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<mods:title id="F5EA1D585C10DDA68DCE6A5BBCDAAB20">Biofouling hydroids (Cnidaria: Hydrozoa) from a Tropical Eastern Pacific island, with remarks on their biogeography</mods:title>
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<emphasis id="B97CD030FFCBFFA0899FFA4C15EAFA6D" bold="true" box="[447,682,1438,1465]" italics="true" pageId="21" pageNumber="586">Obelia microtheca</emphasis>
<bibRefCitation id="EF9971D3FFCBFFA08A92FA4C1424FA6D" author="Fraser CM" box="[690,868,1438,1465]" pageId="21" pageNumber="586" pagination="1 - 105" refId="ref19533" refString="Fraser CM. 1938 a. Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pac Expeditions. 4 (1): 1 - 105." type="journal article" year="1938">Fraser, 1938a</bibRefCitation>
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(
<figureCitation id="133310A7FFCBFFA08A65FA13159DFA08" box="[581,733,1473,1500]" captionStart="Figure 6" captionStartId="19.[160,225,1409,1432]" captionTargetBox="[166,1151,150,1377]" captionTargetId="figure-1@19.[163,1153,142,1377]" captionTargetPageId="19" captionText="Figure 6. Leptothecata: Obeliidae, Haleciidae, Sertularellidae and Sertulariidae. (a) Obelia microtheca: hydrocaulus with two pedicels and hydrothecae, #240607. (b) Obelia microtheca: part of hydrocaulus with long pedicel and a hydrotheca, #240607. (c) Obelia microtheca: part of hydrocaulus and base of a pedicel with a gonotheca, #240607. (d) Halecium sp.: part of hydrocaulus with hydrothecae and base of a branch, #266335. (e) Sertularella affinicostata: stolon and stem with five hydrothecae, #240592. (f) Sertularella affinicostata: part of stem with two hydrothecae, #240592. (g) Tridentata borneensis: base of a stem with two pairs of hydrothecae, #240634. (h) Tridentata borneensis: distal end of a stem with two pairs of hydrothecae, #240634. (i) Tridentata borneensis: gonotheca, #266336. Scale bars: a, i = 0.2 mm; bd, fh = 0.1 mm; e = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012502" httpUri="https://zenodo.org/record/7012502/files/figure.png" pageId="21" pageNumber="586">Figure 6ac</figureCitation>
)
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<taxonomicName id="4C0877A1FFCBFFA08880FA24156DF9DB" authority="Fraser, 1938 a: 37" authorityName="Fraser" authorityPageNumber="37" authorityYear="1938" box="[160,557,1526,1551]" class="Hydrozoa" family="Campanulariidae" genus="Obelia" kingdom="Animalia" order="Leptothecata" pageId="21" pageNumber="586" phylum="Cnidaria" rank="species" species="microtheca">
<emphasis id="B97CD030FFCBFFA08880FA241627F9DB" box="[160,359,1526,1551]" italics="true" pageId="21" pageNumber="586">Obelia microtheca</emphasis>
<bibRefCitation id="EF9971D3FFCBFFA0894EFA24156DF9DB" author="Fraser CM" box="[366,557,1526,1551]" pageId="21" pageNumber="586" pagination="1 - 105" refId="ref19533" refString="Fraser CM. 1938 a. Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pac Expeditions. 4 (1): 1 - 105." type="journal article" year="1938">Fraser, 1938a: 37</bibRefCitation>
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, pl. 9, fig. 40ad.
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locality
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: off Santa Elena
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,
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02°08
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20″S
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,
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81°00
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15″W
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,
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(
<bibRefCitation id="EF9971D3FFCBFFA08B50F9BC1376F953" author="Calder DR &amp; Vervoort W &amp; Hochberg FG" box="[880,1078,1646,1671]" pageId="21" pageNumber="586" pagination="919 - 1058" refId="ref17575" refString="Calder DR, Vervoort W, Hochberg FG. 2009. Lectotype designations of new species of hydroids (Cnidaria, Hydrozoa), described by C. M. Fraser, from Allan Hancock Pacific and Caribbean Sea Expeditions. Zool Mededelingen. 83: 919 - 1058." type="journal article" year="2009">Calder et al. 2009</bibRefCitation>
)
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.
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<emphasis id="B97CD030FFC8FFA38880FF5C163FFF7C" bold="true" box="[160,383,142,168]" italics="true" pageId="22" pageNumber="587">Material examined</emphasis>
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<paragraph id="8BB70C22FFC8FFA38880FF601646FEAC" blockId="22.[160,1156,142,376]" pageId="22" pageNumber="587">
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,
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,
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, 23 colony fragments, to
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high, with gonothecae, coll
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<materialsCitation id="3B60067FFFC8FFA38CA0FF6017F4FF39" ID-GBIF-Occurrence="3891616404" isTailEnumeration="true" latitude="5.54556" location="Wafer Bay" longLatPrecision="1" longitude="-87.06221" pageId="22" pageNumber="587" specimenCount="1">. G</materialsCitation>
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.
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, #240607
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.
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,
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, 5 colony fragments, to
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high, with a gonotheca, coll
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<materialsCitation id="3B60067FFFC8FFA389E6FF2515D6FEC4" ID-GBIF-Occurrence="3891616399" box="[454,662,247,272]" location="Ashton" pageId="22" pageNumber="587" specimenCount="1">
.
<location id="8ED75AF9FFC8FFA389F4FF251567FEC4" LSID="urn:lsid:plazi:treatment:03A1BD34FFCBFFA3899FFA4C132FF914:8ED75AF9FFC8FFA389F4FF251567FEC4" box="[468,551,247,272]" name="Ashton" pageId="22" pageNumber="587">Ashton</location>
, #240611
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.
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<location id="8ED75AF9FFC8FFA38A8EFF251467FEC4" LSID="urn:lsid:plazi:treatment:03A1BD34FFCBFFA3899FFA4C132FF914:8ED75AF9FFC8FFA38A8EFF251467FEC4" box="[686,807,247,272]" latitude="5.54556" longLatPrecision="1" longitude="-87.06221" name="Wafer Bay" pageId="22" pageNumber="587">Wafer Bay</location>
,
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,
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, 9 colony fragments, to
<quantity id="4CF0A1C7FFC8FFA38965FEC816CAFEE7" box="[325,394,282,307]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="587" unit="mm" value="8.0">8 mm</quantity>
high, with gonothecae, coll
</materialsCitation>
<materialsCitation id="3B60067FFFC8FFA38AF6FEC815B9FEE7" ID-GBIF-Occurrence="3891616422" box="[726,761,282,307]" isTailEnumeration="true" latitude="5.54556" location="Wafer Bay" longLatPrecision="1" longitude="-87.06221" pageId="22" pageNumber="587" specimenCount="1">. G</materialsCitation>
<materialsCitation id="3B60067FFFC8FFA38AD8FEC81489FEE7" ID-GBIF-Occurrence="3891616339" box="[760,969,282,307]" location="Ashton" pageId="22" pageNumber="587" specimenCount="1">
.
<location id="8ED75AF9FFC8FFA38B27FEC8141AFEE7" LSID="urn:lsid:plazi:treatment:03A1BD34FFCBFFA3899FFA4C132FF914:8ED75AF9FFC8FFA38B27FEC8141AFEE7" box="[775,858,282,307]" name="Ashton" pageId="22" pageNumber="587">Ashton</location>
, #240605
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.
<materialsCitation id="3B60067FFFC8FFA38BC3FEC81364FE81" ID-GBIF-Occurrence="3891616419" latitude="5.55271" location="Chatham Bay" longLatPrecision="1" longitude="-87.03826" pageId="22" pageNumber="587" specimenCount="1">
<location id="8ED75AF9FFC8FFA38BC3FEC813C1FEE7" LSID="urn:lsid:plazi:treatment:03A1BD34FFCBFFA3899FFA4C132FF914:8ED75AF9FFC8FFA38BC3FEC813C1FEE7" box="[995,1153,282,307]" latitude="5.55271" longLatPrecision="1" longitude="-87.03826" name="Chatham Bay" pageId="22" pageNumber="587">Chatham Bay</location>
,
<geoCoordinate id="EE3C6AE5FFC8FFA38880FEEE17BAFE81" box="[160,250,316,341]" degrees="5.55271" direction="north" orientation="latitude" pageId="22" pageNumber="587" precision="1" value="5.55271">5.55271</geoCoordinate>
,
<geoCoordinate id="EE3C6AE5FFC8FFA38926FEEE163EFE81" box="[262,382,316,341]" degrees="87.03826" direction="west" orientation="longitude" pageId="22" pageNumber="587" precision="1" value="-87.03826">87.03826</geoCoordinate>
, 2 colony fragments, to
<quantity id="4CF0A1C7FFC8FFA38AB5FEEE1597FE81" box="[661,727,316,341]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="587" unit="mm" value="7.0">7 mm</quantity>
high, with a gonotheca, coll
</materialsCitation>
<materialsCitation id="3B60067FFFC8FFA38C03FEEE1375FE81" ID-GBIF-Occurrence="3891616423" box="[1059,1077,316,341]" isTailEnumeration="true" latitude="5.55271" location="Chatham Bay" longLatPrecision="1" longitude="-87.03826" pageId="22" pageNumber="587" specimenCount="1">. I</materialsCitation>
<materialsCitation id="3B60067FFFC8FFA38C15FEEE1642FEAC" ID-GBIF-Occurrence="3891616365" location="Keith" pageId="22" pageNumber="587" specimenCount="1">
.
<location id="8ED75AF9FFC8FFA38C64FEEE13C0FE81" LSID="urn:lsid:plazi:treatment:03A1BD34FFCBFFA3899FFA4C132FF914:8ED75AF9FFC8FFA38C64FEEE13C0FE81" box="[1092,1152,316,341]" name="Keith" pageId="22" pageNumber="587">Keith</location>
, #307695
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.
</paragraph>
</subSubSection>
<subSubSection id="C3125FA9FFC8FFA38880FE76132FF914" pageId="22" pageNumber="587" type="discussion">
<paragraph id="8BB70C22FFC8FFA38880FE761646FE6A" blockId="22.[160,1156,420,1589]" box="[160,262,420,446]" pageId="22" pageNumber="587">
<emphasis id="B97CD030FFC8FFA38880FE761646FE6A" bold="true" box="[160,262,420,446]" italics="true" pageId="22" pageNumber="587">Remarks</emphasis>
</paragraph>
<paragraph id="8BB70C22FFC8FFA38880FE15134BFC77" blockId="22.[160,1156,420,1589]" pageId="22" pageNumber="587">
Material examined here has been identified as
<taxonomicName id="4C0877A1FFC8FFA38A83FE151340FE34" authority="Fraser, 1938 a" authorityName="Fraser" authorityYear="1938" box="[675,1024,455,480]" class="Hydrozoa" family="Campanulariidae" genus="Obelia" kingdom="Animalia" order="Leptothecata" pageId="22" pageNumber="587" phylum="Cnidaria" rank="species" species="microtheca">
<emphasis id="B97CD030FFC8FFA38A83FE151428FE34" box="[675,872,455,480]" italics="true" pageId="22" pageNumber="587">Obelia microtheca</emphasis>
<bibRefCitation id="EF9971D3FFC8FFA38B4EFE151340FE34" author="Fraser CM" box="[878,1024,455,480]" pageId="22" pageNumber="587" pagination="1 - 105" refId="ref19533" refString="Fraser CM. 1938 a. Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pac Expeditions. 4 (1): 1 - 105." type="journal article" year="1938">Fraser, 1938a</bibRefCitation>
</taxonomicName>
by virtue of having the following combination of characters: (1) colonies small (
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high or less), erect, unbranched or mostly so, with sympodial growth; (2) hydrocaulus monosiphonic and only slightly geniculate, divided into internodes; (3) cauline internodes annulated proximally and with alternate apophyses distally; (4) hydrothecal pedicels exceptionally long, annulated either proximally and distally or throughout; (5) hydrothecae small (ca.
<quantity id="4CF0A1C7FFC8FFA38880FD4517B4FD64" box="[160,244,663,688]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="587" unit="mm" value="0.2">0.2 mm</quantity>
deep), funnel-shaped, with depth and diameter at margin essentially equal, walls nearly straight, margin entire; and (6) gonothecae cone-shaped with a terminal collar, axillary in position, arising from proximal ends of hydrothecal pedicels, walls smooth. Hydrothecal pedicels in present material were exceptionally long (
<figureCitation id="133310A7FFC8FFA38B95FD2D1305FCCC" box="[949,1093,767,792]" captionStart="Figure 6" captionStartId="19.[160,225,1409,1432]" captionTargetBox="[166,1151,150,1377]" captionTargetId="figure-1@19.[163,1153,142,1377]" captionTargetPageId="19" captionText="Figure 6. Leptothecata: Obeliidae, Haleciidae, Sertularellidae and Sertulariidae. (a) Obelia microtheca: hydrocaulus with two pedicels and hydrothecae, #240607. (b) Obelia microtheca: part of hydrocaulus with long pedicel and a hydrotheca, #240607. (c) Obelia microtheca: part of hydrocaulus and base of a pedicel with a gonotheca, #240607. (d) Halecium sp.: part of hydrocaulus with hydrothecae and base of a branch, #266335. (e) Sertularella affinicostata: stolon and stem with five hydrothecae, #240592. (f) Sertularella affinicostata: part of stem with two hydrothecae, #240592. (g) Tridentata borneensis: base of a stem with two pairs of hydrothecae, #240634. (h) Tridentata borneensis: distal end of a stem with two pairs of hydrothecae, #240634. (i) Tridentata borneensis: gonotheca, #266336. Scale bars: a, i = 0.2 mm; bd, fh = 0.1 mm; e = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7012502" httpUri="https://zenodo.org/record/7012502/files/figure.png" pageId="22" pageNumber="587">Figure 6a, b</figureCitation>
), but otherwise specimens corresponded with the brief original description of
<taxonomicName id="4C0877A1FFC8FFA38BC9FCF013C3FCEF" authorityName="Fraser" authorityYear="1938" box="[1001,1155,802,827]" class="Hydrozoa" family="Campanulariidae" genus="Obelia" kingdom="Animalia" order="Leptothecata" pageId="22" pageNumber="587" phylum="Cnidaria" rank="species" species="microtheca">
<emphasis id="B97CD030FFC8FFA38BC9FCF013C3FCEF" box="[1001,1155,802,827]" italics="true" pageId="22" pageNumber="587">O. microtheca</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF9971D3FFC8FFA388E6FC961628FC89" author="Fraser CM" box="[198,360,836,861]" pageId="22" pageNumber="587" pagination="1 - 105" refId="ref19533" refString="Fraser CM. 1938 a. Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pac Expeditions. 4 (1): 1 - 105." type="journal article" year="1938">Fraser (1938a)</bibRefCitation>
. While considered conspecific with
<taxonomicName id="4C0877A1FFC8FFA38B29FC961324FC89" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[777,1124,836,861]" class="Hydrozoa" family="Campanulariidae" genus="Obelia" kingdom="Animalia" order="Leptothecata" pageId="22" pageNumber="587" phylum="Cnidaria" rank="species" species="dichotoma">
<emphasis id="B97CD030FFC8FFA38B29FC9614DFFC89" box="[777,927,836,861]" italics="true" pageId="22" pageNumber="587">O. dichotoma</emphasis>
(
<bibRefCitation id="EF9971D3FFC8FFA38B90FC96131CFC89" author="Linnaeus C" box="[944,1116,836,861]" pageId="22" pageNumber="587" refId="ref21295" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Holmiae: Laurentii Salvii. doi: 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
in some earlier works (
<bibRefCitation id="EF9971D3FFC8FFA389A8FCB51573FC54" author="Cornelius PFS" box="[392,563,871,896]" pageId="22" pageNumber="587" pagination="249 - 293" refId="ref18738" refString="Cornelius PFS. 1975. The hydroid species of Obelia (Coelenterata, Hydrozoa: Campanulariidae), with notes on the medusa stage. Bull Br Mus (Nat Hist) Zool. 28: 249 - 293. doi: 10.5962 / p. 314143" type="journal article" year="1975">Cornelius 1975</bibRefCitation>
,
<bibRefCitation id="EF9971D3FFC8FFA38A62FCB51539FC54" author="Cornelius PFS" box="[578,633,871,896]" pageId="22" pageNumber="587" pagination="37 - 148" refId="ref18785" refString="Cornelius PFS. 1982. Hydroids and medusae of the family Campanulariidae recorded from the eastern North Atlantic, with a world synopsis of genera. Bull Br Mus (Nat Hist) Zool. 42: 37 - 148." type="journal article" year="1982">1982</bibRefCitation>
;
<bibRefCitation id="EF9971D3FFC8FFA38AA8FCB51451FC54" author="Calder DR" box="[648,785,871,896]" pageId="22" pageNumber="587" pagination="1 - 140" refId="ref17413" refString="Calder DR. 1991. Shallow-water hydroids of Bermuda. The Thecatae, exclusive of Plumularioidea. Royal Ontario Museum. Life Sci Contrib. 154: 1 - 140." type="journal article" year="1991">Calder 1991</bibRefCitation>
), the species was recognised as valid by Calder et al. (2021) and earlier provisionally so by
<bibRefCitation id="EF9971D3FFC8FFA38B13FC581347FC77" author="Calder DR &amp; Vervoort W &amp; Hochberg FG" box="[819,1031,906,931]" pageId="22" pageNumber="587" pagination="919 - 1058" refId="ref17575" refString="Calder DR, Vervoort W, Hochberg FG. 2009. Lectotype designations of new species of hydroids (Cnidaria, Hydrozoa), described by C. M. Fraser, from Allan Hancock Pacific and Caribbean Sea Expeditions. Zool Mededelingen. 83: 919 - 1058." type="journal article" year="2009">Calder et al. (2009)</bibRefCitation>
.
</paragraph>
<paragraph id="8BB70C22FFC8FFA388E0FC7F142BFB84" blockId="22.[160,1156,420,1589]" pageId="22" pageNumber="587">
<materialsCitation id="3B60067FFFC8FFA388E0FC7F17F4FC3C" ID-GBIF-Occurrence="3891616368" location="Hydroids" municipality="Cocos Island" pageId="22" pageNumber="587" specimenCount="1" typeStatus="lectotype">
<location id="8ED75AF9FFC8FFA388E0FC7F1664FC12" LSID="urn:lsid:plazi:treatment:03A1BD34FFCBFFA3899FFA4C132FF914:8ED75AF9FFC8FFA388E0FC7F1664FC12" box="[192,292,941,966]" municipality="Cocos Island" name="Hydroids" pageId="22" pageNumber="587">Hydroids</location>
from
<collectingMunicipality id="6BD39658FFC8FFA38945FC7F16B0FC12" box="[357,496,941,966]" pageId="22" pageNumber="587">Cocos Island</collectingMunicipality>
were largely indistinguishable from the
<typeStatus id="54B3B280FFC8FFA38B8EFC7F1357FC12" box="[942,1047,941,966]" pageId="22" pageNumber="587" type="lectotype">lectotype</typeStatus>
colony of
<taxonomicName id="4C0877A1FFC8FFA38880FC1D17F4FC3C" authorityName="Fraser" authorityYear="1938" box="[160,180,975,1000]" class="Hydrozoa" family="Campanulariidae" genus="Obelia" kingdom="Animalia" order="Leptothecata" pageId="22" pageNumber="587" phylum="Cnidaria" rank="species" species="microtheca">
<emphasis id="B97CD030FFC8FFA38880FC1D17F4FC3C" box="[160,180,975,1000]" italics="true" pageId="22" pageNumber="587">O</emphasis>
</taxonomicName>
</materialsCitation>
<emphasis id="B97CD030FFC8FFA38894FC1D17F8FC3C" box="[180,184,975,1000]" italics="true" pageId="22" pageNumber="587">.</emphasis>
<materialsCitation id="3B60067FFFC8FFA3889DFC1D1667FBFA" ID-GBIF-Occurrence="3891616356" country="Ecuador" location="Santa Elena Bay" pageId="22" pageNumber="587" specimenCount="1" typeStatus="lectotype">
<emphasis id="B97CD030FFC8FFA3889DFC1D1676FC3C" box="[189,310,975,1000]" italics="true" pageId="22" pageNumber="587">microtheca</emphasis>
(British Columbia Provincial Museum 97600437-002) from
<location id="8ED75AF9FFC8FFA38BEFFC1D13C1FC3C" LSID="urn:lsid:plazi:treatment:03A1BD34FFCBFFA3899FFA4C132FF914:8ED75AF9FFC8FFA38BEFFC1D13C1FC3C" box="[975,1153,975,1000]" country="Ecuador" name="Santa Elena Bay" pageId="22" pageNumber="587">Santa Elena Bay</location>
,
<collectingCountry id="F31F4CB2FFC8FFA38880FC2017BEFBDF" box="[160,254,1010,1035]" name="Ecuador" pageId="22" pageNumber="587">Ecuador</collectingCountry>
, and from specimens identified as the species from the same region by Calder et al. (2021)
</materialsCitation>
. While hydrothecal pedicels reached a greater length in material examined here, the difference is not considered taxonomically significant.
</paragraph>
<paragraph id="8BB70C22FFC8FFA388E0FB881685FAF4" blockId="22.[160,1156,420,1589]" pageId="22" pageNumber="587">
<taxonomicName id="4C0877A1FFC8FFA388E0FB8816C7FBA7" authorityName="Fraser" authorityYear="1938" box="[192,391,1114,1139]" class="Hydrozoa" family="Campanulariidae" genus="Obelia" kingdom="Animalia" order="Leptothecata" pageId="22" pageNumber="587" phylum="Cnidaria" rank="species" species="microtheca">
<emphasis id="B97CD030FFC8FFA388E0FB8816C7FBA7" box="[192,391,1114,1139]" italics="true" pageId="22" pageNumber="587">Obelia microtheca</emphasis>
</taxonomicName>
is a little-known species, originally described from Santa Elena Bay,
<collectingCountry id="F31F4CB2FFC8FFA38880FBAF17BBFB42" box="[160,251,1149,1174]" name="Ecuador" pageId="22" pageNumber="587">Ecuador</collectingCountry>
(
<bibRefCitation id="EF9971D3FFC8FFA3892BFBAF16DEFB42" author="Fraser CM" box="[267,414,1149,1174]" pageId="22" pageNumber="587" pagination="1 - 105" refId="ref19533" refString="Fraser CM. 1938 a. Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pac Expeditions. 4 (1): 1 - 105." type="journal article" year="1938">Fraser 1938a</bibRefCitation>
). Subsequent records of it have come from Independencia Bay,
<collectingCountry id="F31F4CB2FFC8FFA38880FB4D1796FB6C" box="[160,214,1183,1208]" name="Peru" pageId="22" pageNumber="587">Peru</collectingCountry>
,
<collectingRegion id="49CCC2C0FFC8FFA388FFFB4D161EFB6C" box="[223,350,1183,1208]" country="Peru" name="La Libertad" pageId="22" pageNumber="587">La Libertad</collectingRegion>
,
<collectingCountry id="F31F4CB2FFC8FFA38948FB4D1686FB6C" box="[360,454,1183,1208]" name="Ecuador" pageId="22" pageNumber="587">Ecuador</collectingCountry>
, and the Pacific coast of
<collectingCountry id="F31F4CB2FFC8FFA38AF8FB4D1472FB6C" box="[728,818,1183,1208]" name="Panama" pageId="22" pageNumber="587">Panama</collectingCountry>
(
<bibRefCitation id="EF9971D3FFC8FFA38B1EFB4D148CFB6C" author="Fraser CM" box="[830,972,1183,1208]" pageId="22" pageNumber="587" pagination="129 - 153" refId="ref19593" refString="Fraser CM. 1938 c. Hydroids of the 1932, 1933, 1935, and 1938 Allan Hancock Pacific Expeditions. Allan Hancock Pac Expeditions. 4 (3): 129 - 153." type="journal article" year="1938">Fraser 1938c</bibRefCitation>
), and it was one of the commonest species of hydroids in a study of fouling organisms from coastal
<collectingCountry id="F31F4CB2FFC8FFA38880FB3717BBFB2A" box="[160,251,1253,1278]" name="Ecuador" pageId="22" pageNumber="587">Ecuador</collectingCountry>
by Calder et al. (2021). It was not found in samples from the
<collectingRegion id="49CCC2C0FFC8FFA38B99FB3713C3FB2A" box="[953,1155,1253,1278]" country="Ecuador" name="Galapagos" pageId="22" pageNumber="587">Galápagos Islands</collectingRegion>
as part of the same study.
</paragraph>
<paragraph id="8BB70C22FFC8FFA388E0FAF816D0F9E1" blockId="22.[160,1156,420,1589]" pageId="22" pageNumber="587">
Ecologically, hydroids of
<taxonomicName id="4C0877A1FFC8FFA389FCFAF81535FA97" authorityName="Fraser" authorityYear="1938" box="[476,629,1322,1347]" class="Hydrozoa" family="Campanulariidae" genus="Obelia" kingdom="Animalia" order="Leptothecata" pageId="22" pageNumber="587" phylum="Cnidaria" rank="species" species="microtheca">
<emphasis id="B97CD030FFC8FFA389FCFAF81535FA97" box="[476,629,1322,1347]" italics="true" pageId="22" pageNumber="587">O. microtheca</emphasis>
</taxonomicName>
have been reported over a depth range from the shore (
<bibRefCitation id="EF9971D3FFC8FFA38937FA9E16E6FAB1" author="Fraser CM" box="[279,422,1356,1381]" pageId="22" pageNumber="587" pagination="129 - 153" refId="ref19593" refString="Fraser CM. 1938 c. Hydroids of the 1932, 1933, 1935, and 1938 Allan Hancock Pacific Expeditions. Allan Hancock Pac Expeditions. 4 (3): 129 - 153." type="journal article" year="1938">Fraser 1938c</bibRefCitation>
) to
<quantity id="4CF0A1C7FFC8FFA389F2FA9E154BFAB1" box="[466,523,1356,1381]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="587" unit="m" value="18.0">18 m</quantity>
(
<bibRefCitation id="EF9971D3FFC8FFA38A37FA9E15E7FAB1" author="Fraser CM" box="[535,679,1356,1381]" pageId="22" pageNumber="587" pagination="1 - 105" refId="ref19533" refString="Fraser CM. 1938 a. Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pac Expeditions. 4 (1): 1 - 105." type="journal article" year="1938">Fraser 1938a</bibRefCitation>
;
<bibRefCitation id="EF9971D3FFC8FFA38A94FA9E1436FAB1" author="Calder DR &amp; Vervoort W &amp; Hochberg FG" box="[692,886,1356,1381]" pageId="22" pageNumber="587" pagination="919 - 1058" refId="ref17575" refString="Calder DR, Vervoort W, Hochberg FG. 2009. Lectotype designations of new species of hydroids (Cnidaria, Hydrozoa), described by C. M. Fraser, from Allan Hancock Pacific and Caribbean Sea Expeditions. Zool Mededelingen. 83: 919 - 1058." type="journal article" year="2009">Calder et al. 2009</bibRefCitation>
). A substrate generalist, the species has been reported from a rock, large shells, octocorals, bryozoans, algae, a barnacle, a sponge and a hydroid (
<bibRefCitation id="EF9971D3FFC8FFA38A78FA40146AFA7F" author="Calder DR &amp; Vervoort W &amp; Hochberg FG" box="[600,810,1426,1451]" pageId="22" pageNumber="587" pagination="919 - 1058" refId="ref17575" refString="Calder DR, Vervoort W, Hochberg FG. 2009. Lectotype designations of new species of hydroids (Cnidaria, Hydrozoa), described by C. M. Fraser, from Allan Hancock Pacific and Caribbean Sea Expeditions. Zool Mededelingen. 83: 919 - 1058." type="journal article" year="2009">Calder et al. 2009</bibRefCitation>
, 2021). Colony size has been reported to reach
<quantity id="4CF0A1C7FFC8FFA38952FA6616EBFA19" box="[370,427,1460,1485]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="587" unit="cm" value="1.0">1 cm</quantity>
high, although the
<typeStatus id="54B3B280FFC8FFA38AB2FA6615BBFA19" box="[658,763,1460,1485]" pageId="22" pageNumber="587" type="lectotype">lectotype</typeStatus>
specimen at only
<quantity id="4CF0A1C7FFC8FFA38BF0FA661353FA19" box="[976,1043,1460,1485]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="587" unit="mm" value="3.0">3 mm</quantity>
high was fertile (
<bibRefCitation id="EF9971D3FFC8FFA388CEFA0516F4FA24" author="Calder DR &amp; Vervoort W &amp; Hochberg FG" box="[238,436,1495,1520]" pageId="22" pageNumber="587" pagination="919 - 1058" refId="ref17575" refString="Calder DR, Vervoort W, Hochberg FG. 2009. Lectotype designations of new species of hydroids (Cnidaria, Hydrozoa), described by C. M. Fraser, from Allan Hancock Pacific and Caribbean Sea Expeditions. Zool Mededelingen. 83: 919 - 1058." type="journal article" year="2009">Calder et al. 2009</bibRefCitation>
). The species is largely known from the Tropical Eastern Pacific, but its range reportedly extends to southern
<collectingCountry id="F31F4CB2FFC8FFA38A80FA281593F9C7" box="[672,723,1530,1555]" name="Peru" pageId="22" pageNumber="587">Peru</collectingCountry>
in the Warm Temperate Southeastern Pacific (
<bibRefCitation id="EF9971D3FFC8FFA388D4F9CE16C4F9E1" author="Fraser CM" box="[244,388,1564,1589]" pageId="22" pageNumber="587" pagination="129 - 153" refId="ref19593" refString="Fraser CM. 1938 c. Hydroids of the 1932, 1933, 1935, and 1938 Allan Hancock Pacific Expeditions. Allan Hancock Pac Expeditions. 4 (3): 129 - 153." type="journal article" year="1938">Fraser 1938c</bibRefCitation>
).
</paragraph>
<paragraph id="8BB70C22FFC8FFA38880F9B316DCF9AF" blockId="22.[160,1135,1633,1728]" box="[160,412,1633,1659]" pageId="22" pageNumber="587">
<emphasis id="B97CD030FFC8FFA38880F9B316DCF9AF" bold="true" box="[160,412,1633,1659]" italics="true" pageId="22" pageNumber="587">Reported distribution</emphasis>
</paragraph>
<paragraph id="8BB70C22FFC8FFA38880F95716FAF94A" blockId="22.[160,1135,1633,1728]" box="[160,442,1669,1694]" pageId="22" pageNumber="587">Cocos Island: first record.</paragraph>
<paragraph id="8BB70C22FFC8FFA388E0F975132FF914" blockId="22.[160,1135,1633,1728]" box="[192,1135,1703,1728]" pageId="22" pageNumber="587">
Elsewhere:
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(Pacific coast) to Independencia Bay,
<collectingCountry id="F31F4CB2FFC8FFA38B62F9751435F914" box="[834,885,1703,1728]" name="Peru" pageId="22" pageNumber="587">Peru</collectingCountry>
(
<bibRefCitation id="EF9971D3FFC8FFA38BA2F9751352F914" author="Fraser CM" box="[898,1042,1703,1728]" pageId="22" pageNumber="587" pagination="1 - 105" refId="ref19533" refString="Fraser CM. 1938 a. Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pac Expeditions. 4 (1): 1 - 105." type="journal article" year="1938">Fraser 1938a</bibRefCitation>
,
<bibRefCitation id="EF9971D3FFC8FFA38C3FF9751322F914" author="Fraser CM" box="[1055,1122,1703,1728]" pageId="22" pageNumber="587" pagination="129 - 153" refId="ref19593" refString="Fraser CM. 1938 c. Hydroids of the 1932, 1933, 1935, and 1938 Allan Hancock Pacific Expeditions. Allan Hancock Pac Expeditions. 4 (3): 129 - 153." type="journal article" year="1938">1938c</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>