255 lines
32 KiB
XML
255 lines
32 KiB
XML
<document id="F991BF68D261A655282C8ED7A387CA35" ID-CLB-Dataset="36901" ID-DOI="10.11646/zootaxa.3765.1.1" ID-GBIF-Dataset="dc48dc34-f0b6-4715-8044-1aa2d5851737" ID-ISSN="1175-5326" ID-Zenodo-Dep="285551" ID-ZooBank="4776DD1C-9FCC-4F05-8FDF-66AA14B959FC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460384113075" checkinUser="plazi" docAuthor="Kataev, Boris M., Wrase, David W. & Schmidt, Joachim" docDate="2014" docId="232FAA28FFD2021EFF53FBA4FF43FBDD" docLanguage="en" docName="zt03765p028.pdf" docOrigin="Zootaxa 3765 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chydaeus chuliensis, sp. n." docType="treatment" docVersion="15" lastPageNumber="22" masterDocId="DF16D250FFC6020BFFC4FF95FF9EF910" masterDocTitle="New species of the genus Chydaeus from China, Nepal, Myanmar, and Thailand, with remarks on species previously described (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="21" updateTime="1698297231703" updateUser="ExternalLinkService">
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<mods:mods id="A01DABA033C607C954AC4D0D28375B7D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C3DB1334AD4458F9AA9DB6395BD0E0E2">
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<mods:title id="C23662420F804B47132AA705A969FFA7">New species of the genus Chydaeus from China, Nepal, Myanmar, and Thailand, with remarks on species previously described (Coleoptera: Carabidae: Harpalini)</mods:title>
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<mods:name id="AC0C81C9DF80225B79AFED22BDD28A7B" type="personal">
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<mods:roleTerm id="3D6A3F1B66C3A8E82136BE20300155D9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A449A910B7FAD8AB55BBEF0E6F381970">Kataev, Boris M.</mods:namePart>
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<mods:affiliation id="95CFC9E1A557E1CEC1CE5554C15292B3">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<mods:nameIdentifier id="3AD646C6F746517E9987773767D1F550" type="email">harpal@zin.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name id="04EF1B967A4F054BABB8E05796CD5FF6" type="personal">
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<mods:role id="3CC95EA119C9BD2F35CA86FA5A011D11">
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<mods:roleTerm id="4AF8274CFB217964BA56E51C1DD132B6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4D3A287C8A5CD86F343E1573EDA31A50">Wrase, David W.</mods:namePart>
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<mods:affiliation id="F37AE03213C113FBBD3658D7980A46C8">Dunckerstrasse 78, D- 10437 Berlin, Germany.</mods:affiliation>
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<mods:nameIdentifier id="F14E55D059D9AD6EE3FAC4C4B08F7629" type="email">carterus@gmx.de</mods:nameIdentifier>
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<mods:roleTerm id="A3070CAF62367BDE2023DFCC361CD328">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="06D1E5B46CA97D35EF251AF1E337214B">Schmidt, Joachim</mods:namePart>
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<mods:affiliation id="FD2B4307FF5792A517A9CC113A6CF554">University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitätsplatz 2, D – 18055 Rostock, Germany.</mods:affiliation>
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<mods:nameIdentifier id="741DDD6EFB9AB9DE66F7E90155BAE462" type="email">schmidt@agonum.de</mods:nameIdentifier>
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<mods:typeOfResource id="2C207F8E6031DACCB387574C4EBF813F">text</mods:typeOfResource>
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<mods:title id="A7F6FCEC16D34ADC5FE3C30EF1142159">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B8713DA4F8CB546B4D3E3A01A8D7DB85">
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<mods:date id="C3A91B183BE0C03872A9F5DEDB1C9F50">2014</mods:date>
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<mods:detail id="F6B577BEA7C52178689ED6B822EF5C9E" type="pubDate">
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<mods:number id="45DCD2FF0EB2ADEFD018DF9B105B4BD1">2014-02-14</mods:number>
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<mods:detail id="6A646AB009DC3A1A412C4AE5AF307EC5" type="volume">
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<mods:number id="E870E65C5EDC2527F70054D6C2105F73">3765</mods:number>
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<mods:classification id="5D84847CA87EAF23368CCE782420B252">journal article</mods:classification>
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<mods:identifier id="652F6585E27111EBABDE51B5BB442607" type="CLB-Dataset">36901</mods:identifier>
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<mods:identifier id="4663881A45A8C544EF6D964EE66D2725" type="DOI">10.11646/zootaxa.3765.1.1</mods:identifier>
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<mods:identifier id="D7024E3DDBBFD12990E3295778756856" type="GBIF-Dataset">dc48dc34-f0b6-4715-8044-1aa2d5851737</mods:identifier>
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<mods:identifier id="6E6C060AA8AB21F1F1ADE944741D461F" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="232FAA28FFD2021EFF53FBA4FF43FBDD" ID-DOI="http://doi.org/10.5281/zenodo.6138892" ID-GBIF-Taxon="127667017" ID-Zenodo-Dep="6138892" LSID="urn:lsid:plazi:treatment:232FAA28FFD2021EFF53FBA4FF43FBDD" httpUri="http://treatment.plazi.org/id/232FAA28FFD2021EFF53FBA4FF43FBDD" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="E39C48B5FFD2021FFF53FBA4FE47FD5B" box="[151,473,1073,1099]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="AB391B3EFFD2021FFF53FBA4FE47FD5B" blockId="20.[151,473,1073,1133]" box="[151,473,1073,1099]" pageId="20" pageNumber="21">
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<heading id="F071AC52FFD2021FFF53FBA4FE47FD5B" bold="true" box="[151,473,1073,1099]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<emphasis id="99F2C72CFFD2021FFF53FBA4FE47FD5B" bold="true" box="[151,473,1073,1099]" pageId="20" pageNumber="21">
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<taxonomicName id="6C8660BDFFD2021FFF53FBA4FE11FD5B" ID-CoL="V43S" authority="KATAEV ET AL." box="[151,399,1073,1099]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="chuliensis" status="sp. nov.">
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<emphasis id="99F2C72CFFD2021FFF53FBA4FE11FD5B" bold="true" box="[151,399,1073,1099]" italics="true" pageId="20" pageNumber="21">Chydaeus chuliensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="82C17A57FFD2021FFE52FBA4FE47FD5B" box="[406,473,1073,1099]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E39C48B5FFD2021FFF53FBC1FECFFD7D" box="[151,337,1108,1133]" pageId="20" pageNumber="21" type="description">
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<paragraph id="AB391B3EFFD2021FFF53FBC1FECFFD7D" blockId="20.[151,473,1073,1133]" box="[151,337,1108,1133]" pageId="20" pageNumber="21">
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(
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<figureCitation id="33BD07BBFFD2021FFF5BFBC1FF6BFD7D" box="[159,245,1108,1133]" captionStart="FIGURES 49 – 52" captionStartId="18.[151,263,1941,1964]" captionTargetBox="[309,1278,346,1902]" captionTargetId="figure@18.[302,1285,337,1909]" captionTargetPageId="18" captionText="FIGURES 49 – 52. Chydaeus, dorsal habitus (holotypes). 49, C. ganeshensis sp. n.; 50, C. chuliensis sp. n.; 51, C. wuliangensis sp. n.; 52, C. luxiensis sp. n. Scales = 2.0 mm." httpUri="https://zenodo.org/record/285558/files/figure.png" pageId="20" pageNumber="21">Figs 50</figureCitation>
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,
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<figureCitation id="33BD07BBFFD2021FFEC5FBC1FED6FD7D" box="[257,328,1108,1133]" captionStart="FIGURES 53–63" captionStartId="19.[151,263,1044,1067]" captionTargetBox="[183,1405,197,1021]" captionTargetId="figure@19.[177,1410,193,1023]" captionText="FIGURES 53–63. Chydaeus. 53–55, C. ganeshensis sp. n.; 56 – 63, C. chuliensis sp. n. (53–54, 56–61, holotypes; 55, 62–63, paratypes). 53, 57–58, median lobe of aedeagus; 54, 59, left metepisternum; 55, 62, laterotergite IX and stylus; 56, terminal lamella of median lobe; 60, protarsus; 61, mesotarsus; 63, stylus. 53, 56, 58, 60–61, dorsal aspect; 54–55, 59, 62, ventral aspect; 57, 63, lateral aspect. Scales: A = 0.5 mm (Figs 53, 55–56), B = 1.0 mm (Figs 54, 57–63)." figureDoi="http://doi.org/10.5281/zenodo.6114276" httpUri="https://zenodo.org/record/6114276/files/figure.png" pageId="20" pageNumber="21">56–63</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E39C48B5FFD2021FFF53FB09FD05FC30" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="AB391B3EFFD2021FFF53FB09FD2EFDC8" blockId="20.[151,1436,1180,2033]" pageId="20" pageNumber="21">
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<emphasis id="99F2C72CFFD2021FFF53FB09FEDBFDA5" bold="true" box="[151,325,1180,1205]" pageId="20" pageNumber="21">
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<typeStatus id="743DA59CFFD2021FFF53FB09FF4CFDA5" box="[151,210,1180,1205]" pageId="20" pageNumber="21">Type</typeStatus>
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material
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||
</emphasis>
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||
.
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<materialsCitation id="1BEE1163FFD2021FFE98FB09FD2EFDC8" ID-GBIF-Occurrence="3467093325" collectingDate="2006-05-20" collectorName="J. Schmidt" country="Nepal" elevation="1313" latitude="28.149721" location="W Gupchi Danda" longLatPrecision="20" longitude="84.76833" municipality="Manaslu Mts." pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="743DA59CFFD2021FFE98FB09FE57FDA5" box="[348,457,1180,1205]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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:
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||
<specimenCount id="BD80D0B7FFD2021FFE1FFB08FE6CFDA4" box="[475,498,1181,1204]" pageId="20" pageNumber="21" type="male">♂</specimenCount>
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||
, “
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||
<collectingCountry id="D3915BAEFFD2021FFDD4FB09FDF3FDA4" box="[528,621,1180,1204]" name="Nepal" pageId="20" pageNumber="21">NEPAL</collectingCountry>
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||
<collectingMunicipality id="4B5D8144FFD2021FFDB8FB09FC80FDA4" box="[636,798,1180,1205]" pageId="20" pageNumber="21">Manaslu Mts.</collectingMunicipality>
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<locationDeviation id="BA8B021CFFD2021FFCE9FB09FC0BFDA5" box="[813,917,1180,1205]" pageId="20" pageNumber="21">SE-slope</locationDeviation>
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||
,
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||
<location id="AE594DE5FFD2021FFC6CFB08FBE2FDA5" LSID="urn:lsid:plazi:treatment:232FAA28FFD2021EFF53FBA4FF43FBDD:AE594DE5FFD2021FFC6CFB08FBE2FDA5" box="[936,1148,1180,1205]" country="Nepal" latitude="28.149721" longLatPrecision="20" longitude="84.76833" municipality="Manaslu Mts." name="W Gupchi Danda" pageId="20" pageNumber="21">W Gupchi Danda</location>
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<quantity id="6C7EB6DBFFD2021FFB4FFB09FA8AFDA4" box="[1163,1300,1180,1205]" metricMagnitude="3" metricUnit="m" metricValue="1.3125" metricValueMax="2.6" metricValueMin="0.025" pageId="20" pageNumber="21" unit="m" value="1312.5" valueMax="2600.0" valueMin="25.0">
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<elevation id="20ABFC0DFFD2021FFB4FFB09FA8AFDA4" box="[1163,1300,1180,1205]" metricMagnitude="3" metricUnit="m" metricValue="1.3125" metricValueMax="2.6" metricValueMin="0.025" pageId="20" pageNumber="21" unit="m" value="1312.5" valueMax="2600.0" valueMin="25.0">25–2600 m</elevation>
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</quantity>
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<geoCoordinate id="CEB27DF9FFD2021FFAE7FB09FA02FDA5" box="[1315,1436,1180,1205]" degrees="28" direction="north" minutes="08" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="59" value="28.149721">28°08'59N</geoCoordinate>
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<geoCoordinate id="CEB27DF9FFD2021FFF53FB2AFE91FDC8" box="[151,271,1215,1240]" degrees="84" direction="east" minutes="46" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="06" value="84.76833">84°46'06E</geoCoordinate>
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, 19/
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||
<date id="DF383DFEFFD2021FFEFFFB2AFE33FDC8" box="[315,429,1215,1240]" pageId="20" pageNumber="21" value="2006-05-20">
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<collectingDate id="CF7CC416FFD2021FFEFFFB2AFE33FDC8" box="[315,429,1215,1240]" pageId="20" pageNumber="21" value="2006-05-20">20.5.2006</collectingDate>
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</date>
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leg.
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<collectorName id="06737EE8FFD2021FFE21FB55FDC0FDC8" box="[485,606,1215,1240]" pageId="20" pageNumber="21">J. Schmidt</collectorName>
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||
” (cJS).
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</materialsCitation>
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</paragraph>
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<paragraph id="AB391B3EFFD2021FFF03FB70FD05FC30" blockId="20.[151,1436,1180,2033]" pageId="20" pageNumber="21">
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<materialsCitation id="1BEE1163FFD2021FFF03FB70FC33FDEC" ID-GBIF-Occurrence="3467093308" box="[199,941,1252,1277]" collectingDate="2006-05-20" collectorName="J. Schmidt" country="Nepal" elevation="1313" latitude="28.149721" location="Nepal" longLatPrecision="20" longitude="84.76833" municipality="Manaslu Mts." pageId="20" pageNumber="21" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
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||
<typeStatus id="743DA59CFFD2021FFF03FB70FEA6FDED" box="[199,312,1253,1277]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
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||
:
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||
<collectingCountry id="D3915BAEFFD2021FFE83FB71FE10FDED" box="[327,398,1252,1277]" name="Nepal" pageId="20" pageNumber="21">Nepal</collectingCountry>
|
||
:
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||
<specimenCount id="BD80D0B7FFD2021FFE58FB71FE52FDEC" box="[412,460,1252,1276]" pageId="20" pageNumber="21" type="male">2 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="BD80D0B7FFD2021FFE19FB71FD95FDEC" box="[477,523,1252,1276]" pageId="20" pageNumber="21" type="female">2 ♀</specimenCount>
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||
, same data as holotype (cJS, cWr);
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||
</materialsCitation>
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||
<materialsCitation id="1BEE1163FFD2021FFC7DFB71FD05FC30" ID-GBIF-Occurrence="3467093339" collectingDate="2006-05-18" collectionCode="ZIN" collectorName="J. Schmidt" country="Nepal" elevation="2250" latitude="28.143612" location="Nepal" longLatPrecision="20" longitude="84.744995" municipality="Manaslu Mts." pageId="20" pageNumber="21" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
|
||
<specimenCount id="BD80D0B7FFD2021FFC7DFB71FC77FDEC" box="[953,1001,1252,1276]" pageId="20" pageNumber="21" type="male">2 ♂</specimenCount>
|
||
,
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||
<specimenCount id="BD80D0B7FFD2021FFC3DFB71FBB9FDEC" box="[1017,1063,1252,1276]" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
|
||
, same data, but “
|
||
<quantity id="6C7EB6DBFFD2021FFB3CFB71FA02FDEC" box="[1272,1436,1252,1276]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="20" pageNumber="21" unit="m" value="2250.0" valueMax="2300.0" valueMin="2200.0">
|
||
<elevation id="20ABFC0DFFD2021FFB3CFB71FA02FDEC" box="[1272,1436,1252,1276]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="20" pageNumber="21" unit="m" value="2250.0" valueMax="2300.0" valueMin="2200.0">2200–2300 m</elevation>
|
||
</quantity>
|
||
<geoCoordinate id="CEB27DF9FFD2021FFF53FA9DFE91FC30" box="[151,271,1288,1312]" degrees="28" direction="north" minutes="08" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="37" value="28.143612">28°08'37N</geoCoordinate>
|
||
<geoCoordinate id="CEB27DF9FFD2021FFED2FA9DFE10FC30" box="[278,398,1288,1312]" degrees="84" direction="east" minutes="44" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="42" value="84.744995">84°44'42E</geoCoordinate>
|
||
,
|
||
<date id="DF383DFEFFD2021FFE5EFA92FD92FC30" box="[410,524,1287,1312]" pageId="20" pageNumber="21" value="2006-05-18">
|
||
<collectingDate id="CF7CC416FFD2021FFE5EFA92FD92FC30" box="[410,524,1287,1312]" pageId="20" pageNumber="21" value="2006-05-18">18.5.2006</collectingDate>
|
||
</date>
|
||
” (cJS,
|
||
<collectionCode id="CD9783FBFFD2021FFD9FFA9DFD11FC30" box="[603,655,1288,1312]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="20" pageNumber="21">ZIN</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E39C48B5FFD2021EFF03FAB9FDD6F80C" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="AB391B3EFFD2021FFF03FAB9FDF9FC55" blockId="20.[151,1436,1180,2033]" box="[199,615,1324,1349]" pageId="20" pageNumber="21">
|
||
<emphasis id="99F2C72CFFD2021FFF03FAB9FECFFC55" bold="true" box="[199,337,1324,1349]" pageId="20" pageNumber="21">Description</emphasis>
|
||
. Dorsal habitus:
|
||
<figureCitation id="33BD07BBFFD2021FFDD5FAB9FDFDFC55" box="[529,611,1324,1349]" captionStart="FIGURES 49 – 52" captionStartId="18.[151,263,1941,1964]" captionTargetBox="[309,1278,346,1902]" captionTargetId="figure@18.[302,1285,337,1909]" captionTargetPageId="18" captionText="FIGURES 49 – 52. Chydaeus, dorsal habitus (holotypes). 49, C. ganeshensis sp. n.; 50, C. chuliensis sp. n.; 51, C. wuliangensis sp. n.; 52, C. luxiensis sp. n. Scales = 2.0 mm." httpUri="https://zenodo.org/record/285558/files/figure.png" pageId="20" pageNumber="21">Fig. 50</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="AB391B3EFFD2021FFF03FADAFB48FC78" blockId="20.[151,1436,1180,2033]" box="[199,1238,1359,1384]" pageId="20" pageNumber="21">
|
||
Size. Body length
|
||
<quantity id="6C7EB6DBFFD2021FFE52FADAFD81FC77" box="[406,543,1359,1384]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" metricValueMax="9.5" metricValueMin="8.9" pageId="20" pageNumber="21" unit="mm" value="9.2" valueMax="9.5" valueMin="8.9">8.9–9.5 mm</quantity>
|
||
, width
|
||
<quantity id="6C7EB6DBFFD2021FFDB5FAC5FD65FC77" box="[625,763,1360,1384]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="20" pageNumber="21" unit="mm" value="4.0" valueMax="4.2" valueMin="3.8">3.8–4.2 mm</quantity>
|
||
; in
|
||
<typeStatus id="743DA59CFFD2021FFCE7FADAFC1BFC78" box="[803,901,1359,1384]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
|
||
9.1 and
|
||
<quantity id="6C7EB6DBFFD2021FFC22FAC5FBA3FC77" box="[998,1085,1360,1384]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="20" pageNumber="21" unit="mm" value="3.9">3.9 mm</quantity>
|
||
, respectively.
|
||
</paragraph>
|
||
<paragraph id="AB391B3EFFD2021FFF03FAE1FBF8FCA0" blockId="20.[151,1436,1180,2033]" pageId="20" pageNumber="21">Color. Black, shiny on dorsum; palpi and antennae brown or light brown, palpomeres 1–3 and antennomere 1 infuscate, blackish brown; often tarsi and occasionally knees brown or blackish brown.</paragraph>
|
||
<paragraph id="AB391B3EFFD2021FFF03FA28FD34FCE8" blockId="20.[151,1436,1180,2033]" pageId="20" pageNumber="21">
|
||
Very similar in external structural characters to
|
||
<taxonomicName id="6C8660BDFFD2021FFCC5FA29FBC7FCC5" authority="Tschitscherine" authorityName="Tschitscherine" box="[769,1113,1468,1493]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="99F2C72CFFD2021FFCC5FA29FC0BFCC4" box="[769,917,1468,1492]" italics="true" pageId="20" pageNumber="21">C. semenowi</emphasis>
|
||
(Tschitschérine)
|
||
</taxonomicName>
|
||
, which was redescribed by
|
||
<bibRefCitation id="CF1766CFFFD2021FFF53FA75FE25FCE8" author="Kataev" box="[151,443,1503,1528]" pageId="20" pageNumber="21" refString="Kataev, B. M. & Schmidt, J. (2002) Contribution to knowledge of Chydaeus Chaudoir, 1854: Revision of the Chydaeus bedeli (Tschitscherine, 1897), Chydaeus irvinei (Andrewes, 1930), and Chydaeus semenowi (Tschitscherine, 1898) species groups from the Himalaya and China (Coleoptera, Carabidae, Harpalini). Coleoptera, Schwanfelder Coleopterologische Mitteilungen, 5 (2001), 389 - 425." type="journal article" year="2002">Kataev & Schmidt (2002)</bibRefCitation>
|
||
, differing as follows:
|
||
</paragraph>
|
||
<paragraph id="AB391B3EFFD2021FFF03F991FE47FE70" blockId="20.[151,1436,1180,2033]" pageId="20" pageNumber="21">
|
||
Ligular sclerite subtruncate at apex, with apical angles acute and protruding laterad. Antennae slightly slenderer, with antennomeres 5–7 each about 1.7–2.0 times as long as wide. Pronotum relatively narrower than in
|
||
<taxonomicName id="6C8660BDFFD2021FFF53F9D9FEBBFF74" box="[151,293,1612,1636]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="99F2C72CFFD2021FFF53F9D9FEBBFF74" box="[151,293,1612,1636]" italics="true" pageId="20" pageNumber="21">C. semenowi</emphasis>
|
||
</taxonomicName>
|
||
, markedly narrowed basad, in male without microsculpture even along sides, in female with very fine, isodiametric meshes present only latero-basally. Elytra in male with microsculpture present in very narrow area along basal borders, consisting of isodiametric meshes, and on external two or three intervals, consisting of slightly transverse meshes; in female elytral microsculpture present throughout, consisting of fine transverse meshes on disc, and of more distinct, isodiametric meshes in area just at basal borders and in anterior third of three external intervals; meshes in posterior two-thirds of three external intervals in female slightly transverse, much more distinct than on disc and on apex of inner intervals. Metepisterna (
|
||
<figureCitation id="33BD07BBFFD2021FFC59F8B1FC6FFE2D" box="[925,1009,1828,1853]" captionStart="FIGURES 53–63" captionStartId="19.[151,263,1044,1067]" captionTargetBox="[183,1405,197,1021]" captionTargetId="figure@19.[177,1410,193,1023]" captionText="FIGURES 53–63. Chydaeus. 53–55, C. ganeshensis sp. n.; 56 – 63, C. chuliensis sp. n. (53–54, 56–61, holotypes; 55, 62–63, paratypes). 53, 57–58, median lobe of aedeagus; 54, 59, left metepisternum; 55, 62, laterotergite IX and stylus; 56, terminal lamella of median lobe; 60, protarsus; 61, mesotarsus; 63, stylus. 53, 56, 58, 60–61, dorsal aspect; 54–55, 59, 62, ventral aspect; 57, 63, lateral aspect. Scales: A = 0.5 mm (Figs 53, 55–56), B = 1.0 mm (Figs 54, 57–63)." figureDoi="http://doi.org/10.5281/zenodo.6114276" httpUri="https://zenodo.org/record/6114276/files/figure.png" pageId="20" pageNumber="21">Fig. 59</figureCitation>
|
||
) short, about as long as wide, rather strongly narrowed posteriad.
|
||
</paragraph>
|
||
<paragraph id="AB391B3EFFD2021FFF03F8F8FB4CFEE1" blockId="20.[151,1436,1180,2033]" pageId="20" pageNumber="21">
|
||
Pro- and mesotarsi in male (
|
||
<figureCitation id="33BD07BBFFD2021FFDCFF8F9FD10FE95" box="[523,654,1900,1925]" captionStart="FIGURES 53–63" captionStartId="19.[151,263,1044,1067]" captionTargetBox="[183,1405,197,1021]" captionTargetId="figure@19.[177,1410,193,1023]" captionText="FIGURES 53–63. Chydaeus. 53–55, C. ganeshensis sp. n.; 56 – 63, C. chuliensis sp. n. (53–54, 56–61, holotypes; 55, 62–63, paratypes). 53, 57–58, median lobe of aedeagus; 54, 59, left metepisternum; 55, 62, laterotergite IX and stylus; 56, terminal lamella of median lobe; 60, protarsus; 61, mesotarsus; 63, stylus. 53, 56, 58, 60–61, dorsal aspect; 54–55, 59, 62, ventral aspect; 57, 63, lateral aspect. Scales: A = 0.5 mm (Figs 53, 55–56), B = 1.0 mm (Figs 54, 57–63)." figureDoi="http://doi.org/10.5281/zenodo.6114276" httpUri="https://zenodo.org/record/6114276/files/figure.png" pageId="20" pageNumber="21">Figs 60–61</figureCitation>
|
||
) less strongly enlarged, slightly narrower than pro- and mesotibia at apex, respectively. Female genitalia:
|
||
<figureCitation id="33BD07BBFFD2021FFDF2F81AFD28FEB8" box="[566,694,1935,1960]" captionStart="FIGURES 53–63" captionStartId="19.[151,263,1044,1067]" captionTargetBox="[183,1405,197,1021]" captionTargetId="figure@19.[177,1410,193,1023]" captionText="FIGURES 53–63. Chydaeus. 53–55, C. ganeshensis sp. n.; 56 – 63, C. chuliensis sp. n. (53–54, 56–61, holotypes; 55, 62–63, paratypes). 53, 57–58, median lobe of aedeagus; 54, 59, left metepisternum; 55, 62, laterotergite IX and stylus; 56, terminal lamella of median lobe; 60, protarsus; 61, mesotarsus; 63, stylus. 53, 56, 58, 60–61, dorsal aspect; 54–55, 59, 62, ventral aspect; 57, 63, lateral aspect. Scales: A = 0.5 mm (Figs 53, 55–56), B = 1.0 mm (Figs 54, 57–63)." figureDoi="http://doi.org/10.5281/zenodo.6114276" httpUri="https://zenodo.org/record/6114276/files/figure.png" pageId="20" pageNumber="21">Figs 62–63</figureCitation>
|
||
. Median lobe of aedeagus (
|
||
<figureCitation id="33BD07BBFFD2021FFC2CF81AFBF7FEB8" box="[1000,1129,1935,1960]" captionStart="FIGURES 53–63" captionStartId="19.[151,263,1044,1067]" captionTargetBox="[183,1405,197,1021]" captionTargetId="figure@19.[177,1410,193,1023]" captionText="FIGURES 53–63. Chydaeus. 53–55, C. ganeshensis sp. n.; 56 – 63, C. chuliensis sp. n. (53–54, 56–61, holotypes; 55, 62–63, paratypes). 53, 57–58, median lobe of aedeagus; 54, 59, left metepisternum; 55, 62, laterotergite IX and stylus; 56, terminal lamella of median lobe; 60, protarsus; 61, mesotarsus; 63, stylus. 53, 56, 58, 60–61, dorsal aspect; 54–55, 59, 62, ventral aspect; 57, 63, lateral aspect. Scales: A = 0.5 mm (Figs 53, 55–56), B = 1.0 mm (Figs 54, 57–63)." figureDoi="http://doi.org/10.5281/zenodo.6114276" httpUri="https://zenodo.org/record/6114276/files/figure.png" pageId="20" pageNumber="21">Figs 57–58</figureCitation>
|
||
) in lateral aspect with apex straight or slightly directed ventrad; terminal lamella in dorsal aspect (
|
||
<figureCitation id="33BD07BBFFD2021FFC15F821FBB8FEDD" box="[977,1062,1972,1997]" captionStart="FIGURES 53–63" captionStartId="19.[151,263,1044,1067]" captionTargetBox="[183,1405,197,1021]" captionTargetId="figure@19.[177,1410,193,1023]" captionText="FIGURES 53–63. Chydaeus. 53–55, C. ganeshensis sp. n.; 56 – 63, C. chuliensis sp. n. (53–54, 56–61, holotypes; 55, 62–63, paratypes). 53, 57–58, median lobe of aedeagus; 54, 59, left metepisternum; 55, 62, laterotergite IX and stylus; 56, terminal lamella of median lobe; 60, protarsus; 61, mesotarsus; 63, stylus. 53, 56, 58, 60–61, dorsal aspect; 54–55, 59, 62, ventral aspect; 57, 63, lateral aspect. Scales: A = 0.5 mm (Figs 53, 55–56), B = 1.0 mm (Figs 54, 57–63)." figureDoi="http://doi.org/10.5281/zenodo.6114276" httpUri="https://zenodo.org/record/6114276/files/figure.png" pageId="20" pageNumber="21">Fig. 56</figureCitation>
|
||
) much narrower than that in
|
||
<taxonomicName id="6C8660BDFFD2021FFA46F821FE9BFEE0" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="99F2C72CFFD2021FFA46F821FE9BFEE0" italics="true" pageId="20" pageNumber="21">C. semenowi</emphasis>
|
||
</taxonomicName>
|
||
; internal sac with a large spiny patch on left side and a small spiny patch on right side.
|
||
</paragraph>
|
||
<paragraph id="AB391B3EFFD3021EFF03FF02FDD6F80C" blockId="21.[151,1436,151,717]" pageId="21" pageNumber="22">
|
||
Proportions: HWmax/PWmax = 0.70–0.75; HWmin/PWmax = 0.61–0.63; HWmax/HWmin = 1.15–1.19; PWmax/PL = 1.33–1.45; EL/EW = 1.36–1.43 (
|
||
<quantity id="6C7EB6DBFFD3021EFD71FF29FCD8F9C5" box="[693,838,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" metricValueMax="3.6322" metricValueMin="3.4798" pageId="21" pageNumber="22" unit="in" value="1.4" valueMax="1.43" valueMin="1.37">1.37–1.43 in</quantity>
|
||
male and
|
||
<quantity id="6C7EB6DBFFD3021EFC04FF29FBCEF9C5" box="[960,1104,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.5052" metricValueMax="3.5559999999999996" metricValueMin="3.4544" pageId="21" pageNumber="22" unit="in" value="1.38" valueMax="1.4" valueMin="1.36">1.36–1.40 in</quantity>
|
||
female); EL/PL = 2.43–2.57 (
|
||
<quantity id="6C7EB6DBFFD3021EFF5AFF4AFEACF9E8" box="[158,306,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="6.3246" metricValueMax="6.476999999999999" metricValueMin="6.1722" pageId="21" pageNumber="22" unit="in" value="2.49" valueMax="2.55" valueMin="2.43">2.43–2.55 in</quantity>
|
||
male and
|
||
<quantity id="6C7EB6DBFFD3021EFE70FF4AFDDBF9E8" box="[436,581,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.5278" metricValueMin="6.4262" pageId="21" pageNumber="22" unit="in" value="2.55" valueMax="2.57" valueMin="2.53">2.53–2.57 in</quantity>
|
||
female); EW/PWmax = 1.24–1.31 (
|
||
<quantity id="6C7EB6DBFFD3021EFC37FF4AFB17F9E8" box="[1011,1161,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="3.2512" metricValueMax="3.3274" metricValueMin="3.175" pageId="21" pageNumber="22" unit="in" value="1.28" valueMax="1.31" valueMin="1.25">1.25–1.31 in</quantity>
|
||
male and
|
||
<quantity id="6C7EB6DBFFD3021EFACEFF75FA02F9E8" box="[1290,1436,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="3.1877" metricValueMax="3.2258" metricValueMin="3.1496" pageId="21" pageNumber="22" unit="in" value="1.255" valueMax="1.27" valueMin="1.24">1.24–1.27 in</quantity>
|
||
female); PWmax/PWmin = 1.29–1.34.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E39C48B5FFD3021EFF03FEB2FC01F875" pageId="21" pageNumber="22" type="etymology">
|
||
<paragraph id="AB391B3EFFD3021EFF03FEB2FC01F875" blockId="21.[151,1436,151,717]" pageId="21" pageNumber="22">
|
||
<emphasis id="99F2C72CFFD3021EFF03FEB2FED8F850" bold="true" box="[199,326,295,320]" pageId="21" pageNumber="22">Etymology</emphasis>
|
||
. The specific name refers to the Chuli Himal on the southeastern Manaslu mountain range in Central
|
||
<collectingCountry id="D3915BAEFFD3021EFF34FED9FEABF875" box="[240,309,332,357]" name="Nepal" pageId="21" pageNumber="22">Nepal</collectingCountry>
|
||
, on which southern slope the
|
||
<typeStatus id="743DA59CFFD3021EFD46FED8FD2CF875" box="[642,690,333,357]" pageId="21" pageNumber="22">type</typeStatus>
|
||
series was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E39C48B5FFD3021EFF03FEFAFED4FB70" pageId="21" pageNumber="22" type="discussion">
|
||
<paragraph id="AB391B3EFFD3021EFF03FEFAFED4FB70" blockId="21.[151,1436,151,717]" pageId="21" pageNumber="22">
|
||
<emphasis id="99F2C72CFFD3021EFF03FEFAFE4AF898" bold="true" box="[199,468,367,392]" pageId="21" pageNumber="22">Comparative remarks</emphasis>
|
||
. This new species is very similar in habitus and apparently closely related to
|
||
<taxonomicName id="6C8660BDFFD3021EFA46FEE5FE9BF8BC" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="99F2C72CFFD3021EFA46FEE5FE9BF8BC" italics="true" pageId="21" pageNumber="22">C. semenowi</emphasis>
|
||
</taxonomicName>
|
||
, which is widely distributed over the Himalaya.
|
||
<taxonomicName id="6C8660BDFFD3021EFCFEFE01FBBCF8BC" box="[826,1058,404,428]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="chuliensis" status="sp. nov.">
|
||
<emphasis id="99F2C72CFFD3021EFCFEFE01FBBCF8BC" box="[826,1058,404,428]" italics="true" pageId="21" pageNumber="22">Chydaeus chuliensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="82C17A57FFD3021EFBE8FE01FBF3F8BD" box="[1068,1133,404,429]" pageId="21" pageNumber="22" rank="species">
|
||
<emphasis id="99F2C72CFFD3021EFBE8FE01FBF3F8BD" bold="true" box="[1068,1133,404,429]" pageId="21" pageNumber="22">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
distinctly differs from the latter species in having the metepisterna much shorter, the elytral microsculpture in female much finer and consisting largely of transverse meshes, and the median lobe of aedeagus in male with much narrower terminal lamella and with a larger left spiny patch in the internal sac. Additionally, the apex of the median lobe of the new species is straight or slightly directed ventrad, as opposed to that of
|
||
<taxonomicName id="6C8660BDFFD3021EFC6FFDB1FBA2FB2C" box="[939,1084,548,572]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="99F2C72CFFD3021EFC6FFDB1FBA2FB2C" box="[939,1084,548,572]" italics="true" pageId="21" pageNumber="22">C. semenowi</emphasis>
|
||
</taxonomicName>
|
||
, which has the apex distinctly directed dorsad.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E39C48B5FFD3021EFF03FDF9FF43FBDD" pageId="21" pageNumber="22" type="distribution">
|
||
<paragraph id="AB391B3EFFD3021EFF03FDF9FF43FBDD" blockId="21.[151,1436,151,717]" pageId="21" pageNumber="22">
|
||
<emphasis id="99F2C72CFFD3021EFF03FDF9FEC7FB95" bold="true" box="[199,345,620,645]" pageId="21" pageNumber="22">Distribution</emphasis>
|
||
. Known only from the
|
||
<typeStatus id="743DA59CFFD3021EFDAEFDF8FD04FB95" box="[618,666,621,645]" pageId="21" pageNumber="22">type</typeStatus>
|
||
locality, the southeastern slope of the Chuli Himal west of Gupchi Danda, Central
|
||
<collectingCountry id="D3915BAEFFD3021EFE8CFD1AFE11FBB8" box="[328,399,655,680]" name="Nepal" pageId="21" pageNumber="22">Nepal</collectingCountry>
|
||
. This new species seems to be endemic to the southeastern portion of the Manaslu mountain range.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |