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<document id="1187DC9AF3F7AC19C993194F464C7502" ID-CLB-Dataset="226692" ID-DOI="10.11646/phytotaxa.579.1.6" ID-GBIF-Dataset="7526d7b5-488c-40af-ba2a-5a7a9ce6d1bf" ID-ISSN="1179-3163" ID-Zenodo-Dep="7532127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673570700385" checkinUser="plazi" docAuthor="Montesinos-Tubée, Daniel B. &amp; Iamonico, Duilio" docDate="2023" docId="8D770006FFF15203FF42E0AEFB76FC8A" docLanguage="en" docName="phytotaxa.579.1.6.pdf" docOrigin="Phytotaxa 579 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Arenaria humboldtiana Montesinos &amp; Iamonico 2023, nom. nov." docType="treatment" docVersion="5" lastPageNumber="58" masterDocId="714E787EFFF25207FFCAE41CFFE4FFCA" masterDocTitle="Typifications of Andean Arenaria names (Caryophyllaceae), with a nomen novum" masterLastPageNumber="60" masterPageNumber="54" pageNumber="57" updateTime="1699473875558" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="774737A2BBEDBEE1D5B3F661797A2D47">Typifications of Andean Arenaria names (Caryophyllaceae), with a nomen novum</mods:title>
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<mods:namePart id="E356BB20294091ABBAF6915DF43ED02E">Montesinos-Tubée, Daniel B.</mods:namePart>
<mods:nameIdentifier id="CB0287D7579A1F9067F267F12F38777F" type="ORCID">0000-0002-4439-5089</mods:nameIdentifier>
<mods:affiliation id="13E4B8683C75AE2A068191AE6B08A3CD">Botanic Garden and Botanical Museum Berlin (BGBM), Freie Universität Berlin, Königin-Luise-Str. 6 - 8, 14195 Berlin, Germany. &amp; Instituto Científico Michael Owen Dillon, Av. Jorge Chávez 610, Cercado, Arequipa, Peru. &amp; Instituto de Ciencia y Gestión Ambiental de la Universidad Nacional de San Agustín de Arequipa. Calle San Agustín 108, Arequipa- Peru. &amp; dbmtperu @ gmail. com; https: // orcid. org / 0000 - 0002 - 4439 - 5089</mods:affiliation>
<mods:nameIdentifier id="B361DCF7A40CC7D1953611A6E0CBABBB" type="email">dbmtperu@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F9E536442B01442396F8E80E7F2A70BF">Iamonico, Duilio</mods:namePart>
<mods:nameIdentifier id="407801EC30AA3EEEDA862BBC201EE4EF" type="ORCID">0000-0001-5491-7568</mods:nameIdentifier>
<mods:affiliation id="52959F398AA8A3AEA1E57467081D1F74">Ce. R. S. I. Te. S., Viale XXIV Maggio 7, 04100 Latina, Italy. &amp; duilio. iamonico @ uniroma 1. it; https: // orcid. org / 0000 - 0001 - 5491 - 7568</mods:affiliation>
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<mods:title id="39152DB46844B69B7A13B2DF34591AB4">Phytotaxa</mods:title>
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<mods:part id="1CA627D52CF64D4B3EEF4AB7820A0C97">
<mods:date id="A0B96F0A874515D6FEF20A995168866B">2023</mods:date>
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<mods:number id="60001246EAA753ED68F1D251221686CB">2023-01-13</mods:number>
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<mods:number id="1E8CAF3A86A821E4C63E9F11695EA716">579</mods:number>
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<mods:identifier id="F730CFA443F042AC2C3FD2E6C2A64433" type="DOI">10.11646/phytotaxa.579.1.6</mods:identifier>
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<treatment id="8D770006FFF15203FF42E0AEFB76FC8A" ID-DOI="http://doi.org/10.5281/zenodo.7544401" ID-GBIF-Taxon="208979674" ID-Zenodo-Dep="7544401" LSID="urn:lsid:plazi:treatment:8D770006FFF15203FF42E0AEFB76FC8A" httpUri="http://treatment.plazi.org/id/8D770006FFF15203FF42E0AEFB76FC8A" lastPageId="4" lastPageNumber="58" pageId="3" pageNumber="57">
<subSubSection id="4DC4E29BFFF15204FF42E0AEFCF8F82E" pageId="3" pageNumber="57" type="discussion">
<paragraph id="0561B110FFF15204FF42E0AEFE7FFB06" blockId="3.[136,1452,1202,2021]" box="[136,411,1202,1229]" pageId="3" pageNumber="57">
<heading id="5E29067CFFF15204FF42E0AEFE7FFB06" bold="true" box="[136,411,1202,1229]" fontSize="11" level="1" pageId="3" pageNumber="57" reason="1">
<taxonomicName id="C2DECA93FFF15204FF42E0AEFE7FFB06" authorityYear="1911" box="[136,411,1202,1229]" pageId="3" pageNumber="57">
<emphasis id="37AA6D02FFF15204FF42E0AEFE7FFB06" bold="true" box="[136,411,1202,1229]" italics="true" pageId="3" pageNumber="57">Scleranthus peruvianus</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="0561B110FFF15204FF42E0CAFA64F96B" blockId="3.[136,1452,1202,2021]" pageId="3" pageNumber="57">
<bibRefCitation id="614FCCE1FFF15204FF42E0CAFE5CFB3A" author="Muschler, R." box="[136,440,1238,1264]" pageId="3" pageNumber="57" pagination="441 - 461" refId="ref4583" refString="Muschler, R. (1911) Caryophyllaceae andinae. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 45: 441 - 461." type="journal article" year="1911">Muschler (1911: 460–461)</bibRefCitation>
provided a detailed description of of
<taxonomicName id="C2DECA93FFF15204FCA1E0CAFB90FB3A" authorityName="Muschler" authorityYear="1911" box="[875,1140,1238,1264]" class="Magnoliopsida" family="Caryophyllaceae" genus="Scleranthus" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="57" phylum="Tracheophyta" rank="species" species="peruvianus">
<emphasis id="37AA6D02FFF15204FCA1E0CAFB90FB3A" box="[875,1140,1238,1264]" italics="true" pageId="3" pageNumber="57">Scleranthus peruvianus</emphasis>
</taxonomicName>
, as well as the provenance (“
<collectingCountry id="7DC9F180FFF15204FF51E0E6FF31FADE" box="[155,213,1274,1300]" name="Peru" pageId="3" pageNumber="57">Peru</collectingCountry>
: Saltus Coymolache supra Hualgayoc, departimento
<collectingRegion id="C71A7FF2FFF15204FC81E0E6FC22FADE" box="[843,966,1274,1300]" country="Peru" name="Cajamarca" pageId="3" pageNumber="57">Cajamarca</collectingRegion>
, in formationes fruticetis el cactacearum speciminibus composita,
<quantity id="C2261CF5FFF15204FE69E102FDAFFAF3" box="[419,587,1310,1337]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="3" pageNumber="57" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">
4000
<emphasis id="37AA6D02FFF15204FE11E102FE13FAF2" bold="true" box="[475,503,1310,1336]" pageId="3" pageNumber="57">―</emphasis>
4100 m
</quantity>
s.m.”). A single specimen (
<typeStatus id="DA650FB2FFF15204FCB6E102FC31FAF2" box="[892,981,1310,1336]" pageId="3" pageNumber="57" type="syntype">syntype</typeStatus>
according to the Art. 9.6 of ICN) was also reported: “WEBERBAUER n. 3985.
<emphasis id="37AA6D02FFF15204FE33E15EFDF1FA96" bold="true" box="[505,533,1346,1372]" pageId="3" pageNumber="57">―</emphasis>
Specimina florigera 11. Maii 1904.
<emphasis id="37AA6D02FFF15204FC69E15EFC5BFA96" bold="true" box="[931,959,1346,1372]" pageId="3" pageNumber="57">―</emphasis>
Herb. Berol.”. Hence,
<bibRefCitation id="614FCCE1FFF15204FB71E15EFA4FFA96" author="Muschler, R." box="[1211,1451,1346,1372]" pageId="3" pageNumber="57" pagination="441 - 461" refId="ref4583" refString="Muschler, R. (1911) Caryophyllaceae andinae. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 45: 441 - 461." type="journal article" year="1911">Muschler (1911: 461)</bibRefCitation>
explicitly referred to a specimen preserved at the Herbarium Berolinensis (the current acronym is B). However, most of the B specimens were destroyed during the World War II (see e.g.,
<bibRefCitation id="614FCCE1FFF15204FCB8E196FBF5FA6E" author="Sleumer, H." box="[882,1041,1418,1444]" pageId="3" pageNumber="57" pagination="172 - 175" refId="ref4990" refString="Sleumer, H. (1949) The Botanical Gardens and Museum at Berlin-Dahlem. Kew Bulletin 4 (2): 172 - 175. https: // doi. org / 10.2307 / 4113675" type="journal article" year="1949">Sleumer 1949</bibRefCitation>
,
<bibRefCitation id="614FCCE1FFF15204FBD6E196FB55FA6E" author="Hiepko, P." box="[1052,1201,1418,1444]" pageId="3" pageNumber="57" pagination="219 - 252" refId="ref3974" refString="Hiepko, P. (1987) The collections of the Botanical Museum Berlin Dahlem (B) and their history. In: Scholtz, H. (Ed.) Botany in Berlin. Englera 7: 219 - 252." type="journal article" year="1987">Hiepko 1987</bibRefCitation>
). According to
<bibRefCitation id="614FCCE1FFF15204FA96E196FEA6FA02" author="Stafleu, A. F. &amp; Cowan, R. S." pageId="3" pageNumber="57" refId="ref5114" refString="Stafleu, A. F. &amp; Cowan, R. S. (1988) Taxonomic literature, 2 nd ed., vol. 7 (W - Z). Bohn, Scheltema &amp; Holtema, Utrecht, 653 pp." type="book" year="1988">Stafleu &amp; Cowan (1988)</bibRefCitation>
and HUH-Index of Botanists (2013+), a large part of Weberbauer’s Herbarium and types are deposited also at MOL and USM where, therefore, duplicates of the specimen cited by
<bibRefCitation id="614FCCE1FFF15204FBD0E1CEFB3DFA26" author="Muschler, R." box="[1050,1241,1490,1516]" pageId="3" pageNumber="57" pagination="441 - 461" refId="ref4583" refString="Muschler, R. (1911) Caryophyllaceae andinae. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 45: 441 - 461." type="journal article" year="1911">Muschler (1911)</bibRefCitation>
in the protologue could be found there. We traced
<specimenCount id="13D87A99FFF15204FE34E1EBFD7EF9DA" box="[510,666,1526,1553]" pageId="3" pageNumber="57" type="generic">one specimen</specimenCount>
at MOL (barcode MOL00000505) bearing one plant collected by A. Weberbauer in “
<emphasis id="37AA6D02FFF15204FEF5E207FB97F9FF" box="[319,1139,1562,1589]" italics="true" pageId="3" pageNumber="57">
Paso de Coymolache, encima de Hualgayoc (Departimento:
<collectingRegion id="C71A7FF2FFF15204FC26E207FB8EF9FF" box="[1004,1130,1563,1589]" country="Peru" name="Cajamarca" pageId="3" pageNumber="57">Cajamarca</collectingRegion>
)
</emphasis>
|
<emphasis id="37AA6D02FFF15204FB4DE206FAFFF9FE" box="[1159,1307,1562,1588]" italics="true" pageId="3" pageNumber="57">
<quantity id="C2261CF5FFF15204FB4DE206FAFFF9FE" box="[1159,1307,1562,1588]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="3" pageNumber="57" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">4000-4100 m</quantity>
</emphasis>
” as reported on the original label. Although the date of collection is lacking, the specimen was numbered with 3985, as indicated in the protologue. Therefore, we are sure that this MOL specimen is part of the original material. It morphologically matches the Muschler’s description and it is here designated as the
<typeStatus id="DA650FB2FFF15204FCB3E29AFC04F96A" box="[889,992,1670,1696]" pageId="3" pageNumber="57" type="lectotype">lectotype</typeStatus>
of the name
<taxonomicName id="C2DECA93FFF15204FBB8E29AFA9DF96A" authorityName="Muschler" authorityYear="1911" box="[1138,1401,1670,1696]" class="Magnoliopsida" family="Caryophyllaceae" genus="Scleranthus" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="57" phylum="Tracheophyta" rank="species" species="peruvianus">
<emphasis id="37AA6D02FFF15204FBB8E29AFA9DF96A" box="[1138,1401,1670,1696]" italics="true" pageId="3" pageNumber="57">Scleranthus peruvianus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0561B110FFF15204FF77E2B6FCF8F82E" blockId="3.[136,1452,1202,2021]" pageId="3" pageNumber="57">
Concerning the identity of this taxon, note firstly that it was mentioned by
<bibRefCitation id="614FCCE1FFF15204FBC7E2B6FB35F90E" author="MacBride, J. F." box="[1037,1233,1706,1732]" pageId="3" pageNumber="57" pagination="578 - 639" refId="ref4515" refString="MacBride, J. F. (1936) Caryophyllaceae Juss. Flora of Peru. Publications of the Field Museum of Natural History. Botanical series 13 (2): 578 - 639." type="journal article" year="1936">MacBride (1936)</bibRefCitation>
,
<bibRefCitation id="614FCCE1FFF15204FB14E2B6FF31F922" author="Brako, L. &amp; Zarucchi, J." pageId="3" pageNumber="57" pagination="1 - 1286" refId="ref3511" refString="Brako, L. &amp; Zarucchi, J. (1993) Catalogue of the Flowering Plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45: 1 - 1286." type="journal article" year="1993">Brako &amp; Zarucchi (1993)</bibRefCitation>
, and Ulloa-Ulloa
<emphasis id="37AA6D02FFF15204FE50E2D3FE36F922" box="[410,466,1742,1768]" italics="true" pageId="3" pageNumber="57">et al.</emphasis>
(2017) under
<taxonomicName id="C2DECA93FFF15204FDA4E2D2FD16F922" box="[622,754,1742,1768]" class="Magnoliopsida" family="Caryophyllaceae" genus="Scleranthus" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="57" phylum="Tracheophyta" rank="genus">
<emphasis id="37AA6D02FFF15204FDA4E2D2FD16F922" box="[622,754,1742,1768]" italics="true" pageId="3" pageNumber="57">Scleranthus</emphasis>
</taxonomicName>
. Also,
<bibRefCitation id="614FCCE1FFF15204FC8AE2D2FC14F922" author="POWO" box="[832,1008,1742,1768]" pageId="3" pageNumber="57" refId="ref4707" refString="POWO (2022 c) Plants of the World Online. Scleranthus peruvianus Muschl. Available from: https: // powo. science. kew. org / taxon / urn: lsid: ipni. org: names: 230610 - 2 (accessed 29 November 2022)." type="url" year="2022">POWO (2022c)</bibRefCitation>
currently accepts Muschler’s species as belonging to
<taxonomicName id="C2DECA93FFF15204FED1E2EEFE7BF8C6" box="[283,415,1778,1804]" class="Magnoliopsida" family="Caryophyllaceae" genus="Scleranthus" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="57" phylum="Tracheophyta" rank="genus">
<emphasis id="37AA6D02FFF15204FED1E2EEFE7BF8C6" box="[283,415,1778,1804]" italics="true" pageId="3" pageNumber="57">Scleranthus</emphasis>
</taxonomicName>
. According to the detailed description given by
<bibRefCitation id="614FCCE1FFF15204FC7DE2EEFB96F8C6" author="Muschler, R." box="[951,1138,1778,1804]" pageId="3" pageNumber="57" pagination="441 - 461" refId="ref4583" refString="Muschler, R. (1911) Caryophyllaceae andinae. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 45: 441 - 461." type="journal article" year="1911">Muschler (1911)</bibRefCitation>
,
<taxonomicName id="C2DECA93FFF15204FBB4E2EFFAF1F8C6" authorityName="Muschler" authorityYear="1911" box="[1150,1301,1779,1805]" class="Magnoliopsida" family="Caryophyllaceae" genus="Scleranthus" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="57" phylum="Tracheophyta" rank="species" species="peruvianus">
<emphasis id="37AA6D02FFF15204FBB4E2EFFAF1F8C6" box="[1150,1301,1779,1805]" italics="true" pageId="3" pageNumber="57">S. peruvianus</emphasis>
</taxonomicName>
flowers have five petals (“Petala 5, alba, subulata, calycis lacinias subaequantia”), while
<taxonomicName id="C2DECA93FFF15204FC18E30AFBB2F8FA" box="[978,1110,1814,1840]" class="Magnoliopsida" family="Caryophyllaceae" genus="Scleranthus" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="57" phylum="Tracheophyta" rank="genus">
<emphasis id="37AA6D02FFF15204FC18E30AFBB2F8FA" box="[978,1110,1814,1840]" italics="true" pageId="3" pageNumber="57">Scleranthus</emphasis>
</taxonomicName>
is defined as having apetalous flowers (see e.g.,
<bibRefCitation id="614FCCE1FFF15204FE99E327FD7AF89E" author="West, J. G. &amp; Garnock-Jones, P. J." box="[339,670,1850,1877]" pageId="3" pageNumber="57" pagination="435 - 450" refId="ref5694" refString="West, J. G. &amp; Garnock-Jones, P. J. (1986) Evolution and biogeography of Scleranthus (Caryophyllaceae) in Australasia. In: Barlow, B. A (Eds.) Flora and Fauna of Alpine Australasia. Ages and Origins. Sciro in association with Australian Systematic Botany Society. Australia. pp. 435 - 450." type="book chapter" year="1986">West &amp; Garnock-Jones 1986</bibRefCitation>
,
<bibRefCitation id="614FCCE1FFF15204FD61E326FCA1F89E" author="Walters, S. M." box="[683,837,1850,1876]" pageId="3" pageNumber="57" pagination="139" refId="ref5590" refString="Walters, S. M. (Ed.) (1993) Caryophyllaceae Juss. In: Tutin, T. G., Burges, N. A., Chater, A. O, Edmondson, J. R., Heywood, V. H., Moore, D. M., Valentine, D. H., Walters, S. M. &amp; Webb, D. A. (eds.) Flora Europaea, ed. 2, vol. 1. Cambridge University Press, Cambridge, pp. 139." type="journal article" year="1993">Walters 1993</bibRefCitation>
). On the basis of our examination of the
<typeStatus id="DA650FB2FFF15204FAEAE326FA6EF89E" box="[1312,1418,1850,1876]" pageId="3" pageNumber="57" type="lectotype">lectotype</typeStatus>
,
<taxonomicName id="C2DECA93FFF15204FA5CE327FEE0F8B2" authorityName="Muschler" authorityYear="1911" class="Magnoliopsida" family="Caryophyllaceae" genus="Scleranthus" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="57" phylum="Tracheophyta" rank="species" species="peruvianus">
<emphasis id="37AA6D02FFF15204FA5CE327FEE0F8B2" italics="true" pageId="3" pageNumber="57">S. peruvianus</emphasis>
</taxonomicName>
should be transferred to
<taxonomicName id="C2DECA93FFF15204FDE5E343FD77F8B3" authorityName="Linnaeus" authorityYear="1753" box="[559,659,1887,1913]" class="Magnoliopsida" family="Caryophyllaceae" genus="Arenaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="57" phylum="Tracheophyta" rank="genus">
<emphasis id="37AA6D02FFF15204FDE5E343FD77F8B3" box="[559,659,1887,1913]" italics="true" pageId="3" pageNumber="57">Arenaria</emphasis>
</taxonomicName>
due to its lanceolate-linear leaves without stipules, five free petals, two styles, and dehiscent capsules with two teeth, each with 6–10 seeds. Since an “
<taxonomicName id="C2DECA93FFF15204FBFEE39FFAF6F856" authorityName="Poeppig ex Rohrbach" authorityYear="1872" box="[1076,1298,1923,1949]" class="Magnoliopsida" family="Caryophyllaceae" genus="Arenaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="57" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="37AA6D02FFF15204FBFEE39FFAF6F856" box="[1076,1298,1923,1949]" italics="true" pageId="3" pageNumber="57">Arenaria peruviana</emphasis>
</taxonomicName>
” was already validly published by Rohrbach (
<taxonomicName id="C2DECA93FFF15204FE3AE3BBFBDBF80B" authority="Poeppig ex Rohrbach 1872: 275" authorityName="Poeppig ex Rohrbach" authorityPageNumber="275" authorityYear="1872" box="[496,1087,1958,1985]" class="Magnoliopsida" family="Caryophyllaceae" genus="Arenaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="57" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="37AA6D02FFF15204FE3AE3BBFD29F80A" box="[496,717,1959,1985]" italics="true" pageId="3" pageNumber="57">Arenaria peruviana</emphasis>
Poeppig ex Rohrbach 1872: 275
</taxonomicName>
), a
<emphasis id="37AA6D02FFF15204FBACE3BBFAE6F80A" box="[1126,1282,1959,1984]" italics="true" pageId="3" pageNumber="57">nomen novum</emphasis>
is necessary to avoid a later and illegitimate homonym (Art. 53.1 of ICN).
</paragraph>
</subSubSection>
<subSubSection id="4DC4E29BFFF65203FF42E482FCF9FF72" box="[136,797,158,185]" pageId="4" pageNumber="58" type="nomenclature">
<paragraph id="0561B110FFF65203FF42E482FCF9FF72" blockId="4.[136,1452,158,833]" box="[136,797,158,185]" pageId="4" pageNumber="58">
<taxonomicName id="C2DECA93FFF65203FF42E482FD4FFF72" ID-CoL="B3MD4" authority="Montesinos &amp; Iamonico" authorityName="Montesinos &amp; Iamonico" authorityYear="2023" box="[136,683,158,185]" class="Magnoliopsida" family="Caryophyllaceae" genus="Arenaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="58" phylum="Tracheophyta" rank="species" species="humboldtiana" status="nom. nov.">
<emphasis id="37AA6D02FFF65203FF42E482FE73FF73" bold="true" box="[136,407,158,185]" italics="true" pageId="4" pageNumber="58">Arenaria humboldtiana</emphasis>
Montesinos &amp; Iamonico
</taxonomicName>
,
<emphasis id="37AA6D02FFF65203FD79E483FCF9FF72" box="[691,797,159,184]" italics="true" pageId="4" pageNumber="58">
<taxonomicNameLabel id="2C99D079FFF65203FD79E483FCF9FF72" box="[691,797,159,184]" pageId="4" pageNumber="58">nom. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="4DC4E29BFFF65203FCEBE483FA99FF72" box="[801,1405,158,184]" pageId="4" pageNumber="58" type="reference_group">
<paragraph id="0561B110FFF65203FCEBE483FA99FF72" blockId="4.[136,1452,158,833]" box="[801,1405,158,184]" pageId="4" pageNumber="58">
<emphasis id="37AA6D02FFF65203FCEBE483FCA3FF72" box="[801,839,159,184]" italics="true" pageId="4" pageNumber="58">pro</emphasis>
<taxonomicName id="C2DECA93FFF65203FC86E482FA99FF72" ID-CoL="4VNHV" authority="Muschler (1911: 460 - 461)" authorityName="Muschler" authorityPageNumber="460 - 461" authorityYear="1911" box="[844,1405,158,184]" class="Magnoliopsida" family="Caryophyllaceae" genus="Scleranthus" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="58" phylum="Tracheophyta" rank="species" species="peruvianus">
<emphasis id="37AA6D02FFF65203FC86E482FBB5FF72" box="[844,1105,158,184]" italics="true" pageId="4" pageNumber="58">Scleranthus peruvianus</emphasis>
<bibRefCitation id="614FCCE1FFF65203FB9CE482FA99FF72" author="Muschler, R." box="[1110,1405,158,184]" pageId="4" pageNumber="58" pagination="441 - 461" refId="ref4583" refString="Muschler, R. (1911) Caryophyllaceae andinae. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 45: 441 - 461." type="journal article" year="1911">Muschler (1911: 460–461)</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="4DC4E29BFFF65203FA4BE483FCC7FECA" pageId="4" pageNumber="58" type="multiple">
<paragraph id="0561B110FFF65203FA4BE483FCC7FECA" blockId="4.[136,1452,158,833]" pageId="4" pageNumber="58">
non
<taxonomicName id="C2DECA93FFF65203FF42E4DFFD15FF17" authority="Poeppig ex Rohrbach (1872: 275)" authorityName="Poeppig ex Rohrbach" authorityPageNumber="275" authorityYear="1872" box="[136,753,194,221]" class="Magnoliopsida" family="Caryophyllaceae" genus="Arenaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="58" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="37AA6D02FFF65203FF42E4DFFE82FF16" box="[136,358,195,221]" italics="true" pageId="4" pageNumber="58">Arenaria peruviana</emphasis>
Poeppig ex Rohrbach (1872: 275)
</taxonomicName>
, nec
<taxonomicName id="C2DECA93FFF65203FCE7E4DFFEEAFECA" authority="(Muschler 1911: 457 - 458) Molinari (2016: 276)" authorityName="Molinari" authorityPageNumber="276" authorityYear="2016" baseAuthorityName="Muschler" baseAuthorityPageNumber="457 - 458" baseAuthorityYear="1911" class="Magnoliopsida" family="Caryophyllaceae" genus="Arenaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="58" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="37AA6D02FFF65203FCE7E4DFFBE8FF16" box="[813,1036,195,221]" italics="true" pageId="4" pageNumber="58">Arenaria peruviana</emphasis>
(
<bibRefCitation id="614FCCE1FFF65203FBD1E4DEFADDFF17" author="Muschler, R." box="[1051,1337,194,221]" pageId="4" pageNumber="58" pagination="441 - 461" refId="ref4583" refString="Muschler, R. (1911) Caryophyllaceae andinae. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 45: 441 - 461." type="journal article" year="1911">Muschler 1911: 457–458</bibRefCitation>
)
<bibRefCitation id="614FCCE1FFF65203FA83E4DEFEEAFECA" author="Molinari, E." pageId="4" pageNumber="58" pagination="275 - 277" refId="ref4551" refString="Molinari, E. (2016) Further nomenclatural changes regarding Peruvian endemics. Polish Botanical Journal 61: 275 - 277. https: // 10.1515 / pbj- 2016 - 0021" type="journal article" year="2016">Molinari (2016: 276)</bibRefCitation>
</taxonomicName>
,
<emphasis id="37AA6D02FFF65203FED3E4FBFE75FECA" box="[281,401,230,256]" italics="true" pageId="4" pageNumber="58">nom. illeg.</emphasis>
(later homonym; Art. 53.1 of ICN).
</paragraph>
</subSubSection>
<subSubSection id="4DC4E29BFFF65203FF42E512FAD0FE8D" pageId="4" pageNumber="58" type="materials_examined">
<paragraph id="0561B110FFF65203FF42E512FAD0FE8D" blockId="4.[136,1452,158,833]" pageId="4" pageNumber="58">
<materialsCitation id="B5B6BB4DFFF65203FF42E512FAD4FE8D" ID-GBIF-Occurrence="4015708309" collectionCode="MOL" country="Peru" elevation="4050" location="Hualgayoc" municipality="Coymolache Pass" pageId="4" pageNumber="58" specimenCode="MOL00000505" specimenCount="1" stateProvince="Cajamarca" typeStatus="lectotype">
<typeStatus id="DA650FB2FFF65203FF42E512FF0CFEE9" box="[136,232,270,291]" pageId="4" pageNumber="58" type="lectotype">Lectotype</typeStatus>
(designated here):—
<collectingCountry id="7DC9F180FFF65203FE61E512FE08FEEE" box="[427,492,270,292]" name="Peru" pageId="4" pageNumber="58">PERU</collectingCountry>
.
<collectingRegion id="C71A7FF2FFF65203FE39E512FDBAFEEE" box="[499,606,270,292]" country="Peru" name="Cajamarca" pageId="4" pageNumber="58">Cajamarca</collectingRegion>
:
<collectingMunicipality id="E5052B6AFFF65203FDA9E512FCE8FEE9" box="[611,780,270,292]" pageId="4" pageNumber="58">Coymolache Pass</collectingMunicipality>
, above
<location id="0001E7CBFFF65203FC85E512FC5DFEEE" LSID="urn:lsid:plazi:treatment:8D770006FFF15203FF42E0AEFB76FC8A:0001E7CBFFF65203FC85E512FC5DFEEE" box="[847,953,270,292]" country="Peru" municipality="Coymolache Pass" name="Hualgayoc" pageId="4" pageNumber="58" stateProvince="Cajamarca">Hualgayoc</location>
, in grassland steppes,
<quantity id="C2261CF5FFF65203FB48E512FAE2FEE9" box="[1154,1286,270,291]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="4" pageNumber="58" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">
<elevation id="8EF35623FFF65203FB48E512FAE2FEE9" box="[1154,1286,270,291]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="4" pageNumber="58" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">4000–4100 m</elevation>
</quantity>
,
<emphasis id="37AA6D02FFF65203FAC7E512FACBFEE9" box="[1293,1327,270,291]" italics="true" pageId="4" pageNumber="58">s.d.</emphasis>
,
<emphasis id="37AA6D02FFF65203FAFDE512FF09FE8D" italics="true" pageId="4" pageNumber="58">Weberbauer 3985</emphasis>
(
<specimenCode id="5578196BFFF65203FF30E52EFE7CFE82" box="[250,408,306,328]" pageId="4" pageNumber="58">MOL00000505</specimenCode>
!, image available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.mol00000505)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4DC4E29BFFF65203FF42E54EFC8BFE12" pageId="4" pageNumber="58" type="etymology">
<paragraph id="0561B110FFF65203FF42E54EFC8BFE12" blockId="4.[136,1452,158,833]" pageId="4" pageNumber="58">
<emphasis id="37AA6D02FFF65203FF42E54EFECAFEA6" bold="true" box="[136,302,338,364]" pageId="4" pageNumber="58">Etymology:―</emphasis>
The specific epithet honours Alexander von Humboldt (1769–1859), considered as a leading mind in the natural sciences, and who worked widely in botany, but also physics, geology, and oceanology. Humboldt is known for having placed Andean flora and fauna into distinct climatic and topographic contexts and defined human impacts on climate change as potentially affecting the evolution of society.
</paragraph>
</subSubSection>
<subSubSection id="4DC4E29BFFF65203FF77E5FEFDE0FDEA" pageId="4" pageNumber="58" type="distribution">
<paragraph id="0561B110FFF65203FF77E5FEFDE0FDEA" blockId="4.[136,1452,158,833]" pageId="4" pageNumber="58">
<emphasis id="37AA6D02FFF65203FF77E5FEFDE2FE36" bold="true" box="[189,518,482,508]" pageId="4" pageNumber="58">Distribution and ecology:―</emphasis>
<taxonomicName id="C2DECA93FFF65203FDCCE5FFFCE8FE36" authorityName="Montesinos &amp; Iamonico" authorityYear="2023" box="[518,780,482,509]" class="Magnoliopsida" family="Caryophyllaceae" genus="Arenaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="58" phylum="Tracheophyta" rank="species" species="humboldtiana">
<emphasis id="37AA6D02FFF65203FDCCE5FFFCE8FE36" box="[518,780,482,509]" italics="true" pageId="4" pageNumber="58">Arenaria humboldtiana</emphasis>
</taxonomicName>
is an endemic Peruvian species occurring in montane forests at
<quantity id="C2261CF5FFF65203FF69E61AFEDAFDEB" box="[163,318,518,545]" metricMagnitude="3" metricUnit="m" metricValue="3.95" metricValueMax="4.1" metricValueMin="3.8" pageId="4" pageNumber="58" unit="m" value="3950.0" valueMax="4100.0" valueMin="3800.0">3800–4100 m</quantity>
a.s.l. (pers. obs.).
</paragraph>
</subSubSection>
<subSubSection id="4DC4E29BFFF65203FF77E636FAB0FDA2" pageId="4" pageNumber="58" type="discussion">
<paragraph id="0561B110FFF65203FF77E636FAB0FDA2" blockId="4.[136,1452,158,833]" pageId="4" pageNumber="58">
<emphasis id="37AA6D02FFF65203FF77E636FEFFFD8E" bold="true" box="[189,283,554,580]" pageId="4" pageNumber="58">Note:―</emphasis>
By recognizing Muschler’s taxon under the genus
<taxonomicName id="C2DECA93FFF65203FC96E637FC24FD8F" authorityName="Linnaeus" authorityYear="1753" box="[860,960,555,581]" class="Magnoliopsida" family="Caryophyllaceae" genus="Arenaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="58" phylum="Tracheophyta" rank="genus">
<emphasis id="37AA6D02FFF65203FC96E637FC24FD8F" box="[860,960,555,581]" italics="true" pageId="4" pageNumber="58">Arenaria</emphasis>
</taxonomicName>
,
<taxonomicName id="C2DECA93FFF65203FC1AE636FA48FD8E" authority="Linnaeus (1753: 406)" authorityName="Linnaeus" authorityPageNumber="406" authorityYear="1753" box="[976,1452,554,581]" class="Magnoliopsida" family="Caryophyllaceae" genus="Scleranthus" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="58" phylum="Tracheophyta" rank="species" species="annuus">
<emphasis id="37AA6D02FFF65203FC1AE636FB4AFD8E" box="[976,1198,554,580]" italics="true" pageId="4" pageNumber="58">Scleranthus annuus</emphasis>
<bibRefCitation id="614FCCE1FFF65203FB7DE636FA48FD8E" author="Linnaeus, C." box="[1207,1452,554,581]" pageId="4" pageNumber="58" refId="ref4410" refString="Linnaeus, C. (1753) Species plantarum, vol. 1. Laurentii Salvii, Holmiae, 560 pp. https: // dx. doi. org / 10.5962 / bhl. title. 669" type="book" year="1753">Linnaeus (1753: 406)</bibRefCitation>
</taxonomicName>
remains as the only species (alien) of
<taxonomicName id="C2DECA93FFF65203FDE4E652FD56FDA2" box="[558,690,590,616]" class="Magnoliopsida" family="Caryophyllaceae" genus="Scleranthus" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="58" phylum="Tracheophyta" rank="genus">
<emphasis id="37AA6D02FFF65203FDE4E652FD56FDA2" box="[558,690,590,616]" italics="true" pageId="4" pageNumber="58">Scleranthus</emphasis>
</taxonomicName>
known to occur in South America (see e.g.,
<bibRefCitation id="614FCCE1FFF65203FB6EE652FAACFDA2" author="POWO" box="[1188,1352,590,616]" pageId="4" pageNumber="58" refId="ref4758" refString="POWO (2022 d) Plants of the World Online. Scleranthus annuus L. Available from: https: // powo. science. kew. org / taxon / urn: lsid: ipni. org: names: 300494 - 2 (accessed 29 November 2022)." type="url" year="2022">POWO 2022d</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4DC4E29BFFF65203FF77E66EFB76FC8A" pageId="4" pageNumber="58" type="materials_examined">
<paragraph id="0561B110FFF65203FF77E66EFB76FC8A" blockId="4.[136,1452,158,833]" pageId="4" pageNumber="58">
<emphasis id="37AA6D02FFF65203FF77E66EFE38FD46" bold="true" box="[189,476,626,652]" pageId="4" pageNumber="58">Examined specimens:―</emphasis>
<collectingCountry id="7DC9F180FFF65203FE16E66FFDC3FD47" box="[476,551,627,653]" name="Peru" pageId="4" pageNumber="58">PERU</collectingCountry>
.
<collectingRegion id="C71A7FF2FFF65203FDFBE66EFD49FD46" box="[561,685,626,652]" country="Peru" name="Cajamarca" pageId="4" pageNumber="58">Cajamarca</collectingRegion>
; Celendín; Jalca de Kumulca, mountain pass on the road Cajamarca-Celendín,
<quantity id="C2261CF5FFF65203FF36E68AFEB0FD7B" box="[252,340,662,689]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="58" unit="m" value="3700.0">3700 m</quantity>
,
<date id="716097D0FFF65203FEABE68AFDF3FD7A" box="[353,535,662,688]" pageId="4" pageNumber="58" value="2004-08-26">26 August 2004</date>
,
<emphasis id="37AA6D02FFF65203FDE9E68AFC89FD7A" box="[547,877,662,689]" italics="true" pageId="4" pageNumber="58">Sklenar &amp; Zapata Cruz 8712</emphasis>
(B101104856!);
<collectingRegion id="C71A7FF2FFF65203FBFAE68AFB48FD7A" box="[1072,1196,662,688]" country="Peru" name="Cajamarca" pageId="4" pageNumber="58">Cajamarca</collectingRegion>
; Celendín; Sorochuco, Minasconga, W of the lake, swamp areas,
<quantity id="C2261CF5FFF65203FD98E6A6FD42FD1F" box="[594,678,698,725]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="58" unit="m" value="3900.0">3900 m</quantity>
,
<date id="716097D0FFF65203FD7AE6A6FC67FD1E" box="[688,899,698,724]" pageId="4" pageNumber="58" value="1997-12-13">13 December 1997</date>
,
<emphasis id="37AA6D02FFF65203FC46E6A6FB84FD1E" box="[908,1120,698,724]" italics="true" pageId="4" pageNumber="58">Sánchez Vega 9226</emphasis>
(CPUN013316!);
<collectingRegion id="C71A7FF2FFF65203FAE6E6A6FA4CFD1E" box="[1324,1448,698,724]" country="Peru" name="Cajamarca" pageId="4" pageNumber="58">Cajamarca</collectingRegion>
; Celendín; Sorochuco, Minasconga, E of the lake Lucmacocha or Perol, following the path that sorrounds the lake, Jalca grasslands con rocky outcrops,
<quantity id="C2261CF5FFF65203FDEBE71EFD9EFCD7" box="[545,634,770,797]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="58" unit="m" value="3800.0">3800 m</quantity>
.,
<date id="716097D0FFF65203FD42E71EFCB4FCD6" box="[648,848,770,796]" pageId="4" pageNumber="58" value="2003-12-04">4 December 2003</date>
,
<emphasis id="37AA6D02FFF65203FC90E71EFB88FCD6" box="[858,1132,770,796]" italics="true" pageId="4" pageNumber="58">Sánchez Vega et al. 2497</emphasis>
(CPUN16750!);
<collectingRegion id="C71A7FF2FFF65203FAE6E71EFA4CFCD6" box="[1324,1448,770,796]" country="Peru" name="Cajamarca" pageId="4" pageNumber="58">Cajamarca</collectingRegion>
; Hualgayoc; Coymolache,
<quantity id="C2261CF5FFF65203FE66E73AFDE7FC8B" box="[428,515,806,833]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="58" unit="m" value="4000.0">4000 m</quantity>
,
<date id="716097D0FFF65203FDC4E73AFD43FC8A" box="[526,679,806,833]" pageId="4" pageNumber="58" value="2001-06-15">15 June 2001</date>
,
<emphasis id="37AA6D02FFF65203FD78E73AFC33FC8A" box="[690,983,806,832]" italics="true" pageId="4" pageNumber="58">Sánchez Vega et al. 10786</emphasis>
(CPUN14898!).
</paragraph>
</subSubSection>
</treatment>
</document>
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