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<document id="3C8017EA32F436FEA9E1BDDBB290A627" ID-DOI="10.5281/zenodo.5740174" ID-ISSN="0374-1036" ID-Zenodo-Dep="5740174" ID-ZooBank="157F415D-B3B2-4248-A24A-79AAB723D20B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638254161608" checkinUser="marcus" docAuthor="Fery, Hans &amp; Hájek, Jiří" docDate="2013" docId="03ECF1056C6CFF9AFE28EBCAFC6DFB71" docLanguage="en" docName="ActaEntMusNatPra.2013.53.2.601-607.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Hydroporus toubkal Fery &amp; Hájek 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="606" masterDocId="FFD5897D6C6DFF9FFF94EF22FFDDFFA2" masterDocTitle="Hydroporus toubkal sp. nov. from the High Atlas, Morocco, a new member of the H. longulus group (Coleoptera: Dytiscidae: Hydroporinae)" masterLastPageNumber="607" masterPageNumber="601" pageNumber="602" updateTime="1699435705148" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="C731B0F3F70F0AFA8D62EADC55C7A807" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B2CEC1012CC08C74D2F417B18B3641E2">
<mods:title id="90C87EDE62EE0B58D0DD6FC5EE88AFCB">Hydroporus toubkal sp. nov. from the High Atlas, Morocco, a new member of the H. longulus group (Coleoptera: Dytiscidae: Hydroporinae)</mods:title>
</mods:titleInfo>
<mods:name id="D6127944578EEEECE6F6E7A89F1F39EB" type="personal">
<mods:role id="D7BFC32E252A362AA55735300D2F3C88">
<mods:roleTerm id="482F106884398846BB42C1E72F1F23EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B5BCF90E9A11410367E0BBC638E7704">Fery, Hans</mods:namePart>
<mods:affiliation id="217D2C88F873AFDA3494B6B6853B9330">Räuschstrasse 73, D- 13509 Berlin, Germany; e-mail: hanfry @ aol. com</mods:affiliation>
</mods:name>
<mods:name id="0DA46FC7AD3576CF317D6FA66A35FA8E" type="personal">
<mods:role id="0BDEA01A614913E931B9050D040FFDC5">
<mods:roleTerm id="C119A5B0454805E11FDEC8B09B061936">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CE26E88395D6AD1D80961A02A4794E6">Hájek, Jiří</mods:namePart>
<mods:affiliation id="6B8DBDA7FAF84D7585D6B39AB7D0D5B5">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: jiri _ hajek @ nm. cz</mods:affiliation>
</mods:name>
<mods:typeOfResource id="65935F99776701212BBC660C9E94559D">text</mods:typeOfResource>
<mods:relatedItem id="2F197837B326D4ED64EFE23325E3680A" type="host">
<mods:titleInfo id="38E47EAA1DFD49B1B52F55086D2EC69B">
<mods:title id="9A4646215C3801C3041E5EFEA7B861C7">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="E9BF22C38594D4A13405A0BCDFD11842">
<mods:date id="504ED3B99AA6CBEB1E304169FD90C71A">2013</mods:date>
<mods:detail id="61C15C7979D7525BA12DE01D99479BE5" type="pubDate">
<mods:number id="8A806092ED4888A7D7AD1F99361AB182">2013-11-15</mods:number>
</mods:detail>
<mods:detail id="B18EE6AA832D877600CE2062DD6EC737" type="volume">
<mods:number id="03D1A080C26950CB1D0BCAC9B823513C">53</mods:number>
</mods:detail>
<mods:detail id="BBD3F705F594569FC516BF932ECA38F3" type="issue">
<mods:number id="1F8B22F2369DC77E4A5923404E446263">2</mods:number>
</mods:detail>
<mods:extent id="DBE78B6F9DB8456CC2EE3D23AA8F73B6" unit="page">
<mods:start id="5C207D9E765A46643D3B10D5BD946AC7">601</mods:start>
<mods:end id="B911113625512125F98EADA2CC41440F">607</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6321B778FB5CB4423259ED0EB05F5572">journal article</mods:classification>
<mods:identifier id="B771831B87C0A881ADFCBE8481C306BE" type="DOI">10.5281/zenodo.5740174</mods:identifier>
<mods:identifier id="475EB10F1EC08B190F2BEBD0FDE540E4" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="B7905B2C2FF65F579B6976FBB8FC442F" type="Zenodo-Dep">5740174</mods:identifier>
<mods:identifier id="C221872CB8F1D2ADF9264FA098199509" type="ZooBank">157F415D-B3B2-4248-A24A-79AAB723D20B</mods:identifier>
</mods:mods>
<treatment id="03ECF1056C6CFF9AFE28EBCAFC6DFB71" LSID="urn:lsid:plazi:treatment:03ECF1056C6CFF9AFE28EBCAFC6DFB71" httpUri="http://treatment.plazi.org/id/03ECF1056C6CFF9AFE28EBCAFC6DFB71" lastPageId="5" lastPageNumber="606" pageId="1" pageNumber="602">
<subSubSection id="C35F13986C6CFF9EFE28EBCAFD47FAB8" pageId="1" pageNumber="602" type="nomenclature">
<paragraph id="8BFA40136C6CFF9EFE28EBCAFD2BFAA3" blockId="1.[444,758,1256,1281]" box="[444,758,1256,1281]" pageId="1" pageNumber="602">
<heading id="D0B2F77F6C6CFF9EFE28EBCAFD2BFAA3" bold="true" box="[444,758,1256,1281]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="602" reason="3">
<emphasis id="B9319C016C6CFF9EFE28EBCAFD2BFAA3" bold="true" box="[444,758,1256,1281]" pageId="1" pageNumber="602">
<taxonomicName id="4C453B906C6CFF9EFE28EBCAFD47FAA3" ID-CoL="8LZBR" authority="Fery &amp; Hájek, 2013" authorityName="Fery &amp; Hájek" authorityYear="2013" box="[444,666,1256,1281]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="602" phylum="Arthropoda" rank="species" species="toubkal" status="sp. nov.">
<emphasis id="B9319C016C6CFF9EFE28EBCAFD47FAA3" bold="true" box="[444,666,1256,1281]" italics="true" pageId="1" pageNumber="602">Hydroporus toubkal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202217A6C6CFF9EFD34EBCBFD2BFAA3" box="[672,758,1257,1281]" pageId="1" pageNumber="602" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA40136C6CFF9EFD83EA25FD47FAB8" blockId="1.[113,1088,1287,1367]" box="[535,666,1287,1306]" pageId="1" pageNumber="602">(Figs 1–4, 8–9)</paragraph>
</subSubSection>
<subSubSection id="C35F13986C6CFF9EFFE5EA0BFED5FAF5" pageId="1" pageNumber="602" type="reference_group">
<paragraph id="8BFA40136C6CFF9EFFE5EA0BFED5FAF5" blockId="1.[113,1088,1287,1367]" pageId="1" pageNumber="602">
<treatmentCitationGroup id="AB55673D6C6CFF9EFFE5EA0BFED5FAF5" pageId="1" pageNumber="602">
?
<taxonomicName id="4C453B906C6CFF9EFF15EA08FE27FA9E" ID-CoL="6MJFM" authority="Mulsant &amp; Rey, 1861" authorityName="Mulsant &amp; Rey" authorityYear="1861" box="[129,506,1321,1341]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="602" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B9319C016C6CFF9EFF15EA08FEEBFA9E" box="[129,310,1322,1340]" italics="true" pageId="1" pageNumber="602">Hydroporus longulus</emphasis>
Mulsant &amp; Rey, 1861
</taxonomicName>
:
<treatmentCitation id="0AE466026C6CFF9EFD96EA08FD79FA9E" author="KOCHER L." box="[514,676,1321,1341]" page="18" pageId="1" pageNumber="602" year="1958">
<bibRefCitation id="EFD43DE26C6CFF9EFD96EA08FD5AFA9E" author="KOCHER L." box="[514,647,1321,1341]" pageId="1" pageNumber="602" pagination="1 - 244" refId="ref2847" refString="KOCHER L. 1958: Catalogue commente des coleopteres du Maroc. Fascicule II. Hydrocanthares, palpicornes, brachelytres. Travaux de l'Institut Scientifique Cherifien (Societe des Sciences Naturelles et Physiques du Maroc), Serie Zoologique 14: 1 - 244." type="journal article" year="1958">KOCHER (1958)</bibRefCitation>
: 18
</treatmentCitation>
(misidentification);
<treatmentCitation id="0AE466026C6CFF9EFCC0EA0BFBD5FA9E" author="GUIGNOT F." box="[852,1032,1321,1340]" page="395" pageId="1" pageNumber="602" year="1959">
<bibRefCitation id="EFD43DE26C6CFF9EFCC0EA0BFC02FA9E" author="GUIGNOT F." box="[852,991,1321,1340]" pageId="1" pageNumber="602" pagination="321 - 648" refId="ref2623" refString="GUIGNOT F. 1959: Revision des hydrocanthares d'Afrique (Coleoptera Dytiscoidea), deuxieme partie. Annales du Musee Royal du Congo Belge, Serie 8 vo (Sciences Zoologiques) 78: 321 - 648." type="journal article" year="1959">GUIGNOT (1959)</bibRefCitation>
: 395
</treatmentCitation>
(misidentification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35F13986C6CFF9BFFE5EA4CFD2DF9F1" lastPageId="4" lastPageNumber="605" pageId="1" pageNumber="602" type="materials_examined">
<paragraph id="8BFA40136C6CFF9EFFE5EA4CFE49FA39" blockId="1.[113,1089,1389,1622]" pageId="1" pageNumber="602">
<materialsCitation id="3B2D4A4E6C6CFF9EFFE5EA4CFBE0FA23" box="[113,1085,1389,1409]" collectorName="Refuge du Toubkal" country="Morocco" elevation="3100" location="Toubkal Mt." pageId="1" pageNumber="602" specimenCount="1" typeStatus="holotype">
<emphasis id="B9319C016C6CFF9EFFE5EA4CFF31FA23" bold="true" box="[113,236,1389,1409]" pageId="1" pageNumber="602">
<typeStatus id="54FEFEB16C6CFF9EFFE5EA4CFF43FA23" box="[113,158,1390,1409]" pageId="1" pageNumber="602">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F35200836C6CFF9EFF66EA4CFE9CFA23" box="[242,321,1390,1409]" name="Morocco" pageId="1" pageNumber="602">Morocco</collectingCountry>
, High Atlas, [Jebel]
<location id="8E9A16C86C6CFF9EFE61EA4CFDBFFA23" LSID="urn:lsid:plazi:treatment:03ECF1056C6CFF9AFE28EBCAFC6DFB71:8E9A16C86C6CFF9EFE61EA4CFDBFFA23" box="[501,610,1390,1409]" country="Morocco" name="Toubkal Mt." pageId="1" pageNumber="602">Toubkal Mt.</location>
, near
<collectorName id="26B025C56C6CFF9EFD03EA4CFCE0FA23" box="[663,829,1390,1409]" pageId="1" pageNumber="602">Refuge du Toubkal</collectorName>
“Les Mouflons”, ca.
<quantity id="4CBDEDF66C6CFF9EFC6DEA4CFBE0FA23" box="[1017,1085,1390,1409]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="602" unit="m" value="3100.0">
<elevation id="0068A7206C6CFF9EFC6DEA4CFBE0FA23" box="[1017,1085,1390,1409]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="602" unit="m" value="3100.0">3100 m</elevation>
</quantity>
</materialsCitation>
; coordinates ca.
<geoCoordinate id="EE7126D46C6CFF9EFF63EAAAFE9EFA39" box="[247,323,1416,1435]" degrees="31.067" direction="north" orientation="latitude" pageId="1" pageNumber="602" precision="55" value="31.067">31.067N</geoCoordinate>
,
<geoCoordinate id="EE7126D46C6CFF9EFEDFEAAAFE4DFA39" box="[331,400,1416,1435]" degrees="7.936" direction="west" orientation="longitude" pageId="1" pageNumber="602" precision="55" value="-7.936">7.936W</geoCoordinate>
.
</paragraph>
<paragraph id="8BFA40136C6CFF9EFFE5EA81FD5AFA49" blockId="1.[113,1089,1389,1622]" pageId="1" pageNumber="602">
<emphasis id="B9319C016C6CFF9EFFE5EA81FF2EFA14" bold="true" box="[113,243,1442,1462]" pageId="1" pageNumber="602">Type material.</emphasis>
<typeStatus id="54FEFEB16C6CFF9EFF62EA81FE95FA17" box="[246,328,1443,1462]" pageId="1" pageNumber="602" type="holotype">HOLOTYPE</typeStatus>
: J: ‘
<collectingCountry id="F35200836C6CFF9EFEFAEA81FE00FA14" box="[366,477,1443,1462]" name="Morocco" pageId="1" pageNumber="602">MOROCCO</collectingCountry>
/ Toubkal
<quantity id="4CBDEDF66C6CFF9EFDA6EA81FD43FA14" box="[562,670,1443,1462]" metricMagnitude="3" metricUnit="m" metricValue="3.0" metricValueMax="3.4" metricValueMin="2.6" pageId="1" pageNumber="602" unit="m" value="3000.0" valueMax="3400.0" valueMin="2600.0">2600-3400m</quantity>
/
<date id="FFFB66D36C6CFF9EFD3FEA81FD29FA14" box="[683,756,1443,1462]" pageId="1" pageNumber="602" value="2008-05-05">5.5.2008</date>
/ lgt.Orszulik [printed]’, ‘
<emphasis id="B9319C016C6CFF9EFC5DEA80FB9CFA14" bold="true" box="[969,1089,1442,1462]" pageId="1" pageNumber="602">
<typeStatus id="54FEFEB16C6CFF9EFC5DEA80FB9CFA14" box="[969,1089,1442,1462]" pageId="1" pageNumber="602" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
/
<emphasis id="B9319C016C6CFF9EFFE9EA9DFE85FA73" box="[125,344,1471,1489]" italics="true" pageId="1" pageNumber="602">
<taxonomicName id="4C453B906C6CFF9EFFE9EA9DFED5FA73" authorityName="Clairville" authorityYear="1806" box="[125,264,1471,1489]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="602" phylum="Arthropoda" rank="genus">HYDROPORUS</taxonomicName>
/
<taxonomicName id="4C453B906C6CFF9EFE8CEA9DFE85FA73" authority="Fery &amp; Hájek, 2013" authorityName="Fery &amp; Hájek" authorityYear="2013" box="[280,344,1471,1489]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="602" phylum="Arthropoda" rank="species" species="toubkal" status="sp. nov.">toubkal</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A202217A6C6CFF9EFEC9EA9CFE7DFA73" box="[349,416,1470,1489]" pageId="1" pageNumber="602" rank="species">sp. nov.</taxonomicNameLabel>
/ H. Fery &amp; J. Hájek des. 2013 [red label, printed]’ (
<collectionCode id="ED54D8D66C6CFF9EFCCBEA9CFC7DFA73" box="[863,928,1470,1489]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="602">NMPC</collectionCode>
).
<typeStatus id="54FEFEB16C6CFF9EFC3BEA9CFBDBFA6D" box="[943,1030,1470,1489]" pageId="1" pageNumber="602" type="paratype">PARATYPES</typeStatus>
: 2 JJ
<specimenCount id="9D438B9A6C6CFF9EFFE5EAFAFF7DFA49" box="[113,160,1490,1515]" pageId="1" pageNumber="602" type="female">4 ♀♀</specimenCount>
, same label data as the
<typeStatus id="54FEFEB16C6CFF9EFEFFEAFAFE6BFA49" box="[363,438,1496,1515]" pageId="1" pageNumber="602" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED54D8D66C6CFF9EFE54EAFAFE36FA49" box="[448,491,1496,1515]" pageId="1" pageNumber="602">CHF</collectionCode>
,
<collectionCode id="ED54D8D66C6CFF9EFE60EAFAFDE9FA49" box="[500,564,1496,1515]" pageId="1" pageNumber="602">KOFM</collectionCode>
,
<collectionCode id="ED54D8D66C6CFF9EFDA8EAFAFDA1FA49" box="[572,636,1496,1515]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="602">NMPC</collectionCode>
).
</paragraph>
<paragraph id="8BFA40136C6CFF9EFFE5EAD0FD02F9F4" blockId="1.[113,1089,1389,1622]" pageId="1" pageNumber="602">
<materialsCitation id="3B2D4A4E6C6CFF9EFFE5EAD0FEA0F983" collectingDate="1985-08-09" collectorName="H. Fery" country="Morocco" elevation="2100" latitude="31.23" location="Haute Atlas" longLatPrecision="731" longitude="-7.81" pageId="1" pageNumber="602" specimenCount="1" specimenCount-female="1">
<emphasis id="B9319C016C6CFF9EFFE5EAD0FE5BF9A4" bold="true" box="[113,390,1522,1542]" pageId="1" pageNumber="602">Additional material examined.</emphasis>
<specimenCount id="9D438B9A6C6CFF9EFE1EEAD1FE74F9A4" box="[394,425,1517,1542]" pageId="1" pageNumber="602" type="female">1 ♀</specimenCount>
,
<collectingCountry id="F35200836C6CFF9EFE26EAD1FDDCF9A4" box="[434,513,1523,1542]" name="Morocco" pageId="1" pageNumber="602">Morocco</collectingCountry>
,
<location id="8E9A16C86C6CFF9EFD9DEAD1FDB2F9A4" LSID="urn:lsid:plazi:treatment:03ECF1056C6CFF9AFE28EBCAFC6DFB71:8E9A16C86C6CFF9EFD9DEAD1FDB2F9A4" box="[521,623,1523,1542]" country="Morocco" latitude="31.23" longLatPrecision="731" longitude="-7.81" name="Haute Atlas" pageId="1" pageNumber="602">Haute Atlas</location>
,
<locationDeviation id="9A4859316C6CFF9EFDE2EAD1FCE8F9A4" box="[630,821,1523,1542]" location="Oukaimeden" pageId="1" pageNumber="602">
<quantity id="4CBDEDF66C6CFF9EFDE2EAD1FD7DF9A4" box="[630,672,1523,1542]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="602" unit="km" value="4.0">4 km</quantity>
NE Oukaïmeden
</locationDeviation>
, ca.
<geoCoordinate id="EE7126D46C6CFF9EFCCEEAD1FC44F9A4" box="[858,921,1523,1542]" degrees="31.23" direction="north" orientation="latitude" pageId="1" pageNumber="602" precision="555" value="31.23">31.23N</geoCoordinate>
<geoCoordinate id="EE7126D46C6CFF9EFC09EAD1FC0AF9A4" box="[925,983,1523,1542]" degrees="7.81" direction="west" orientation="longitude" pageId="1" pageNumber="602" precision="555" value="-7.81">7.81W</geoCoordinate>
, ca.
<quantity id="4CBDEDF66C6CFF9EFC68EAD1FBE3F9A4" box="[1020,1086,1523,1542]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="602" unit="m" value="2100.0">
<elevation id="0068A7206C6CFF9EFC68EAD1FBE3F9A4" box="[1020,1086,1523,1542]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="602" unit="m" value="2100.0">2100 m</elevation>
</quantity>
, brook,
<date id="FFFB66D36C6CFF9EFF39E92CFED1F983" box="[173,268,1550,1569]" pageId="1" pageNumber="602" value="1985-08-09">
<collectingDate id="EFBF9F3B6C6CFF9EFF39E92CFED1F983" box="[173,268,1550,1569]" pageId="1" pageNumber="602" value="1985-08-09">9.viii.1985</collectingDate>
</date>
,
<collectorName id="26B025C56C6CFF9EFE80E92CFE88F983" box="[276,341,1550,1569]" pageId="1" pageNumber="602">H. Fery</collectorName>
leg.
</materialsCitation>
; so far the specimen has been determined as “
<taxonomicName id="4C453B906C6CFF9EFC95E92DFCB9F983" authorityName="Mulsant &amp; Rey" authorityYear="1861" box="[769,868,1551,1569]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="602" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B9319C016C6CFF9EFC95E92DFCB9F983" box="[769,868,1551,1569]" italics="true" pageId="1" pageNumber="602">H. longulus</emphasis>
</taxonomicName>
?”;
<materialsCitation id="3B2D4A4E6C6CFF9EFC13E92CFD06F9F4" country="Morocco" location="Marrakesh. As" pageId="1" pageNumber="602" specimenCount="1">
the locality is situated ca.
<quantity id="4CBDEDF66C6CFF9EFF1BE90AFF19F999" box="[143,196,1576,1595]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="602" unit="km" value="15.0">15 km</quantity>
NNE of the locus typicus of the new species and ca.
<quantity id="4CBDEDF66C6CFF9EFD1DE90AFD63F999" box="[649,702,1576,1595]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="602" unit="km" value="50.0">50 km</quantity>
SSE
<location id="8E9A16C86C6CFF9EFD79E90AFCB7F99E" LSID="urn:lsid:plazi:treatment:03ECF1056C6CFF9AFE28EBCAFC6DFB71:8E9A16C86C6CFF9EFD79E90AFCB7F99E" box="[749,874,1576,1596]" country="Morocco" name="Marrakesh. As" pageId="1" pageNumber="602">Marrakesh.As</location>
single females cannot be determined with certainty, we refrain from including it in the
<typeStatus id="54FEFEB16C6CFF9EFDE9E961FD7FF9F4" box="[637,674,1603,1622]" pageId="1" pageNumber="602">type</typeStatus>
series
</materialsCitation>
.
</paragraph>
<caption id="DF3A109B6C6EFF9CFFE5EAF7FE62F9F0" ID-DOI="http://doi.org/10.5281/zenodo.5740178" ID-Zenodo-Dep="5740178" httpUri="https://zenodo.org/record/5740178/files/figure.png" pageId="3" pageNumber="604" startId="3.[113,150,1493,1512]" targetBox="[120,1077,240,1439]" targetPageId="3">
<paragraph id="8BFA40136C6EFF9CFFE5EAF7FE62F9F0" blockId="3.[113,1089,1493,1619]" pageId="3" pageNumber="604">
Figs 2–9. 2–4, 8–9 –
<taxonomicName id="4C453B906C6EFF9CFEBEEAF4FE0BFA4A" authority="Fery &amp; Hájek, 2013" authorityName="Fery &amp; Hájek" authorityYear="2013" box="[298,470,1494,1512]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="604" phylum="Arthropoda" rank="species" species="toubkal" status="sp. nov.">
<emphasis id="B9319C016C6EFF9CFEBEEAF4FE0BFA4A" box="[298,470,1494,1512]" italics="true" pageId="3" pageNumber="604">Hydroporus toubkal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202217A6C6EFF9CFE48EAF7FDFEFA4A" box="[476,547,1493,1512]" pageId="3" pageNumber="604" rank="species">sp. nov.</taxonomicNameLabel>
: 2 – median lobe of aedeagus in ventral view; 3 – median lobe in lateral view; 4 – left paramere; 8 – gonocoxosternum; 9 – gonocoxae. 5 –
<taxonomicName id="4C453B906C6EFF9CFC80EAD2FBE3F9A0" authority="Mulsant &amp; Rey, 1861" authorityName="Mulsant &amp; Rey" authorityYear="1861" box="[788,1086,1519,1539]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="604" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B9319C016C6EFF9CFC80EAD2FCA5F9A0" box="[788,888,1520,1538]" italics="true" pageId="3" pageNumber="604">H. longulus</emphasis>
Mulsant &amp; Rey, 1861
</taxonomicName>
, median lobe of aedeagus in lateral view (adapted from
<bibRefCitation id="EFD43DE26C6EFF9CFDC7E928FD6DF9BF" author="FERY H." box="[595,688,1546,1565]" pageId="3" pageNumber="604" pagination="217 - 269" refId="ref2474" refString="FERY H. 1999: Revision of a part of the memnonius-group of Hydroporus Clairville, 1806 (Insecta: Coleoptera: Dytiscidae) with the description of nine new taxa, and notes on other species of the genus. Annalen des Naturhistorischen Museums in Wien 101 B: 217 - 269." type="journal article" year="1999">FERY 1999</bibRefCitation>
). 6 –
<taxonomicName id="4C453B906C6EFF9CFD76E929FC10F9BF" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[738,973,1546,1565]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="604" phylum="Arthropoda" rank="species" species="nevadensis">
<emphasis id="B9319C016C6EFF9CFD76E929FC81F9BF" box="[738,860,1547,1565]" italics="true" pageId="3" pageNumber="604">H. nevadensis</emphasis>
Sharp, 1882
</taxonomicName>
, median lobe of aedeagus in lateral view (adapted from
<bibRefCitation id="EFD43DE26C6EFF9CFE4EE907FDEBF99A" author="FERY H." box="[474,566,1573,1592]" pageId="3" pageNumber="604" pagination="217 - 269" refId="ref2474" refString="FERY H. 1999: Revision of a part of the memnonius-group of Hydroporus Clairville, 1806 (Insecta: Coleoptera: Dytiscidae) with the description of nine new taxa, and notes on other species of the genus. Annalen des Naturhistorischen Museums in Wien 101 B: 217 - 269." type="journal article" year="1999">FERY 1999</bibRefCitation>
). 7 –
<taxonomicName id="4C453B906C6EFF9CFDF1E904FC93F99A" authority="Zaitzev, 1927" authorityName="Zaitzev" authorityYear="1927" box="[613,846,1573,1592]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="604" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B9319C016C6EFF9CFDF1E904FD0FF99A" box="[613,722,1574,1592]" italics="true" pageId="3" pageNumber="604">H. jacobsoni</emphasis>
Zaitzev, 1927
</taxonomicName>
, median lobe of aedeagus in lateral view (adapted from FERY 2009).
</paragraph>
</caption>
<paragraph id="8BFA40136C69FF9BFF03EFF9FEC7FED1" blockId="4.[151,1127,219,1619]" pageId="4" pageNumber="605">diameter. Discal puncture lines not very distinct but perceptible, marked by somewhat denser normal punctures and a few interspersed coarser punctures; sutural puncture line lacking. Margin of elytra slightly ascending towards humeral angle in lateral view. Lateral elytral beading distinct, narrower than pronotal beading. Epipleuron visible to humeral angle in lateral view.</paragraph>
<paragraph id="8BFA40136C69FF9BFF21EE5EFCABFD91" blockId="4.[151,1127,219,1619]" pageId="4" pageNumber="605">Legs. Protibia rather strongly broadened distally. Pro- and mesotarsomeres I–III slightly dilated and with sucking setae, tarsomeres I with small oval sucker discs in addition to sucking setae. Protarsal claws short, more or less of same length, proximally broadened; posterior (outer) claw strongly curved near base, almost evenly curved over rest of length; anterior (inner) claw more strongly curved near base and almost straight in distal two thirds; mesotarsal claws of same length and evenly curved, longer than protarsal claws.</paragraph>
<paragraph id="8BFA40136C69FF9BFF21ED1EFC7DFCD1" blockId="4.[151,1127,219,1619]" pageId="4" pageNumber="605">
Venter with most parts weakly microreticulate; gula, prosternal process, medial part of metaventrite and metacoxal processes smooth; genae smooth close to gula, laterally distinctly reticulated. Meshes regularly polygonal on genae, sides of metaventrite, metacoxal plates, epipleura, and last (
<collectionCode id="ED54D8D66C69FF9BFEF6EDBEFE5DFD16" box="[354,384,668,692]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4" pageNumber="605">VI</collectionCode>
) abdominal ventrite; meshes on abdominal ventrites I–V very elongate, on sides more or less longitudinally oriented, in middle transverse. Head posterior to eyes with distinct crease, near mouthparts with several distinct wrinkles. Gula on sides with a few coarse punctures, genae impunctate. Punctation on sides of metaventrite, on metacoxal plates, and on sides of abdominal ventrites I–II rather coarse; on epipleura, centre of metaventrite, metacoxal processes, and rest of abdominal ventrites less coarse. More or less all punctures with a seta.
<collectingRegion id="49818EF16C69FF9BFE83EC7EFE83FCD6" box="[279,350,860,884]" country="France" name="Centre" pageId="4" pageNumber="605">Centre</collectingRegion>
of abdominal ventrites III–V with a tuft of long setae.
</paragraph>
<paragraph id="8BFA40136C69FF9BFF21EC59FE62FB91" blockId="4.[151,1127,219,1619]" pageId="4" pageNumber="605">Prosternal process lanceolate, very elongate; longitudinal carina narrow, more or less tectiform in cross-section, but ridge rounded; sides broadened and margin provided with very coarse punctures and long setae. Process reaching between mesocoxae and onto furrow of anteromedial metaventral process; thus, mesocoxae not contiguous. Base of prosternum strongly impressed; file broad, smooth, transverse ridges very weak; between procoxae with one strong transverse ridge.</paragraph>
<paragraph id="8BFA40136C69FF9BFF21EB19FD66FB31" blockId="4.[151,1127,219,1619]" pageId="4" pageNumber="605">Posterior margins of metacoxal processes medially protruded backwards, laterally sinuate; metacoxal lines diverging anteriorly, not reaching posterior margin of metaventrite; intralinear space posteriorly with numerous long golden setae.</paragraph>
<paragraph id="8BFA40136C69FF9BFF21EBB8FE18FAB6" blockId="4.[151,1127,219,1619]" pageId="4" pageNumber="605">
<emphasis id="B9319C016C69FF9BFF21EBB8FEFEFB16" bold="true" box="[181,291,1178,1204]" pageId="4" pageNumber="605">
<emphasis id="B9319C016C69FF9BFF21EBB8FEC6FB11" bold="true" box="[181,283,1178,1203]" italics="true" pageId="4" pageNumber="605">Genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus asymmetric; in ventral view only slightly convergent in basal two thirds, more evenly tapering to apex in apical third, apex very shortly rounded (
<figureCitation id="137E5C966C69FF9BFF0BEBF9FF3FFB56" box="[159,226,1243,1268]" captionStart="Figs 2–9. 2–4, 8–9" captionStartId="3.[113,150,1493,1512]" captionTargetBox="[120,1077,240,1439]" captionTargetId="figure-15@3.[113,1088,219,1454]" captionTargetPageId="3" captionText="Figs 2–9. 2–4, 8–9 – Hydroporus toubkal sp. nov.: 2 – median lobe of aedeagus in ventral view; 3 – median lobe in lateral view; 4 – left paramere; 8 – gonocoxosternum; 9 – gonocoxae. 5 – H. longulus Mulsant &amp; Rey, 1861, median lobe of aedeagus in lateral view (adapted from FERY 1999). 6 – H. nevadensis Sharp, 1882, median lobe of aedeagus in lateral view (adapted from FERY 1999). 7 – H. jacobsoni Zaitzev, 1927, median lobe of aedeagus in lateral view (adapted from FERY 2009)." figureDoi="http://doi.org/10.5281/zenodo.5740178" httpUri="https://zenodo.org/record/5740178/files/figure.png" pageId="4" pageNumber="605">Fig. 2</figureCitation>
); in lateral view distinctly narrowed in apical third to pointed apex (
<figureCitation id="137E5C966C69FF9BFC50EBF9FBDBFB56" box="[964,1030,1243,1268]" captionStart="Figs 2–9. 2–4, 8–9" captionStartId="3.[113,150,1493,1512]" captionTargetBox="[120,1077,240,1439]" captionTargetId="figure-15@3.[113,1088,219,1454]" captionTargetPageId="3" captionText="Figs 2–9. 2–4, 8–9 – Hydroporus toubkal sp. nov.: 2 – median lobe of aedeagus in ventral view; 3 – median lobe in lateral view; 4 – left paramere; 8 – gonocoxosternum; 9 – gonocoxae. 5 – H. longulus Mulsant &amp; Rey, 1861, median lobe of aedeagus in lateral view (adapted from FERY 1999). 6 – H. nevadensis Sharp, 1882, median lobe of aedeagus in lateral view (adapted from FERY 1999). 7 – H. jacobsoni Zaitzev, 1927, median lobe of aedeagus in lateral view (adapted from FERY 2009)." figureDoi="http://doi.org/10.5281/zenodo.5740178" httpUri="https://zenodo.org/record/5740178/files/figure.png" pageId="4" pageNumber="605">Fig. 3</figureCitation>
). Lateral lobe (paramere) as in
<figureCitation id="137E5C966C69FF9BFE14EBD9FE1CFAB6" box="[384,449,1275,1300]" captionStart="Figs 2–9. 2–4, 8–9" captionStartId="3.[113,150,1493,1512]" captionTargetBox="[120,1077,240,1439]" captionTargetId="figure-15@3.[113,1088,219,1454]" captionTargetPageId="3" captionText="Figs 2–9. 2–4, 8–9 – Hydroporus toubkal sp. nov.: 2 – median lobe of aedeagus in ventral view; 3 – median lobe in lateral view; 4 – left paramere; 8 – gonocoxosternum; 9 – gonocoxae. 5 – H. longulus Mulsant &amp; Rey, 1861, median lobe of aedeagus in lateral view (adapted from FERY 1999). 6 – H. nevadensis Sharp, 1882, median lobe of aedeagus in lateral view (adapted from FERY 1999). 7 – H. jacobsoni Zaitzev, 1927, median lobe of aedeagus in lateral view (adapted from FERY 2009)." figureDoi="http://doi.org/10.5281/zenodo.5740178" httpUri="https://zenodo.org/record/5740178/files/figure.png" pageId="4" pageNumber="605">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="8BFA40136C69FF9BFF21EA38FCCBFAD6" blockId="4.[151,1127,219,1619]" pageId="4" pageNumber="605">
<emphasis id="B9319C016C69FF9BFF21EA38FED2FA96" bold="true" box="[181,271,1306,1332]" pageId="4" pageNumber="605">
<emphasis id="B9319C016C69FF9BFF21EA38FED5FA91" bold="true" box="[181,264,1306,1331]" italics="true" pageId="4" pageNumber="605">Female</emphasis>
.
</emphasis>
Females without conspicuous external differences to males except slightly narrower pro- and especially mesotarsomeres, lack of sucker discs, and evenly curved protarsal claws. Gonocoxosternum and gonocoxae as in
<figureCitation id="137E5C966C69FF9BFD19EA79FCCFFAD6" box="[653,786,1371,1396]" captionStart="Figs 2–9. 2–4, 8–9" captionStartId="3.[113,150,1493,1512]" captionTargetBox="[120,1077,240,1439]" captionTargetId="figure-15@3.[113,1088,219,1454]" captionTargetPageId="3" captionText="Figs 2–9. 2–4, 8–9 – Hydroporus toubkal sp. nov.: 2 – median lobe of aedeagus in ventral view; 3 – median lobe in lateral view; 4 – left paramere; 8 – gonocoxosternum; 9 – gonocoxae. 5 – H. longulus Mulsant &amp; Rey, 1861, median lobe of aedeagus in lateral view (adapted from FERY 1999). 6 – H. nevadensis Sharp, 1882, median lobe of aedeagus in lateral view (adapted from FERY 1999). 7 – H. jacobsoni Zaitzev, 1927, median lobe of aedeagus in lateral view (adapted from FERY 2009)." figureDoi="http://doi.org/10.5281/zenodo.5740178" httpUri="https://zenodo.org/record/5740178/files/figure.png" pageId="4" pageNumber="605">Figs 8 and 9</figureCitation>
.
</paragraph>
<paragraph id="8BFA40136C69FF9BFF21EA58FD2DF9F1" blockId="4.[151,1127,219,1619]" pageId="4" pageNumber="605">
<emphasis id="B9319C016C69FF9BFF21EA58FEF0FA31" bold="true" box="[181,301,1402,1427]" italics="true" pageId="4" pageNumber="605">Variability.</emphasis>
Specimens of the type series vary in the extent of the brownish colouration of the lateral parts of elytra – in
<specimenCount id="9D438B9A6C69FF9BFE60EABEFD4FFA11" box="[500,658,1435,1459]" pageId="4" pageNumber="605" type="generic">two specimens</specimenCount>
the brown colour shines through the lateral fourth of elytra. The antennomeres can be almost unicolorous. In some
<typeStatus id="54FEFEB16C69FF9BFCE1EA9EFC00FA76" box="[885,989,1468,1492]" pageId="4" pageNumber="605" type="paratype">paratypes</typeStatus>
, the puncture line along the anterior margin of the pronotum is not really interrupted in the middle and the setation on the pronotal disc is indistinct, probably worn out. One male
<typeStatus id="54FEFEB16C69FF9BFC50EADEFBFDF9B6" box="[964,1056,1532,1556]" pageId="4" pageNumber="605" type="paratype">paratype</typeStatus>
varies slightly also in the shape of the male genitalia: the apical part of the median lobe is slightly shorter and in ventral view broader than in the
<typeStatus id="54FEFEB16C69FF9BFD1FE919FD31F9F1" box="[651,748,1595,1619]" pageId="4" pageNumber="605" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F13986C68FF9AFF04EFFEFE2BFEB6" pageId="5" pageNumber="606" type="description">
<paragraph id="8BFA40136C68FF9AFF04EFFEFE2BFEB6" blockId="5.[113,1089,219,1236]" pageId="5" pageNumber="606">
<emphasis id="B9319C016C68FF9AFF04EFFEFEE5FF51" bold="true" box="[144,312,220,243]" italics="true" pageId="5" pageNumber="606">Measurements.</emphasis>
TL
<quantity id="4CBDEDF66C68FF9AFEF5EFFEFE02FF51" box="[353,479,220,244]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="3.9" metricValueMin="3.7" pageId="5" pageNumber="606" unit="mm" value="3.8" valueMax="3.9" valueMin="3.7">3.7–3.9 mm</quantity>
(
<typeStatus id="54FEFEB16C68FF9AFE78EFF9FD96FF51" box="[492,587,219,243]" pageId="5" pageNumber="606" type="holotype">holotype</typeStatus>
<quantity id="4CBDEDF66C68FF9AFDC4EFFEFD78FF56" box="[592,677,220,244]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="606" unit="mm" value="3.8">3.8 mm</quantity>
); TL-h
<quantity id="4CBDEDF66C68FF9AFD65EFF9FCB2FF51" box="[753,879,219,243]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.6" metricValueMin="3.2" pageId="5" pageNumber="606" unit="mm" value="3.4" valueMax="3.6" valueMin="3.2">3.2–3.6 mm</quantity>
(
<typeStatus id="54FEFEB16C68FF9AFCE8EFF9FC06FF51" box="[892,987,219,243]" pageId="5" pageNumber="606" type="holotype">holotype</typeStatus>
<quantity id="4CBDEDF66C68FF9AFC74EFFEFBE8FF56" box="[992,1077,220,244]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="606" unit="mm" value="3.4">3.4 mm</quantity>
); TW 1.8–2.0 mm (
<typeStatus id="54FEFEB16C68FF9AFEBBEFD9FE53FEB1" box="[303,398,251,275]" pageId="5" pageNumber="606" type="holotype">holotype</typeStatus>
<quantity id="4CBDEDF66C68FF9AFE01EFDEFE36FEB6" box="[405,491,252,276]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="606" unit="mm" value="1.9">1.9 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F13986C68FF9AFFE5EE39FDC0FD51" pageId="5" pageNumber="606" type="diagnosis">
<paragraph id="8BFA40136C68FF9AFFE5EE39FDC0FD51" blockId="5.[113,1089,219,1236]" pageId="5" pageNumber="606">
<emphasis id="B9319C016C68FF9AFFE5EE39FEB5FE91" bold="true" box="[113,360,283,307]" pageId="5" pageNumber="606">Differential diagnosis.</emphasis>
The new species is most similar to
<taxonomicName id="4C453B906C68FF9AFD46EE3FFC6CFE96" authorityName="Mulsant &amp; Rey" authorityYear="1861" box="[722,945,285,308]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B9319C016C68FF9AFD46EE3FFC6CFE96" box="[722,945,285,308]" italics="true" pageId="5" pageNumber="606">Hydroporus longulus</emphasis>
</taxonomicName>
from western Central Europe and
<taxonomicName id="4C453B906C68FF9AFEDCEE1FFDB5FEF6" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[328,616,315,340]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="nevadensis">
<emphasis id="B9319C016C68FF9AFEDCEE1FFE03FEF6" box="[328,478,317,340]" italics="true" pageId="5" pageNumber="606">H. nevadensis</emphasis>
Sharp, 1882
</taxonomicName>
, which is distributed in the mountain ranges of
<collectingCountry id="F35200836C68FF9AFF1BEE79FF11FED1" box="[143,204,347,371]" name="Spain" pageId="5" pageNumber="606">Spain</collectingCountry>
and
<collectingCountry id="F35200836C68FF9AFE95EE79FE82FED1" box="[257,351,347,371]" name="Portugal" pageId="5" pageNumber="606">Portugal</collectingCountry>
.
<taxonomicName id="4C453B906C68FF9AFEFDEE7FFD9DFED6" authority="Fery &amp; Hájek, 2013" authorityName="Fery &amp; Hájek" authorityYear="2013" box="[361,576,349,372]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="toubkal" status="sp. nov.">
<emphasis id="B9319C016C68FF9AFEFDEE7FFD9DFED6" box="[361,576,349,372]" italics="true" pageId="5" pageNumber="606">Hydroporus toubkal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202217A6C68FF9AFDD3EE7EFD46FED1" box="[583,667,348,371]" pageId="5" pageNumber="606" rank="species">sp. nov.</taxonomicNameLabel>
can hardly be distinguished from these species by extern features alone; the shape of the median lobe of the aedeagus is the only reliable character which allows for separation of the three species (compare
<figureCitation id="137E5C966C68FF9AFC38EEB9FBD0FE11" box="[940,1037,411,436]" captionStart="Figs 2–9. 2–4, 8–9" captionStartId="3.[113,150,1493,1512]" captionTargetBox="[120,1077,240,1439]" captionTargetId="figure-15@3.[113,1088,219,1454]" captionTargetPageId="3" captionText="Figs 2–9. 2–4, 8–9 – Hydroporus toubkal sp. nov.: 2 – median lobe of aedeagus in ventral view; 3 – median lobe in lateral view; 4 – left paramere; 8 – gonocoxosternum; 9 – gonocoxae. 5 – H. longulus Mulsant &amp; Rey, 1861, median lobe of aedeagus in lateral view (adapted from FERY 1999). 6 – H. nevadensis Sharp, 1882, median lobe of aedeagus in lateral view (adapted from FERY 1999). 7 – H. jacobsoni Zaitzev, 1927, median lobe of aedeagus in lateral view (adapted from FERY 2009)." figureDoi="http://doi.org/10.5281/zenodo.5740178" httpUri="https://zenodo.org/record/5740178/files/figure.png" pageId="5" pageNumber="606">Figs 3, 5</figureCitation>
, and 6), a statement which also applies to most other species in the group. The median lobe of the Caucasian
<taxonomicName id="4C453B906C68FF9AFE9AEEFFFDF3FE56" authority="Zaitzev, 1927" authorityName="Zaitzev" authorityYear="1927" box="[270,558,475,500]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B9319C016C68FF9AFE9AEEFFFE4AFE56" box="[270,407,477,500]" italics="true" pageId="5" pageNumber="606">H. jacobsoni</emphasis>
Zaitzev, 1927
</taxonomicName>
resembles that of the new species, particularly in ventral view; in lateral view the differences are more obvious: the apex is distinctly broader in
<taxonomicName id="4C453B906C68FF9AFF1AED3FFEC5FD96" authorityName="Zaitzev" authorityYear="1927" box="[142,280,541,564]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B9319C016C68FF9AFF1AED3FFEC5FD96" box="[142,280,541,564]" italics="true" pageId="5" pageNumber="606">H. jacobsoni</emphasis>
</taxonomicName>
and by no means pointed (
<figureCitation id="137E5C966C68FF9AFDAAED39FD5CFD96" box="[574,641,539,564]" captionStart="Figs 2–9. 2–4, 8–9" captionStartId="3.[113,150,1493,1512]" captionTargetBox="[120,1077,240,1439]" captionTargetId="figure-15@3.[113,1088,219,1454]" captionTargetPageId="3" captionText="Figs 2–9. 2–4, 8–9 – Hydroporus toubkal sp. nov.: 2 – median lobe of aedeagus in ventral view; 3 – median lobe in lateral view; 4 – left paramere; 8 – gonocoxosternum; 9 – gonocoxae. 5 – H. longulus Mulsant &amp; Rey, 1861, median lobe of aedeagus in lateral view (adapted from FERY 1999). 6 – H. nevadensis Sharp, 1882, median lobe of aedeagus in lateral view (adapted from FERY 1999). 7 – H. jacobsoni Zaitzev, 1927, median lobe of aedeagus in lateral view (adapted from FERY 2009)." figureDoi="http://doi.org/10.5281/zenodo.5740178" httpUri="https://zenodo.org/record/5740178/files/figure.png" pageId="5" pageNumber="606">Fig. 7</figureCitation>
). Finally we want to mention two further members of the
<taxonomicName id="4C453B906C68FF9AFE89ED1FFEA4FDF6" authorityName="Mulsant &amp; Rey" authorityYear="1861" box="[285,377,573,596]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B9319C016C68FF9AFE89ED1FFEA4FDF6" box="[285,377,573,596]" italics="true" pageId="5" pageNumber="606">longulus</emphasis>
</taxonomicName>
group:
<taxonomicName id="4C453B906C68FF9AFE5FED1FFCA0FDF1" authority="Regimbart, 1895" authorityName="Regimbart" authorityYear="1895" box="[459,893,571,596]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="jurjurensis">
<emphasis id="B9319C016C68FF9AFE5FED1FFD19FDF6" box="[459,708,573,596]" italics="true" pageId="5" pageNumber="606">Hydroporus jurjurensis</emphasis>
Régimbart, 1895
</taxonomicName>
from the Algerian Djurjura mountain range (the only other species of the group from northern Africa) which can be easily separated by the sharply cut apex of the median lobe (cf.
<bibRefCitation id="EFD43DE26C68FF9AFCAEED5EFC09FD31" author="GUIGNOT F." box="[826,980,635,660]" pageId="5" pageNumber="606" pagination="321 - 648" refId="ref2623" refString="GUIGNOT F. 1959: Revision des hydrocanthares d'Afrique (Coleoptera Dytiscoidea), deuxieme partie. Annales du Musee Royal du Congo Belge, Serie 8 vo (Sciences Zoologiques) 78: 321 - 648." type="journal article" year="1959">GUIGNOT 1959</bibRefCitation>
: Fig. 348) and
<taxonomicName id="4C453B906C68FF9AFF0AEDBFFD16FD11" authority="Hernando &amp; Fresneda, 1996" authorityName="Hernando &amp; Fresneda" authorityYear="1996" box="[158,715,667,692]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="constantini">
<emphasis id="B9319C016C68FF9AFF0AEDBFFE47FD16" box="[158,410,669,692]" italics="true" pageId="5" pageNumber="606">Hydroporus constantini</emphasis>
<bibRefCitation id="EFD43DE26C68FF9AFE34EDB9FD16FD11" author="HERNANDO C. &amp; FRESNEDA J." box="[416,715,667,692]" pageId="5" pageNumber="606" pagination="155 - 161" refId="ref2762" refString="HERNANDO C. &amp; FRESNEDA J. 1996: Hydroporus constantini n. sp. de la Peninsula Iberica (Coleoptera, Dytiscidae). Nouvelle Revue d'Entomologie (Nouvelle Serie) 13: 155 - 161." type="journal article" year="1996">Hernando &amp; Fresneda, 1996</bibRefCitation>
</taxonomicName>
from central northern
<collectingCountry id="F35200836C68FF9AFC2EEDB9FC2AFD11" box="[954,1015,667,691]" name="Spain" pageId="5" pageNumber="606">Spain</collectingCountry>
whose median lobe shape is distinctly different from all species considered in the present work (cf.
<bibRefCitation id="EFD43DE26C68FF9AFFE5EDFEFE7BFD51" author="HERNANDO C. &amp; FRESNEDA J." box="[113,422,731,756]" pageId="5" pageNumber="606" pagination="155 - 161" refId="ref2762" refString="HERNANDO C. &amp; FRESNEDA J. 1996: Hydroporus constantini n. sp. de la Peninsula Iberica (Coleoptera, Dytiscidae). Nouvelle Revue d'Entomologie (Nouvelle Serie) 13: 155 - 161." type="journal article" year="1996">HERNANDO &amp; FRESNEDA 1996</bibRefCitation>
:
<figureCitation id="137E5C966C68FF9AFE25EDF9FDD2FD51" box="[433,527,731,756]" captionStart="Figs 2–9. 2–4, 8–9" captionStartId="3.[113,150,1493,1512]" captionTargetBox="[120,1077,240,1439]" captionTargetId="figure-15@3.[113,1088,219,1454]" captionTargetPageId="3" captionText="Figs 2–9. 2–4, 8–9 – Hydroporus toubkal sp. nov.: 2 – median lobe of aedeagus in ventral view; 3 – median lobe in lateral view; 4 – left paramere; 8 – gonocoxosternum; 9 – gonocoxae. 5 – H. longulus Mulsant &amp; Rey, 1861, median lobe of aedeagus in lateral view (adapted from FERY 1999). 6 – H. nevadensis Sharp, 1882, median lobe of aedeagus in lateral view (adapted from FERY 1999). 7 – H. jacobsoni Zaitzev, 1927, median lobe of aedeagus in lateral view (adapted from FERY 2009)." figureDoi="http://doi.org/10.5281/zenodo.5740178" httpUri="https://zenodo.org/record/5740178/files/figure.png" pageId="5" pageNumber="606">Figs 2, 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F13986C68FF9AFFE5EDDEFDD6FC51" pageId="5" pageNumber="606" type="etymology">
<paragraph id="8BFA40136C68FF9AFFE5EDDEFF12FCF1" blockId="5.[113,1089,219,1236]" pageId="5" pageNumber="606">
<emphasis id="B9319C016C68FF9AFFE5EDDEFF2FFCB6" bold="true" box="[113,242,764,788]" pageId="5" pageNumber="606">Etymology.</emphasis>
The new species is named after the
<typeStatus id="54FEFEB16C68FF9AFD16EDDEFD6DFCB6" box="[642,688,764,788]" pageId="5" pageNumber="606">type</typeStatus>
locality, Toubkal massif of the High Atlas mountain range; the specific epithet is a noun in the nominative case standing in apposition.
</paragraph>
<paragraph id="8BFA40136C68FF9AFFE5EC79FDD6FC51" blockId="5.[113,1089,219,1236]" pageId="5" pageNumber="606">
<emphasis id="B9319C016C68FF9AFFE5EC79FE47FCD1" bold="true" box="[113,410,859,883]" pageId="5" pageNumber="606">Collecting circumstances.</emphasis>
The specimens of the
<typeStatus id="54FEFEB16C68FF9AFD0AEC7EFD11FCD6" box="[670,716,860,884]" pageId="5" pageNumber="606">type</typeStatus>
series were collected in a spring with mosses, ca.
<quantity id="4CBDEDF66C68FF9AFEBEEC59FEA8FC31" box="[298,373,891,915]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="606" unit="km" value="0.5">0.5 km</quantity>
below the Refuge du Toubkal “Les Mouflons” (K. Orszulik, pers. comm.). The single female from Oukaïmeden was collected in a small stream together with
<taxonomicName id="4C453B906C68FF9AFFE5EC9FFE01FC71" authority="(Olivier, 1795)" baseAuthorityName="Olivier" baseAuthorityYear="1795" box="[113,476,955,980]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="biguttatus">
<emphasis id="B9319C016C68FF9AFFE5EC9FFEE9FC76" box="[113,308,957,980]" italics="true" pageId="5" pageNumber="606">Agabus biguttatus</emphasis>
(Olivier, 1795)
</taxonomicName>
and
<taxonomicName id="4C453B906C68FF9AFD85EC9FFE9EFC51" authority="discretus Fairmaire &amp; Brisout de Barneville, 1859" authorityName="Fairmaire &amp; Brisout de Barneville" authorityYear="1859" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="subSpecies" species="discretus" subSpecies="discretus">
<emphasis id="B9319C016C68FF9AFD85EC9FFC83FC76" box="[529,862,957,980]" italics="true" pageId="5" pageNumber="606">Hydroporus discretus discretus</emphasis>
Fairmaire &amp; Brisout de Barneville, 1859
</taxonomicName>
(both
<taxonomicName id="4C453B906C68FF9AFE1DECF9FDDDFC51" authorityName="Leach" authorityYear="1815" box="[393,512,987,1011]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F13986C68FF9AFFE5ECD9FC6DFB71" pageId="5" pageNumber="606" type="distribution">
<paragraph id="8BFA40136C68FF9AFFE5ECD9FED7FBF1" blockId="5.[113,1089,219,1236]" pageId="5" pageNumber="606">
<emphasis id="B9319C016C68FF9AFFE5ECD9FED9FBB1" bold="true" box="[113,260,1019,1043]" pageId="5" pageNumber="606">Distribution.</emphasis>
Currently only known from the
<typeStatus id="54FEFEB16C68FF9AFDF6ECDEFD4DFBB6" box="[610,656,1020,1044]" pageId="5" pageNumber="606">type</typeStatus>
locality, Toubkal massif (
<collectingCountry id="F35200836C68FF9AFC32ECDEFBD1FBB6" box="[934,1036,1020,1044]" name="Morocco" pageId="5" pageNumber="606">Morocco</collectingCountry>
), but possibly more widespread in the High Atlas mountain range (see the single female from Oukaïmeden).
</paragraph>
<paragraph id="8BFA40136C68FF9AFFE5EB7EFC6DFB71" blockId="5.[113,1089,219,1236]" pageId="5" pageNumber="606">
<emphasis id="B9319C016C68FF9AFFE5EB7EFF65FBD6" bold="true" box="[113,184,1116,1140]" pageId="5" pageNumber="606">Notes.</emphasis>
<bibRefCitation id="EFD43DE26C68FF9AFF55EB7EFE47FBD6" author="GUIGNOT F." box="[193,410,1116,1140]" pageId="5" pageNumber="606" pagination="1 - 287" refId="ref2606" refString="GUIGNOT F. 1947: Coleopteres hydrocanthares. Faune de France 48: 1 - 287." type="journal article" year="1947">GUIGNOT (1947: 111</bibRefCitation>
and
<bibRefCitation id="EFD43DE26C68FF9AFE46EB79FD9CFBD1" author="GUIGNOT F." box="[466,577,1115,1139]" pageId="5" pageNumber="606" pagination="321 - 648" refId="ref2623" refString="GUIGNOT F. 1959: Revision des hydrocanthares d'Afrique (Coleoptera Dytiscoidea), deuxieme partie. Annales du Musee Royal du Congo Belge, Serie 8 vo (Sciences Zoologiques) 78: 321 - 648." type="journal article" year="1959">1959: 396</bibRefCitation>
) gave records of
<taxonomicName id="4C453B906C68FF9AFC91EB7FFC59FBD6" authorityName="Mulsant &amp; Rey" authorityYear="1861" box="[773,900,1117,1140]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B9319C016C68FF9AFC91EB7FFC59FBD6" box="[773,900,1117,1140]" italics="true" pageId="5" pageNumber="606">H. longulus</emphasis>
</taxonomicName>
from the Middle Atlas and the High Atlas respectively;
<bibRefCitation id="EFD43DE26C68FF9AFD87EB5EFD00FB36" author="KOCHER L." box="[531,733,1147,1172]" pageId="5" pageNumber="606" pagination="1 - 244" refId="ref2847" refString="KOCHER L. 1958: Catalogue commente des coleopteres du Maroc. Fascicule II. Hydrocanthares, palpicornes, brachelytres. Travaux de l'Institut Scientifique Cherifien (Societe des Sciences Naturelles et Physiques du Maroc), Serie Zoologique 14: 1 - 244." type="journal article" year="1958">KOCHER (1958: 18)</bibRefCitation>
also gave records from the High Atlas. We have not been able to study any material belonging to the records given by these two authors, but presume that these records may belong to
<taxonomicName id="4C453B906C68FF9AFD72EB9FFC8AFB76" authority="Fery &amp; Hájek, 2013" authorityName="Fery &amp; Hájek" authorityYear="2013" box="[742,855,1213,1236]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="toubkal" status="sp. nov.">
<emphasis id="B9319C016C68FF9AFD72EB9FFC8AFB76" box="[742,855,1213,1236]" italics="true" pageId="5" pageNumber="606">H. toubkal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202217A6C68FF9AFCC9EB9EFC6DFB71" box="[861,944,1212,1235]" pageId="5" pageNumber="606" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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