303 lines
26 KiB
XML
303 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1026.63749" ID-GBIF-Dataset="09411512-24e4-4f35-90f9-b19a85ea0535" ID-PMC="PMC8018941" ID-Pensoft-Pub="1313-2970-1026-69" ID-Pensoft-UUID="762AC1314DE05514BFD79A8DC8F34E2F" ID-PubMed="33850419" ID-ZooBank="05EFF10E62144368BE471AA57A2C38D7" ModsDocID="1313-2970-1026-69" checkinTime="1616886939438" checkinUser="pensoft" docAuthor="Kurina, Olavi" docDate="2021" docId="B92821CC83945DDE8E2F3199603BAFBC" docLanguage="en" docName="ZooKeys 1026: 69-142" docOrigin="ZooKeys 1026" docPubDate="2021-03-26" docSource="http://dx.doi.org/10.3897/zookeys.1026.63749" docTitle="Sciophila georgei Kurina 2021, sp. nov." docType="treatment" docUuid="D9E0ED72-E487-480C-A89F-4E6DDD98C406" docUuidSource="ZooBank" docVersion="6" id="762AC1314DE05514BFD79A8DC8F34E2F" lastPageNumber="69" masterDocId="762AC1314DE05514BFD79A8DC8F34E2F" masterDocTitle="A contribution towards checklist of fungus gnats (Diptera, Diadocidiidae, Ditomyiidae, Bolitophilidae, Keroplatidae, Mycetophilidae) in Georgia, Transcaucasia" masterLastPageNumber="142" masterPageNumber="69" pageNumber="69" updateTime="1668149993877" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A contribution towards checklist of fungus gnats (Diptera, Diadocidiidae, Ditomyiidae, Bolitophilidae, Keroplatidae, Mycetophilidae) in Georgia, Transcaucasia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kurina, Olavi</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4858-4629</mods:nameIdentifier>
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<mods:affiliation>Institute of Agricultural and Environmental Sciences, Estonian University of Life Sciences, Kreutzwaldi st 5 D, 51006 Tartu, Estonia</mods:affiliation>
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<mods:nameIdentifier type="email">olavi.kurina@emu.ee</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-03-26</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1026</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>69</mods:start>
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<mods:end>142</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1026.63749</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1026.63749</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1026-69</mods:identifier>
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<mods:identifier type="ZooBank">05EFF10E62144368BE471AA57A2C38D7</mods:identifier>
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<mods:identifier type="Pensoft-UUID">762AC1314DE05514BFD79A8DC8F34E2F</mods:identifier>
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||
</mods:mods>
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<treatment ID-GBIF-Taxon="180747847" LSID="urn:lsid:zoobank.org:act:D9E0ED72-E487-480C-A89F-4E6DDD98C406" httpUri="http://treatment.plazi.org/id/B92821CC83945DDE8E2F3199603BAFBC" lastPageNumber="69" pageId="0" pageNumber="69">
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<subSubSection pageId="0" pageNumber="69" type="nomenclature">
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<paragraph pageId="0" pageNumber="69">
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<taxonomicName LSID="http://zoobank.org/D9E0ED72-E487-480C-A89F-4E6DDD98C406" authority="Kurina, 2021" authorityName="Kurina" authorityYear="2021" class="Insecta" family="Mycetophilidae" genus="Sciophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sciophila georgei" order="Diptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="georgei" status="sp. nov.">Sciophila georgei</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="69">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus photos of new species A Sciophila georgei sp. nov., paratype B Anatella metae sp. nov., paratype C Leia katae sp. nov., holotype, terminalia detached. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1026.63749.figure3" httpUri="https://binary.pensoft.net/fig/524674" pageId="0" pageNumber="69">Figs 3A</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Sciophila georgei sp. nov., male terminalia A dorsal view B ventral view C lateral view D apical part of tergite IX, dorsal view E dorsal view, tergite IX removed F aedeagal complex, dorsal view G gonostylus, internal view H gonostylus, posterior view. Abbreviations: aed = aedeagus, cer = cercus, gc = gonocoxite, gc ap = gonocoxal apodeme, gst lbr = lateral branch of gonostylus, gst mbr = medial branch of gonostylus, gst sibr = small internal branch of gonostylus, par = paramere, tg 9 = tergite IX. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1026.63749.figure4" httpUri="https://binary.pensoft.net/fig/524675" pageId="0" pageNumber="69">, 4A-G</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="type material">
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<paragraph pageId="0" pageNumber="69">
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<typeStatus>Type</typeStatus>
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material.
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</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<emphasis bold="true" pageId="0" pageNumber="69">
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<emphasis italics="true" pageId="0" pageNumber="69">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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.
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3062025344" collectingDate="2011-05-17" collectingDateMax="2011-05-27" collectingDateMin="2011-05-17" collectionCode="IZBE" collectorName="Lagotekhi NR & G. Japoshvili" country="Georgia" elevation="350" latitude="41.795834" location="Matsimi river" longLatPrecision="11" longitude="46.28533" municipality="Male" specimenCount="2" stateProvince="Kakheti" typeStatus="holotype">
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<collectingMunicipality>Male</collectingMunicipality>
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,
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<collectingCountry name="Georgia">Georgia</collectingCountry>
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,
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<collectingRegion country="Georgia" name="Kakheti">Kakheti</collectingRegion>
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,
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<collectorName>Lagotekhi NR</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:B92821CC83945DDE8E2F3199603BAFBC:A348C6FB8F39D861BBFD7B8E43A70F29" country="Georgia" latitude="41.795834" longLatPrecision="11" longitude="46.28533" municipality="Male" name="Matsimi river" stateProvince="Kakheti">Matsimi river</location>
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gorge,
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<geoCoordinate degrees="41" direction="north" minutes="47.75" orientation="latitude" precision="9" value="41.795834">41°47.75'N</geoCoordinate>
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,
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<geoCoordinate degrees="46" direction="east" minutes="17.12" orientation="longitude" precision="9" value="46.28533">46°17.12'E</geoCoordinate>
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,
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<elevation metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">
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<quantity metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</quantity>
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a.s.l.
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</elevation>
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,
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<date value="2011-05-17" valueMax="2011-05-27" valueMin="2011-05-17">
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<collectingDate value="2011-05-17" valueMax="2011-05-27" valueMin="2011-05-17">17-27.v.2011</collectingDate>
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</date>
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,
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<collectingMethod>Malaise trap</collectingMethod>
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, leg.
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<collectorName>G. Japoshvili</collectorName>
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[see Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Collecting data of fungus gnats in Georgia arranged by administrative regions of the country. Codes for the collecting events are used on the Figure 1 and within the list of species. The asterisk (*) indicates collecting with a Malaise trap for which the exact collecting dates are provided in the text." httpUri="http://table.plazi.org/id/D86DE75E4C228A3694D56414FF2DF34E" pageId="0" pageNumber="69" tableUuid="D86DE75E4C228A3694D56414FF2DF34E">1</tableCitation>
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: K-5] (mounted from alcohol,
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<collectionCode country="Estonia" httpUri="http://grbio.org/cool/9bph-kj5p" name="Institute of Zoology and Botany" type="">IZBE</collectionCode>
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).
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<emphasis bold="true" pageId="0" pageNumber="69">
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<emphasis italics="true" pageId="0" pageNumber="69">
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<typeStatus>Paratype</typeStatus>
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</emphasis>
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.
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</emphasis>
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Male, same as
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<typeStatus>holotype</typeStatus>
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(mounted from alcohol,
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<collectionCode country="Estonia" httpUri="http://grbio.org/cool/9bph-kj5p" name="Institute of Zoology and Botany" type="">IZBE</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="diagnosis">
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<paragraph pageId="0" pageNumber="69">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<taxonomicName authorityName="Kurina" authorityYear="2021" class="Insecta" family="Mycetophilidae" genus="Sciophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sciophila georgei" order="Diptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="georgei">
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<emphasis italics="true" pageId="0" pageNumber="69">Sciophila georgei</emphasis>
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</taxonomicName>
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sp. nov. can be distinguished by combination of the characters of the male terminalia as follows: lateral branch of gonostylus ventrally with two apical spine-like setae, small internal branch of gonostylus with one spine-like seta, tergite 9 large with medially rising apical margin that bears two large and simple setae, parameres straight and long, extending over tergite 9 apically, aedeagus small, star-shaped.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="description">
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<paragraph pageId="0" pageNumber="69">Description.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<emphasis bold="true" pageId="0" pageNumber="69">Male.</emphasis>
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Body length
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.7-2.8 mm</quantity>
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(n = 2).
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<emphasis bold="true" pageId="0" pageNumber="69">
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<emphasis italics="true" pageId="0" pageNumber="69">Coloration</emphasis>
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.
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</emphasis>
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Head with vertex and frons dark brown, face and clypeus brown and mouthparts including palpus pale yellow. Scape and pedicel yellow. First three or four flagellomeres yellowish, rest of flagellomeres light brown. Scutum entirely dark brown, antepronotum and proepisternum yellowish, anepisternum, anepimeron and katepisternum light brown, laterotergite and mediotergite brown, scutellum brown. Thoracic setae all yellowish. Wing hyaline, all veins brown including radial veins somewhat darker. Halter with stem and knob pale yellow. All coxae, femora and tibiae yellow, tarsi yellow but seem darker because of dense brown setae. Tibial setae brown, spurs yellowish. Abdomen with tergites light brown, 1-3 tergites somewhat lighter, all sternites yellowish. Abdominal vestiture yellow. Terminalia brown.
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<emphasis bold="true" pageId="0" pageNumber="69">
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<emphasis italics="true" pageId="0" pageNumber="69">Head</emphasis>
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.
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</emphasis>
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Ocelli in a shallow triangular arrangement. Medial
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<taxonomicName authorityName="Walker" authorityYear="1848" class="Insecta" family="Mycetophilidae" genus="Mycetophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mycetophila ocellus" order="Diptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="ocellus">Mycetophila ocellus</taxonomicName>
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somewhat smaller than laterals. Frontal furrow complete. Clypeus subrounded, about as long as broad. Fourth flagellar segment about as long as wide, apical flagellar segment 2.25 times as long as wide basally. Flagellar segments with dense yellowish short setae.
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<emphasis bold="true" pageId="0" pageNumber="69">
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<emphasis italics="true" pageId="0" pageNumber="69">Thorax</emphasis>
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.
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</emphasis>
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Scutum covered with short setae, with marginal and prescutellar setae stronger. Antepronotum with 8-9 setae. Proepisternum with 6-7 setae. Anepisternum with 5-6 setae on upper part, katepisternum and anepimeron non-setose. Laterotergite with 7-9 setae on posterior half. Mediotergite with 10-15 setae on lower part. Metepisternum with setulae. Scutellum with setulae and marginal setae not arranged in pairs.
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<emphasis bold="true" pageId="0" pageNumber="69">
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<emphasis italics="true" pageId="0" pageNumber="69">Wing</emphasis>
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.
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</emphasis>
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Length
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5-2.8 mm</quantity>
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, length to width 2.4-2.7. Wing membrane uniformly covered with micro- and macrothichia. All veins setose, except
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<emphasis italics="true" pageId="0" pageNumber="69">sc-r</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="69">Rs</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="69">
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<collectionCode country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
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2+3
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</emphasis>
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. Costa reaches about one fifth from
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<emphasis italics="true" pageId="0" pageNumber="69">
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<collectionCode country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
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4+5
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</emphasis>
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to
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<emphasis italics="true" pageId="0" pageNumber="69">M1</emphasis>
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.
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<emphasis italics="true" pageId="0" pageNumber="69">Sc</emphasis>
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ending on
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<emphasis italics="true" pageId="0" pageNumber="69">C</emphasis>
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before level of furcation of posterior fork.
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<emphasis italics="true" pageId="0" pageNumber="69">Sc-r</emphasis>
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located slightly before
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<emphasis italics="true" pageId="0" pageNumber="69">Rs. r-m</emphasis>
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about two times as long as
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<emphasis italics="true" pageId="0" pageNumber="69">m-stem</emphasis>
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.
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<emphasis italics="true" pageId="0" pageNumber="69">M4</emphasis>
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basally very faint or shortly interrupted at base.
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<emphasis bold="true" pageId="0" pageNumber="69">
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<emphasis italics="true" pageId="0" pageNumber="69">Legs</emphasis>
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.
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</emphasis>
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Ratio of femur to tibia for fore, mid and hind legs: 0.83-0.93; 0.89-0.97; 0.84-0.92. Ratio of tibia to basitarsus for fore, mid and hind legs: 1.26; 1.42-1.65; 1.33-1.37. Fore tibia with a spur 2.29-2.81 times of tibial maximum width. Mid tibia with anterior spur 3.08-3.15 times and posterior spur 3.42-3.69 times of tibial maximum width. Hind tibia with anterior spur 2.50-2.60 times and posterior spur 3.47-3.57 times of tibial maximum width.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Terminalia</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Sciophila georgei sp. nov., male terminalia A dorsal view B ventral view C lateral view D apical part of tergite IX, dorsal view E dorsal view, tergite IX removed F aedeagal complex, dorsal view G gonostylus, internal view H gonostylus, posterior view. Abbreviations: aed = aedeagus, cer = cercus, gc = gonocoxite, gc ap = gonocoxal apodeme, gst lbr = lateral branch of gonostylus, gst mbr = medial branch of gonostylus, gst sibr = small internal branch of gonostylus, par = paramere, tg 9 = tergite IX. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1026.63749.figure4" httpUri="https://binary.pensoft.net/fig/524675" pageId="0" pageNumber="69">4A-G</figureCitation>
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). Gonocoxites fused for short distance ventrobasally forming medial triangular lobe with medial more sclerotized longitudinal ridge internally. Ventromedial margin of gonocoxite with a membranous flange drawn medially out into digitate apically hooked lobe. Gonocoxite covered with uniform setae except non-setose lateroapical and dorsomedial marginal areas. Dorsoposterior margin of gonocoxite with two prominent medially directed setae. Gonocoxal apodeme anteriorly enlarged, shoe-shaped, well discernible in dorsal view. Tergite 9 large, slightly convergent posteriorly, extending over gonocoxites, subapically constricted with two prominent simple setae apically, apical margin medially rising. Parameres long and straight, apically slightly widening, extending over tergite 9 apically. Aedeagus small, star-shaped, medially with posteriorly projecting digitate process. Lateral branch of the gonostylus laterally setose with aggregation of spine like setae along posterior margin; ventral part extended with two prominent apical spine-like setae. Medial branch of gonostylus with 25-30 furcated megasetae. Small internal branch of gonostylus with one prominent medially directed seta.
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</paragraph>
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<caption doi="10.3897/zookeys.1026.63749.figure3" httpUri="https://binary.pensoft.net/fig/524674" pageId="0" pageNumber="69" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="69">
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<emphasis bold="true" pageId="0" pageNumber="69">Figure 3.</emphasis>
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Habitus photos of new species
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<emphasis bold="true" pageId="0" pageNumber="69">A</emphasis>
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<taxonomicName authorityName="Kurina" authorityYear="2021" class="Insecta" family="Mycetophilidae" genus="Sciophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sciophila georgei" order="Diptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="georgei">
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<emphasis italics="true" pageId="0" pageNumber="69">Sciophila georgei</emphasis>
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</taxonomicName>
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sp. nov.,
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<typeStatus>paratype</typeStatus>
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<emphasis bold="true" pageId="0" pageNumber="69">B</emphasis>
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<taxonomicName authorityName="Kurina" authorityYear="2021" class="Insecta" family="Mycetophilidae" genus="Anatella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anatella metae" order="Diptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="metae">
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<emphasis italics="true" pageId="0" pageNumber="69">Anatella metae</emphasis>
|
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</taxonomicName>
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sp. nov.,
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<typeStatus>paratype</typeStatus>
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<emphasis bold="true" pageId="0" pageNumber="69">C</emphasis>
|
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<taxonomicName authorityName="Kurina" authorityYear="2021" class="Insecta" family="Mycetophilidae" genus="Leia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leia katae" order="Diptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="katae">
|
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<emphasis italics="true" pageId="0" pageNumber="69">Leia katae</emphasis>
|
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</taxonomicName>
|
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sp. nov.,
|
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<typeStatus>holotype</typeStatus>
|
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, terminalia detached. Scale bars:
|
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
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.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="69">
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<emphasis bold="true" pageId="0" pageNumber="69">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="69" type="etymology">
|
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<paragraph pageId="0" pageNumber="69">Etymology.</paragraph>
|
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<paragraph pageId="0" pageNumber="69">
|
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<materialsCitation ID-GBIF-Occurrence="3062025367" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Georgia" location="The" municipality="He" specimenCount="1" stateProvince="Tbilisi">
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<location LSID="urn:lsid:plazi:treatment:B92821CC83945DDE8E2F3199603BAFBC:7388691030835949BD5126EB671BE569" country="Georgia" municipality="He" name="The" stateProvince="Tbilisi">The</location>
|
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species is named in honour of Prof. George Japoshvili (
|
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<collectingRegion country="Georgia" name="Tbilisi">Tbilisi</collectingRegion>
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,
|
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<collectingCountry name="Georgia">Georgia</collectingCountry>
|
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) in recognition of his contribution to study of the
|
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<normalizedToken originalValue="insects’">insects'</normalizedToken>
|
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diversity in
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<collectingCountry name="Georgia">Georgia</collectingCountry>
|
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and his invaluable help in collecting the fungus gnat material that underlies the current communication.
|
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<collectingMunicipality>He</collectingMunicipality>
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was also the collector of the
|
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<typeStatus>type</typeStatus>
|
||
material of this species
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1026.63749.figure4" httpUri="https://binary.pensoft.net/fig/524675" pageId="0" pageNumber="69" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Kurina" authorityYear="2021" class="Insecta" family="Mycetophilidae" genus="Sciophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sciophila georgei" order="Diptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="georgei">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Sciophila georgei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male terminalia
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="69">B</emphasis>
|
||
ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="69">C</emphasis>
|
||
lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="69">D</emphasis>
|
||
apical part of tergite IX, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="69">E</emphasis>
|
||
dorsal view, tergite IX removed
|
||
<emphasis bold="true" pageId="0" pageNumber="69">F</emphasis>
|
||
aedeagal complex, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="69">G</emphasis>
|
||
gonostylus, internal view
|
||
<emphasis bold="true" pageId="0" pageNumber="69">H</emphasis>
|
||
gonostylus, posterior view. Abbreviations: aed = aedeagus, cer = cercus, gc = gonocoxite, gc ap = gonocoxal apodeme, gst lbr = lateral branch of gonostylus, gst mbr = medial branch of gonostylus, gst sibr = small internal branch of gonostylus, par = paramere, tg 9 = tergite IX. Scale bars:
|
||
<quantity metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="remarks">
|
||
<paragraph pageId="0" pageNumber="69">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
More than 190 species of
|
||
<taxonomicName class="Insecta" family="Mycetophilidae" genus="Sciophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sciophila" order="Diptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Sciophila</emphasis>
|
||
</taxonomicName>
|
||
Meigen are known wordwide (
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4722.4.3" author="Kurina, O" journalOrPublisher="Zootaxa" pageId="0" pageNumber="69" pagination="339 - 351" refId="B30" refString="Kurina, O, 2020a. New species and new records of Sciophila Meigen (Diptera: Mycetophilidae) from the Afrotropical region. Zootaxa 4722 (4): 339 - 351, DOI: https://doi.org/10.11646/zootaxa.4722.4.3" title="New species and new records of Sciophila Meigen (Diptera: Mycetophilidae) from the Afrotropical region." url="https://doi.org/10.11646/zootaxa.4722.4.3" volume="4722" year="2020 a">Kurina 2020a</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.3958/059.045.0428" author="Taber, SW" journalOrPublisher="Biometrics" pageId="0" pageNumber="69" refId="B65" refString="Taber, SW, 2021. Eleven new Nearctic species of Sciophila Meigen fungus gnats (Diptera: Mycetophilidae), Southwestern Entomologist 45(4) (2020): 1121-1148. https://doi.org/10.3958/059.045.0428" title="Eleven new Nearctic species of Sciophila Meigen fungus gnats (Diptera: Mycetophilidae), Southwestern Entomologist 45 (4) (2020): 1121 - 1148." url="https://doi.org/10.3958/059.045.0428" year="2021">Taber 2021</bibRefCitation>
|
||
); the most comprehensive key to the Holarctic species is still that by
|
||
<bibRefCitation author="Zaitzev, AI" journalOrPublisher="Nauka, Moscow" pageId="0" pageNumber="69" refId="B70" refString="Zaitzev, AI, 1982. Fungus gnats of the genus Sciophila Meig. of the Holarctic. Nauka, Moscow" title="Fungus gnats of the genus Sciophila Meig. of the Holarctic." year="1982">Zaitzev (1982)</bibRefCitation>
|
||
. Fortunately, all subsequently described species are supplemented with appropriate illustrations of the male terminalia (e.g.
|
||
<bibRefCitation author="Polevoi, AV" journalOrPublisher="Entomological Review" pageId="0" pageNumber="69" pagination="207 - 214" refId="B49" refString="Polevoi, AV, 2001. New and little known species of the fungus gnat subfamilies Mycomyinae and Sciophilinae (Diptera, Mycetophilidae) from Eastern Fennoscandia. Entomological Review 81 (2): 207 - 214" title="New and little known species of the fungus gnat subfamilies Mycomyinae and Sciophilinae (Diptera, Mycetophilidae) from Eastern Fennoscandia." volume="81" year="2001">Polevoi 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.3897/BDJ.5.e11760" author="Salmela, J" journalOrPublisher="Entomologica Fennica" pageId="0" pageNumber="69" refId="B54" refString="Salmela, J, Kolcsar, L-P, 2017. New and poorly known Palaearctic fungus gnats (Diptera, Sciaroidea). Biodiversity Data Journal 5: e11760. https://doi.org/10.3897/BDJ.5.e11760" title="New and poorly known Palaearctic fungus gnats (Diptera, Sciaroidea). Biodiversity Data Journal 5: e 11760." url="https://doi.org/10.3897/BDJ.5.e11760" year="2017">
|
||
Salmela and
|
||
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
|
||
2017
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.3958/059.045.0428" author="Taber, SW" journalOrPublisher="Biometrics" pageId="0" pageNumber="69" refId="B65" refString="Taber, SW, 2021. Eleven new Nearctic species of Sciophila Meigen fungus gnats (Diptera: Mycetophilidae), Southwestern Entomologist 45(4) (2020): 1121-1148. https://doi.org/10.3958/059.045.0428" title="Eleven new Nearctic species of Sciophila Meigen fungus gnats (Diptera: Mycetophilidae), Southwestern Entomologist 45 (4) (2020): 1121 - 1148." url="https://doi.org/10.3958/059.045.0428" year="2021">Taber 2021</bibRefCitation>
|
||
) that provides an adequate compendium of the morphological distinctions. Following the key by
|
||
<bibRefCitation author="Zaitzev, AI" journalOrPublisher="Nauka, Moscow" pageId="0" pageNumber="69" refId="B70" refString="Zaitzev, AI, 1982. Fungus gnats of the genus Sciophila Meig. of the Holarctic. Nauka, Moscow" title="Fungus gnats of the genus Sciophila Meig. of the Holarctic." year="1982">Zaitzev (1982)</bibRefCitation>
|
||
, the new species runs to couplet 31 because of (1) wing with both macro- and microtrichia, (2) gonostylus without additional branches, (3) lateral branch of the gonostylus with two large setae ventroapically, (4) small internal branch of the gonostylus with one large seta, and (5) gonocoxites dorsoapically without protruding appendages. However,
|
||
<taxonomicName lsidName="S. georgei" pageId="0" pageNumber="69" rank="species" species="georgei">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. georgei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from the species included in this couplet by details in the male terminalia. Notably, by the characters of tergite 9 (large, posteriorly convergent, extends over gonocoxites, bears two prominent simple setae apically, apical margin medially rising) and parameres (long, straight, extending over tergite 9 apically).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |