is largely defined by plesiomorphous characters: the body macropterous, the pronotum with both inner and outer basolateral sulcus well-developed, the tarsi distinctly bisulcate, the elytral striation and setation complete,
|
||
<emphasis id="959CEAC7FFDA167C7C4EFECD24257363" box="[1320,1350,343,364]" italics="true" pageId="17" pageNumber="33">i.e.</emphasis>
|
||
, the parascutellar striole long, combined with the parascutellar seta and three discal setae present. Furthermore,
|
||
<emphasis id="959CEAC7FFDA167C7C58FE1122F173B4" italics="true" pageId="17" pageNumber="33">bursa copulatrix</emphasis>
|
||
and endophallus are moderate in length and spermatheca seems to be of both shape and structure primary for the lineage as well.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDA167C7A34FE6C229770A3" blockId="17.[812,1399,315,684]" pageId="17" pageNumber="33">
|
||
<taxonomicName id="60E84D56FFDA167C7A34FE6C22FD7004" authorityName="Dejean" authorityYear="1821" box="[850,926,502,523]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFDA167C7A34FE6C22FD7004" box="[850,926,502,523]" italics="true" pageId="17" pageNumber="33">Argutor</emphasis>
|
||
</taxonomicName>
|
||
is nearly the same, but for certain additional symplesiomorphies such as the apically beaded prosternal process and the laterally setose metatibia, combined with a few synapomorphies such as the dorsally trisulcate tarsi, the parascutellar striole missing (modified), and spermatheca differentiated. The remaining subgenera or groups appear to be farther advanced.
|
||
</paragraph>
|
||
<caption id="F397665DFFDA167C7984F81B25E67A1E" ID-DOI="http://doi.org/10.5281/zenodo.10949116" ID-Zenodo-Dep="10949116" httpUri="https://zenodo.org/record/10949116/files/figure.png" pageId="17" pageNumber="33" startId="17.[226,263,1921,1940]" targetBox="[193,1395,703,1901]" targetPageId="17" targetType="figure">
|
||
<paragraph id="A75736D5FFDA167C7984F81B25E375CA" blockId="17.[188,1398,1921,2065]" pageId="17" pageNumber="33">
|
||
Figs 69–79. Aedeagus: 69–70 —
|
||
<taxonomicName id="60E84D56FFDA167C7B6CF81B23B0759B" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[522,723,1921,1940]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="species" species="laevibasis" status="sp. nov.">
|
||
<emphasis id="959CEAC7FFDA167C7B6CF81B23B0759B" box="[522,723,1921,1940]" italics="true" pageId="17" pageNumber="33">Pterostichus laevibasis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFDA167C7BBCF81B2266759B" bold="true" box="[730,773,1921,1940]" pageId="17" pageNumber="33">
|
||
<taxonomicNameLabel id="8EAF57BCFFDA167C7BBCF81B2266759B" box="[730,773,1921,1940]" pageId="17" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 71–79 —
|
||
<taxonomicName id="60E84D56FFDA167C7A0DF8182284759B" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[875,999,1921,1940]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="species" species="batdaiensis" status="sp.n.">
|
||
<emphasis id="959CEAC7FFDA167C7A0DF8182284759B" box="[875,999,1921,1940]" italics="true" pageId="17" pageNumber="33">P. batdaiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFDA167C7A88F81B257A759B" bold="true" box="[1006,1049,1921,1940]" pageId="17" pageNumber="33">
|
||
<taxonomicNameLabel id="8EAF57BCFFDA167C7A88F81B257A759B" box="[1006,1049,1921,1940]" pageId="17" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 71–72, 75, 77 — paratype from Sa Pa env.; 73–74, 76, 78–79 — paratype from type locality; 69–76 — median lobe; 77–78 — right paramere; 79 — left paramere; 69, 71, 73, 77– 78 — left lateral aspect; 70, 72, 74, 79 — right lateral aspect; 75–76 — ventrobasal aspect. Scale bars 1 mm.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFDA167C7984F85425E67A1E" blockId="17.[188,1398,1921,2065]" pageId="17" pageNumber="33">
|
||
Рис. 69–79. Эдеагус: 69–70 —
|
||
<taxonomicName id="60E84D56FFDA167C7890F85723DF75EF" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[502,700,1997,2016]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="species" species="laevibasis" status="sp. nov.">
|
||
<emphasis id="959CEAC7FFDA167C7890F85723DF75EF" box="[502,700,1997,2016]" italics="true" pageId="17" pageNumber="33">Pterostichus laevibasis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFDA167C7BA7F857238F75EF" bold="true" box="[705,748,1997,2016]" pageId="17" pageNumber="33">
|
||
<taxonomicNameLabel id="8EAF57BCFFDA167C7BA7F857238F75EF" box="[705,748,1997,2016]" pageId="17" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 71–79 —
|
||
<taxonomicName id="60E84D56FFDA167C7A2AF85422A675EF" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[844,965,1997,2016]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="33" phylum="Arthropoda" rank="species" species="batdaiensis" status="sp.n.">
|
||
<emphasis id="959CEAC7FFDA167C7A2AF85422A675EF" box="[844,965,1997,2016]" italics="true" pageId="17" pageNumber="33">P. batdaiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="959CEAC7FFDA167C7AACF857229675EF" bold="true" box="[970,1013,1997,2016]" pageId="17" pageNumber="33">
|
||
<taxonomicNameLabel id="8EAF57BCFFDA167C7AACF857229675EF" box="[970,1013,1997,2016]" pageId="17" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 71–72, 75, 77 — паратип иЗ окрестностей Sa Pa env.; 73–74, 76, 78–79 — паратип иЗ типового местонахоЖдениЯ; 71–76 — среднЯЯ долЯ; 77–78 — праваЯ парамера; 79 — леваЯ парамера; 69, 71, 73, 77–78 — слева; 70, 72, 74, 79 — справа; 75–76 — вентробаЗально. Масштаб 1 мм.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A75736D5FFD9167F7984FEA62301704E" blockId="18.[189,775,315,577]" pageId="18" pageNumber="34">
|
||
In sum, the
|
||
<taxonomicName id="60E84D56FFD9167F783EFEA620C7735E" authorityName="Dejean" authorityYear="1821" box="[344,420,316,337]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFD9167F783EFEA620C7735E" box="[344,420,316,337]" italics="true" pageId="18" pageNumber="34">Argutor</emphasis>
|
||
</taxonomicName>
|
||
lineage is currently recognized as a monophyletic group [Sasakawa, Kubota, 2007] well distinctive from the other
|
||
<taxonomicName id="60E84D56FFD9167F78E7FEEB20997389" ID-CoL="8KVYP" authorityName="Bonelli" authorityYear="1810" box="[385,506,369,390]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFD9167F78E7FEEB20997389" box="[385,506,369,390]" italics="true" pageId="18" pageNumber="34">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
. With this circumstance in mind, one could treat it as the separate genus
|
||
<taxonomicName id="60E84D56FFD9167F7B0FFE1623D673AE" authorityName="Dejean" authorityYear="1821" box="[617,693,396,417]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFD9167F7B0FFE1623D673AE" box="[617,693,396,417]" italics="true" pageId="18" pageNumber="34">Argutor</emphasis>
|
||
</taxonomicName>
|
||
while at least some subgenera it includes seem to be species groups rather than subgenera. On the other hand, some characters observed in
|
||
<taxonomicName id="60E84D56FFD9167F7855FE46201C73FE" authorityName="Dejean" authorityYear="1821" box="[307,383,476,497]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFD9167F7855FE46201C73FE" box="[307,383,476,497]" italics="true" pageId="18" pageNumber="34">Argutor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFD9167F78C8FE4623B473FF" ID-CoL="4Q3PX" authorityName="Reitter" authorityYear="1887" box="[430,727,475,497]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="34" phylum="Arthropoda" rank="species" species="pseudopedius" subGenus="Falsargutor">
|
||
<emphasis id="959CEAC7FFD9167F78C8FE4620A073FE" box="[430,451,476,497]" italics="true" pageId="18" pageNumber="34">P.</emphasis>
|
||
(
|
||
<emphasis id="959CEAC7FFD9167F78B7FE41232573FF" box="[465,582,475,496]" italics="true" pageId="18" pageNumber="34">Falsargutor</emphasis>
|
||
)
|
||
<emphasis id="959CEAC7FFD9167F7B32FE4123B473FF" box="[596,727,475,496]" italics="true" pageId="18" pageNumber="34">pseudopedius</emphasis>
|
||
</taxonomicName>
|
||
may argue against this point of view I am decided to share, as they link the
|
||
<taxonomicName id="60E84D56FFD9167F7875FD8B203C7029" authorityName="Dejean" authorityYear="1821" box="[275,351,529,550]" class="Insecta" family="Carabidae" genus="Argutor" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFD9167F7875FD8B203C7029" box="[275,351,529,550]" italics="true" pageId="18" pageNumber="34">Argutor</emphasis>
|
||
</taxonomicName>
|
||
lineage with other
|
||
<taxonomicName id="60E84D56FFD9167F7B41FD8B23C37029" ID-CoL="8KVYP" authorityName="Bonelli" authorityYear="1810" box="[551,672,529,550]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFD9167F7B41FD8B23C37029" box="[551,672,529,550]" italics="true" pageId="18" pageNumber="34">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
by filling some morphological gaps with themselves.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |