treatments-xml/data/BF/E2/9D/BFE29D80FFCA5227A9BA392EC8A5133F.xml
2024-06-21 12:50:06 +02:00

191 lines
20 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" ID-Pensoft-Pub="2511-6428-1-13" ID-Pensoft-UUID="F4C1C6A2DFD95D3BAF5AF1CE09226837" ID-ZooBank="589077DFF8BB45ABB30D7E0DE1C307FE" ModsDocID="2511-6428-74-1-13" checkinTime="1706792588976" checkinUser="pensoft" docAuthor="Irmler, Ulrich" docDate="2024" docId="BFE29D80FFCA5227A9BA392EC8A5133F" docLanguage="en" docName="ContribEntomol 74(1): 13-34" docOrigin="Contributions to Entomology 74 (1)" docPubDate="2024-01-31" docSource="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" docTitle="Somoleptus densiceps Irmler 2024, sp. nov." docType="treatment" docUuid="F5F35C18-A269-4EEA-9A43-C1EAD7A46E47" docUuidSource="ZooBank" docVersion="1" id="F4C1C6A2DFD95D3BAF5AF1CE09226837" lastPageNumber="13" masterDocId="F4C1C6A2DFD95D3BAF5AF1CE09226837" masterDocTitle="Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)" masterLastPageNumber="34" masterPageNumber="13" pageNumber="13" updateTime="1706792588976" updateUser="pensoft">
<mods:mods id="8EC658803F1975F4E2BF067B9296C60E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="33F28812137FC75A0958D58257D6880F">
<mods:title id="BE213F0BE960CAC5B97E80475F4118DB">Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)</mods:title>
</mods:titleInfo>
<mods:name id="8AD445D5BF727A83DBAA18FE3B6C37B4" type="personal">
<mods:role id="156F90FB5E75ECFFE529742AB683EC6B">
<mods:roleTerm id="2BAD0FB6F4C39AA9615D61666EE59513">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="818DD8D3E86A9A5C778301EE19C1AFF3">Irmler, Ulrich</mods:namePart>
<mods:affiliation id="0DDB9E93AC43E606152FC6315C1E94F9">Institute for Ecosystem Research, University of Kiel, Olshausenstrasse 40, 24098 Kiel, Germany</mods:affiliation>
<mods:nameIdentifier id="E898FD75CE8A767E79E5BAFFF930A363" type="email">uirmler@ecology.uni-kiel.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7CDF0D12EA012ACB3C99960B6ADCA167">text</mods:typeOfResource>
<mods:relatedItem id="C007956E9C97FC5387E860DED7CE2CF3" type="host">
<mods:titleInfo id="C8346E7E2917F6AB365CB87BC07E25EE">
<mods:title id="8C7DDAADF36BC707AC14573DAB09EB83">Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="D16820A3995E7C79C79105C08BCC1F58">
<mods:date id="446443B203FD3F8C07931ED7A70362CF">2024</mods:date>
<mods:detail id="91767D8E70005CDB103E89DE2DF72BA4" type="pubDate">
<mods:number id="CE4D073715CBFBB80DEC6DF3E87A5E7D">2024-01-31</mods:number>
</mods:detail>
<mods:detail id="EE69D252CBFC3F0B1073E7DDD1ADD75C" type="volume">
<mods:number id="14110F42B9A45BF05DBABBA6AD5A68F2">74</mods:number>
</mods:detail>
<mods:detail id="7A3FBFB9EF7E66B9E6E35244415F253D" type="issue">
<mods:number id="63BB20B089DE34707F6E52C10CEDF309">1</mods:number>
</mods:detail>
<mods:extent id="F41369E2CF25B3F0320ACDC30DA18D2C" unit="page">
<mods:start id="172C3C3998E78F231312417460B00995">13</mods:start>
<mods:end id="A4881BE33827BA1E3CD1B30E54B4EF8B">34</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="BB917DE0EF3FD30B3614BAA6CED0B5B9">
<mods:url id="EE7B44510E62DB27DFB4A4F0BC3439D8">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:url>
</mods:location>
<mods:classification id="3190FFD9B9D6DE442A09BE07A744651A">journal article</mods:classification>
<mods:identifier id="ACF6EFC5634593913F756F946DE2B9A7" type="DOI">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:identifier>
<mods:identifier id="7F4285E809CCDF9885851BC88494F6D7" type="Pensoft-Pub">2511-6428-1-13</mods:identifier>
<mods:identifier id="376B264D66A664BED24B85FF289F5AE9" type="ZooBank">589077DFF8BB45ABB30D7E0DE1C307FE</mods:identifier>
<mods:identifier id="4CEBF62E4698FCA477ED2916613B4A68" type="Pensoft-UUID">F4C1C6A2DFD95D3BAF5AF1CE09226837</mods:identifier>
</mods:mods>
<treatment id="BFE29D80FFCA5227A9BA392EC8A5133F" LSID="urn:lsid:zoobank.org:act:F5F35C18-A269-4EEA-9A43-C1EAD7A46E47" httpUri="http://treatment.plazi.org/id/BFE29D80FFCA5227A9BA392EC8A5133F" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection id="C76A40A8E85704D079D2366C233EB387" pageId="0" pageNumber="13" type="nomenclature">
<paragraph id="1E4748729D235FBFCFDB294366C8AE33" pageId="0" pageNumber="13">
<taxonomicName id="926E83DE26013F2875E581A25A79B7DB" LSID="https://zoobank.org/F5F35C18-A269-4EEA-9A43-C1EAD7A46E47" authority="Irmler, 2024" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus densiceps" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="densiceps" status="sp. nov.">Somoleptus densiceps</taxonomicName>
<taxonomicNameLabel id="1F9A0775925CF446D9AA2C4BF83EE48C" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="4B1D229CD118EE67A17B59EBBA03EB1E" pageId="0" pageNumber="13" type="description">
<paragraph id="71C3BE96D776450FDA301F4C8AB635DD" pageId="0" pageNumber="13">
<figureCitation id="F8CAB83F4E4822F475C774D286CFE40C" captionStart="Figures 1316" captionStartId="F4" captionText="Figures 13 - 16. 13. Somoleptus densiceps; 14. S. guianensis; 15. S. luteicornis; 16. S. paramocola. a. Aedeagus; b. Paramere; c. Sternite VII; d. Tergite VII. Scale bars: 0.5 mm (a, c, d); 0.1 mm (b)." figureDoi="10.3897/contrib.entomol.74.e114543.figures13-16" httpUri="https://binary.pensoft.net/fig/975718" pageId="0" pageNumber="13">Figs 13a-d</figureCitation>
<figureCitation id="88D9A1A93B3D5AE914E1215FAE7FACF4" captionStart="Figure 21" captionStartId="F8" captionText="Figure 21. A. Fore-body of Somoleptus densiceps; B. S. guianensis; C. S. luteicornis; D. S. paramocola; E. S. puntarenae; F. S. umicola. Scale bars: 0.5 mm." figureDoi="10.3897/contrib.entomol.74.e114543.figure21" httpUri="https://binary.pensoft.net/fig/975722" pageId="0" pageNumber="13">, 21A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BB0F4126F4243B12CC930CAC6C26A931" pageId="0" pageNumber="13" type="materials_examined">
<paragraph id="D2658B1B8F206A89D539978B926B30AA" pageId="0" pageNumber="13">Material examined.</paragraph>
<paragraph id="9191B4B4D1DA789630B4A91D78784417" pageId="0" pageNumber="13">
<materialsCitation id="B59F7F136D5671750B1823A5764CE7B1" collectingDate="1996-06-19" collectorName="La Culebra Trail, R. Anderson" country="PANAMA" elevation="1450" latitude="8.823055" location="La Culebra Trail" longLatPrecision="21" longitude="-82.42166" specimenCount="1" stateProvince="Chiriqui" typeStatus="Holotype">
<emphasis id="7CE2878322CBC1D17FC303FB7FF09B2F" bold="true" italics="true" pageId="0" pageNumber="13">
<typeStatus id="93671C76E3DA3959A034A8D881E35A2D" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="DC7587F4EFD0C3193A22A82572172381" name="Panama">PANAMA</collectingCountry>
male;
<collectingRegion id="2E7E07858FBC59527930C888368A2E44" country="Panama" name="Chiriqui">Chiriqui</collectingRegion>
;
<locationDeviation id="B33EF3184024DE5C3F59CF7B26891054" location="Boquete">
<quantity id="B42722C8965D66AD9FB21C4E5153CCB1" metricMagnitude="3" metricUnit="m" metricValue="5.6" unit="km" value="5.6">5.6 km</quantity>
N Boquete
</locationDeviation>
;
<collectorName id="785EB07030227D5280F29C6A5AF913FB">
<location id="F29C4010B39A233C7944BE1641E0B13E" LSID="urn:lsid:plazi:treatment:BFE29D80FFCA5227A9BA392EC8A5133F:F29C4010B39A233C7944BE1641E0B13E" country="PANAMA" latitude="8.823055" longLatPrecision="21" longitude="-82.42166" name="La Culebra Trail" stateProvince="Chiriqui">La Culebra Trail</location>
</collectorName>
;
<geoCoordinate id="55EA10CE3D00B6E9E07675D1FBE85645" degrees="8" direction="north" minutes="49" orientation="latitude" precision="15" seconds="23" value="8.823055">8°49'23&quot;N</geoCoordinate>
,
<geoCoordinate id="A8EAC7A845D1297CC97B43330A132906" degrees="82" direction="west" minutes="25" orientation="longitude" precision="15" seconds="18" value="-82.42166">82°25'18&quot;W</geoCoordinate>
;
<quantity id="9FC3427BE80311A26896002B0439651F" metricMagnitude="3" metricUnit="m" metricValue="1.45" unit="m" value="1450.0">
<elevation id="986B18D8197B81C0B79F95D82D2821CE" metricMagnitude="3" metricUnit="m" metricValue="1.45" unit="m" value="1450.0">1450 m</elevation>
</quantity>
elevation; oak forest litter;
<collectingDate id="68021436A982285926931217B73E269B" value="1996-06-19">19 VI 1996</collectingDate>
;
<collectorName id="0CC58B1DA60DD5FD8C1C023AFCC3AF2E">R. Anderson</collectorName>
leg.; KNHM PAN2A96 96-141B.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1BCC2FECC0228D2E330E6EBD7D79177F" pageId="0" pageNumber="13" type="diagnosis">
<paragraph id="611249B171679DBFAA997BBE61E1408C" pageId="0" pageNumber="13">Diagnosis.</paragraph>
<paragraph id="11034B4996896167FD08EA3309F3D56B" pageId="0" pageNumber="13">
<taxonomicName id="01167408A1A8C3B2ADE305DCA0646593" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus densiceps" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="densiceps">
<emphasis id="CBB86240A718E0786179F09CF3D1BE9B" italics="true" pageId="0" pageNumber="13">Somoleptus densiceps</emphasis>
</taxonomicName>
is part of the
<taxonomicName id="02A7DBAFD16852487FB042C139F4976F" baseAuthorityName="LeConte" baseAuthorityYear="1863" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus longicollis" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="longicollis">Somoleptus longicollis</taxonomicName>
-group due to the brush-like cones of the aedeagus and the shape of sternite VII of male. Amongst the Neotropical
<taxonomicName id="637B01A6C68BB3561741541C58A526FF" authorityName="Sharp" authorityYear="1885" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="E477FFEE0A80042C938FBE084016A4E8" italics="true" pageId="0" pageNumber="13">Somoleptus</emphasis>
</taxonomicName>
species,
<taxonomicName id="CCE2F7F40CE1B16264815F2D56FB406F" lsidName="S. densiceps" pageId="0" pageNumber="13" rank="species" species="densiceps">
<emphasis id="26F0A81147F1D687C2C81A602101A77C" italics="true" pageId="0" pageNumber="13">S. densiceps</emphasis>
</taxonomicName>
mostly resembles
<taxonomicName id="B54B6DCFFEC833B80824235647CEA40D" lsidName="S. mexicanus" pageId="0" pageNumber="13" rank="species" species="mexicanus">
<emphasis id="D748D37418DF35195F0F1A9AED4F74F2" italics="true" pageId="0" pageNumber="13">S. mexicanus</emphasis>
</taxonomicName>
<bibRefCitation id="465AC33EB11FF2BD56DECE1A30C6C817" DOI="https://doi.org/10.3897/contrib.entomol.72.e87211" author="Irmler, U" journalOrPublisher="Contributions to Entolology" pageId="0" pageNumber="13" pagination="123 - 178" refId="B4" refString="Irmler, U, 2022. The Neotropical species of the genus Somoleptus Sharp, 1885 (Coleoptera: Staphylininae: Xantholinini). Contributions to Entolology 72: 123 - 178, DOI: https://doi.org/10.3897/contrib.entomol.72.e87211" title="The Neotropical species of the genus Somoleptus Sharp, 1885 (Coleoptera: Staphylininae: Xantholinini)." url="https://doi.org/10.3897/contrib.entomol.72.e87211" volume="72" year="2022">Irmler 2022</bibRefCitation>
by the deep and dense punctation, the short eyes and the slightly triangular shape of the head. Parameres are also very similar between both species. In comparison with
<taxonomicName id="C54D5938594EECA7E010B3368A2E420F" lsidName="S. mexicanus" pageId="0" pageNumber="13" rank="species" species="mexicanus">
<emphasis id="E4A3BA516C54A79BC22CBC610B045143" italics="true" pageId="0" pageNumber="13">S. mexicanus</emphasis>
</taxonomicName>
, punctation on total fore-body in
<taxonomicName id="612A3DB58C5A9C03699C8DCA00D620C6" lsidName="S. densiceps" pageId="0" pageNumber="13" rank="species" species="densiceps">
<emphasis id="4F200490F4F1DC8E6CB1B384A1EC0ADE" italics="true" pageId="0" pageNumber="13">S. densiceps</emphasis>
</taxonomicName>
is much denser, nearly coriaceous.
</paragraph>
<caption id="6C8CF8B3EFFEDABC01367EEBDFB8DB95" doi="10.3897/contrib.entomol.74.e114543.figures9-12" httpUri="https://binary.pensoft.net/fig/975717" pageId="0" pageNumber="13" start="Figures 912" startId="F3">
<paragraph id="D2379ABDFC314A94324E35C6C2A67705" pageId="0" pageNumber="13">
<emphasis id="386DA2A01A26503B43557422848FD0B1" bold="true" pageId="0" pageNumber="13">Figures 9-12.</emphasis>
<emphasis id="524DDE39ACCB72F1472696051C1000D5" bold="true" pageId="0" pageNumber="13">9.</emphasis>
<taxonomicName id="388507E648B328F38BE67E8167386CED" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Lithocharodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithocharodes parallelus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="99A2BFFF2B99049C0147A57965138E3F" italics="true" pageId="0" pageNumber="13">Lithocharodes parallelus</emphasis>
</taxonomicName>
;
<emphasis id="2D626F50457727E937A670B7B0C95780" bold="true" pageId="0" pageNumber="13">10.</emphasis>
<taxonomicName id="4B7FBA77C00861704A9983B2D708B0B6" lsidName="L. zamorae" pageId="0" pageNumber="13" rank="species" species="zamorae">
<emphasis id="27558AD4BC17FB7DA96B8B9E2913FBB4" italics="true" pageId="0" pageNumber="13">L. zamorae</emphasis>
</taxonomicName>
;
<emphasis id="01AFDBEB5CFD8B94DBA37D3D0391BB92" bold="true" pageId="0" pageNumber="13">11.</emphasis>
<taxonomicName id="366DDC1EF6E57B5E11DFD5AA318C6AA2" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus alajuelae" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="alajuelae">
<emphasis id="D9EF1B26D18FA88BD838D4E8A6BEF3AA" italics="true" pageId="0" pageNumber="13">Somoleptus alajuelae</emphasis>
</taxonomicName>
;
<emphasis id="22CD2476DD9B17DBB6DD060613A9716E" bold="true" pageId="0" pageNumber="13">12.</emphasis>
<taxonomicName id="0CEF146C43B698B5DB2B193F03BED61B" lsidName="S. curtioculatus" pageId="0" pageNumber="13" rank="species" species="curtioculatus">
<emphasis id="22463E5C3FB8C39254F3F259CD72E7C5" italics="true" pageId="0" pageNumber="13">S. curtioculatus</emphasis>
</taxonomicName>
.
<emphasis id="9549484A72423281649FB22C01831FDD" bold="true" pageId="0" pageNumber="13">a.</emphasis>
Aedeagus;
<emphasis id="39D051C618ADE7D017EBF5C1FA755ACD" bold="true" pageId="0" pageNumber="13">b.</emphasis>
Paramere;
<emphasis id="E7EB7B38CC949FF637B3E4A208C9378A" bold="true" pageId="0" pageNumber="13">c.</emphasis>
Sternite VII;
<emphasis id="C35E895D4BC6EBA4D21044BE0B18BE93" bold="true" pageId="0" pageNumber="13">d.</emphasis>
Tergite VII. Scale bars: 0.5 mm (
<emphasis id="970A96628910D5C34832FCAB2383E629" bold="true" pageId="0" pageNumber="13">a, c, d</emphasis>
); 0.1 mm (
<emphasis id="636E67120FFA36F43DB2358E39F14FFB" bold="true" pageId="0" pageNumber="13">b</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E831ADFF4664764453D24D727F2D416B" pageId="0" pageNumber="13" type="description">
<paragraph id="5F0EA2CD61169BB8C91107AFC51E1BC1" pageId="0" pageNumber="13">Description.</paragraph>
<paragraph id="64983CDDB094C4ABFFAFA66A148038D5" pageId="0" pageNumber="13">Length: 4.0 mm; Colouration: completely black; antennae dark brown, legs lighter brown.</paragraph>
<paragraph id="599C320B83CD5D12E0093D65C4D357F2" pageId="0" pageNumber="13">Head: 0.68 mm long, 0.54 mm wide; eyes moderately large; PS:E ratio 3.8; postocular sides slightly divergent; posterior part combined with posterior edge semi-circular; inter-antennal furrows indistinct; setiferous punctation deep and extremely dense; on average, interstices shorter than one fourth of diameter of punctures; at sides, partly coriaceous; in apical part, between eyes with indistinct impunctate mid-line; surface without microsculpture; due to dense punctation scarcely shiny. Antennae with first antennomere elongate; half as long as total head-length; second and third antennomere conical; longer than wide; combined half as long as first antennomere; following antennomere at least twice as wide as long; apically, increasing in width. Pronotum: 0.80 mm long, 0.53 mm wide; widest at anterior third; strongly convergent to neck; posteriorly slightly convergent to sub-rectangular angles; setiferous punctation dense and extremely dense; partly coriaceous; on average, interstices between punctures one fourth as wide as diameter of punctures; wide mid-line impunctate with three single punctures; surface without microsculpture; polished. Elytra: 0.74 mm long, 0.67 mm wide; humeral angles obtuse; sides approximately parallel; posterior angles sub-rectangular; posterior margin slightly retreated to suture; setiferous punctation deep and dense; on average, interstices as wide as half-length of diameter of punctures; surface without microsculpture; shiny. Abdomen with setiferous punctation dense, but much finer than on fore-body; surface without microsculpture; shiny; posterior margin of sternite VII of male slightly triangular with short central knob; tergite VII of male with straight posterior margin; meso-tibia with 5, meta-tibia with 2 ctenidia. Aedeagus oval with sub-rectangular anterior margin; dorsal plate covering nearly total central lobe; endophallus elongate with one torsion; covered by short lobes; cones at apical orifice long; C:A ratio 0.34; paramere bilobed; inner lobe elongate with several long setae at inner edge; outer lobe circular; transparent.</paragraph>
</subSubSection>
<subSubSection id="5E1A934B3E6DFC2605EC9B917F7EB609" pageId="0" pageNumber="13" type="etymology">
<paragraph id="652B824668EF660DAA3F9E2903418370" pageId="0" pageNumber="13">Etymology.</paragraph>
<paragraph id="C73D044C70BAFF7844AF9D433182A8C6" pageId="0" pageNumber="13">
The species name is a combination of the Latin word
<emphasis id="267F683886DDF356F331E71FEB0EB3B2" italics="true" pageId="0" pageNumber="13">densus</emphasis>
meaning dense and
<emphasis id="6F690E228C3F660DCF8AECA08656539F" italics="true" pageId="0" pageNumber="13">ceps</emphasis>
(short form of
<emphasis id="3DB6A0F1424A54E37D33D6B1354705AC" italics="true" pageId="0" pageNumber="13">caput</emphasis>
) meaning
<normalizedToken id="5F69E3B4354F5A17117D3021C63257DC" originalValue="“head”">&quot;head&quot;</normalizedToken>
and refers to the dense punctation of the head.
</paragraph>
</subSubSection>
<subSubSection id="F440FB3007772B796443B98D85446E61" pageId="0" pageNumber="13" type="geography">
<paragraph id="649201BD3F37AC914042802C3FDA66F6" pageId="0" pageNumber="13">Geography.</paragraph>
<paragraph id="F91CB82925198673E3FFF73D5739F330" pageId="0" pageNumber="13">Panama.</paragraph>
</subSubSection>
<subSubSection id="F366B3E5333DCEB555CD147F35942C21" pageId="0" pageNumber="13" type="ecology">
<paragraph id="75ABDC99745DAC3BB1D6B45BBADFAB58" pageId="0" pageNumber="13">Ecology.</paragraph>
<paragraph id="BDAA193A1210A9A192508D0A70A22940" pageId="0" pageNumber="13">Forest floor of montane forests at approximately 1500 m elevation.</paragraph>
</subSubSection>
</treatment>
</document>