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<document id="587CF298E0FCC42E3C876F71A5A57AA8" ID-DOI="10.15298/rusentj.32.1.03" ID-ISSN="0132-8069" ID-Zenodo-Dep="10949096" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712198792598" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2023" docId="2F4187C3FFC0166179A0FEA125747143" docLanguage="en" docName="RussEntomolJ.32.1.16-39.pdf" docOrigin="Russian Entomological Journal 32 (1)" docSource="http://dx.doi.org/10.15298/rusentj.32.1.03" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.4:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="4" docTitle="Pterostichus (Vietoderus) laevibasis Fedorenko 2023, sp.n." docType="treatment" docVersion="4" lastPageNumber="28" masterDocId="D378FFBBFFCB166D7966FF9A2163720F" masterDocTitle="New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam" masterLastPageNumber="39" masterPageNumber="16" pageNumber="27" updateTime="1731592677940" updateUser="julia">
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<mods:title id="13807953CBD4C9784F7C44581C4B6505">New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam</mods:title>
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<mods:title id="97624C1ECB2D472BBED8773AA0DDD807">Russian Entomological Journal</mods:title>
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<mods:date id="E33EA1F4E86414779EBDE7BEB931A01C">2023</mods:date>
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<mods:number id="7B1E0C4352FBD759A3C9F271029334FF">2023-03-31</mods:number>
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<treatment id="2F4187C3FFC0166179A0FEA125747143" LSID="urn:lsid:plazi:treatment:2F4187C3FFC0166179A0FEA125747143" httpUri="http://treatment.plazi.org/id/2F4187C3FFC0166179A0FEA125747143" lastPageId="12" lastPageNumber="28" pageId="11" pageNumber="27">
<subSubSection id="EFF2655EFFC0166679A0FEA1239E735C" box="[198,765,315,339]" pageId="11" pageNumber="27" type="nomenclature">
<paragraph id="A75736D5FFC0166679A0FEA1239E735C" blockId="11.[189,775,315,593]" box="[198,765,315,339]" pageId="11" pageNumber="27">
<heading id="FC1F81B9FFC0166679A0FEA1239E735C" box="[198,765,315,339]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="27" reason="8">
<taxonomicName id="60E84D56FFC0166679A0FEA12320735C" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[198,579,315,339]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="laevibasis" subGenus="Vietoderus">
<emphasis id="959CEAC7FFC0166679A0FEA1202A735C" box="[198,329,315,339]" italics="true" pageId="11" pageNumber="27">Pterostichus</emphasis>
(
<emphasis id="959CEAC7FFC01666783FFEA120A9735C" box="[345,458,315,339]" italics="true" pageId="11" pageNumber="27">Vietoderus</emphasis>
)
<emphasis id="959CEAC7FFC0166678BCFEA12320735C" box="[474,579,315,339]" italics="true" pageId="11" pageNumber="27">laevibasis</emphasis>
</taxonomicName>
Fedorenko,
<emphasis id="959CEAC7FFC016667BAEFEA1239E735C" bold="true" box="[712,765,315,339]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC016667BAEFEA1239E735C" box="[712,765,315,339]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EFF2655EFFC016667997FEC323B0737E" box="[241,723,344,369]" pageId="11" pageNumber="27" type="description">
<paragraph id="A75736D5FFC016667997FEC323B0737E" blockId="11.[189,775,315,593]" box="[241,723,344,369]" pageId="11" pageNumber="27">
<figureCitation id="3FD32A50FFC016667997FEC3203D737E" box="[241,350,345,369]" captionStart="Figs 611" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[202,1388,618,1932]" captionTargetId="figure-261@3.[190,1398,609,1945]" captionTargetPageId="3" captionText="Figs 611. Pterostichus: 6, 9 — P. tonkinensis, ♂ from Phia Oac Mt; 7, 10 — P. austrinus sp.n., holotype; 8, 11 — P. laevibasis sp.n., paratype ♂; 68 — dorsal habitus; 911 — apices of elytra and abdomen, left aspect. Рис. 611. Pterostichus: 6, 9 — P. tonkinensis, ♂ с горы Phia Oac; 7, 10 — P. austrinus sp.n., голотип; 8, 11 — P. laevibasis sp.n., паратип ♂; 68 — габитус дорЗально; 911 — вершина надкрылий и брюшка слева." figureDoi="http://doi.org/10.5281/zenodo.10949100" httpUri="https://zenodo.org/record/10949100/files/figure.png" pageId="11" pageNumber="27">Figs 8, 11</figureCitation>
,
<figureCitation id="3FD32A50FFC01666780DFEC320D2737E" box="[363,433,345,369]" captionStart="Figs 2234" captionStartId="7.[226,263,1921,1940]" captionTargetBox="[190,1399,697,1904]" captionTargetId="figure-163@7.[189,1399,694,1904]" captionText="Figs 2234. Aedeagus: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 2829 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 2226 — median lobe; 2834 — right paramere, scale as for respective median lobe; 2225, 27 — ventrobasal aspect; 2934 — left lateral aspect; 26, 28 — dorsal aspect. Scale bars 1 mm. Рис. 2234. Эдеагус: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 2829 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 2226 — среднЯЯ долЯ; 2834 — праваЯ парамера в масштабе соответствуюЩей средней доли; 2225, 27 — вентробаЗально; 2934 — слева; 26, 28 — сверху. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949106" httpUri="https://zenodo.org/record/10949106/files/figure.png" pageId="11" pageNumber="27">27, 34</figureCitation>
, 3940, 49, 5860, 6970.
</paragraph>
</subSubSection>
<subSubSection id="EFF2655EFFC016667984FEE425E77368" pageId="11" pageNumber="27" type="materials_examined">
<paragraph id="A75736D5FFC016667984FEE423007036" blockId="11.[189,775,315,593]" pageId="11" pageNumber="27">
<materialsCitation id="17803C88FFC016667984FEE4203F73D6" collectingDate="2018-10-03" collectingDateMax="2018-10-11" collectingDateMin="2018-10-03" collectionCode="ZMMU" collectorName="D. Fedorenko'" elevation="1800" latitude="22.61389" location="Phia Oac Mt." longLatPrecision="21" longitude="105.86667" pageId="11" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang" typeStatus="holotype">
MATERIAL.
<typeStatus id="78538877FFC01666783BFEE420CE739E" box="[349,429,382,401]" pageId="11" pageNumber="27" type="holotype">Holotype</typeStatus>
<specimenCount id="B1EEFD5CFFC0166678D2FEE120AB739E" box="[436,456,379,401]" count="1" pageId="11" pageNumber="27" type="male"></specimenCount>
(
<collectionCode id="C1F9AE10FFC0166678B3FEE4237F739E" box="[469,540,382,401]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="11" pageNumber="27" type="Museum">ZMMU</collectionCode>
), labelled: N-Vietnam,
<quantity id="60109B30FFC016667B97FEE421BA73A6" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="27" unit="km" value="40.0">40 km</quantity>
W of/
<collectingRegion id="652CF837FFC01666784DFE0C20ED73A6" box="[299,398,406,425]" country="Vietnam" name="Cao Bang" pageId="11" pageNumber="27">Cao Bang</collectingRegion>
,
<location id="A237600EFFC0166678F8FE0C234773A6" LSID="urn:lsid:plazi:treatment:2F4187C3FFC0166179A0FEA125747143:A237600EFFC0166678F8FE0C234773A6" box="[414,548,406,425]" latitude="22.61389" longLatPrecision="21" longitude="105.86667" name="Phia Oac Mt." pageId="11" pageNumber="27" stateProvince="Cao Bang">Phia Oac Mt.</location>
/ 3-11.X./
<geoCoordinate id="C2DC5012FFC016667BF6FE0F226573A7" box="[656,774,405,424]" degrees="22" direction="north" minutes="36" orientation="latitude" pageId="11" pageNumber="27" precision="15" seconds="50" value="22.61389">22°36´50´´N</geoCoordinate>
<geoCoordinate id="C2DC5012FFC0166679DBFE37204573CF" box="[189,294,429,448]" degrees="105" direction="east" minutes="52" orientation="longitude" pageId="11" pageNumber="27" precision="15" seconds="0" value="105.86667">105°52´0´´E</geoCoordinate>
/ h~
<quantity id="60109B30FFC01666782DFE3420F373CE" box="[331,400,430,449]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="27" unit="m" value="1800.0">
<elevation id="2CC5D1E6FFC01666782DFE3420F373CE" box="[331,400,430,449]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="27" unit="m" value="1800.0">1800 m</elevation>
</quantity>
, deciduous/ ba[m]boo forest,
<date id="D3561015FFC016667BF2FE34239C73CE" box="[660,767,430,449]" pageId="11" pageNumber="27" value="2018-10-03" valueMax="2018-10-11" valueMin="2018-10-03">
<collectingDate id="C312E9FDFFC016667BF2FE34239C73CE" box="[660,767,430,449]" pageId="11" pageNumber="27" value="2018-10-03" valueMax="2018-10-11" valueMin="2018-10-03">3-11.X.2018</collectingDate>
</date>
/ leg.
<collectorName id="0A1D5303FFC0166679BAFE5C203F73D6" box="[220,348,454,473]" pageId="11" pageNumber="27">D.Fedorenko</collectorName>
</materialsCitation>
.
<materialsCitation id="17803C88FFC01666780DFE5C20D87006" collectingDate="2018-05-22" collectingDateMax="2018-06-06" collectingDateMin="2018-05-22" collectionCode="SIEE" collectorName="A. Abramov'" elevation="1675" latitude="22.6075" location="Phia Oac Mt." longLatPrecision="21" longitude="105.86667" pageId="11" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cao Bang" typeStatus="paratype">
<typeStatus id="78538877FFC01666780DFE5C20A273D6" box="[363,449,454,473]" pageId="11" pageNumber="27" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="C1F9AE10FFC0166678B5FE5C236473D6" box="[467,519,454,473]" pageId="11" pageNumber="27">SIEE</collectionCode>
):
<specimenCount id="B1EEFD5CFFC016667B7AFE59234B73D6" box="[540,552,451,473]" count="1" pageId="11" pageNumber="27" type="female"></specimenCount>
, same data, except for
<geoCoordinate id="C2DC5012FFC0166679DBFE47204973FF" box="[189,298,477,496]" degrees="22" direction="north" minutes="36" orientation="latitude" pageId="11" pageNumber="27" precision="15" seconds="25" value="22.606945">22°36´25´´N</geoCoordinate>
<geoCoordinate id="C2DC5012FFC016667855FE4720C473FF" box="[307,423,477,496]" degrees="105" direction="east" minutes="52" orientation="longitude" pageId="11" pageNumber="27" precision="15" seconds="08" value="105.86889">105°52´08´´E</geoCoordinate>
/ h~
<quantity id="60109B30FFC0166678ABFE47232673FE" box="[461,581,477,497]" metricMagnitude="3" metricUnit="m" metricValue="1.675" metricValueMax="1.7" metricValueMin="1.65" pageId="11" pageNumber="27" unit="m" value="1675.0" valueMax="1700.0" valueMin="1650.0">
<elevation id="2CC5D1E6FFC0166678ABFE47232673FE" box="[461,581,477,497]" metricMagnitude="3" metricUnit="m" metricValue="1.675" metricValueMax="1.7" metricValueMin="1.65" pageId="11" pageNumber="27" unit="m" value="1675.0" valueMax="1700.0" valueMin="1650.0">1650-1700 m</elevation>
</quantity>
,/ deciduous forest, 3- 11./ leg.D.
<bibRefCitation id="C3794B24FFC016667871FE6C20D77006" author="Fedorenko D. N." box="[279,436,502,521]" pageId="11" pageNumber="27" pagination="107 - 121" refId="ref19508" refString="Fedorenko D. N. 2018. Notes on Pterostichus subgenus Steropanus (Coleoptera: Carabidae: Pterostichini), with descriptions of new species // Russian Entomol. J. Vol. 27. No. 2. P. 107 - 121. doi: 10.15298 / rusentj. 27.2.01." type="journal article" year="2018">Fedorenko 2018</bibRefCitation>
;
</materialsCitation>
<materialsCitation id="17803C88FFC0166678A3FE69233C7036" collectingDate="2018-05-22" collectingDateMax="2018-06-06" collectingDateMin="2018-05-22" collectionCode="SIEE" collectorName="A. Abramov'" elevation="1675" latitude="22.6075" location="Phia Oac Mt." longLatPrecision="21" longitude="105.86667" pageId="11" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cao Bang" typeStatus="paratype">
<specimenCount id="B1EEFD5CFFC0166678A3FE6920BA7006" box="[453,473,499,521]" count="1" pageId="11" pageNumber="27" type="male"></specimenCount>
, same data, except for ‘…/
<collectingRegion id="652CF837FFC016667B82FE6C218C702E" country="Vietnam" name="Cao Bang" pageId="11" pageNumber="27">Cao Bang</collectingRegion>
,
<location id="A237600EFFC01666799BFD9420E2702E" LSID="urn:lsid:plazi:treatment:2F4187C3FFC0166179A0FEA125747143:A237600EFFC01666799BFD9420E2702E" box="[253,385,526,545]" latitude="22.6075" longLatPrecision="21" longitude="105.86667" name="Phia Oac Mt." pageId="11" pageNumber="27" stateProvince="Cao Bang">Phia Oac Mt.</location>
/ E-slope, h=
<quantity id="60109B30FFC016667B64FD9423E2702E" box="[514,641,526,545]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="11" pageNumber="27" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">
<elevation id="2CC5D1E6FFC016667B64FD9423E2702E" box="[514,641,526,545]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="11" pageNumber="27" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">1600-1800 m</elevation>
</quantity>
/
<geoCoordinate id="C2DC5012FFC016667BF5FD942265702E" box="[659,774,526,545]" degrees="22" direction="north" minutes="36" orientation="latitude" pageId="11" pageNumber="27" precision="15" seconds="27" value="22.6075">22°36´27´´N</geoCoordinate>
<geoCoordinate id="C2DC5012FFC0166679DBFDBF20457037" box="[189,294,549,568]" degrees="105" direction="east" minutes="52" orientation="longitude" pageId="11" pageNumber="27" precision="15" seconds="0" value="105.86667">105°52´0´´E</geoCoordinate>
/
<date id="D3561015FFC016667851FDBC20DC7036" box="[311,447,550,569]" pageId="11" pageNumber="27" value="2018-05-22" valueMax="2018-06-06" valueMin="2018-05-22">
<collectingDate id="C312E9FDFFC016667851FDBC20DC7036" box="[311,447,550,569]" pageId="11" pageNumber="27" value="2018-05-22" valueMax="2018-06-06" valueMin="2018-05-22">22.V-6.VI.2018</collectingDate>
</date>
/ leg.
<collectorName id="0A1D5303FFC016667888FDBC233C7036" box="[494,607,550,569]" pageId="11" pageNumber="27">A.Abramov</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="A75736D5FFC016667984FDA425E77368" blockId="11.[189,775,315,593]" lastBlockId="11.[812,1398,315,599]" pageId="11" pageNumber="27">
Aedeagus examined in the
<typeStatus id="78538877FFC0166678B2FDA4237C705E" box="[468,543,574,593]" pageId="11" pageNumber="27" type="holotype">holotype</typeStatus>
(integuments of aedeagus are soft following slightly teneral condition of the specimen); internal sac examined in the male
<typeStatus id="78538877FFC016667D55FECE25E37368" box="[1075,1152,340,359]" pageId="11" pageNumber="27" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="EFF2655EFFC016617A34FEF1250D70C8" lastPageId="12" lastPageNumber="28" pageId="11" pageNumber="27" type="description">
<paragraph id="A75736D5FFC016617A34FEF1206273FE" blockId="11.[812,1398,315,599]" lastBlockId="12.[189,776,315,2071]" lastPageId="12" lastPageNumber="28" pageId="11" pageNumber="27">
DESCRIPTION. With characters of the subgenus. BL 11.312.8 mm. Body (
<figureCitation id="3FD32A50FFC016667D73FE1C25307393" box="[1045,1107,390,412]" captionStart="Figs 611" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[202,1388,618,1932]" captionTargetId="figure-261@3.[190,1398,609,1945]" captionTargetPageId="3" captionText="Figs 611. Pterostichus: 6, 9 — P. tonkinensis, ♂ from Phia Oac Mt; 7, 10 — P. austrinus sp.n., holotype; 8, 11 — P. laevibasis sp.n., paratype ♂; 68 — dorsal habitus; 911 — apices of elytra and abdomen, left aspect. Рис. 611. Pterostichus: 6, 9 — P. tonkinensis, ♂ с горы Phia Oac; 7, 10 — P. austrinus sp.n., голотип; 8, 11 — P. laevibasis sp.n., паратип ♂; 68 — габитус дорЗально; 911 — вершина надкрылий и брюшка слева." figureDoi="http://doi.org/10.5281/zenodo.10949100" httpUri="https://zenodo.org/record/10949100/files/figure.png" pageId="11" pageNumber="27">Fig. 8</figureCitation>
) shiny black, pronotum and elytra slightly iridescent; antennae, palps,
<emphasis id="959CEAC7FFC016667DB6FE3B25BE73B9" box="[1232,1245,417,438]" italics="true" pageId="11" pageNumber="27">ti</emphasis>
1 and tarsi reddish brown, antennae toward apices increasingly pale, red (
<typeStatus id="78538877FFC016667A55FE4C22EA73E3" box="[819,905,470,492]" pageId="11" pageNumber="27" type="holotype">holotype</typeStatus>
and female
<typeStatus id="78538877FFC016667D63FE4C253B73E4" box="[1029,1112,470,491]" pageId="11" pageNumber="27" type="paratype">paratype</typeStatus>
slightly paler coloured, with tarsi red, tibiae reddish brown to red, and reflexed lateral margins of both pronotum and elytra reddish, which is due to somewhat teneral condition; left elytral intervals 14 with an occasional, preapical, red patch in the female). Microsculpture isodiametric, superficial yet distinct on head, very superficial on pronotum and elytra, consisting of almost indistinct transverse meshes on the former and very dense transverse lines on the latter; reflexed lateral margin of elytra with very coarse microsculpture, consisting of minute and very dense punctures (apparently formed by isodiametric microgranules fused).
</paragraph>
<caption id="F397665DFFC016667984F81B22117A1E" ID-DOI="http://doi.org/10.5281/zenodo.10949110" ID-Zenodo-Dep="10949110" httpUri="https://zenodo.org/record/10949110/files/figure.png" pageId="11" pageNumber="27" startId="11.[226,263,1921,1940]" targetBox="[188,1398,626,1889]" targetPageId="11" targetType="figure">
<paragraph id="A75736D5FFC016667984F81B255975CA" blockId="11.[189,1399,1921,2065]" pageId="11" pageNumber="27">
Figs 4351. Aedeagus with everted and inflated internal sac: 4345 —
<taxonomicName id="60E84D56FFC016667A36F81B2572759B" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[848,1041,1921,1940]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="glabellus" status="sp. nov.">
<emphasis id="959CEAC7FFC016667A36F81B2572759B" box="[848,1041,1921,1940]" italics="true" pageId="11" pageNumber="27">Pterostichus glabellus</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC016667D71F81B2521759B" bold="true" box="[1047,1090,1921,1940]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC016667D71F81B2521759B" box="[1047,1090,1921,1940]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 46 —
<taxonomicName id="60E84D56FFC016667DE0F818258D759B" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[1158,1262,1922,1940]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="austrinus" status="sp.n.">
<emphasis id="959CEAC7FFC016667DE0F818258D759B" box="[1158,1262,1922,1940]" italics="true" pageId="11" pageNumber="27">P. austrinus</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC016667D93F81B2443759B" bold="true" box="[1269,1312,1921,1940]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC016667D93F81B2443759B" box="[1269,1312,1921,1940]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 47 —
<taxonomicName id="60E84D56FFC016667C02F818207D75A3" authorityName="Fedorenko" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="959CEAC7FFC016667C02F818207D75A3" italics="true" pageId="11" pageNumber="27">P. tonkinensis</emphasis>
</taxonomicName>
; 48 —
<taxonomicName id="60E84D56FFC016667807F80020B775A3" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[353,468,1945,1964]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="subirideus" status="sp.n.">
<emphasis id="959CEAC7FFC016667807F80020B775A3" box="[353,468,1945,1964]" italics="true" pageId="11" pageNumber="27">P. subirideus</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC0166678BCF803236675A3" bold="true" box="[474,517,1945,1964]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC0166678BCF803236675A3" box="[474,517,1945,1964]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, dorsal aspect; 49 —
<taxonomicName id="60E84D56FFC016667BA3F800225775A3" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[709,820,1945,1964]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="laevibasis" status="sp.n.">
<emphasis id="959CEAC7FFC016667BA3F800225775A3" box="[709,820,1945,1964]" italics="true" pageId="11" pageNumber="27">P. laevibasis</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC016667A5DF803220675A3" bold="true" box="[827,869,1945,1964]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC016667A5DF803220675A3" box="[827,869,1945,1964]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 5051 —
<taxonomicName id="60E84D56FFC016667AACF800252775A3" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[970,1092,1945,1964]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="batdaiensis" status="sp.n.">
<emphasis id="959CEAC7FFC016667AACF800252775A3" box="[970,1092,1945,1964]" italics="true" pageId="11" pageNumber="27">P. batdaiensis</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC016667D2DF803251575A3" bold="true" box="[1099,1142,1945,1964]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC016667D2DF803251575A3" box="[1099,1142,1945,1964]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 43, 50 — left lateral aspect; 44 — apical aspect; 45, 51 — right lateral aspect; 4647, 49 — ventrobasal aspect. Scale bars 1 mm.
</paragraph>
<paragraph id="A75736D5FFC016667984F85422117A1E" blockId="11.[189,1399,1921,2065]" pageId="11" pageNumber="27">
Рис. 4351. Эдеагус с вывернутым и раЗдутым внутренним мешком: 4345 —
<taxonomicName id="60E84D56FFC016667ACCF857250975EF" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[938,1130,1997,2016]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="glabellus" status="sp. nov.">
<emphasis id="959CEAC7FFC016667ACCF857250975EF" box="[938,1130,1997,2016]" italics="true" pageId="11" pageNumber="27">Pterostichus glabellus</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC016667D09F85725F975EF" bold="true" box="[1135,1178,1997,2016]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC016667D09F85725F975EF" box="[1135,1178,1997,2016]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 46 —
<taxonomicName id="60E84D56FFC016667DBFF854242375EF" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[1241,1344,1998,2016]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="austrinus" status="sp.n.">
<emphasis id="959CEAC7FFC016667DBFF854242375EF" box="[1241,1344,1998,2016]" italics="true" pageId="11" pageNumber="27">P. austrinus</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC016667C23F857241375EF" bold="true" box="[1349,1392,1997,2016]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC016667C23F857241375EF" box="[1349,1392,1997,2016]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 47 —
<taxonomicName id="60E84D56FFC016667993F87C200C75F7" authorityName="Fedorenko" authorityYear="2023" box="[245,367,2021,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="959CEAC7FFC016667993F87C200C75F7" box="[245,367,2021,2040]" italics="true" pageId="11" pageNumber="27">P. tonkinensis</emphasis>
</taxonomicName>
; 48 —
<taxonomicName id="60E84D56FFC0166678D3F87C234B75F7" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[437,552,2021,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="subirideus" status="sp.n.">
<emphasis id="959CEAC7FFC0166678D3F87C234B75F7" box="[437,552,2021,2040]" italics="true" pageId="11" pageNumber="27">P. subirideus</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC016667B49F87F233975F7" bold="true" box="[559,602,2021,2040]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC016667B49F87F233975F7" box="[559,602,2021,2040]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, сверху; 49 —
<taxonomicName id="60E84D56FFC016667B8FF87C223A75F7" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[745,857,2021,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="laevibasis" status="sp.n.">
<emphasis id="959CEAC7FFC016667B8FF87C223A75F7" box="[745,857,2021,2040]" italics="true" pageId="11" pageNumber="27">P. laevibasis</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC016667A06F87F22E875F7" bold="true" box="[864,907,2021,2040]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC016667A06F87F22E875F7" box="[864,907,2021,2040]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 5051 —
<taxonomicName id="60E84D56FFC016667A97F87C250F75F7" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[1009,1132,2021,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="27" phylum="Arthropoda" rank="species" species="batdaiensis" status="sp.n.">
<emphasis id="959CEAC7FFC016667A97F87C250F75F7" box="[1009,1132,2021,2040]" italics="true" pageId="11" pageNumber="27">P. batdaiensis</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC016667D12F87F25FC75F7" bold="true" box="[1140,1183,2021,2040]" pageId="11" pageNumber="27">
<taxonomicNameLabel id="8EAF57BCFFC016667D12F87F25FC75F7" box="[1140,1183,2021,2040]" pageId="11" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 43, 50 — слева; 44 — апикально; 45, 51 — справа; 4647, 49 — вентробаЗально. Масштаб 1 мм.
</paragraph>
</caption>
<paragraph id="A75736D5FFC716617984FE6C204E709E" blockId="12.[189,776,315,2071]" pageId="12" pageNumber="28">
Body otherwise as for
<taxonomicName id="60E84D56FFC7166178BEFE6D22647003" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[472,775,502,524]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="28" phylum="Arthropoda" rank="species" species="austrinus" status="sp.n." subGenus="Neohaptoderus">
<emphasis id="959CEAC7FFC7166178BEFE6D208D7003" box="[472,494,503,524]" italics="true" pageId="12" pageNumber="28">P.</emphasis>
(
<emphasis id="959CEAC7FFC716617B64FE6C23F47004" box="[514,663,502,523]" italics="true" pageId="12" pageNumber="28">Neohaptoderus</emphasis>
)
<emphasis id="959CEAC7FFC716617BCAFE6D22647003" box="[684,775,503,524]" italics="true" pageId="12" pageNumber="28">austrinus</emphasis>
</taxonomicName>
<emphasis id="959CEAC7FFC7166179DBFD8B218E7029" bold="true" box="[189,237,529,550]" pageId="12" pageNumber="28">
<taxonomicNameLabel id="8EAF57BCFFC7166179DBFD8B218E7029" box="[189,237,529,550]" pageId="12" pageNumber="28" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
except the following characters: frons and vertex slightly flattened, neck constriction indistinct, almost so on sides; frontal sulci narrow, more impressed, and shorter, rather abruptly disappearing before the level of anterior supra-ocular seta.
</paragraph>
<paragraph id="A75736D5FFC716617984FD0C23AB76A8" blockId="12.[189,776,315,2071]" pageId="12" pageNumber="28">Pronotum subcircular and rather flat; sides strongly and evenly rounded, broadest just before middle. Base truncate, subsinuate at middle; basal angles very obtuse, blunt or rounded; two basolateral sulci on each side deep, straight, crenulate or with a few fine punctures at bottom, connected by a shallow lateral section of basal bead; inner sulci running parallel to each other on basal two fifths, outher ones about two thirds as long, slightly diverging apicad and slightly separated from lateral grooves; basolateral fovea smooth and flat, barely more convex anteriorly. Apex evenly sinuate between apical angles; these projecting, slightly acute to right, rather sharp or blunted at the very tips; apical bead almost complete, obliterate just medially. Lateral bead fine, lateral groove narrow at apex and at base, slightly broadened in between, as wide as lateral bead at middle, finely and densely punctate in apical three quarters. Median line nearly complete, crenulate, fine yet distinct. Basal and apical transverse impressions indistinct. Each side with a vague sublateral line, indistinct in apical half, very shallow behind and curved toward mid-length of inner basolateral sulcus.</paragraph>
<paragraph id="A75736D5FFC716617984FB3123A87439" blockId="12.[189,776,315,2071]" pageId="12" pageNumber="28">
Elytra oblong-oval, nearly parallel-sided in male, slightly more rounded on sides in female, more rounded toward humeri and toward apices; these rounded combined or slightly separate and blunted; preapical sinuation rather shallow, preapical plica traceable in lateral view. Base moderately wide, humeral tooth very minute and invisible in dorsal view; basal ridge straight and transverse, curved laterally to humerus, humeral angle obtuse. Striae deep, finely punctate, those 16 or 17 inside humeral angle; parascutellar striole very short (short, oblique and almost adjoining stria 1 on left elytron in female
<typeStatus id="78538877FFC71661780FFA2C20A277C4" box="[361,449,1462,1483]" pageId="12" pageNumber="28" type="paratype">paratype</typeStatus>
). Intervals convex, those 7, 5 and 3 merging preapically in succession; intervals 7 and 8 subequally wide. Discal seta d2 just medial, D2/EL 0.450.52 (0.50, n = 3×2). Stria 7 bisetose preapically, with posterior seta adjoining marginal groove (stria 8). USS: 619.
</paragraph>
<paragraph id="A75736D5FFC716617984F9A123C87554" blockId="12.[189,776,315,2071]" pageId="12" pageNumber="28">
Ventral side. Propleura finely punctate in inner half, mesepisterna in anterior half, metepisterna and sides of abdominal sternites IIIV moderately and unevenly punctate. Prosternum with indistinct median groove, prosternal process in dorsal view truncate apically, with slight apical bead, in lateral view subrectangular; inclination wide, flat, laterally edged and slightly beaded. Lateral-to-apical abdominal bead entire. Abdominal sternite VII with two apical setae and a large, subtriangular, median tubercle in male (
<figureCitation id="3FD32A50FFC716617B11F88B23DE7528" box="[631,701,1809,1831]" captionStart="Figs 611" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[202,1388,618,1932]" captionTargetId="figure-261@3.[190,1398,609,1945]" captionTargetPageId="3" captionText="Figs 611. Pterostichus: 6, 9 — P. tonkinensis, ♂ from Phia Oac Mt; 7, 10 — P. austrinus sp.n., holotype; 8, 11 — P. laevibasis sp.n., paratype ♂; 68 — dorsal habitus; 911 — apices of elytra and abdomen, left aspect. Рис. 611. Pterostichus: 6, 9 — P. tonkinensis, ♂ с горы Phia Oac; 7, 10 — P. austrinus sp.n., голотип; 8, 11 — P. laevibasis sp.n., паратип ♂; 68 — габитус дорЗально; 911 — вершина надкрылий и брюшка слева." figureDoi="http://doi.org/10.5281/zenodo.10949100" httpUri="https://zenodo.org/record/10949100/files/figure.png" pageId="12" pageNumber="28">Fig. 11</figureCitation>
), quadrisetose in female; apical setae in male and inner setae in female about 1/3 sternite length distant from apex.
</paragraph>
<paragraph id="A75736D5FFC716617984F8FB232C7578" blockId="12.[189,776,315,2071]" box="[226,591,1888,1911]" pageId="12" pageNumber="28">Legs: tarsomere 5 glabrous ventrally.</paragraph>
<paragraph id="A75736D5FFC716617984F8E124397363" blockId="12.[189,776,315,2071]" lastBlockId="12.[812,1399,315,844]" pageId="12" pageNumber="28">
Aedeagus (
<figureCitation id="3FD32A50FFC716617836F8E120A2759F" box="[336,449,1915,1937]" captionStart="Figs 2234" captionStartId="7.[226,263,1921,1940]" captionTargetBox="[190,1399,697,1904]" captionTargetId="figure-163@7.[189,1399,694,1904]" captionText="Figs 2234. Aedeagus: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 2829 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 2226 — median lobe; 2834 — right paramere, scale as for respective median lobe; 2225, 27 — ventrobasal aspect; 2934 — left lateral aspect; 26, 28 — dorsal aspect. Scale bars 1 mm. Рис. 2234. Эдеагус: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 2829 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 2226 — среднЯЯ долЯ; 2834 — праваЯ парамера в масштабе соответствуюЩей средней доли; 2225, 27 — вентробаЗально; 2934 — слева; 26, 28 — сверху. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949106" httpUri="https://zenodo.org/record/10949106/files/figure.png" pageId="12" pageNumber="28">Figs 27, 34</figureCitation>
, 3940, 49, 6970): median lobe in lateral view arcuate; apex in dorsal view fairly long, lamellate, apically rounded, with a small preapical tooth on right side. Right paramere short, triangular, with apex pointed, round or elliptic in cross-section. Everted and inflated internal sac in dorsal view bent to the left and then ventrad, inner curve with six preapical vesicles, which are hypertrophied and merged into a common, large, transverse bulb.
</paragraph>
<paragraph id="A75736D5FFC716617A34FEEB250D70C8" blockId="12.[812,1399,315,844]" pageId="12" pageNumber="28">
Female urite VIII (
<figureCitation id="3FD32A50FFC716617D6FFEEB251F7389" box="[1033,1148,368,391]" captionStart="Figs 5262" captionStartId="13.[226,263,1873,1892]" captionTargetBox="[193,1399,824,1838]" captionTargetId="figure-208@13.[189,1399,820,1839]" captionTargetPageId="13" captionText="Figs 5262. Abdominal urites VIIIIX in female: 5253, 61 — Pterostichus austrinus sp.n.; 5455 — P. glabellus sp.n.; 5657 — P. unisetosus sp.n.; 5860 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — tergite VIII; 53, 55, 57, 59 — sternite VIII; 60 — urite IX and reproductive tract, dorsal (a) and ventral (b) aspects; 6162 — tergite IX, ventral aspect; bc — bursa copulatrix; gsc — gonosubcoxite; LT — laterotergite; ov — common oviduct; sc — seminal canal. Scale bars 1 mm. Рис. 5262. Уриты VIIIIX брюшка самки: 5253, 61 — Pterostichus austrinus sp.n.; 5455 — P. glabellus sp.n.; 5657 — P. unisetosus sp.n.; 5860 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — тергит VIII; 53, 55, 57, 59 — стернит VIII; 60 — урит IX и репродуктивный тракт, дорЗально (a) и вентрально (b); 6162 — тергит IX, вентрально; bc — копулЯтивнаЯ сумка; gsc — гоносубкоксит; LT — латеротергит; ov — непарный Яйцевод; sc — семепровод. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949112" httpUri="https://zenodo.org/record/10949112/files/figure.png" pageId="12" pageNumber="28">Figs 5859</figureCitation>
): tergite and sternite wide (long), hemisternite rounded laterally and apically, evenly sclerotized, except for a small, rounded, latero-apical region, basolateral apodemes triangular, wide, slightly longer than wide at bases. Tergite IX (
<figureCitation id="3FD32A50FFC716617D4AFE41251173FE" box="[1068,1138,475,497]" captionStart="Figs 5262" captionStartId="13.[226,263,1873,1892]" captionTargetBox="[193,1399,824,1838]" captionTargetId="figure-208@13.[189,1399,820,1839]" captionTargetPageId="13" captionText="Figs 5262. Abdominal urites VIIIIX in female: 5253, 61 — Pterostichus austrinus sp.n.; 5455 — P. glabellus sp.n.; 5657 — P. unisetosus sp.n.; 5860 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — tergite VIII; 53, 55, 57, 59 — sternite VIII; 60 — urite IX and reproductive tract, dorsal (a) and ventral (b) aspects; 6162 — tergite IX, ventral aspect; bc — bursa copulatrix; gsc — gonosubcoxite; LT — laterotergite; ov — common oviduct; sc — seminal canal. Scale bars 1 mm. Рис. 5262. Уриты VIIIIX брюшка самки: 5253, 61 — Pterostichus austrinus sp.n.; 5455 — P. glabellus sp.n.; 5657 — P. unisetosus sp.n.; 5860 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — тергит VIII; 53, 55, 57, 59 — стернит VIII; 60 — урит IX и репродуктивный тракт, дорЗально (a) и вентрально (b); 6162 — тергит IX, вентрально; bc — копулЯтивнаЯ сумка; gsc — гоносубкоксит; LT — латеротергит; ov — непарный Яйцевод; sc — семепровод. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949112" httpUri="https://zenodo.org/record/10949112/files/figure.png" pageId="12" pageNumber="28">Fig. 60</figureCitation>
) more or less ground plan; laterotergite rather narrow rounded and setulose at apex; gonocoxite crescent, with apical part elongated, preapical nematiform seta, one inner and two outer ensiform setae in basal two fifths; gonosubcoxite with 23, very short, ensiform setae at latero-apical margin. Reproductive tract (
<figureCitation id="3FD32A50FFC716617C29FDFB222B709E" captionStart="Figs 5262" captionStartId="13.[226,263,1873,1892]" captionTargetBox="[193,1399,824,1838]" captionTargetId="figure-208@13.[189,1399,820,1839]" captionTargetPageId="13" captionText="Figs 5262. Abdominal urites VIIIIX in female: 5253, 61 — Pterostichus austrinus sp.n.; 5455 — P. glabellus sp.n.; 5657 — P. unisetosus sp.n.; 5860 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — tergite VIII; 53, 55, 57, 59 — sternite VIII; 60 — urite IX and reproductive tract, dorsal (a) and ventral (b) aspects; 6162 — tergite IX, ventral aspect; bc — bursa copulatrix; gsc — gonosubcoxite; LT — laterotergite; ov — common oviduct; sc — seminal canal. Scale bars 1 mm. Рис. 5262. Уриты VIIIIX брюшка самки: 5253, 61 — Pterostichus austrinus sp.n.; 5455 — P. glabellus sp.n.; 5657 — P. unisetosus sp.n.; 5860 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — тергит VIII; 53, 55, 57, 59 — стернит VIII; 60 — урит IX и репродуктивный тракт, дорЗально (a) и вентрально (b); 6162 — тергит IX, вентрально; bc — копулЯтивнаЯ сумка; gsc — гоносубкоксит; LT — латеротергит; ov — непарный Яйцевод; sc — семепровод. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949112" httpUri="https://zenodo.org/record/10949112/files/figure.png" pageId="12" pageNumber="28">Fig. 60</figureCitation>
): seminal canal at least moderately long (receptacle lost through dissection), with a well-developed basal sclerite and a bulbous structure near its base.
</paragraph>
</subSubSection>
<subSubSection id="EFF2655EFFC716617A34FD5122F07119" pageId="12" pageNumber="28" type="distribution">
<paragraph id="A75736D5FFC716617A34FD51240270EE" blockId="12.[812,1399,315,844]" box="[850,1377,715,737]" pageId="12" pageNumber="28">
DISTRIBUTION. Known from the
<typeStatus id="78538877FFC716617DD6FD5625B970EE" box="[1200,1242,716,737]" pageId="12" pageNumber="28">type</typeStatus>
locality only.
</paragraph>
<paragraph id="A75736D5FFC716617A34FD7C22F07119" blockId="12.[812,1399,315,844]" pageId="12" pageNumber="28">NAME. Noun that refers to the smooth base of the pronotum.</paragraph>
</subSubSection>
<subSubSection id="EFF2655EFFC716617A34FC8125747143" pageId="12" pageNumber="28" type="description">
<paragraph id="A75736D5FFC716617A34FC8125747143" blockId="12.[812,1399,315,844]" pageId="12" pageNumber="28">HABITATS AND HABITS. All the specimens were collected in cloudy forests.</paragraph>
</subSubSection>
</treatment>
</document>