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<document id="BB96F090E990E144F67A5E33D5794413" ID-DOI="10.11646/zootaxa.4027.4.1" ID-GBIF-Dataset="7e77f875-e8de-478e-a96b-19d376e00e3a" ID-ISSN="1175-5326" ID-Zenodo-Dep="234517" ID-ZooBank="3B32D183-45BA-41F9-82A9-C9C108D53899" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461283140705" checkinUser="plazi" docAuthor="Galil, Bella S. &amp; Ng, Peter K. L." docDate="2015" docId="8870B305E33A4C1C9699FEB288AC8BB5" docLanguage="en" docName="zt04027p486.pdf" docOrigin="Zootaxa 4027 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ryphila bertrandi Galil &amp; Ng, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="462" masterDocId="7449CB7DE3314C17960EFFDF8C528D68" masterDocTitle="Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="486" masterPageNumber="451" pageNumber="462" updateTime="1698614571376" updateUser="plazi">
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<mods:title id="FA0F2AE108367B278AECE09073CC6DF4">Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:name id="77B7EC2E07C4293C08F8E948DFE0FA22" type="personal">
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<mods:namePart id="72AAB01C7DDD11BC7FB8F724CADCEB44">Galil, Bella S.</mods:namePart>
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<mods:namePart id="0D70E0D2E5B9719705DFF5FFA9EC3940">Ng, Peter K. L.</mods:namePart>
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<mods:title id="CE8B6451497EC1C3D679013082F97BDF">Zootaxa</mods:title>
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<mods:date id="01875E62ECA2754B185883CB1D22849D">2015</mods:date>
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<treatment id="8870B305E33A4C1C9699FEB288AC8BB5" ID-DOI="http://doi.org/10.5281/zenodo.6102262" ID-GBIF-Taxon="119640942" ID-Zenodo-Dep="6102262" LSID="urn:lsid:plazi:treatment:8870B305E33A4C1C9699FEB288AC8BB5" httpUri="http://treatment.plazi.org/id/8870B305E33A4C1C9699FEB288AC8BB5" lastPageNumber="462" pageId="11" pageNumber="462">
<subSubSection id="48C35198E33A4C1C9699FEB28D1F8CC0" pageId="11" pageNumber="462" type="nomenclature">
<paragraph id="00660213E33A4C1C9699FEB28D838CEF" blockId="11.[151,465,365,424]" box="[151,465,365,391]" pageId="11" pageNumber="462">
<heading id="5B2EB57FE33A4C1C9699FEB28D838CEF" bold="true" box="[151,465,365,391]" fontSize="11" level="1" pageId="11" pageNumber="462" reason="1">
<emphasis id="32ADDE01E33A4C1C9699FEB28D838CEF" bold="true" box="[151,465,365,391]" pageId="11" pageNumber="462">
<taxonomicName id="C7D97990E33A4C1C9699FEB28D3E8CEF" ID-CoL="6WZ7K" box="[151,364,365,391]" class="Malacostraca" family="Leucosiidae" genus="Ryphila" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="bertrandi" status="sp. nov.">
<emphasis id="32ADDE01E33A4C1C9699FEB28D3E8CEF" bold="true" box="[151,364,365,391]" italics="true" pageId="11" pageNumber="462">Ryphila bertrandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="299E637AE33A4C1C977DFEB28D838CEF" box="[371,465,365,391]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="00660213E33A4C1C9699FE508D1F8CC0" blockId="11.[151,465,365,424]" box="[151,333,399,424]" pageId="11" pageNumber="462">
(
<figureCitation id="98E21E96E33A4C1C9691FE508CBF8CC0" box="[159,237,399,424]" captionStart="FIGURE 4" captionStartId="10.[151,250,1786,1808]" captionTargetBox="[151,1436,193,1762]" captionTargetId="figure@10.[151,1436,193,1764]" captionTargetPageId="10" captionText="FIGURE 4. Colours in life. A, Nursilia dentata, male (7.2 mm) (ZRC 2014.0165), stn PD 75; B, Parilia major, immature male (35.1 mm) (MNHN-IU- 2013 - 1044), stn CP 3968; C, Raylilia coniculifera, ovigerous female (10.5 mm) (ZRC 2014.0169), stn PD 45; D, Raylilia uenoi, young female (10.4 mm) (MNHN-IU- 2013 - 117), stn PR 07; E, Ryphila bertrandi sp. nov., paratype ovigerous female (12.5 mm) (MNHN-IU- 2013 - 12750), stn PD 68; F, Tanaoa kuka sp. nov., holotype male (33.0 mm) (MNHN- IU- 2013 - 7963), stn CP 3984." httpUri="https://zenodo.org/record/234521/files/figure.png" pageId="11" pageNumber="462">Figs. 4</figureCitation>
E, 9, 10)
</paragraph>
</subSubSection>
<subSubSection id="48C35198E33A4C1C9699FE088F618F34" pageId="11" pageNumber="462" type="materials_examined">
<paragraph id="00660213E33A4C1C9699FE088F618F34" blockId="11.[151,1437,471,1757]" pageId="11" pageNumber="462">
<emphasis id="32ADDE01E33A4C1C9699FE088D288C98" bold="true" box="[151,378,471,496]" pageId="11" pageNumber="462">Material examined</emphasis>
.
<typeStatus id="DF62BCB1E33A4C1C9784FE088DA58C98" box="[394,503,471,496]" pageId="11" pageNumber="462" type="holotype">Holotype</typeStatus>
: female (
<quantity id="C721AFF6E33A4C1C9450FE088E978C98" box="[606,709,471,496]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="11" pageNumber="462" unit="mm" value="11.5">11.5 mm</quantity>
) (MNHN-IU-2013-12749), stn PD72, between Yabob &amp; Bil Bil I.,
<geoCoordinate id="65ED64D4E33A4C1C96BBFE238D708F7D" box="[181,290,508,533]" direction="south" orientation="latitude" pageId="11" pageNumber="462" precision="92" value="-5.27">05°16.2'S</geoCoordinate>
<geoCoordinate id="65ED64D4E33A4C1C9726FE238DF58F7D" box="[296,423,508,533]" direction="east" orientation="longitude" pageId="11" pageNumber="462" precision="92" value="145.775">145°46.5'E</geoCoordinate>
,
<quantity id="C721AFF6E33A4C1C97BCFE238E458F7C" box="[434,535,508,533]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="11" pageNumber="462" unit="m" value="22.5" valueMax="25.0" valueMin="20.0">2025 m</quantity>
,
<date id="746724D3E33A4C1C942DFE238EC48F7D" box="[547,662,508,533]" pageId="11" pageNumber="462" value="2012-06-12">6.12.2012</date>
.
<typeStatus id="DF62BCB1E33A4C1C94AEFE228F408F7D" box="[672,786,509,533]" pageId="11" pageNumber="462" type="paratype">Paratypes</typeStatus>
: 1 ovigerous female (
<quantity id="C721AFF6E33A4C1C920EFE23880B8F7C" box="[1024,1113,508,533]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="11" pageNumber="462" unit="mm" value="9.6">9.6 mm</quantity>
) (
<collectionCode id="66C89AD6E33A4C1C9260FE2388F48F7C" box="[1134,1190,508,532]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="462">ZRC</collectionCode>
2015.270), same data as
<typeStatus id="DF62BCB1E33A4C1C96B9FDC08D4F8F50" box="[183,285,543,568]" pageId="11" pageNumber="462" type="holotype">holotype</typeStatus>
; 1 ovigerous female (
<quantity id="C721AFF6E33A4C1C941BFDC08E2F8F50" box="[533,637,543,568]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="462" unit="mm" value="12.5">12.5 mm</quantity>
) (MNHN-IU-2013-12750), stn PD68, south Kesup Gun River mouth,
<geoCoordinate id="65ED64D4E33A4C1C9699FD9B8CBC8F35" box="[151,238,580,605]" direction="south" orientation="latitude" pageId="11" pageNumber="462" precision="925" value="-5.266667">05°16'S</geoCoordinate>
<geoCoordinate id="65ED64D4E33A4C1C96FBFD9B8D278F35" box="[245,373,580,605]" direction="east" orientation="longitude" pageId="11" pageNumber="462" precision="92" value="145.775">145°46.5'E</geoCoordinate>
,
<quantity id="C721AFF6E33A4C1C978EFD9B8D9B8F34" box="[384,457,580,604]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="11" pageNumber="462" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">23 m</quantity>
,
<collectingCountry id="78CE4283E33A4C1C97DBFD9A8EE28F35" box="[469,688,580,605]" name="Papua New Guinea" pageId="11" pageNumber="462">Papua New Guinea</collectingCountry>
,
<date id="746724D3E33A4C1C94B5FD9B8F7D8F34" box="[699,815,580,604]" pageId="11" pageNumber="462" value="2012-04-12">4.12.2012</date>
.
</paragraph>
</subSubSection>
<subSubSection id="48C35198E33A4C1C96C9FDB8886A88D4" pageId="11" pageNumber="462" type="description">
<paragraph id="00660213E33A4C1C96C9FDB88DF18EAD" blockId="11.[151,1437,471,1757]" pageId="11" pageNumber="462">
<emphasis id="32ADDE01E33A4C1C96C9FDB88D038FE8" bold="true" box="[199,337,615,640]" pageId="11" pageNumber="462">Description</emphasis>
. Carapace discoidal, with shallow branchio-cardiac grooves (
<figureCitation id="98E21E96E33A4C1C9220FDB888258FE8" box="[1070,1143,615,640]" captionStart="FIGURE 9" captionStartId="17.[151,250,1554,1576]" captionTargetBox="[151,1436,193,1531]" captionTargetId="figure@17.[151,1436,193,1533]" captionTargetPageId="17" captionText="FIGURE 9. Ryphila bertrandi sp. nov. A, C F, holotype female (11.5 mm) (MNHN-IU- 2013 - 12749), stn PD 72; B, paratype ovigerous female (9.6 mm) (ZRC 2015.270), stn PD 72. A, B, overall habitus; C, frontal margin and orbits; D, frontal view of cephalothorax and chelipeds; E, left lateral view of cephalothorax; F, third maxillipeds." httpUri="https://zenodo.org/record/234526/files/figure.png" pageId="11" pageNumber="462">Fig. 9</figureCitation>
A, B). Dorsal surface of carapace minutely granular anteriorly, with closely-spaced pearliform granules on branchial, cardial, intestinal regions. Frontal margin horizontal, setose (
<figureCitation id="98E21E96E33A4C1C9493FD708EBA8FA0" box="[669,744,687,712]" captionStart="FIGURE 9" captionStartId="17.[151,250,1554,1576]" captionTargetBox="[151,1436,193,1531]" captionTargetId="figure@17.[151,1436,193,1533]" captionTargetPageId="17" captionText="FIGURE 9. Ryphila bertrandi sp. nov. A, C F, holotype female (11.5 mm) (MNHN-IU- 2013 - 12749), stn PD 72; B, paratype ovigerous female (9.6 mm) (ZRC 2015.270), stn PD 72. A, B, overall habitus; C, frontal margin and orbits; D, frontal view of cephalothorax and chelipeds; E, left lateral view of cephalothorax; F, third maxillipeds." httpUri="https://zenodo.org/record/234526/files/figure.png" pageId="11" pageNumber="462">Fig. 9</figureCitation>
C). Margins of carapace closely beaded, granules smaller anteriorly (
<figureCitation id="98E21E96E33A4C1C9717FD0B8D318F84" box="[281,355,724,749]" captionStart="FIGURE 9" captionStartId="17.[151,250,1554,1576]" captionTargetBox="[151,1436,193,1531]" captionTargetId="figure@17.[151,1436,193,1533]" captionTargetPageId="17" captionText="FIGURE 9. Ryphila bertrandi sp. nov. A, C F, holotype female (11.5 mm) (MNHN-IU- 2013 - 12749), stn PD 72; B, paratype ovigerous female (9.6 mm) (ZRC 2015.270), stn PD 72. A, B, overall habitus; C, frontal margin and orbits; D, frontal view of cephalothorax and chelipeds; E, left lateral view of cephalothorax; F, third maxillipeds." httpUri="https://zenodo.org/record/234526/files/figure.png" pageId="11" pageNumber="462">Fig. 9</figureCitation>
E). Hepatic facet distinct. Pterygostomian region prominently granular (
<figureCitation id="98E21E96E33A4C1C92B7FD0B89508F84" box="[1209,1282,724,749]" captionStart="FIGURE 9" captionStartId="17.[151,250,1554,1576]" captionTargetBox="[151,1436,193,1531]" captionTargetId="figure@17.[151,1436,193,1533]" captionTargetPageId="17" captionText="FIGURE 9. Ryphila bertrandi sp. nov. A, C F, holotype female (11.5 mm) (MNHN-IU- 2013 - 12749), stn PD 72; B, paratype ovigerous female (9.6 mm) (ZRC 2015.270), stn PD 72. A, B, overall habitus; C, frontal margin and orbits; D, frontal view of cephalothorax and chelipeds; E, left lateral view of cephalothorax; F, third maxillipeds." httpUri="https://zenodo.org/record/234526/files/figure.png" pageId="11" pageNumber="462">Fig. 9</figureCitation>
F). Epimeral, posterior surfaces minutely granular. Antennular fossa sealed by basal antennular article. Antennae short, inserted between antennular fossa, orbit. Eyes retractable. Epistome continuous with expanded, beaded subhepatic margin, projecting beyond frontal margin, medially notched. Anterior margin of efferent branchial channel forms part of lower orbital margin, entire, fissure situated ventrally to orbit. Third maxilliped exopod semicircular (
<figureCitation id="98E21E96E33A4C1C9338FCBB892E8E14" box="[1334,1404,868,893]" captionStart="FIGURE 9" captionStartId="17.[151,250,1554,1576]" captionTargetBox="[151,1436,193,1531]" captionTargetId="figure@17.[151,1436,193,1533]" captionTargetPageId="17" captionText="FIGURE 9. Ryphila bertrandi sp. nov. A, C F, holotype female (11.5 mm) (MNHN-IU- 2013 - 12749), stn PD 72; B, paratype ovigerous female (9.6 mm) (ZRC 2015.270), stn PD 72. A, B, overall habitus; C, frontal margin and orbits; D, frontal view of cephalothorax and chelipeds; E, left lateral view of cephalothorax; F, third maxillipeds." httpUri="https://zenodo.org/record/234526/files/figure.png" pageId="11" pageNumber="462">Fig. 9</figureCitation>
F), expanded, densely setose; endopod triangular, merus densely setose, shorter than ischium along inner margin, apex invisible in dorsal view.
</paragraph>
<paragraph id="00660213E33A4C1C96C9FC10894B893D" blockId="11.[151,1437,471,1757]" pageId="11" pageNumber="462">
Chelipeds subequal. Cheliped merus 0.6 as long as carapace in female; entirely covered with pearliform granules, decreasing in size distally (
<figureCitation id="98E21E96E33A4C1C9434FC2B8ED98964" box="[570,651,1012,1037]" captionStart="FIGURE 10" captionStartId="18.[151,250,1938,1960]" captionTargetBox="[171,1414,403,1896]" captionTargetId="figure@18.[171,1415,403,1897]" captionTargetPageId="18" captionText="FIGURE 10. Ryphila bertrandi sp. nov., holotype female (11.5 mm) (MNHN-IU- 2013 - 12749), stn PD 72. A, inner view of right chela; B, dorsal view of right cheliped merus; C, anterior view of left cheliped merus; D, thoracic sternum and vulvae; E, ventral surfaces, thoracic sternum and abdomen." httpUri="https://zenodo.org/record/234527/files/figure.png" pageId="11" pageNumber="462">Fig. 10</figureCitation>
AC). Carpus with row of minute granules on inner margin; patch of minute granules distally on dorsal margin. Dorsal margin of chela cristate, with 2 parallel rows of granules, ventral margin minutely granular. Fingers slightly longer than palm, cutting edges cristate, distally denticulate.
</paragraph>
<paragraph id="00660213E33A4C1C96C9FB808C9589A8" blockId="11.[151,1437,471,1757]" pageId="11" pageNumber="462">Ambulatory legs slender, short, meri subcylindrical, longer than carpi, propodi, ventral margin minutely granular; dorsal, ventral propodal margins carinate; dactyli longer than propodi, lanceolate, terminating in cornute tips.</paragraph>
<paragraph id="00660213E33A4C1C96C9FB138DB58838" blockId="11.[151,1437,471,1757]" pageId="11" pageNumber="462">
Female abdomen with first 2 somites transversely narrow, yoke-like, granular; somites 36 fused, greatly enlarged, shield-like, proximally granular; telson laciniate (
<figureCitation id="98E21E96E33A4C1C955AFB308FFE8860" box="[852,940,1263,1288]" captionStart="FIGURE 10" captionStartId="18.[151,250,1938,1960]" captionTargetBox="[171,1414,403,1896]" captionTargetId="figure@18.[171,1415,403,1897]" captionTargetPageId="18" captionText="FIGURE 10. Ryphila bertrandi sp. nov., holotype female (11.5 mm) (MNHN-IU- 2013 - 12749), stn PD 72. A, inner view of right chela; B, dorsal view of right cheliped merus; C, anterior view of left cheliped merus; D, thoracic sternum and vulvae; E, ventral surfaces, thoracic sternum and abdomen." httpUri="https://zenodo.org/record/234527/files/figure.png" pageId="11" pageNumber="462">Fig. 10</figureCitation>
E). Thoracic sternites with closely spaced pearliform granules. Vulvae submedially near suture 6/5; small, rounded, opening interiorly, hooded by calcified pear-shaped dome (
<figureCitation id="98E21E96E33A4C1C977CFAE88D918838" box="[370,451,1335,1360]" captionStart="FIGURE 10" captionStartId="18.[151,250,1938,1960]" captionTargetBox="[171,1414,403,1896]" captionTargetId="figure@18.[171,1415,403,1897]" captionTargetPageId="18" captionText="FIGURE 10. Ryphila bertrandi sp. nov., holotype female (11.5 mm) (MNHN-IU- 2013 - 12749), stn PD 72. A, inner view of right chela; B, dorsal view of right cheliped merus; C, anterior view of left cheliped merus; D, thoracic sternum and vulvae; E, ventral surfaces, thoracic sternum and abdomen." httpUri="https://zenodo.org/record/234527/files/figure.png" pageId="11" pageNumber="462">Fig. 10</figureCitation>
D).
</paragraph>
<paragraph id="00660213E33A4C1C96C9FA83886A88D4" blockId="11.[151,1437,471,1757]" pageId="11" pageNumber="462">
<emphasis id="32ADDE01E33A4C1C96C9FA838D3A881D" bold="true" box="[199,360,1372,1397]" pageId="11" pageNumber="462">Colour in life</emphasis>
. Dorsal surface of carapace and chelipeds brown, granules bright orange-red, pale blotches on post orbital regions, and on both sides of cardiac region. Rounded knob distally on cheliped merus bright red, finger tips white. Ambulatory legs pale, spotted red on merus and carpus (
<figureCitation id="98E21E96E33A4C1C95DAFA7B884A88D4" box="[980,1048,1444,1469]" captionStart="FIGURE 4" captionStartId="10.[151,250,1786,1808]" captionTargetBox="[151,1436,193,1762]" captionTargetId="figure@10.[151,1436,193,1764]" captionTargetPageId="10" captionText="FIGURE 4. Colours in life. A, Nursilia dentata, male (7.2 mm) (ZRC 2014.0165), stn PD 75; B, Parilia major, immature male (35.1 mm) (MNHN-IU- 2013 - 1044), stn CP 3968; C, Raylilia coniculifera, ovigerous female (10.5 mm) (ZRC 2014.0169), stn PD 45; D, Raylilia uenoi, young female (10.4 mm) (MNHN-IU- 2013 - 117), stn PR 07; E, Ryphila bertrandi sp. nov., paratype ovigerous female (12.5 mm) (MNHN-IU- 2013 - 12750), stn PD 68; F, Tanaoa kuka sp. nov., holotype male (33.0 mm) (MNHN- IU- 2013 - 7963), stn CP 3984." httpUri="https://zenodo.org/record/234521/files/figure.png" pageId="11" pageNumber="462">Fig. 4</figureCitation>
E).
</paragraph>
</subSubSection>
<subSubSection id="48C35198E33A4C1C96C9FA1888D78B18" pageId="11" pageNumber="462" type="discussion">
<paragraph id="00660213E33A4C1C96C9FA1888D78B18" blockId="11.[151,1437,471,1757]" pageId="11" pageNumber="462">
<emphasis id="32ADDE01E33A4C1C96C9FA188D668888" bold="true" box="[199,308,1479,1504]" pageId="11" pageNumber="462">Remarks</emphasis>
.
<taxonomicName id="C7D97990E33A4C1C974AFA188E5F8888" box="[324,525,1479,1504]" class="Malacostraca" family="Leucosiidae" genus="Ryphila" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="bertrandi" status="sp. nov.">
<emphasis id="32ADDE01E33A4C1C974AFA188E5F8888" box="[324,525,1479,1504]" italics="true" pageId="11" pageNumber="462">Ryphila bertrandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="299E637AE33A4C1C9418FA188E208888" box="[534,626,1479,1504]" pageId="11" pageNumber="462" rank="species">
<emphasis id="32ADDE01E33A4C1C9418FA188E208888" bold="true" box="[534,626,1479,1504]" pageId="11" pageNumber="462">sp. nov.</emphasis>
</taxonomicNameLabel>
shares with it congeners
<taxonomicName id="C7D97990E33A4C1C9599FA1688828888" authority="Herbst, 1783" authorityName="Herbst" authorityYear="1783" box="[919,1232,1479,1504]" class="Malacostraca" family="Leucosiidae" genus="Ryphila" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="cancellus">
<emphasis id="32ADDE01E33A4C1C9599FA1688708888" box="[919,1058,1479,1504]" italics="true" pageId="11" pageNumber="462">R. cancellus</emphasis>
(Herbst, 1783)
</taxonomicName>
and
<taxonomicName id="C7D97990E33A4C1C9305FA168D3A8B6C" authority="Henderson, 1893" authorityName="Henderson" authorityYear="1893" class="Malacostraca" family="Leucosiidae" genus="Ryphila" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="verrucosa">
<emphasis id="32ADDE01E33A4C1C9305FA1689CE8888" box="[1291,1436,1481,1504]" italics="true" pageId="11" pageNumber="462">R. verrucosa</emphasis>
(Henderson, 1893)
</taxonomicName>
a projecting epistome and ventral margin of hepatic facet and reniform third maxilliped exopod. It is clearly distinguished from both, however, in having cheliped merus entirely covered with pearliform granules dorsally rather than with several rows of pearliform granules, and the carapace dorsally minutely granular anteriorly, with larger pearliform granules on the branchial, cardiac and intestinal regions.
</paragraph>
</subSubSection>
<subSubSection id="48C35198E33A4C1C96C9F9A388AC8BB5" pageId="11" pageNumber="462" type="etymology">
<paragraph id="00660213E33A4C1C96C9F9A38E788BD0" blockId="11.[151,1437,471,1757]" pageId="11" pageNumber="462">
<emphasis id="32ADDE01E33A4C1C96C9F9A38D148BFD" bold="true" box="[199,326,1660,1685]" pageId="11" pageNumber="462">Etymology</emphasis>
. In honour of our good friend, Bertrand Richer de Forges, intrepid explorer and collector of crabs from all over the Indo-West Pacific.
</paragraph>
<paragraph id="00660213E33A4C1C96C9F91B88AC8BB5" blockId="11.[151,1437,471,1757]" box="[199,1278,1732,1757]" pageId="11" pageNumber="462">
<emphasis id="32ADDE01E33A4C1C96C9F91B8DAF8BB5" bold="true" box="[199,509,1732,1757]" pageId="11" pageNumber="462">Geographical distribution</emphasis>
. This species is known only from
<typeStatus id="DF62BCB1E33A4C1C9571F91A8FFD8BB5" box="[895,943,1733,1757]" pageId="11" pageNumber="462">type</typeStatus>
location,
<collectingCountry id="78CE4283E33A4C1C9210F91A88A98BB5" box="[1054,1275,1732,1757]" name="Papua New Guinea" pageId="11" pageNumber="462">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>