464 lines
53 KiB
XML
464 lines
53 KiB
XML
<document id="2EE4C058AD308255EF8A6E0674319EA4" ID-DOI="10.5281/zenodo.189074" ID-GBIF-Dataset="ab03e35b-2874-407a-9071-9d887e0322ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="189074" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461099212284" checkinUser="plazi" docAuthor="Tan, Heok Hui" docDate="2009" docId="5C0887D37F4BE25A61C1FADF590AF821" docLanguage="en" docName="zt02165p068.pdf" docOrigin="Zootaxa 2165" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Betta midas Tan, 2009, new species" docType="treatment" docVersion="7" lastPageNumber="67" masterDocId="A031FFAB7F4EE2526156FFB25B64FD1B" masterDocTitle="Redescription of Betta anabatoides Bleeker, and a new species of Betta from West Kalimantan, Borneo (Teleostei: Osphronemidae)" masterLastPageNumber="68" masterPageNumber="59" pageNumber="64" updateTime="1698590524541" updateUser="plazi">
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<mods:titleInfo id="FBBF46C8FC3E3745B1FF82702FD02F9C">
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<mods:title id="F24861AD2827227F0FE00AB643404281">Redescription of Betta anabatoides Bleeker, and a new species of Betta from West Kalimantan, Borneo (Teleostei: Osphronemidae)</mods:title>
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<mods:name id="55A4F6C321C90EA5FABC1FD3DC20DCCF" type="personal">
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<mods:roleTerm id="E7844F6631AEF53B77BEF7492BCB0EA6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D8D23C6F872E4D15E323F95918B7EBB1">Tan, Heok Hui</mods:namePart>
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<mods:typeOfResource id="3BA4B2E75BA4CC39DC93BEABA24B0E58">text</mods:typeOfResource>
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<mods:title id="4C72A0106365303E360242A16B6975E8">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A3736D8CC1E26AD64C4DC0286B59142E">
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<mods:date id="4DCDF7024B110A8101A09A06AA160BE4">2009</mods:date>
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<mods:detail id="B3C525C6A18B1F11D719E45B595B1AF2" type="volume">
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<mods:number id="D1ECBCA48B9CCCC8C39F035D089B7131">2165</mods:number>
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<mods:start id="83F00DC132F2A7C16F799361DBAD7516">59</mods:start>
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<mods:classification id="1FC90FEE6C8E57BFB385AF579F4B816C">journal article</mods:classification>
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<mods:identifier id="9FA48D99DF02C3F9513D6D5578DC718B" type="DOI">10.5281/zenodo.189074</mods:identifier>
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<mods:identifier id="7858087D671AA4F13C09CF6FC8437CBB" type="GBIF-Dataset">ab03e35b-2874-407a-9071-9d887e0322ba</mods:identifier>
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<mods:identifier id="C245775B38A6CE8865F52C1682ABBB12" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8D033FC3189AA09E08EBEFDA498FB5A2" type="Zenodo-Dep">189074</mods:identifier>
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<treatment id="5C0887D37F4BE25A61C1FADF590AF821" ID-DOI="http://doi.org/10.5281/zenodo.6220716" ID-GBIF-Taxon="119621265" ID-Zenodo-Dep="6220716" LSID="urn:lsid:plazi:treatment:5C0887D37F4BE25A61C1FADF590AF821" httpUri="http://treatment.plazi.org/id/5C0887D37F4BE25A61C1FADF590AF821" lastPageId="8" lastPageNumber="67" pageId="5" pageNumber="64">
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<subSubSection id="9CBB654E7F4BE25761C1FADF5A7CF8B1" pageId="5" pageNumber="64" type="nomenclature">
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<paragraph id="D41E36C57F4BE25761C1FADF5AA7F89C" blockId="5.[151,451,1389,1450]" box="[151,451,1389,1415]" pageId="5" pageNumber="64">
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<heading id="8F5681A97F4BE25761C1FADF5AA7F89C" bold="true" box="[151,451,1389,1415]" fontSize="11" level="1" pageId="5" pageNumber="64" reason="1">
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<emphasis id="E6D5EAD77F4BE25761C1FADF5AA7F89C" bold="true" box="[151,451,1389,1415]" pageId="5" pageNumber="64">
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<taxonomicName id="13A14D467F4BE25761C1FADF5A40F89C" ID-CoL="LNZY" box="[151,292,1389,1415]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="midas" status="sp. nov.">
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<emphasis id="E6D5EAD77F4BE25761C1FADF5A40F89C" bold="true" box="[151,292,1389,1415]" italics="true" pageId="5" pageNumber="64">Betta midas</emphasis>
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="FDE657AC7F4BE2576062FADF5AA7F89C" box="[308,451,1389,1415]" pageId="5" pageNumber="64" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="D41E36C57F4BE25761C1FA225A7CF8B1" blockId="5.[151,451,1389,1450]" box="[151,280,1424,1450]" pageId="5" pageNumber="64">
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||
(
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<figureCitation id="4C9A2A407F4BE25761C9FA225A6BF8B1" box="[159,271,1424,1450]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStartId-0="6.[151,258,713,737]" captionStartId-1="6.[151,255,1377,1401]" captionStartId-2="6.[151,255,1978,2002]" captionStartId-3="7.[151,255,764,788]" captionStartId-4="7.[151,255,1233,1257]" captionTargetBox-0="[151,1434,197,686]" captionTargetBox-1="[151,1434,825,1352]" captionTargetBox-2="[151,1434,1455,1954]" captionTargetBox-3="[151,1434,180,739]" captionTargetBox-4="[151,1435,841,1208]" captionTargetId-0="figure@6.[151,1436,197,688]" captionTargetId-1="figure@6.[151,1436,825,1354]" captionTargetId-2="figure@6.[151,1436,1455,1954]" captionTargetId-3="figure@7.[151,1436,180,740]" captionTargetId-4="figure@7.[151,1436,841,1209]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="7" captionText-0="FIGURE 4. Betta midas, ZRC 46190, 48.3 mm SL, paratype from Sungai Kepayan, West Kalimantan (right side, laterally inverted)." captionText-1="FIGURE 5. Betta midas, not preserved, ca. 35 mm SL, Sungai Kepayan, West Kalimantan." captionText-2="FIGURE 6. Betta midas, ZRC 51527, 66.0 mm SL, paratype from Tayan." captionText-3="FIGURE 7. Betta midas, ZRC 51527, 66.0 mm SL, live paratype from Tayan." captionText-4="FIGURE 8. Betta midas, ZRC 51526, 61.4 mm SL, Sarawak, Lundu." httpUri-0="https://zenodo.org/record/189078/files/figure.png" httpUri-1="https://zenodo.org/record/189079/files/figure.png" httpUri-2="https://zenodo.org/record/189080/files/figure.png" httpUri-3="https://zenodo.org/record/189081/files/figure.png" httpUri-4="https://zenodo.org/record/189082/files/figure.png" pageId="5" pageNumber="64">Figs. 4–8</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<paragraph id="D41E36C57F4BE25761C1FA6F59C9FB2F" pageId="5" pageNumber="64">
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||
<table id="A6A1C4657F4B1DAD61C1FA6F5EE5FB2F" box="[151,1409,1501,1588]" gridcols="8" gridrows="3" pageId="5" pageNumber="64">
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||
<tr id="6A9134877F4B1DAD61C1FA6F5EE5F8EF" box="[151,1409,1501,1524]" gridrow="0" pageId="5" pageNumber="64">
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||
<th id="29405DFB7F4B1DAD61C1FA6F5BABF8EF" box="[151,207,1501,1524]" gridcol="0" gridrow="0" pageId="5" pageNumber="64">
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||
<taxonomicName id="13A14D467F4BE25761C1FA6F5BABF8EF" box="[151,207,1501,1524]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="genus">
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<emphasis id="E6D5EAD77F4BE25761C1FA6F5BABF8EF" box="[151,207,1501,1524]" italics="true" pageId="5" pageNumber="64">Betta</emphasis>
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</taxonomicName>
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||
</th>
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<th id="29405DFB7F4B1DAD6180FA6F584AF8EF" box="[214,814,1501,1524]" gridcol="1" gridrow="0" pageId="5" pageNumber="64">
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<taxonomicName id="13A14D467F4BE2576180FA6F5A3CF8EF" box="[214,344,1501,1524]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="anabatoides">
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<emphasis id="E6D5EAD77F4BE2576180FA6F5A3CF8EF" box="[214,344,1501,1524]" italics="true" pageId="5" pageNumber="64">anabatoides</emphasis>
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</taxonomicName>
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(non Bleeker) — Roberts, 1989: 172 (part),
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</th>
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<th id="29405DFB7F4B1DAD6263FA6F58FEF8EF" box="[821,922,1501,1524]" gridcol="2" gridrow="0" pageId="5" pageNumber="64">fig. 129a;</th>
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||
<th id="29405DFB7F4B1DAD62F4FA6F5F75F8EF" box="[930,1041,1501,1524]" gridcol="3" gridrow="0" pageId="5" pageNumber="64">
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Kottelat
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<emphasis id="E6D5EAD77F4BE25762A8FA6F5F75F8EF" box="[1022,1041,1501,1524]" italics="true" pageId="5" pageNumber="64">et</emphasis>
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||
</th>
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||
<th id="29405DFB7F4B1DAD654EFA6F5F5DF8EF" box="[1048,1081,1501,1524]" gridcol="4" gridrow="0" pageId="5" pageNumber="64">
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<emphasis id="E6D5EAD77F4BE257654EFA6F5F50F8EF" box="[1048,1076,1501,1524]" italics="true" pageId="5" pageNumber="64">al.</emphasis>
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,
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</th>
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<th id="29405DFB7F4B1DAD6516FA6F5F90F8EF" box="[1088,1268,1501,1524]" gridcol="5" gridrow="0" pageId="5" pageNumber="64">1993: 222 (part),</th>
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<th id="29405DFB7F4B1DAD65ADFA6F5E52F8EF" box="[1275,1334,1501,1524]" gridcol="6" gridrow="0" pageId="5" pageNumber="64">cover</th>
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||
<th id="29405DFB7F4B1DAD646BFA6F5EE5F8EF" box="[1341,1409,1501,1524]" gridcol="7" gridrow="0" pageId="5" pageNumber="64">photo.</th>
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||
</tr>
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||
<tr id="6A9134877F4B1DAD61C1FA4F5EE5FB0F" box="[151,1409,1533,1556]" gridrow="1" pageId="5" pageNumber="64" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
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||
<th id="29405DFB7F4B1DAD61C1FA4F5BABFB0F" box="[151,207,1533,1556]" gridcol="0" gridrow="1" pageId="5" pageNumber="64">
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||
<taxonomicName id="13A14D467F4BE25761C1FA4F5BABFB0F" box="[151,207,1533,1556]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="genus">
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||
<emphasis id="E6D5EAD77F4BE25761C1FA4F5BABFB0F" box="[151,207,1533,1556]" italics="true" pageId="5" pageNumber="64">Betta</emphasis>
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||
</taxonomicName>
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||
</th>
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||
<td id="29405DFB7F4B1DAD6180FA4F584AFB0F" box="[214,814,1533,1556]" gridcol="1" gridrow="1" pageId="5" pageNumber="64">
|
||
<taxonomicName id="13A14D467F4BE2576180FA4F5A1EFB0F" box="[214,378,1533,1556]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="anabatoides">
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||
cf.
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||
<emphasis id="E6D5EAD77F4BE25761AEFA4F5A1EFB0F" box="[248,378,1533,1556]" italics="true" pageId="5" pageNumber="64">anabatoides</emphasis>
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||
</taxonomicName>
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||
(non Bleeker) — Kottelat, 1991: 276.
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||
</td>
|
||
</tr>
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||
<tr id="6A9134877F4B1DAD61C1F9AF5EE5FB2F" box="[151,1409,1565,1588]" gridrow="2" pageId="5" pageNumber="64" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||
<th id="29405DFB7F4B1DAD61C1F9AF5BABFB2F" box="[151,207,1565,1588]" gridcol="0" gridrow="2" pageId="5" pageNumber="64">
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||
<taxonomicName id="13A14D467F4BE25761C1F9AF5BABFB2F" box="[151,207,1565,1588]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="genus">
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||
<emphasis id="E6D5EAD77F4BE25761C1F9AF5BABFB2F" box="[151,207,1565,1588]" italics="true" pageId="5" pageNumber="64">Betta</emphasis>
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||
</taxonomicName>
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||
</th>
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||
<td id="29405DFB7F4B1DAD6180F9AF584AFB2F" box="[214,814,1565,1588]" gridcol="1" gridrow="2" pageId="5" pageNumber="64">
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||
spec. affin.
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<taxonomicName id="13A14D467F4BE2576004F9AF5AB1FB2F" box="[338,469,1565,1588]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="anabatoides">
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||
<emphasis id="E6D5EAD77F4BE2576004F9AF5AB1FB2F" box="[338,469,1565,1588]" italics="true" pageId="5" pageNumber="64">anabatoides</emphasis>
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||
</taxonomicName>
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||
— Linke, 1991: 49.
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</td>
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||
</tr>
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||
</table>
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||
</paragraph>
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||
<subSubSection id="9CBB654E7F4BE25561C1F9D15FA3F8BE" lastPageId="7" lastPageNumber="66" pageId="5" pageNumber="64" type="materials_examined">
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||
<paragraph id="D41E36C57F4BE25761C1F9D159C8FBD0" blockId="5.[151,1437,1635,2010]" pageId="5" pageNumber="64">
|
||
<emphasis id="E6D5EAD77F4BE25761C1F9D15AE8FB66" bold="true" box="[151,396,1635,1661]" pageId="5" pageNumber="64">Material examined</emphasis>
|
||
.
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||
<emphasis id="E6D5EAD77F4BE25760CBF9D1597EFB66" bold="true" box="[413,538,1635,1661]" pageId="5" pageNumber="64">
|
||
<typeStatus id="0B1A88677F4BE25760CBF9D15971FB66" box="[413,533,1635,1661]" pageId="5" pageNumber="64" type="holotype">Holotype</typeStatus>
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||
.
|
||
</emphasis>
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||
<collectionCode id="B2B0AE007F4BE2576375F9D65900FB65" box="[547,612,1636,1662]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="64">MZB</collectionCode>
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||
3846, 58.0 mm
|
||
<collectionCode id="B2B0AE007F4BE2576278F9D65831FB65" box="[814,853,1636,1662]" pageId="5" pageNumber="64">SL</collectionCode>
|
||
; West Kalimantan, Kabupaten Pontianak: small stream draining into Sungai Sepatah, a tributary to Sungai
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||
<taxonomicName id="13A14D467F4BE257622AF9385884FBBF" box="[892,992,1674,1700]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="59" phylum="Chordata" rank="species" species="mandor">Mandor</taxonomicName>
|
||
,
|
||
<quantity id="13599B207F4BE25762BBF9385F58FBBF" box="[1005,1084,1674,1700]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="64" unit="km" value="24.0">24 km</quantity>
|
||
NE of Pontianak (
|
||
<geoCoordinate id="B19550027F4BE257647EF9385EF8FBBF" box="[1320,1436,1674,1700]" direction="north" orientation="latitude" pageId="5" pageNumber="64" precision="92" value="0.125">0°07.5’N</geoCoordinate>
|
||
<geoCoordinate id="B19550027F4BE25761C1F9035A68FBD0" box="[151,268,1713,1739]" direction="east" orientation="longitude" pageId="5" pageNumber="64" precision="925" value="109.5">109°30’E</geoCoordinate>
|
||
); T. R. Roberts
|
||
<emphasis id="E6D5EAD77F4BE2576092F9035A9AFBD1" box="[452,510,1713,1738]" italics="true" pageId="5" pageNumber="64">et al.</emphasis>
|
||
,
|
||
<date id="A01F10057F4BE257635AF90359C3FBD0" box="[524,679,1713,1739]" pageId="5" pageNumber="64" value="1976-07-10">10 July 1976</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D41E36C57F4BE2576190F9655F10FAC1" blockId="5.[151,1437,1635,2010]" pageId="5" pageNumber="64">
|
||
<typeStatus id="0B1A88677F4BE2576190F9655A27FBEA" box="[198,323,1751,1777]" pageId="5" pageNumber="64" type="paratype">
|
||
<emphasis id="E6D5EAD77F4BE2576190F9655A27FBEA" bold="true" box="[198,323,1751,1777]" pageId="5" pageNumber="64">Paratypes</emphasis>
|
||
</typeStatus>
|
||
(all from West Kalimantan).
|
||
<collectionCode id="B2B0AE007F4BE25763EAF96A5991FBE9" box="[700,757,1752,1778]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="64">ZRC</collectionCode>
|
||
43711, 8 ex.,
|
||
<collectionCode id="B2B0AE007F4BE25762E3F96A5891FBE9" box="[949,1013,1752,1778]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="64">MZB</collectionCode>
|
||
9334, 2 ex., CMK 20838, 2 ex., 35.7–51.0 mm
|
||
<collectionCode id="B2B0AE007F4BE257600EF94C5AE5FA03" box="[344,385,1790,1816]" pageId="5" pageNumber="64">SL</collectionCode>
|
||
; Kabupaten Pontianak: Sungai Kepayan, blackwater brook at km 58 Pontianak on Pontianak–Anjungan road, ca.
|
||
<quantity id="13599B207F4BE2576340F8975931FA24" box="[534,597,1829,1855]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="64" unit="km" value="7.0">7 km</quantity>
|
||
before Kampung Anjungan (
|
||
<geoCoordinate id="B19550027F4BE2576297F8975F36FA24" box="[961,1106,1829,1855]" direction="north" orientation="latitude" pageId="5" pageNumber="64" precision="9" value="0.314">00°18.84’N</geoCoordinate>
|
||
<geoCoordinate id="B19550027F4BE257650DF8975F99FA24" box="[1115,1277,1829,1855]" direction="east" orientation="longitude" pageId="5" pageNumber="64" precision="9" value="109.134834">109°08.09’E</geoCoordinate>
|
||
); donated by Patrick Yap,
|
||
<date id="A01F10057F4BE2576078F8FE5ADFFA7D" box="[302,443,1868,1894]" pageId="5" pageNumber="64" value="1998-07-08">8 July 1998</date>
|
||
. —
|
||
<collectionCode id="B2B0AE007F4BE25760BDF8FE5940FA7D" box="[491,548,1868,1894]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="64">ZRC</collectionCode>
|
||
46191, 4 ex., 49.0–
|
||
<quantity id="13599B207F4BE2576259F8FE5813FA7D" box="[783,887,1868,1894]" metricMagnitude="-2" metricUnit="m" metricValue="6.14" pageId="5" pageNumber="64" unit="mm" value="61.4">61.4 mm</quantity>
|
||
<collectionCode id="B2B0AE007F4BE2576229F8FE58C1FA7D" box="[895,933,1868,1894]" pageId="5" pageNumber="64">SL</collectionCode>
|
||
; Kabupaten Pontianak: Anjungan; donated by F. Yuwono,
|
||
<date id="A01F10057F4BE2576019F8C05AA7FA97" box="[335,451,1906,1932]" pageId="5" pageNumber="64" value="1999-10">Oct 1999</date>
|
||
. — CMK 16320, 7 ex.,
|
||
<quantity id="13599B207F4BE25763BEF8C058F2FA97" box="[744,918,1906,1932]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="5.15" metricValueMin="3.65" pageId="5" pageNumber="64" unit="mm" value="44.0" valueMax="51.5" valueMin="36.5">36.5–51.5 mm</quantity>
|
||
<collectionCode id="B2B0AE007F4BE25762C9F8C058A2FA97" box="[927,966,1906,1932]" pageId="5" pageNumber="64">SL</collectionCode>
|
||
; Anjungan; don. P. Yap,
|
||
<date id="A01F10057F4BE25765A5F8C05E16FA97" box="[1267,1394,1906,1932]" pageId="5" pageNumber="64" value="2000-05">May 2000</date>
|
||
. —
|
||
<collectionCode id="B2B0AE007F4BE25761C1F82B5BB4FAA8" box="[151,208,1945,1971]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="64">ZRC</collectionCode>
|
||
46190, 12 ex.,
|
||
<quantity id="13599B207F4BE25760DEF82B5957FAA8" box="[392,563,1945,1971]" metricMagnitude="-2" metricUnit="m" metricValue="5.205" metricValueMax="5.85" metricValueMin="4.56" pageId="5" pageNumber="64" unit="mm" value="52.05" valueMax="58.5" valueMin="45.6">45.6–58.5 mm</quantity>
|
||
<collectionCode id="B2B0AE007F4BE257636DF82B5905FAA8" box="[571,609,1945,1971]" pageId="5" pageNumber="64">SL</collectionCode>
|
||
; Kabupaten Pontianak: Kapuas basin; donated by P. Yap,
|
||
<date id="A01F10057F4BE257645CF82B5EFCFAA8" box="[1290,1432,1945,1971]" pageId="5" pageNumber="64" value="2000-03-03">3 Mar 2000</date>
|
||
. —
|
||
<collectionCode id="B2B0AE007F4BE25761EAF8725B91FAC1" box="[188,245,1984,2010]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="64">ZRC</collectionCode>
|
||
51527, 1 ex., 66.0 mm
|
||
<collectionCode id="B2B0AE007F4BE257635DF8725954FAC1" box="[523,560,1984,2010]" pageId="5" pageNumber="64">SL</collectionCode>
|
||
; Kapuas basin,
|
||
<collectingCountry id="ACB676557F4BE25763BEF872584BFAC1" box="[744,815,1984,2010]" name="Thailand" pageId="5" pageNumber="64">Tayan</collectingCountry>
|
||
area; H. H. Tan,
|
||
<date id="A01F10057F4BE25762AEF8725F14FAC1" box="[1016,1136,1984,2010]" pageId="5" pageNumber="64" value="2007-08">Aug 2007</date>
|
||
.
|
||
</paragraph>
|
||
<caption id="80DE664D7F48E25461C1FD7B5A39FE18" httpUri="https://zenodo.org/record/189078/files/figure.png" pageId="6" pageNumber="65" targetBox="[151,1434,197,686]" targetPageId="6">
|
||
<paragraph id="D41E36C57F48E25461C1FD7B5A39FE18" blockId="6.[151,1436,713,771]" pageId="6" pageNumber="65">
|
||
<emphasis id="E6D5EAD77F48E25461C1FD7B5A7BFFFA" bold="true" box="[151,287,713,737]" pageId="6" pageNumber="65">FIGURE 4.</emphasis>
|
||
<taxonomicName id="13A14D467F48E2546071FD7B5AC9FFFB" box="[295,429,713,736]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="midas">
|
||
<emphasis id="E6D5EAD77F48E2546071FD7B5AC9FFFB" box="[295,429,713,736]" italics="true" pageId="6" pageNumber="65">Betta midas</emphasis>
|
||
</taxonomicName>
|
||
, ZRC 46190, 48.3 mm SL, paratype from Sungai Kepayan, West Kalimantan (right side, laterally inverted).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="80DE664D7F48E25461C1FAD35F0CF862" box="[151,1128,1377,1401]" httpUri="https://zenodo.org/record/189079/files/figure.png" pageId="6" pageNumber="65" targetBox="[151,1434,825,1352]" targetPageId="6">
|
||
<paragraph id="D41E36C57F48E25461C1FAD35F0CF862" blockId="6.[151,1128,1377,1401]" box="[151,1128,1377,1401]" pageId="6" pageNumber="65">
|
||
<emphasis id="E6D5EAD77F48E25461C1FAD35B9BF862" bold="true" box="[151,255,1377,1401]" pageId="6" pageNumber="65">FIGURE</emphasis>
|
||
5.
|
||
<taxonomicName id="13A14D467F48E2546076FAD05AFBF862" box="[288,415,1378,1401]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="midas">
|
||
<emphasis id="E6D5EAD77F48E2546076FAD05AFBF862" box="[288,415,1378,1401]" italics="true" pageId="6" pageNumber="65">Betta midas</emphasis>
|
||
</taxonomicName>
|
||
, not preserved, ca. 35 mm SL, Sungai Kepayan, West Kalimantan.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="80DE664D7F48E25461C1F80858C9FACA" box="[151,941,1978,2002]" httpUri="https://zenodo.org/record/189080/files/figure.png" pageId="6" pageNumber="65" targetBox="[151,1434,1455,1954]" targetPageId="6">
|
||
<paragraph id="D41E36C57F48E25461C1F80858C9FACA" blockId="6.[151,941,1978,2002]" box="[151,941,1978,2002]" pageId="6" pageNumber="65">
|
||
<emphasis id="E6D5EAD77F48E25461C1F8085A7DFAC9" bold="true" box="[151,281,1978,2002]" pageId="6" pageNumber="65">FIGURE 6.</emphasis>
|
||
<taxonomicName id="13A14D467F48E2546076F8085AFBFACA" box="[288,415,1978,2001]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="midas">
|
||
<emphasis id="E6D5EAD77F48E2546076F8085AFBFACA" box="[288,415,1978,2001]" italics="true" pageId="6" pageNumber="65">Betta midas</emphasis>
|
||
</taxonomicName>
|
||
, ZRC 51527, 66.0 mm SL, paratype from Tayan.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="80DE664D7F49E25561C1FD4E58B8FE08" box="[151,988,764,788]" httpUri="https://zenodo.org/record/189081/files/figure.png" pageId="7" pageNumber="66" targetBox="[151,1434,180,739]" targetPageId="7">
|
||
<paragraph id="D41E36C57F49E25561C1FD4E58B8FE08" blockId="7.[151,988,764,788]" box="[151,988,764,788]" pageId="7" pageNumber="66">
|
||
<emphasis id="E6D5EAD77F49E25561C1FD4E5B9BFE0F" bold="true" box="[151,255,764,788]" pageId="7" pageNumber="66">FIGURE</emphasis>
|
||
7.
|
||
<taxonomicName id="13A14D467F49E2556076FD4E5AFBFE08" box="[288,415,764,787]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="midas">
|
||
<emphasis id="E6D5EAD77F49E2556076FD4E5AFBFE08" box="[288,415,764,787]" italics="true" pageId="7" pageNumber="66">Betta midas</emphasis>
|
||
</taxonomicName>
|
||
, ZRC 51527, 66.0 mm SL, live paratype from Tayan.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="80DE664D7F49E25561C1FB63581AF9F2" box="[151,894,1233,1257]" httpUri="https://zenodo.org/record/189082/files/figure.png" pageId="7" pageNumber="66" targetBox="[151,1435,841,1208]" targetPageId="7">
|
||
<paragraph id="D41E36C57F49E25561C1FB63581AF9F2" blockId="7.[151,894,1233,1257]" box="[151,894,1233,1257]" pageId="7" pageNumber="66">
|
||
<emphasis id="E6D5EAD77F49E25561C1FB635A7CF9F2" bold="true" box="[151,280,1233,1257]" pageId="7" pageNumber="66">FIGURE 8.</emphasis>
|
||
<taxonomicName id="13A14D467F49E2556076FB605AFBF9F2" box="[288,415,1234,1257]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="midas">
|
||
<emphasis id="E6D5EAD77F49E2556076FB605AFBF9F2" box="[288,415,1234,1257]" italics="true" pageId="7" pageNumber="66">Betta midas</emphasis>
|
||
</taxonomicName>
|
||
, ZRC 51526, 61.4 mm SL, Sarawak, Lundu.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D41E36C57F49E2556190FAA55FA3F8BE" blockId="7.[151,1437,1303,2025]" pageId="7" pageNumber="66">
|
||
<emphasis id="E6D5EAD77F49E2556190FAA55AD5F82A" bold="true" box="[198,433,1303,1329]" pageId="7" pageNumber="66">Non-type material</emphasis>
|
||
<collectionCode id="B2B0AE007F49E25560E9FAA55A9CF82A" box="[447,504,1303,1329]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="66">ZRC</collectionCode>
|
||
43712, 4 ex., 35.0–
|
||
<quantity id="13599B207F49E25563ACFAA5580CF82A" box="[762,872,1303,1329]" metricMagnitude="-2" metricUnit="m" metricValue="4.33" pageId="7" pageNumber="66" unit="mm" value="43.3">43.3 mm</quantity>
|
||
<collectionCode id="B2B0AE007F49E2556223FAA558FFF82A" box="[885,923,1303,1329]" pageId="7" pageNumber="66">SL</collectionCode>
|
||
; West Kalimantan, Kabupaten Pontianak: Sungai Kepayan, blackwater brook at km 58 Pontianak on Pontianak–Anjungan road, ca.
|
||
<quantity id="13599B207F49E2556452FA8C5E20F843" box="[1284,1348,1342,1368]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="66" unit="km" value="7.0">7 km</quantity>
|
||
before Kampung Anjungan (
|
||
<geoCoordinate id="B19550027F49E25560CEFAD65947F865" box="[408,547,1380,1406]" direction="north" orientation="latitude" pageId="7" pageNumber="66" precision="9" value="0.314">00°18.84’N</geoCoordinate>
|
||
<geoCoordinate id="B19550027F49E255637AFAD659A3F865" box="[556,711,1380,1406]" direction="east" orientation="longitude" pageId="7" pageNumber="66" precision="9" value="109.134834">109°08.09’E</geoCoordinate>
|
||
); donated by Patrick Yap,
|
||
<date id="A01F10057F49E2556556FAD65FEBF865" box="[1024,1167,1380,1406]" pageId="7" pageNumber="66" value="1998-07-08">8 July 1998</date>
|
||
. —
|
||
<collectionCode id="B2B0AE007F49E2556596FAD65F9DF865" box="[1216,1273,1380,1406]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="66">ZRC</collectionCode>
|
||
51526, 1 ex.,
|
||
<quantity id="13599B207F49E25561C1FA395B9BF8BE" box="[151,255,1419,1445]" metricMagnitude="-2" metricUnit="m" metricValue="6.14" pageId="7" pageNumber="66" unit="mm" value="61.4">61.4 mm</quantity>
|
||
<collectionCode id="B2B0AE007F49E2556050FA395A4FF8BE" box="[262,299,1419,1445]" pageId="7" pageNumber="66">SL</collectionCode>
|
||
;
|
||
<collectingCountry id="ACB676557F49E255606EFA395AC5F8BE" box="[312,417,1419,1445]" name="Malaysia" pageId="7" pageNumber="66">Sarawak</collectingCountry>
|
||
: Lundu, Batang Kayan basin, Sungei Temelan; M. Lo,
|
||
<date id="A01F10057F49E2556570FA395FA7F8BE" box="[1062,1219,1419,1445]" pageId="7" pageNumber="66" value="2005-08-28">28 Aug 2005</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CBB654E7F49E2556190FA035896FB5B" pageId="7" pageNumber="66" type="diagnosis">
|
||
<paragraph id="D41E36C57F49E2556190FA035896FB5B" blockId="7.[151,1437,1303,2025]" pageId="7" pageNumber="66">
|
||
<emphasis id="E6D5EAD77F49E2556190FA035A22F8D0" bold="true" box="[198,326,1457,1483]" pageId="7" pageNumber="66">Diagnosis.</emphasis>
|
||
<taxonomicName id="13A14D467F49E2556019FA005AB8F8D0" box="[335,476,1458,1483]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="midas">
|
||
<emphasis id="E6D5EAD77F49E2556019FA005AB8F8D0" box="[335,476,1458,1483]" italics="true" pageId="7" pageNumber="66">Betta midas</emphasis>
|
||
</taxonomicName>
|
||
is unique in having the following combination of characters: relatively thick-set body (body depth 24.3–29.0 % SL); head devoid of pattern except for black stripe that runs from black lower lip, through eye to edge of opercle; opercle with gold iridescent scales when live; belly and body with gold iridescent scales; some median caudal rays projecting beyond fin margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CBB654E7F49E25A6190F9FE5831FC81" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66" type="description">
|
||
<paragraph id="D41E36C57F49E2556190F9FE5F80FA1A" blockId="7.[151,1437,1303,2025]" pageId="7" pageNumber="66">
|
||
<emphasis id="E6D5EAD77F49E2556190F9FE5A3AFB7D" bold="true" box="[198,350,1612,1638]" pageId="7" pageNumber="66">Description.</emphasis>
|
||
General appearance as illustrated in
|
||
<figureCitation id="4C9A2A407F49E2556271F9FE58F8FB7D" box="[807,924,1612,1638]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStartId-0="6.[151,258,713,737]" captionStartId-1="6.[151,255,1377,1401]" captionStartId-2="6.[151,255,1978,2002]" captionStartId-3="7.[151,255,764,788]" captionStartId-4="7.[151,255,1233,1257]" captionTargetBox-0="[151,1434,197,686]" captionTargetBox-1="[151,1434,825,1352]" captionTargetBox-2="[151,1434,1455,1954]" captionTargetBox-3="[151,1434,180,739]" captionTargetBox-4="[151,1435,841,1208]" captionTargetId-0="figure@6.[151,1436,197,688]" captionTargetId-1="figure@6.[151,1436,825,1354]" captionTargetId-2="figure@6.[151,1436,1455,1954]" captionTargetId-3="figure@7.[151,1436,180,740]" captionTargetId-4="figure@7.[151,1436,841,1209]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="7" captionText-0="FIGURE 4. Betta midas, ZRC 46190, 48.3 mm SL, paratype from Sungai Kepayan, West Kalimantan (right side, laterally inverted)." captionText-1="FIGURE 5. Betta midas, not preserved, ca. 35 mm SL, Sungai Kepayan, West Kalimantan." captionText-2="FIGURE 6. Betta midas, ZRC 51527, 66.0 mm SL, paratype from Tayan." captionText-3="FIGURE 7. Betta midas, ZRC 51527, 66.0 mm SL, live paratype from Tayan." captionText-4="FIGURE 8. Betta midas, ZRC 51526, 61.4 mm SL, Sarawak, Lundu." httpUri-0="https://zenodo.org/record/189078/files/figure.png" httpUri-1="https://zenodo.org/record/189079/files/figure.png" httpUri-2="https://zenodo.org/record/189080/files/figure.png" httpUri-3="https://zenodo.org/record/189081/files/figure.png" httpUri-4="https://zenodo.org/record/189082/files/figure.png" pageId="7" pageNumber="66">Figs. 4–8</figureCitation>
|
||
. Meristic and morphometric data listed in Table 1. Head blunt, relatively long (HL 31.9–34.4 % SL); body relatively thick-set (body depth 24.3–29.0 % SL); pectoral fin rounded; pelvic fin filamentous; dorsal fin situated far back (predorsal length 65.5–67.4 % SL); dorsal and anal fins pointed with elongated posterior rays; caudal peduncle deep (caudal peduncle depth 16.8–19.0 % SL); caudal fin lanceolate, with some median rays projecting beyond fin margin.
|
||
</paragraph>
|
||
<paragraph id="D41E36C57F49E25A6190F8BF5907FDC3" blockId="7.[151,1437,1303,2025]" lastBlockId="8.[151,1437,152,1338]" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66">
|
||
<emphasis id="E6D5EAD77F49E2556190F8BF5AC7FA3C" bold="true" box="[198,419,1805,1831]" pageId="7" pageNumber="66">Coloration in life.</emphasis>
|
||
Refer to
|
||
<bibRefCitation id="B0304B347F49E2556347F8BC59B3FA33" author="Linke" box="[529,727,1806,1832]" pageId="7" pageNumber="68" refString="Linke, H. (1991) Labyrinth fish. The Bubble-nest-builders. Their identification, care and breeding. Tetra-Press, Melle, Germany, 174 pp." type="book" year="1991">Linke (1991: 49)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="B0304B347F49E2556247F8BC5F66FA33" author="Kottelat" box="[785,1026,1806,1832]" pageId="7" pageNumber="68" refString="Kottelat, M., Whitten, A. J., Kartikasari, S. N. & Wirjoatmodjo, S. (1993) Freshwater Fishes of Western Indonesia and Sulawesi. Periplus Editions, Hong Kong, 259 pp., 84 pls." type="book" year="1993">
|
||
Kottelat
|
||
<emphasis id="E6D5EAD77F49E2556221F8BC58D5FA3C" box="[887,945,1806,1831]" italics="true" pageId="7" pageNumber="66">et al.</emphasis>
|
||
(1993
|
||
</bibRefCitation>
|
||
: cover photo) and
|
||
<figureCitation id="4C9A2A407F49E255658CF8BC5E4FFA33" box="[1242,1323,1806,1832]" captionStart="FIGURE 5" captionStartId="6.[151,255,1377,1401]" captionTargetBox="[151,1434,825,1352]" captionTargetId="figure@6.[151,1436,825,1354]" captionTargetPageId="6" captionText="FIGURE 5. Betta midas, not preserved, ca. 35 mm SL, Sungai Kepayan, West Kalimantan." httpUri="https://zenodo.org/record/189079/files/figure.png" pageId="7" pageNumber="66">Figs. 5</figureCitation>
|
||
and
|
||
<figureCitation id="4C9A2A407F49E2556435F8BC5E16FA33" box="[1379,1394,1806,1832]" captionStart="FIGURE 7" captionStartId="7.[151,255,764,788]" captionTargetBox="[151,1434,180,739]" captionTargetId="figure@7.[151,1436,180,740]" captionTargetPageId="7" captionText="FIGURE 7. Betta midas, ZRC 51527, 66.0 mm SL, live paratype from Tayan." httpUri="https://zenodo.org/record/189081/files/figure.png" pageId="7" pageNumber="66">7</figureCitation>
|
||
for colour illustration. There appears to be no discernible external characters to differentiate the sexes. Dorsal of body brown, ventral lighter brown speckled with gold iridescent scales. Dorsal of head brown with blotches of darker brown behind eye, opercle with distinct greenish-gold iridescence. Distinct pre- and postorbital black stripes on head, lower lip black, chin-bar absent. Body sometimes with central, lower central dark brown stripes and caudal peduncle spot. Fins brownish, dorsal fin with 5–8 dark brown dorsal transverse bars, anal fin sometimes with dark distal edge, caudal fin with 10–12 brown transverse bars. Pectoral fin yellowish. Pelvic fin with bright whitish distal tip.
|
||
</paragraph>
|
||
<paragraph id="D41E36C57F46E25A6190FF575831FC81" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="67">
|
||
<emphasis id="E6D5EAD77F46E25A6190FF57597FFDE4" bold="true" box="[198,539,229,255]" pageId="8" pageNumber="67">Coloration in preservative.</emphasis>
|
||
See
|
||
<figureCitation id="4C9A2A407F46E25A630EFF5759C9FDE4" box="[600,685,229,255]" captionStart="FIGURE 4" captionStartId="6.[151,258,713,737]" captionTargetBox="[151,1434,197,686]" captionTargetId="figure@6.[151,1436,197,688]" captionTargetPageId="6" captionText="FIGURE 4. Betta midas, ZRC 46190, 48.3 mm SL, paratype from Sungai Kepayan, West Kalimantan (right side, laterally inverted)." httpUri="https://zenodo.org/record/189078/files/figure.png" pageId="8" pageNumber="67">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="4C9A2A407F46E25A63EAFF5759AEFDE4" box="[700,714,229,255]" captionStart="FIGURE 6" captionStartId="6.[151,255,1978,2002]" captionTargetBox="[151,1434,1455,1954]" captionTargetId="figure@6.[151,1436,1455,1954]" captionTargetPageId="6" captionText="FIGURE 6. Betta midas, ZRC 51527, 66.0 mm SL, paratype from Tayan." httpUri="https://zenodo.org/record/189080/files/figure.png" pageId="8" pageNumber="67">6</figureCitation>
|
||
and
|
||
<figureCitation id="4C9A2A407F46E25A625EFF57587DFDE4" box="[776,793,229,255]" captionStart="FIGURE 8" captionStartId="7.[151,255,1233,1257]" captionTargetBox="[151,1435,841,1208]" captionTargetId="figure@7.[151,1436,841,1209]" captionTargetPageId="7" captionText="FIGURE 8. Betta midas, ZRC 51526, 61.4 mm SL, Sarawak, Lundu." httpUri="https://zenodo.org/record/189082/files/figure.png" pageId="8" pageNumber="67">8</figureCitation>
|
||
. Dorsal of body dark brown, ventral lighter brown to whitish. Dorsal of head dark brown, opercle area whitish. Distinct pre- and postorbital black stripes on head, chin-bar absent. Body with faint central, lower central stripes and caudal peduncle spot. Fins brownish, dorsal fin with 5–8 dorsal transverse black bars, caudal fin with 10–12 faint transverse bars. Pectoral fin yellowish. Pelvic fin light brown, with hyaline filamentous second ray.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CBB654E7F46E25A6190FE1459BBFE8B" pageId="8" pageNumber="67" type="distribution">
|
||
<paragraph id="D41E36C57F46E25A6190FE145B92FF2F" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="67">
|
||
<emphasis id="E6D5EAD77F46E25A6190FE145A03FCDB" bold="true" box="[198,359,422,448]" pageId="8" pageNumber="67">Distribution.</emphasis>
|
||
<taxonomicName id="13A14D467F46E25A6038FE155A9EFCDB" box="[366,506,423,448]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="midas">
|
||
<emphasis id="E6D5EAD77F46E25A6038FE155A9EFCDB" box="[366,506,423,448]" italics="true" pageId="8" pageNumber="67">Betta midas</emphasis>
|
||
</taxonomicName>
|
||
is currently known from the lower Kapuas basin near the village of Anjungan, about
|
||
<quantity id="13599B207F46E25A61B7FE7F5A49FCFC" box="[225,301,461,487]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="67" unit="km" value="60.0">60 km</quantity>
|
||
away from the town of Pontianak towards
|
||
<taxonomicName id="13A14D467F46E25A626EFE7F58FCFCFC" box="[824,920,461,487]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="mandor">Mandor</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A62F3FE7F5F60FCFC" box="[933,1028,461,487]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="mandor">Mandor</taxonomicName>
|
||
area and
|
||
<collectingCountry id="ACB676557F46E25A652FFE7F5FA6FCFC" box="[1145,1218,461,487]" name="Thailand" pageId="8" pageNumber="67">Tayan</collectingCountry>
|
||
area; possibly the Sambas river basin; and the extreme western part of
|
||
<collectingCountry id="ACB676557F46E25A6250FE46580AFF15" box="[774,878,500,526]" name="Malaysia" pageId="8" pageNumber="67">Sarawak</collectingCountry>
|
||
, in heathforest streams near the town of Lundu (
|
||
<figureCitation id="4C9A2A407F46E25A61C9FDA85B82FF2F" box="[159,230,538,564]" captionStart="FIGURE 3" captionStartId="4.[151,255,1776,1800]" captionTargetBox="[156,1436,197,1745]" captionTargetId="figure@4.[151,1436,194,1750]" captionTargetPageId="4" captionText="FIGURE 3. Map showing known distribution of B. anabatoides (circle) and B. midas (square) in Borneo." httpUri="https://zenodo.org/record/189077/files/figure.png" pageId="8" pageNumber="67">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D41E36C57F46E25A6190FDF259BBFE8B" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="67">
|
||
<emphasis id="E6D5EAD77F46E25A6190FDF25A3EFF41" bold="true" box="[198,346,576,602]" pageId="8" pageNumber="67">Field notes.</emphasis>
|
||
The
|
||
<typeStatus id="0B1A88677F46E25A60F6FDF35961FF40" box="[416,517,577,603]" pageId="8" pageNumber="67" type="paratype">paratype</typeStatus>
|
||
series (ZRC 43711) is from a blackwater river (Sungai Kepayan) running through remnant and intact peat swamp forest, with a
|
||
<collectingCountry id="ACB676557F46E25A6244FDDA5852FF99" box="[786,822,616,642]" name="Philippines" pageId="8" pageNumber="67">pH</collectingCountry>
|
||
of 4.1. Depth ranged from
|
||
<quantity id="13599B207F46E25A652CFDDA5FA7FF99" box="[1146,1219,616,642]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="67" unit="cm" value="20.0">20 cm</quantity>
|
||
to more than
|
||
<specimenCount id="C2A7FD4C7F46E25A6431FDDA5EF8FF99" box="[1383,1436,616,642]" pageId="8" pageNumber="67" type="generic">
|
||
<quantity id="13599B207F46E25A6431FDDA5EFCFF99" box="[1383,1432,616,642]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="67" unit="m" value="2.0">2 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Syntopic species include:
|
||
<taxonomicName id="13A14D467F46E25A609DFD3D59D9FFB3" box="[459,701,655,680]" class="Actinopterygii" family="Cyprinidae" genus="Osteochilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="spilurus">
|
||
<emphasis id="E6D5EAD77F46E25A609DFD3D59D9FFB3" box="[459,701,655,680]" italics="true" pageId="8" pageNumber="67">Osteochilus spilurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A639AFD3D5897FFB3" box="[716,1011,655,680]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="rhomboocellatus">
|
||
<emphasis id="E6D5EAD77F46E25A639AFD3D5897FFB3" box="[716,1011,655,680]" italics="true" pageId="8" pageNumber="67">Puntius rhomboocellatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A6555FD3D5E6EFFB3" box="[1027,1290,655,680]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="dorsiocellata">
|
||
<emphasis id="E6D5EAD77F46E25A6555FD3D5E6EFFB3" box="[1027,1290,655,680]" italics="true" pageId="8" pageNumber="67">Rasbora dorsiocellata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A644FFD3D5EF1FFB3" box="[1305,1429,655,680]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="E6D5EAD77F46E25A644FFD3D5EF1FFB3" box="[1305,1429,655,680]" italics="true" pageId="8" pageNumber="67">R. gracilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E6D5EAD77F46E25A61C1FD075962FFD5" box="[151,518,693,718]" italics="true" pageId="8" pageNumber="67">
|
||
<taxonomicName id="13A14D467F46E25A61C1FD075A08FFD5" box="[151,364,693,718]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="pauciperforata">R. pauciperforata</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A602FFD075962FFD5" box="[377,518,693,718]" class="Actinopterygii" family="Cyprinidae" genus="Sundadanio" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="axelrodi">Sundadanio</taxonomicName>
|
||
</emphasis>
|
||
cf.
|
||
<emphasis id="E6D5EAD77F46E25A6362FD0759F1FFD5" box="[564,661,693,718]" italics="true" pageId="8" pageNumber="67">axelrodi</emphasis>
|
||
(
|
||
<taxonomicName id="13A14D467F46E25A63F0FD075854FFD4" box="[678,816,693,719]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
|
||
),
|
||
<taxonomicName id="13A14D467F46E25A6212FD075FADFFD4" box="[836,1225,693,719]" class="Actinopterygii" family="Cobitidae" genus="Kottelatlimia" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="pristes">
|
||
<emphasis id="E6D5EAD77F46E25A6212FD075F5CFFD5" box="[836,1080,693,718]" italics="true" pageId="8" pageNumber="67">Kottelatlimia pristes</emphasis>
|
||
(Cobitidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A6580FD075AA2FFED" class="Actinopterygii" family="Balitoridae" genus="Neohomaloptera" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="johorensis">
|
||
<emphasis id="E6D5EAD77F46E25A6580FD075A7CFFEE" italics="true" pageId="8" pageNumber="67">Neohomaloptera johorensis</emphasis>
|
||
(Balitoridae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A6085FD6E5831FFED" box="[467,853,732,758]" class="Actinopterygii" family="Bagridae" genus="Nanobagrus" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="fuscus">
|
||
<emphasis id="E6D5EAD77F46E25A6085FD6E59A7FFEE" box="[467,707,732,757]" italics="true" pageId="8" pageNumber="67">Nanobagrus fuscus</emphasis>
|
||
(Bagridae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A6235FD6E5FA9FFEE" box="[867,1229,732,757]" class="Actinopterygii" family="Siluridae" genus="Kryptopterus" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="E6D5EAD77F46E25A6235FD6E5FA9FFEE" box="[867,1229,732,757]" italics="true" pageId="8" pageNumber="67">Kryptopterus macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A6588FD6E5EF1FFEE" box="[1246,1429,732,757]" class="Actinopterygii" family="Siluridae" genus="Ompok" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="weberi">
|
||
<emphasis id="E6D5EAD77F46E25A6588FD6E5EF1FFEE" box="[1246,1429,732,757]" italics="true" pageId="8" pageNumber="67">Ompok weberi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A61C1FCB0595BFE07" box="[151,575,770,796]" class="Actinopterygii" family="Siluridae" genus="Silurichthys" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="phaiosoma">
|
||
<emphasis id="E6D5EAD77F46E25A61C1FCB05AD7FE00" box="[151,435,770,795]" italics="true" pageId="8" pageNumber="67">Silurichthys phaiosoma</emphasis>
|
||
(Siluridae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A631AFCB058D6FE00" box="[588,946,770,795]" class="Actinopterygii" family="Zenarchopteridae" genus="Hemirhamphodon" kingdom="Animalia" order="Beloniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="phaiosoma">
|
||
<emphasis id="E6D5EAD77F46E25A631AFCB058D6FE00" box="[588,946,770,795]" italics="true" pageId="8" pageNumber="67">Hemirhamphodon phaiosoma</emphasis>
|
||
</taxonomicName>
|
||
[often mis-identified as
|
||
<taxonomicName id="13A14D467F46E25A6588FCB05EF0FE00" box="[1246,1428,770,795]" class="Actinopterygii" family="Zenarchopteridae" genus="Hemirhamphodon" kingdom="Animalia" order="Beloniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="kapuasensis">
|
||
<emphasis id="E6D5EAD77F46E25A6588FCB05EF0FE00" box="[1246,1428,770,795]" italics="true" pageId="8" pageNumber="67">H. kapuasensis</emphasis>
|
||
</taxonomicName>
|
||
, which has a restricted distribution in middle Kapuas] (
|
||
<taxonomicName id="13A14D467F46E25A626BFC9B5F6EFE58" box="[829,1034,809,835]" class="Actinopterygii" family="Hemiramphidae" kingdom="Animalia" order="Beloniformes" pageId="8" pageNumber="67" phylum="Chordata" rank="family">Hemiramphidae</taxonomicName>
|
||
),
|
||
<taxonomicName id="13A14D467F46E25A6576FC9B5EF3FE58" box="[1056,1431,809,835]" class="Actinopterygii" family="Nandidae" genus="Nandus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="nebulosus">
|
||
<emphasis id="E6D5EAD77F46E25A6576FC9B5E65FE59" box="[1056,1281,809,834]" italics="true" pageId="8" pageNumber="67">Nandus nebulosus</emphasis>
|
||
(Nandidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A61C1FCE25A0FFE72" box="[151,363,848,873]" class="Actinopterygii" family="Osphronemidae" genus="Belontia" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="hasseltii">
|
||
<emphasis id="E6D5EAD77F46E25A61C1FCE25A0FFE72" box="[151,363,848,873]" italics="true" pageId="8" pageNumber="67">Belontia hasseltii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A602DFCE25978FE72" box="[379,540,848,873]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="edithae">
|
||
<emphasis id="E6D5EAD77F46E25A602DFCE25978FE72" box="[379,540,848,873]" italics="true" pageId="8" pageNumber="67">Betta edithae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A637AFCE2585BFE72" box="[556,831,848,873]" class="Actinopterygii" family="Osphronemidae" genus="Luciocephalus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="pulcher">
|
||
<emphasis id="E6D5EAD77F46E25A637AFCE2585BFE72" box="[556,831,848,873]" italics="true" pageId="8" pageNumber="67">Luciocephalus pulcher</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A6206FCE25FB6FE72" box="[848,1234,848,873]" class="Actinopterygii" family="Osphronemidae" genus="Parosphromenus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="anjunganensis">
|
||
<emphasis id="E6D5EAD77F46E25A6206FCE25FB6FE72" box="[848,1234,848,873]" italics="true" pageId="8" pageNumber="67">Parosphromenus anjunganensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A65B4FCE25EF1FE72" box="[1250,1429,848,873]" class="Actinopterygii" family="Osphronemidae" genus="Parosphromenus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="ornaticauda">
|
||
<emphasis id="E6D5EAD77F46E25A65B4FCE25EF1FE72" box="[1250,1429,848,873]" italics="true" pageId="8" pageNumber="67">P. ornaticauda</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13A14D467F46E25A61C1FCC559BFFE8B" box="[151,731,886,912]" class="Actinopterygii" family="Osphronemidae" genus="Sphaerichthys" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="osphromenoides">
|
||
<emphasis id="E6D5EAD77F46E25A61C1FCC55967FE8B" box="[151,515,887,912]" italics="true" pageId="8" pageNumber="67">Sphaerichthys osphromenoides</emphasis>
|
||
(Osphronemidae)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CBB654E7F46E25A6190FC2F5FD4FEC5" pageId="8" pageNumber="67" type="etymology">
|
||
<paragraph id="D41E36C57F46E25A6190FC2F5FD4FEC5" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="67">
|
||
<emphasis id="E6D5EAD77F46E25A6190FC2F5A37FEAC" bold="true" box="[198,339,925,951]" pageId="8" pageNumber="67">Etymology.</emphasis>
|
||
An allusion to the Greek
|
||
<taxonomicName id="13A14D467F46E25A63CFFC2F5986FEAD" box="[665,738,925,950]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="midas">
|
||
<emphasis id="E6D5EAD77F46E25A63CFFC2F5986FEAD" box="[665,738,925,950]" italics="true" pageId="8" pageNumber="67">Midas</emphasis>
|
||
</taxonomicName>
|
||
, a mythical king whose touch turned everything to gold, arising from the iridescent gold opercle and body scales. Applied as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CBB654E7F46E25A6190FC58590AF821" pageId="8" pageNumber="67" type="discussion">
|
||
<paragraph id="D41E36C57F46E25A6190FC585A09F949" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="67">
|
||
<emphasis id="E6D5EAD77F46E25A6190FC585A24F91F" bold="true" box="[198,320,1002,1028]" pageId="8" pageNumber="67">Remarks.</emphasis>
|
||
Roberts’ (1989: 172) material from other than the
|
||
<typeStatus id="0B1A88677F46E25A62CAFC5858AAF91F" box="[924,974,1002,1028]" pageId="8" pageNumber="67">type</typeStatus>
|
||
locality identified as
|
||
<taxonomicName id="13A14D467F46E25A6599FC585EE4F918" box="[1231,1408,1002,1027]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="anabatoides">
|
||
<emphasis id="E6D5EAD77F46E25A6599FC585EE4F918" box="[1231,1408,1002,1027]" italics="true" pageId="8" pageNumber="67">B. anabatoides</emphasis>
|
||
</taxonomicName>
|
||
is in fact another species belonging to the
|
||
<taxonomicName id="13A14D467F46E25A6323FBA3586CF931" box="[629,776,1041,1066]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="akarensis">
|
||
<emphasis id="E6D5EAD77F46E25A6323FBA3586CF931" box="[629,776,1041,1066]" italics="true" pageId="8" pageNumber="67">B. akarensis</emphasis>
|
||
</taxonomicName>
|
||
group: this exhibits a chin-bar and has a slimmer body (e.g., MZB 3848).
|
||
</paragraph>
|
||
<paragraph id="D41E36C57F46E25A6190FBED590AF821" blockId="8.[151,1437,152,1338]" pageId="8" pageNumber="67">
|
||
<taxonomicName id="13A14D467F46E25A6190FBED5A37F963" box="[198,339,1119,1144]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="midas">
|
||
<emphasis id="E6D5EAD77F46E25A6190FBED5A37F963" box="[198,339,1119,1144]" italics="true" pageId="8" pageNumber="67">Betta midas</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from
|
||
<taxonomicName id="13A14D467F46E25A63CCFBED582FF963" box="[666,843,1119,1144]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="67" phylum="Chordata" rank="species" species="anabatoides">
|
||
<emphasis id="E6D5EAD77F46E25A63CCFBED582FF963" box="[666,843,1119,1144]" italics="true" pageId="8" pageNumber="67">B. anabatoides</emphasis>
|
||
</taxonomicName>
|
||
by the following characters: a less thick-set body (
|
||
<figureCitation id="4C9A2A407F46E25A61F6FB375B8BF984" box="[160,239,1157,1183]" captionStart="FIGURE 2" captionStartId="2.[151,255,1805,1829]" captionTargetBox="[151,1434,1144,1780]" captionTargetId="figure@2.[151,1436,1144,1781]" captionTargetPageId="2" captionText="FIGURE 2. Betta anabatoides, not preserved, ca. 35 mm SL, Central Kalimantan." httpUri="https://zenodo.org/record/189076/files/figure.png" pageId="8" pageNumber="67">Fig. 2</figureCitation>
|
||
; body depth 24.3–29.0% SL, vs. 30.5–33.7); a less deep caudal peduncle (16.8–19.0 % SL, vs. 19.8–22.1); the presence of caudal transverse bars (vs. absence); the presenc of distinct iridescent gold opercle scales (vs. faint); the presence of gold body and belly scales (vs. absence); a blunt lateral head profile (vs. pointed); a slightly lesser predorsal length (65.5–67.4 % SL, vs. 66.8–69.7); slightly smaller anal-fin base length (52.4–57.9 % SL, vs. 55.2–61.1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |